236 lines
53 KiB
XML
236 lines
53 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3804004" ID-GBIF-Dataset="4d505549-2f45-49c2-b388-8fc8bfdbd5f3" ID-GBIF-Taxon="163737024" ID-Zenodo-Dep="3804004" checkinTime="1587658244385" checkinUser="plazi" docAuthor="Jałoszynski, Paweł, Luo, Xiao-Zhu, Hammel, Jorg U., Yamamoto, Sh ^ uhei & Beutel, Rolf G." docDate="2020" docId="03E7BF21FF80C80AFF39F9C1FA37FD7E" docLanguage="en" docName="14772019.2020.1747561.pdf" docOrigin="Journal of Systematic Palaeontology 18" docStyle="DocumentStyle{}" docTitle="Lepiceratus ankylosaurus Jaloszynski, Luo, Yamamoto & Beutel 2020, sp. nov." docType="treatment" docVersion="6" lastPageId="8" lastPageNumber="9" masterDocId="FFDEC759FF87C802FFE7FFC1FF8DFFDA" masterDocTitle="The mid-Cretaceous † Lepiceratus Gen. nov. and the Evolution of the Relict Beetle Family Lepiceridae (Insecta: Coleoptera: Myxophaga)" masterLastPageNumber="14" masterPageNumber="1" pageId="7" pageNumber="8" updateTime="1643446050710" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The mid-Cretaceous † Lepiceratus Gen. nov. and the Evolution of the Relict Beetle Family Lepiceridae (Insecta: Coleoptera: Myxophaga)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jałoszynski, Paweł</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Luo, Xiao-Zhu</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hammel, Jorg U.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yamamoto, Sh ^ uhei</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Beutel, Rolf G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Systematic Palaeontology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-04-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>18</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>14</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1080/14772019.2020.1747561</mods:identifier>
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<mods:identifier type="GBIF-Dataset">4d505549-2f45-49c2-b388-8fc8bfdbd5f3</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3763694</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3804004" ID-GBIF-Taxon="163737024" ID-Zenodo-Dep="3804004" LSID="urn:lsid:plazi:treatment:03E7BF21FF80C80AFF39F9C1FA37FD7E" httpUri="http://treatment.plazi.org/id/03E7BF21FF80C80AFF39F9C1FA37FD7E" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<subSubSection pageId="7" pageNumber="8" type="nomenclature">
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<paragraph blockId="7.[222,718,1536,1627]" pageId="7" pageNumber="8">
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†
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<taxonomicName authority="Jaloszynski, Luo, Yamamoto & Beutel" authorityName="Jaloszynski, Luo, Yamamoto & Beutel" authorityYear="2020" class="Insecta" family="Lepiceridae" genus="Lepiceratus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ankylosaurus" status="sp. nov.">
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<emphasis bold="true" box="[235,521,1536,1560]" italics="true" pageId="7" pageNumber="8">Lepiceratus ankylosaurus</emphasis>
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Jałoszynski, Luo, Yamamoto & Beutel
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</taxonomicName>
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,
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<taxonomicNameLabel box="[548,635,1569,1592]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection box="[413,526,1603,1627]" pageId="7" pageNumber="8" type="description">
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<paragraph blockId="7.[222,718,1536,1627]" box="[413,526,1603,1627]" pageId="7" pageNumber="8">
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(
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<figureCitation box="[422,517,1603,1627]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[159,226,1858,1879]" captionStartId-1="4.[159,226,1321,1342]" captionStartId-2="5.[159,226,1539,1560]" captionTargetBox-0="[175,1455,196,1830]" captionTargetBox-1="[159,1454,189,1304]" captionTargetBox-2="[159,1467,188,1517]" captionTargetId-0="figure@3.[159,1467,188,1840]" captionTargetId-1="figure@4.[159,1467,188,1304]" captionTargetId-2="figure@5.[159,1467,188,1517]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." captionText-1="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." captionText-2="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi-0="http://doi.org/10.5281/zenodo.3763696" figureDoi-1="http://doi.org/10.5281/zenodo.3763698" figureDoi-2="http://doi.org/10.5281/zenodo.3763700" httpUri-0="https://zenodo.org/record/3763696/files/figure.png" httpUri-1="https://zenodo.org/record/3763698/files/figure.png" httpUri-2="https://zenodo.org/record/3763700/files/figure.png" pageId="7" pageNumber="8">Figs 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="8" type="materials_examined">
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<paragraph blockId="7.[159,781,1684,1772]" pageId="7" pageNumber="8">
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<materialsCitation ID-GBIF-Occurrence="2608526856" collectionCode="FMNH" country="Myanmar" location="Hukawng Valley, Kachin State of northern" pageId="7" pageNumber="8" specimenCode="FMNHINS-3966034" specimenCount="1" typeStatus="holotype">
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<emphasis bold="true" box="[159,269,1684,1707]" pageId="7" pageNumber="8">
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<typeStatus box="[159,265,1684,1707]" pageId="7" pageNumber="8">Holotype</typeStatus>
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.
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</emphasis>
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Sex
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<heading box="[399,506,1684,1707]" centered="true" fontSize="10" level="5" pageId="7" pageNumber="8" reason="4">unknown;</heading>
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<specimenCode box="[551,777,1684,1707]" pageId="7" pageNumber="8">FMNHINS-3966034</specimenCode>
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;
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<location LSID="urn:lsid:plazi:treatment:03E7BF21FF80C80AFF39F9C1FA37FD7E:8E9158ECFF80C805FF78F974FC81F916" box="[159,780,1717,1740]" country="Myanmar" name="Hukawng Valley, Kachin State of northern" pageId="7" pageNumber="8">Hukawng Valley, Kachin State of northern</location>
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<collectingCountry box="[159,263,1749,1772]" name="Myanmar" pageId="7" pageNumber="8">Myanmar</collectingCountry>
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(
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<collectionCode box="[279,362,1749,1772]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" pageId="7" pageNumber="8">FMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection box="[159,432,1797,1820]" pageId="7" pageNumber="8" type="diagnosis">
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<paragraph blockId="7.[159,432,1797,1820]" box="[159,432,1797,1820]" pageId="7" pageNumber="8">
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<emphasis bold="true" box="[159,276,1797,1820]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
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As for genus.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
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<paragraph blockId="7.[159,781,1845,1901]" lastBlockId="7.[844,1467,1044,1901]" pageId="7" pageNumber="8">
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<emphasis bold="true" box="[159,298,1845,1868]" pageId="7" pageNumber="8">Description.</emphasis>
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Body (
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<figureCitation box="[391,515,1845,1869]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="7" pageNumber="8">Figs 1A–D</figureCitation>
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,
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<figureCitation box="[535,645,1845,1868]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="7" pageNumber="8">2A, B, D</figureCitation>
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,
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<figureCitation box="[665,771,1845,1868]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="7" pageNumber="8">3A, B, E</figureCitation>
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) stout, length 1.71 mm, region from anterior prothorax to posterior third of elytra approximately parallel-sided; pigmentation light brown with diffuse pattern of darker areas on most convex structures (pronotal and elytral tubercles and ridges); body surface coarsely sculptured and matte. Entire body covered with thin crust of presumably mud particles and glandular secretion, best visible in concave areas on pronotum and elytra and on ventral body side.
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</paragraph>
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<paragraph blockId="7.[844,1467,1044,1901]" pageId="7" pageNumber="8">
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Head (
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<figureCitation box="[950,1074,1300,1324]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="7" pageNumber="8">Figs 1A–D</figureCitation>
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;
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<figureCitation box="[1096,1161,1300,1324]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="7" pageNumber="8">2A–D</figureCitation>
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;
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<figureCitation box="[1182,1282,1300,1324]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="7" pageNumber="8">3A–C, E</figureCitation>
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) strongly transverse, length 0.21 mm, width 0.63 mm; with very large and strongly convex compound eyes, in dorsal view occupying nearly entire lateral margin of head; tempora vestigial; vertex and frons convex medially and impressed sub-laterally, with paired oblique dorsal protuberances or ridges.
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</paragraph>
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<paragraph blockId="7.[844,1467,1044,1901]" pageId="7" pageNumber="8">
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Antennae short, length 0.45 mm; scapus (
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<figureCitation box="[1325,1411,1525,1548]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="7" pageNumber="8">Figs 2C</figureCitation>
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,
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<figureCitation box="[1426,1458,1525,1548]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="7" pageNumber="8">3C</figureCitation>
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) large, elongate; pedicellus (
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<figureCitation box="[1148,1234,1557,1580]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="7" pageNumber="8">Figs 2C</figureCitation>
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,
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<figureCitation box="[1250,1281,1557,1580]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="7" pageNumber="8">3C</figureCitation>
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) sub-cylindrical, about as broad as long; antennomere 3 (
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<figureCitation box="[1316,1407,1589,1612]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="7" pageNumber="8">Figs 2C</figureCitation>
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,
|
||
<figureCitation box="[1426,1458,1589,1612]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="7" pageNumber="8">3C</figureCitation>
|
||
) about as long as broad; antennomeres 4–6 (
|
||
<figureCitation box="[1325,1411,1621,1644]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="7" pageNumber="8">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation box="[1426,1458,1621,1644]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="7" pageNumber="8">3C</figureCitation>
|
||
) each slightly broader than 3 and slightly transverse; antennal club (
|
||
<figureCitation box="[1011,1098,1686,1709]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="7" pageNumber="8">Figs 2C</figureCitation>
|
||
,
|
||
<figureCitation box="[1115,1146,1686,1709]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="7" pageNumber="8">3C</figureCitation>
|
||
) slightly flattened dorsoventrally, slightly shorter than all proximal antennomeres combined, with anterior margin nearly straight and posterior margin slightly rounded; oblique apical surface with densely arranged short setae.
|
||
</paragraph>
|
||
<paragraph blockId="7.[844,1467,1044,1901]" lastBlockId="8.[159,782,189,1901]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
|
||
Pronotum (
|
||
<figureCitation box="[996,1118,1846,1869]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="7" pageNumber="8">Fig. 2A, C</figureCitation>
|
||
) about 0.7 as broad as long, length 0.55 mm, width 0.79 mm; anterior margin sub- trapezoidal and forming short blunt projection overhanging posterodorsal region of head; anterior margin between anterior projection and anterior corner slightly concave; lateral margins irregularly undulate and convergent posterad; posterior margin shallowly and broadly emarginate. Antennal pocket (
|
||
<figureCitation box="[618,715,351,374]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Figs 1D</figureCitation>
|
||
,
|
||
<figureCitation box="[741,772,351,374]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="8" pageNumber="9">3C</figureCitation>
|
||
) behind each eye open anteroventrally, deep, large and sharply demarcated, with its concave surface smooth. Pronotal disc with convex median area bearing longitudinal rows of several tubercles of various diameters and heights, elongate median region impressed and relatively smooth; impressed lateral portions of pronotal disc with sub-median transverse row of diffuse and weakly elevated tubercles. Setae on pronotum indiscernible.
|
||
</paragraph>
|
||
<paragraph blockId="8.[159,782,189,1901]" pageId="8" pageNumber="9">
|
||
Elytra (
|
||
<figureCitation box="[274,400,643,667]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Figs 1A–D</figureCitation>
|
||
,
|
||
<figureCitation box="[422,535,643,666]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="8" pageNumber="9">2A, B, D</figureCitation>
|
||
,
|
||
<figureCitation box="[557,666,643,666]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="8" pageNumber="9">3A, B, E</figureCitation>
|
||
) broadest shortly behind humeri; length 0.95 mm, width 0.91 mm. In lateral view (
|
||
<figureCitation box="[350,441,708,731]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="8" pageNumber="9">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation box="[461,492,708,731]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="8" pageNumber="9">3B</figureCitation>
|
||
) dorsal surface of elytra nearly parallel to longitudinal body axis in anterior half, in posterior half strongly rounded, thus elytral apex not visible in dorsal view. Humeral angles distinctly projecting anterad, anterior margin of each elytron thus slightly concave. Elytral tubercles of various heights and diameters, longest tubercles situated on posterior elytral surface and directed posterad; most tubercles bear one apical bristle (best visible on posterior half of elytra). In lateral view, sub-median ‘tooth’ of elytral edge obtuseangled, concavity behind it rounded.
|
||
</paragraph>
|
||
<paragraph blockId="8.[159,782,189,1901]" lastBlockId="8.[845,1467,189,436]" pageId="8" pageNumber="9">
|
||
Three pairs of legs similar. Procoxae (
|
||
<figureCitation box="[637,725,1065,1088]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
|
||
) triangular, anterior portion separated by prosternal process, posteromesal edges nearly adjacent at midline. Articulation of procoxae with prosternum situated laterally, with notch in nearly straight anterior coxal margin and small sternal condyle. Mesocoxae (
|
||
<figureCitation box="[680,772,1228,1251]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
|
||
) widely separated and obliquely inserted, flat and distinctly smaller than procoxae; anterior and posterior margins rounded, posterior edge slightly overlapping with trochanteral base. Metacoxae (
|
||
<figureCitation box="[563,650,1358,1381]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
|
||
) flat, more transverse but otherwise similar to mesocoxae, separated by posteromedian katepisternal excavation, which is narrower than each metacoxa. Protrochanters (
|
||
<figureCitation box="[643,729,1455,1478]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
|
||
) triangular and about half as long as procoxae. Mesotrochanters (
|
||
<figureCitation box="[392,513,1520,1543]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
|
||
) slightly smaller. Metatrochanters (
|
||
<figureCitation box="[350,435,1553,1576]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
|
||
) about as large as protrochanters. Profemora (
|
||
<figureCitation box="[346,435,1585,1608]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
|
||
) about twice as long as procoxae; their anterior margins slightly curved and ventral margins slightly concave. Meso- and metafemora (
|
||
<figureCitation captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
|
||
) slightly shorter than profemora, fitting into elytral excavations (
|
||
<figureCitation box="[303,392,1716,1739]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Figs 1D</figureCitation>
|
||
,
|
||
<figureCitation box="[408,441,1716,1739]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="8" pageNumber="9">2D</figureCitation>
|
||
). Protibia (
|
||
<figureCitation box="[569,654,1716,1739]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
|
||
) sub-cylindrical, shorter than profemora, with fine spines along anterior and posterior margins, lacking distal tibial spur but with posterolaterally directed robust spine at apex. Meso- and metatibiae (
|
||
<figureCitation box="[435,526,1846,1869]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
|
||
) similar to protibiae. Protarsus (
|
||
<figureCitation box="[278,364,1878,1901]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
|
||
) including claw slightly shorter than protibia; showing indistinct subdivision into four incompletely delimited subunits and with spines along anterior and posterior edges; single curved claw about as long as apical protarsal subunit. Meso- and metatarsi (
|
||
<figureCitation box="[1371,1458,285,308]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
|
||
) gradually tapering distad, not showing discernible subdivision.
|
||
</paragraph>
|
||
<paragraph blockId="8.[845,1467,189,436]" pageId="8" pageNumber="9">
|
||
Abdomen (
|
||
<figureCitation box="[995,1118,381,404]" captionStart="Figure 1" captionStartId="3.[159,226,1858,1879]" captionTargetBox="[175,1455,196,1830]" captionTargetId="figure@3.[159,1467,188,1840]" captionTargetPageId="3" captionText="Figure 1. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, A, B, macrophotographs; C, artistic reconstruction; and D, schematic morphological details observable under light microscope. A, C, dorsal habitus; B, D, ventral habitus. Abbreviations: anf, antennal furrow; anp, antennal pocket; ce, compound eye; cx1–3, coxa; elex, elytral excavation; fm1–3, femur; hyp, hypomeron; kes, katepisternum; lb, labrum; mnt, mentum; ppr, prosternal process; r1–5, elytral rows of tubercles; st, abdominal sternite 2; tb1–3, tibia; tch1–3, trochanter; tr1–3, tarsus; v2, mesoventrite; v3, metaventrite." figureDoi="http://doi.org/10.5281/zenodo.3763696" httpUri="https://zenodo.org/record/3763696/files/figure.png" pageId="8" pageNumber="9">Figs 1B, D</figureCitation>
|
||
,
|
||
<figureCitation box="[1135,1168,381,404]" captionStart="Figure 2" captionStartId="4.[159,226,1321,1342]" captionTargetBox="[159,1454,189,1304]" captionTargetId="figure@4.[159,1467,188,1304]" captionTargetPageId="4" captionText="Figure 2. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, surface reconstructions. A, dorsal habitus; B, lateral habitus; C, right antenna and eye in ventral view; D, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; an3–6, antennomere 3–6; ce, compound eye; elex, elytral excavation;?mxp, presumable maxillary palp; pd, pedicel; sc, scape; v2, mesoventrite." figureDoi="http://doi.org/10.5281/zenodo.3763698" httpUri="https://zenodo.org/record/3763698/files/figure.png" pageId="8" pageNumber="9">2D</figureCitation>
|
||
,
|
||
<figureCitation box="[1185,1214,381,404]" captionStart="Figure 3" captionStartId="5.[159,226,1539,1560]" captionTargetBox="[159,1467,188,1517]" captionTargetId="figure@5.[159,1467,188,1517]" captionTargetPageId="5" captionText="Figure 3. †Lepiceratus ankylosaurus gen. and sp. nov., holotype, synchrotron microtomography, volume renderings. A, dorsal habitus; B, submedian sagittal virtual section; C, right antenna, eye and anterolateral portion of prothorax in ventral view; D, distal surface of right antennal club; E, ventral habitus. Abbreviations: anc, antennal club; anf, antennal furrow; anp, antennal pocket; an3–6, antennomere 3–6; ce, compound eye; els, elytral suture; epi, elytral epipleura; pd, pedicel; ppr, prosternal process; sc, scape; scs, scutellar shield; wg, wings." figureDoi="http://doi.org/10.5281/zenodo.3763700" httpUri="https://zenodo.org/record/3763700/files/figure.png" pageId="8" pageNumber="9">3E</figureCitation>
|
||
) covered with punctures, setae indiscernible.
|
||
</paragraph>
|
||
<paragraph blockId="8.[845,1467,461,580]" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[845,1085,461,484]" pageId="8" pageNumber="9">Derivation of name.</emphasis>
|
||
After ankylosaurs, the dinosaurs with longitudinal rows of armour plates resembling the rows of tubercles in the new species (noun in apposition).
|
||
</paragraph>
|
||
<paragraph blockId="8.[845,1433,605,629]" box="[845,1433,605,629]" pageId="8" pageNumber="9">
|
||
<emphasis bold="true" box="[845,1078,606,629]" pageId="8" pageNumber="9">Stratigraphic range.</emphasis>
|
||
Late Albian–early Cenomanian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="9" type="distribution">
|
||
<paragraph blockId="8.[845,1097,653,709]" box="[845,1097,653,676]" pageId="8" pageNumber="9">
|
||
<heading bold="true" box="[845,1097,653,676]" centered="true" fontSize="10" level="4" pageId="8" pageNumber="9" reason="4">
|
||
<emphasis bold="true" box="[845,1097,653,676]" pageId="8" pageNumber="9">Geographical range.</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="8.[845,1097,653,709]" box="[845,1036,686,709]" pageId="8" pageNumber="9">Hukawng Valley.</paragraph>
|
||
<paragraph blockId="8.[1123,1219,653,676]" box="[1123,1219,653,676]" pageId="8" pageNumber="9">
|
||
<heading box="[1123,1219,653,676]" centered="true" fontSize="10" level="5" pageId="8" pageNumber="9" reason="4">Northern</heading>
|
||
</paragraph>
|
||
<paragraph blockId="8.[1246,1357,653,676]" box="[1246,1357,653,676]" pageId="8" pageNumber="9">
|
||
<heading box="[1246,1357,653,676]" centered="true" fontSize="10" level="5" pageId="8" pageNumber="9" reason="4">Myanmar,</heading>
|
||
</paragraph>
|
||
<paragraph blockId="8.[1383,1466,653,676]" box="[1383,1466,653,676]" pageId="8" pageNumber="9">
|
||
<heading box="[1383,1466,653,676]" centered="true" fontSize="10" level="5" pageId="8" pageNumber="9" reason="4">Kachin,</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |