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<document id="8FCEB4C073B23D6E8A1554E23F48ECEB" ID-DOI="10.22073/pja.v13i4.85812" ID-ISSN="2251-8169" ID-Zenodo-Dep="14668778" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737003594861" checkinUser="felipe" docAuthor="Durucan, Furkan" docDate="2024" docId="03DDE642FFC3EF1FFB3A9603D82F22A2" docLanguage="en" docName="PersJourAcarology.13.4.771-796.pdf" docOrigin="Persian Journal of Acarology 13" docStyle="DocumentStyle:B3904D664642C9C0DBE4CD09E74105CD.9:PersJourAcarology.2017-.journal_article" docStyleId="B3904D664642C9C0DBE4CD09E74105CD" docStyleName="PersJourAcarology.2017-.journal_article" docStyleVersion="9" docTitle="Copidognathus cribosoma" docType="treatment" docVersion="2" lastPageNumber="774" masterDocId="FFE49E3AFFC1EF1CFA789436DA4D2363" masterDocTitle="Three new records and some juveniles of the genus Copidognathus (Acari, Halacaridae) from Türkiye" masterLastPageNumber="796" masterPageNumber="771" pageNumber="773" updateTime="1737010000186" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-2.0">
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<mods:title id="5964110E3F6D3A20E5A2AFF11B581C95">Three new records and some juveniles of the genus Copidognathus (Acari, Halacaridae) from Türkiye</mods:title>
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<mods:namePart id="B4DC6878422DD9D7B2983A3F74329BD9">Durucan, Furkan</mods:namePart>
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<mods:title id="B61D35AC6C3CBD6EF357E8AE32A1BFD1">Persian Journal of Acarology</mods:title>
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<mods:date id="05C4AF952E10772FC99DD6A3E8B49D99">2024</mods:date>
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<treatment id="03DDE642FFC3EF1FFB3A9603D82F22A2" LSID="urn:lsid:plazi:treatment:03DDE642FFC3EF1FFB3A9603D82F22A2" httpUri="http://treatment.plazi.org/id/03DDE642FFC3EF1FFB3A9603D82F22A2" lastPageId="3" lastPageNumber="774" pageId="2" pageNumber="773">
<subSubSection id="C36E04DFFFC3EF1EFB3A9603DEBF2131" box="[322,1266,565,594]" pageId="2" pageNumber="773" type="nomenclature">
<paragraph id="8BCB5754FFC3EF1EFB3A9603DEBF2131" blockId="2.[322,1266,565,594]" box="[322,1266,565,594]" pageId="2" pageNumber="773">
<heading id="D083E038FFC3EF1EFB3A9603DEBF2131" bold="true" box="[322,1266,565,594]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="773" reason="2">
<emphasis id="B9008B46FFC3EF1EFB3A9603DEBF2131" bold="true" box="[322,1266,565,594]" pageId="2" pageNumber="773">
<taxonomicName id="4C742CD7FFC3EF1EFB3A9603D9162131" authority="(Police, 1909)" baseAuthorityName="Police" baseAuthorityYear="1909" box="[322,859,565,594]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="773" phylum="Arthropoda" rank="species" species="cribosoma">
<emphasis id="B9008B46FFC3EF1EFB3A9603D8D12131" bold="true" box="[322,668,565,594]" italics="true" pageId="2" pageNumber="773">Copidognathus cribosoma</emphasis>
(
<bibRefCitation id="EFE52AA5FFC3EF1EF8D49603D91F2131" author="Police, G." box="[684,850,565,594]" pageId="2" pageNumber="773" pagination="409 - 443" refId="ref9433" refString="Police, G. (1909) Alcune nuove specie di Halacaridae del Golfe di Napoli. ArchIvIo ZoologIco, NapolI, 3: 409 - 443." type="journal article" year="1909">Police, 1909</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="134F4BD1FFC3EF1EF9159603DE4B2131" box="[877,1030,565,594]" captionStart="Figure 2" captionStartId="12.[151,226,1924,1948]" captionTargetBox="[220,1378,250,1883]" captionTargetId="figure-239@12.[206,1382,229,1892]" captionTargetPageId="12" captionText="Figure 2. Copidognathus cribosoma (Police, 1909) (adults) A. Idiosoma, dorsal view (female); B. Idiosoma, ventral view (female); C. Ocular plate; D. Epimera II; E. Ovipositor; F. Genitoanal plate (male); G. Spermatopositor; H. Gnathosoma, dorsal view (female); I. Leg I, medial view (female); J. Leg II, medial view (female); K. Leg III, medial view (female); L. Leg IV, medial view (female); M. Tarsus I, dorsomedial view (female); N. Tibia I, medial view (female); O. Tibia II, medial view (female). Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14668784" httpUri="https://zenodo.org/record/14668784/files/figure.png" pageId="2" pageNumber="773">Figs. 2AO</figureCitation>
,
<figureCitation id="134F4BD1FFC3EF1EFE6E9603DE392131" box="[1046,1140,565,594]" captionStart="Figure 11" captionStartId="21.[151,226,1921,1945]" captionTargetBox="[215,1377,237,1881]" captionTargetId="figure-280@21.[206,1381,229,1890]" captionTargetPageId="21" captionText="Figure 11. Line art illustrations of juveniles and larva of Copidognathus species (AC. C. brachyostomus, PN; DG. C. cribosoma, PN; HK. C. humerosus, PN; LM, C. humerosus, L) A. Idiosoma, dorsal view; B. Idiosoma, ventral view; C. Gnathosoma, ventral view; D. Idiosoma, dorsal view; E. Idiosoma, ventral view; F. Gnathosoma, ventral view; G. Leg I, medial view; H. Idiosoma, dorsal view; I. Idiosoma, ventral view; J. Gnathosoma, ventral view; K. Tarsus I, medial view; L. Idiosoma, dorsal view; M. Leg III, medial view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14668806" httpUri="https://zenodo.org/record/14668806/files/figure.png" pageId="2" pageNumber="773">11DG</figureCitation>
,
<tableCitation id="C6F662EFFFC3EF1EFEFC9603DEAA2131" box="[1156,1255,565,594]" captionStart="Table 1" captionStartId="23.[151,216,236,260]" captionText="Table 1. Measurements for CopIdognathus crIbosoma specimens." pageId="2" pageNumber="773">Table 1</tableCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36E04DFFFC3EF1FFAFC9648D82F22A2" lastPageId="3" lastPageNumber="774" pageId="2" pageNumber="773" type="description">
<paragraph id="8BCB5754FFC3EF1EFAFC9648D96221F8" blockId="2.[132,815,638,667]" box="[132,815,638,667]" pageId="2" pageNumber="773">
<emphasis id="B9008B46FFC3EF1EFAFC9648DB1721F8" bold="true" box="[132,346,638,667]" pageId="2" pageNumber="773">Redescription </emphasis>
Measurements presented in
<tableCitation id="C6F662EFFFC3EF1EF8B09648D96A21F8" box="[712,807,638,667]" captionStart="Table 1" captionStartId="23.[151,216,236,260]" captionText="Table 1. Measurements for CopIdognathus crIbosoma specimens." pageId="2" pageNumber="773">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BCB5754FFC3EF1EFAEF96F1DBF42186" blockId="2.[151,1437,711,2065]" box="[151,441,711,741]" pageId="2" pageNumber="773">
<emphasis id="B9008B46FFC3EF1EFAEF96F1DBF42186" box="[151,441,711,741]" italics="true" pageId="2" pageNumber="773">Morphology and notes</emphasis>
</paragraph>
<paragraph id="8BCB5754FFC3EF1EFAA896DBDB2B27A0" blockId="2.[151,1437,711,2065]" pageId="2" pageNumber="773">
<emphasis id="B9008B46FFC3EF1EFAA896DBDB002069" bold="true" box="[208,333,749,778]" italics="true" pageId="2" pageNumber="773">Female </emphasis>
All dorsal and ventral plates divided by narrow bands of densely striated cuticle and the plates with uniformly rosette pores numerous canaliculi (
<figureCitation id="134F4BD1FFC3EF1EF9E99727DE69204D" box="[913,1060,785,814]" captionStart="Figure 2" captionStartId="12.[151,226,1924,1948]" captionTargetBox="[220,1378,250,1883]" captionTargetId="figure-239@12.[206,1382,229,1892]" captionTargetPageId="12" captionText="Figure 2. Copidognathus cribosoma (Police, 1909) (adults) A. Idiosoma, dorsal view (female); B. Idiosoma, ventral view (female); C. Ocular plate; D. Epimera II; E. Ovipositor; F. Genitoanal plate (male); G. Spermatopositor; H. Gnathosoma, dorsal view (female); I. Leg I, medial view (female); J. Leg II, medial view (female); K. Leg III, medial view (female); L. Leg IV, medial view (female); M. Tarsus I, dorsomedial view (female); N. Tibia I, medial view (female); O. Tibia II, medial view (female). Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14668784" httpUri="https://zenodo.org/record/14668784/files/figure.png" pageId="2" pageNumber="773">Figs. 2A, B</figureCitation>
). AD longer than wide (1.10); anterior margin ovate and with small frontal lamella. Dorsal setae small. Pair of ds-1 on AD posterior to pair of glp-1; ds-
<quantity id="4C8CFAB1FFC3EF1EFBF0976DDBFA201B" box="[392,439,859,888]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="773" unit="in" value="2.0">2 in</quantity>
anteromedial margin of OC. Posterior end of OC pointed and almost extending to level of leg III. Gland pores (glp-2) small and lateral to OC. Pore canaliculus posterior to gland pore (
<figureCitation id="134F4BD1FFC3EF1EFA9B9792DB0620A2" box="[227,331,932,961]" captionStart="Figure 2" captionStartId="12.[151,226,1924,1948]" captionTargetBox="[220,1378,250,1883]" captionTargetId="figure-239@12.[206,1382,229,1892]" captionTargetPageId="12" captionText="Figure 2. Copidognathus cribosoma (Police, 1909) (adults) A. Idiosoma, dorsal view (female); B. Idiosoma, ventral view (female); C. Ocular plate; D. Epimera II; E. Ovipositor; F. Genitoanal plate (male); G. Spermatopositor; H. Gnathosoma, dorsal view (female); I. Leg I, medial view (female); J. Leg II, medial view (female); K. Leg III, medial view (female); L. Leg IV, medial view (female); M. Tarsus I, dorsomedial view (female); N. Tibia I, medial view (female); O. Tibia II, medial view (female). Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14668784" httpUri="https://zenodo.org/record/14668784/files/figure.png" pageId="2" pageNumber="773">Fig. 2C</figureCitation>
). PD wide, anteriorly truncate, posteriorly ovate. Pair of ds-3 just below the anterior margin of PD, ds-4 and ds-5 at 0.31 and 0.64 of PD's total length, respectively. Pair of ds-6 on anal cone. AE with three pairs of ventral setae and a pair of epimeral pores. Epimeral pores 34 wide. Epimeral processes I and II are small and lamellar (
<figureCitation id="134F4BD1FFC3EF1EF9329025D9F82753" box="[842,949,1043,1072]" captionStart="Figure 2" captionStartId="12.[151,226,1924,1948]" captionTargetBox="[220,1378,250,1883]" captionTargetId="figure-239@12.[206,1382,229,1892]" captionTargetPageId="12" captionText="Figure 2. Copidognathus cribosoma (Police, 1909) (adults) A. Idiosoma, dorsal view (female); B. Idiosoma, ventral view (female); C. Ocular plate; D. Epimera II; E. Ovipositor; F. Genitoanal plate (male); G. Spermatopositor; H. Gnathosoma, dorsal view (female); I. Leg I, medial view (female); J. Leg II, medial view (female); K. Leg III, medial view (female); L. Leg IV, medial view (female); M. Tarsus I, dorsomedial view (female); N. Tibia I, medial view (female); O. Tibia II, medial view (female). Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14668784" httpUri="https://zenodo.org/record/14668784/files/figure.png" pageId="2" pageNumber="773">Fig. 2D</figureCitation>
). PE with one dorsolateral and three ventral setae. GA anterior part truncate with uniformly arranged with canaliculi. Ovipositor extending beyond GO but not reaching pgs-1. Three pairs of pgs arranged around GO. GO 54 long, 33 wide. Distance between edges of GA and GO 90, equalling 0.55 of GO length. Genital sclerites with a pair of sgs (
<figureCitation id="134F4BD1FFC3EF1EFA8C9090DB1527A0" box="[244,344,1190,1219]" captionStart="Figure 2" captionStartId="12.[151,226,1924,1948]" captionTargetBox="[220,1378,250,1883]" captionTargetId="figure-239@12.[206,1382,229,1892]" captionTargetPageId="12" captionText="Figure 2. Copidognathus cribosoma (Police, 1909) (adults) A. Idiosoma, dorsal view (female); B. Idiosoma, ventral view (female); C. Ocular plate; D. Epimera II; E. Ovipositor; F. Genitoanal plate (male); G. Spermatopositor; H. Gnathosoma, dorsal view (female); I. Leg I, medial view (female); J. Leg II, medial view (female); K. Leg III, medial view (female); L. Leg IV, medial view (female); M. Tarsus I, dorsomedial view (female); N. Tibia I, medial view (female); O. Tibia II, medial view (female). Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14668784" httpUri="https://zenodo.org/record/14668784/files/figure.png" pageId="2" pageNumber="773">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="8BCB5754FFC3EF1EFAA890FDDF772652" blockId="2.[151,1437,711,2065]" pageId="2" pageNumber="773">
Gnathosoma (
<figureCitation id="134F4BD1FFC3EF1EFBFF90FDDBA2278B" box="[391,495,1227,1256]" captionStart="Figure 2" captionStartId="12.[151,226,1924,1948]" captionTargetBox="[220,1378,250,1883]" captionTargetId="figure-239@12.[206,1382,229,1892]" captionTargetPageId="12" captionText="Figure 2. Copidognathus cribosoma (Police, 1909) (adults) A. Idiosoma, dorsal view (female); B. Idiosoma, ventral view (female); C. Ocular plate; D. Epimera II; E. Ovipositor; F. Genitoanal plate (male); G. Spermatopositor; H. Gnathosoma, dorsal view (female); I. Leg I, medial view (female); J. Leg II, medial view (female); K. Leg III, medial view (female); L. Leg IV, medial view (female); M. Tarsus I, dorsomedial view (female); N. Tibia I, medial view (female); O. Tibia II, medial view (female). Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14668784" httpUri="https://zenodo.org/record/14668784/files/figure.png" pageId="2" pageNumber="773">Fig. 2H</figureCitation>
) 1.35 times longer than wide. Rostrum triangular, reaching end of P2. Tectum truncate, palps are slender and consist of four segments. P1 to P4 lengths 12, 36, 9, 34 respectively. P1 and P3 without setae, P2 with single seta, P4 with three setae at basal whorl.
</paragraph>
<paragraph id="8BCB5754FFC3EF1EFAA8910FDBF02550" blockId="2.[151,1437,711,2065]" pageId="2" pageNumber="773">
All legs (
<figureCitation id="134F4BD1FFC3EF1EFB31910FDBEF2635" box="[329,418,1337,1366]" captionStart="Figure 2" captionStartId="12.[151,226,1924,1948]" captionTargetBox="[220,1378,250,1883]" captionTargetId="figure-239@12.[206,1382,229,1892]" captionTargetPageId="12" captionText="Figure 2. Copidognathus cribosoma (Police, 1909) (adults) A. Idiosoma, dorsal view (female); B. Idiosoma, ventral view (female); C. Ocular plate; D. Epimera II; E. Ovipositor; F. Genitoanal plate (male); G. Spermatopositor; H. Gnathosoma, dorsal view (female); I. Leg I, medial view (female); J. Leg II, medial view (female); K. Leg III, medial view (female); L. Leg IV, medial view (female); M. Tarsus I, dorsomedial view (female); N. Tibia I, medial view (female); O. Tibia II, medial view (female). Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14668784" httpUri="https://zenodo.org/record/14668784/files/figure.png" pageId="2" pageNumber="773">Figs. 2</figureCitation>
I-O) slender and long but shorter than idiosoma. On the surface of telofemora, genua and tibiae I and II foveate with rosette pores. Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 2, 5, 4, 7(2), 6 5; leg II: 1, 2, 5, 4, 6(2), 3; leg III: 1, 2, 3, 3, 5(1), 4; leg IV: 0, 2, 3, 3, 5(1), 3. Tibiae I to IV with 2, 2, 1, 1 bipectinate setae. Tarsi I to IV with 3, 3, 4, 3 dorsal setae and 3, 0, 0, 0 ventral setae. All legs with two lateral claws with accessory process but large pectines (oil immersion) and a small bidentate median claw.
</paragraph>
<paragraph id="8BCB5754FFC3EF1EFAA8920DDE7B25A5" blockId="2.[151,1437,711,2065]" pageId="2" pageNumber="773">
<emphasis id="B9008B46FFC3EF1EFAA8920DDB66253B" bold="true" box="[208,299,1595,1624]" pageId="2" pageNumber="773">
<emphasis id="B9008B46FFC3EF1EFAA8920DDB59253B" bold="true" box="[208,276,1595,1624]" italics="true" pageId="2" pageNumber="773">Male</emphasis>
</emphasis>
Male is similar to female except for the genital plate. GO surrounded by 5557 pgs (
<figureCitation id="134F4BD1FFC3EF1EFF10920DDAF0251F" captionStart="Figure 2" captionStartId="12.[151,226,1924,1948]" captionTargetBox="[220,1378,250,1883]" captionTargetId="figure-239@12.[206,1382,229,1892]" captionTargetPageId="12" captionText="Figure 2. Copidognathus cribosoma (Police, 1909) (adults) A. Idiosoma, dorsal view (female); B. Idiosoma, ventral view (female); C. Ocular plate; D. Epimera II; E. Ovipositor; F. Genitoanal plate (male); G. Spermatopositor; H. Gnathosoma, dorsal view (female); I. Leg I, medial view (female); J. Leg II, medial view (female); K. Leg III, medial view (female); L. Leg IV, medial view (female); M. Tarsus I, dorsomedial view (female); N. Tibia I, medial view (female); O. Tibia II, medial view (female). Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14668784" httpUri="https://zenodo.org/record/14668784/files/figure.png" pageId="2" pageNumber="773">Fig. 2F</figureCitation>
). Spermatopositor large (7885), but not extending anterior margin of GA (
<figureCitation id="134F4BD1FFC3EF1EFECE9269DF6F251F" box="[1206,1314,1631,1660]" captionStart="Figure 2" captionStartId="12.[151,226,1924,1948]" captionTargetBox="[220,1378,250,1883]" captionTargetId="figure-239@12.[206,1382,229,1892]" captionTargetPageId="12" captionText="Figure 2. Copidognathus cribosoma (Police, 1909) (adults) A. Idiosoma, dorsal view (female); B. Idiosoma, ventral view (female); C. Ocular plate; D. Epimera II; E. Ovipositor; F. Genitoanal plate (male); G. Spermatopositor; H. Gnathosoma, dorsal view (female); I. Leg I, medial view (female); J. Leg II, medial view (female); K. Leg III, medial view (female); L. Leg IV, medial view (female); M. Tarsus I, dorsomedial view (female); N. Tibia I, medial view (female); O. Tibia II, medial view (female). Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14668784" httpUri="https://zenodo.org/record/14668784/files/figure.png" pageId="2" pageNumber="773">Fig. 2G</figureCitation>
). Genital sclerites with four pairs of sgs; anterior pairs hair-like, posterior ones spur-like. GO 40 long, 17 wide. Distance between edges of GA and GO 90, equalling 2.25 of GO length.
</paragraph>
<paragraph id="8BCB5754FFC3EF1EFAA892F8DEB12B72" blockId="2.[151,1437,711,2065]" pageId="2" pageNumber="773">
<emphasis id="B9008B46FFC3EF1EFAA892F8DBC12588" bold="true" box="[208,396,1742,1771]" italics="true" pageId="2" pageNumber="773">Protonymph </emphasis>
AD with three porose areolae and glp-1 on lateral margin of the plate. OC longer than wide. PD anteriorly ovate. Ds-1 on anterior edge of porose areaolae on AD. Pairs of ds-2 and ds- 3 on striated integument. Ds-4 and ds-5 on PD. Ds-6 on anal cone (
<figureCitation id="134F4BD1FFC3EF1EFE74932EDECD2456" box="[1036,1152,1816,1845]" captionStart="Figure 11" captionStartId="21.[151,226,1921,1945]" captionTargetBox="[215,1377,237,1881]" captionTargetId="figure-280@21.[206,1381,229,1890]" captionTargetPageId="21" captionText="Figure 11. Line art illustrations of juveniles and larva of Copidognathus species (AC. C. brachyostomus, PN; DG. C. cribosoma, PN; HK. C. humerosus, PN; LM, C. humerosus, L) A. Idiosoma, dorsal view; B. Idiosoma, ventral view; C. Gnathosoma, ventral view; D. Idiosoma, dorsal view; E. Idiosoma, ventral view; F. Gnathosoma, ventral view; G. Leg I, medial view; H. Idiosoma, dorsal view; I. Idiosoma, ventral view; J. Gnathosoma, ventral view; K. Tarsus I, medial view; L. Idiosoma, dorsal view; M. Leg III, medial view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14668806" httpUri="https://zenodo.org/record/14668806/files/figure.png" pageId="2" pageNumber="773">Fig. 11D</figureCitation>
). AE with three setae and pair of epimeral pore. GP large almost quadrangular with a pair of genital acetabula and pair of setae (
<figureCitation id="134F4BD1FFC3EF1EFA919357DB11241D" box="[233,348,1889,1918]" captionStart="Figure 11" captionStartId="21.[151,226,1921,1945]" captionTargetBox="[215,1377,237,1881]" captionTargetId="figure-280@21.[206,1381,229,1890]" captionTargetPageId="21" captionText="Figure 11. Line art illustrations of juveniles and larva of Copidognathus species (AC. C. brachyostomus, PN; DG. C. cribosoma, PN; HK. C. humerosus, PN; LM, C. humerosus, L) A. Idiosoma, dorsal view; B. Idiosoma, ventral view; C. Gnathosoma, ventral view; D. Idiosoma, dorsal view; E. Idiosoma, ventral view; F. Gnathosoma, ventral view; G. Leg I, medial view; H. Idiosoma, dorsal view; I. Idiosoma, ventral view; J. Gnathosoma, ventral view; K. Tarsus I, medial view; L. Idiosoma, dorsal view; M. Leg III, medial view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14668806" httpUri="https://zenodo.org/record/14668806/files/figure.png" pageId="2" pageNumber="773">Fig. 11E</figureCitation>
). Gnathosoma longer than wide (
<figureCitation id="134F4BD1FFC3EF1EF9709357D934241D" box="[776,889,1889,1918]" captionStart="Figure 11" captionStartId="21.[151,226,1921,1945]" captionTargetBox="[215,1377,237,1881]" captionTargetId="figure-280@21.[206,1381,229,1890]" captionTargetPageId="21" captionText="Figure 11. Line art illustrations of juveniles and larva of Copidognathus species (AC. C. brachyostomus, PN; DG. C. cribosoma, PN; HK. C. humerosus, PN; LM, C. humerosus, L) A. Idiosoma, dorsal view; B. Idiosoma, ventral view; C. Gnathosoma, ventral view; D. Idiosoma, dorsal view; E. Idiosoma, ventral view; F. Gnathosoma, ventral view; G. Leg I, medial view; H. Idiosoma, dorsal view; I. Idiosoma, ventral view; J. Gnathosoma, ventral view; K. Tarsus I, medial view; L. Idiosoma, dorsal view; M. Leg III, medial view. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14668806" httpUri="https://zenodo.org/record/14668806/files/figure.png" pageId="2" pageNumber="773">Fig. 11F</figureCitation>
). Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 2, 3, 4, 5(1), 6; leg II: 1, 2, 3, 4, 4(1), 3; leg III: 1, 2, 2, 3, 5(1), 4; leg IV: 0, 1+2 (basifemur + telofemur) 3, 5(1), 3. Each of tibiae I to IV with one bipectinate setae. All legs with two lateral claws with accessory process but large pectines (oil immersion) and a small bidentate median claw.
</paragraph>
<paragraph id="8BCB5754FFC2EF1FFAEF94DADB4A226A" blockId="3.[151,1436,236,449]" box="[151,263,236,265]" pageId="3" pageNumber="774">
<emphasis id="B9008B46FFC2EF1FFAEF94DADB4A226A" box="[151,263,236,265]" italics="true" pageId="3" pageNumber="774">Remarks</emphasis>
</paragraph>
<paragraph id="8BCB5754FFC2EF1FFAA89527D82F22A2" blockId="3.[151,1436,236,449]" pageId="3" pageNumber="774">
This species was originally described by
<bibRefCitation id="EFE52AA5FFC2EF1FF8879527D9FC224D" author="Police, G." box="[767,945,273,302]" pageId="3" pageNumber="774" pagination="409 - 443" refId="ref9433" refString="Police, G. (1909) Alcune nuove specie di Halacaridae del Golfe di Napoli. ArchIvIo ZoologIco, NapolI, 3: 409 - 443." type="journal article" year="1909">Police (1909)</bibRefCitation>
from Naples,
<collectingCountry id="F36317C4FFC2EF1FFE019527DEF9224D" box="[1145,1204,273,302]" name="Italy" pageId="3" pageNumber="774">Italy</collectingCountry>
(Tyrrhenian Sea, Mediterranean). Later,
<bibRefCitation id="EFE52AA5FFC2EF1FFBA99500D97E2230" author="Morselli, I. &amp; Mari, M." box="[465,819,310,339]" pageId="3" pageNumber="774" pagination="229 - 247" refId="ref9387" refString="Morselli, I. &amp; Mari, M. (1982) Alacaridi (Acari, Prostigmata) di fondi sabbiosi della Costa Ionica del Salento. AttI della SocIeta Toscana dI ScIenze NaturalI, MemorIe, SerIe B, 88: 229 - 247." type="journal article" year="1982">Morselli and Mari (1982)</bibRefCitation>
reported it from Salento,
<collectingCountry id="F36317C4FFC2EF1FFED49500DEAB2230" box="[1196,1254,310,339]" name="Italy" pageId="3" pageNumber="774">Italy</collectingCountry>
(Ionian Sea, Mediterranean), in sandy sediment habitats. In this study, it is recorded for the first time from
<collectingRegion id="49B099B6FFC2EF1FFF56956DDFDA221B" box="[1326,1431,347,376]" country="Turkey" name="Antalya" pageId="3" pageNumber="774">Antalya</collectingRegion>
,
<collectingCountry id="F36317C4FFC2EF1FFAEF9549DAB122FF" box="[151,252,383,412]" name="Turkey" pageId="3" pageNumber="774">Türkiye</collectingCountry>
(Levantine Sea, Mediterranean). The external morphological characteristics of the specimens align with those from Salento,
<collectingCountry id="F36317C4FFC2EF1FF85A9592D81222A2" box="[546,607,420,449]" name="Italy" pageId="3" pageNumber="774">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>