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<document id="1CB07A6FC0CFB30E0E5007696BD384DC" ID-CLB-Dataset="29397" ID-DOI="10.11646/zootaxa.4462.3.1" ID-GBIF-Dataset="ee3428d1-9e3b-495d-99a5-e61ea679b5e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="1441611" ID-ZooBank="75D121A7-F718-4F69-9E33-E82AB371403A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538403914940" checkinUser="plazi" docAuthor="Kompier, Tom" docDate="2018" docId="03C887DB60678C06FF67D52B1BA8FDBE" docLanguage="en" docName="zootaxa.4462.3.1.pdf" docOrigin="Zootaxa 4462 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Asiagomphus auricolor" docType="treatment" docVersion="8" lastPageNumber="308" masterDocId="FFF1FFA360658C01FFF0D13A1956FF82" masterDocTitle="The genus Asiagomphus in Vietnam, with descriptions of three new species, and first descriptions of the male of Asiagomphus auricolor (Fraser, 1926) and of the female of Asiagomphus reinhardti Kosterin &amp; Yokoi, 2016 (Odonata: Gomphidae)" masterLastPageNumber="330" masterPageNumber="301" pageNumber="303" updateTime="1698561160019" updateUser="ExternalLinkService">
<mods:mods id="F9BF640952C1E8B347A2B4C246DCF88C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="50F64D880C0D7F703F27034A40D6D48A">
<mods:title id="27E2B9F7AEF7F7661A91EC38A209F046">The genus Asiagomphus in Vietnam, with descriptions of three new species, and first descriptions of the male of Asiagomphus auricolor (Fraser, 1926) and of the female of Asiagomphus reinhardti Kosterin &amp; Yokoi, 2016 (Odonata: Gomphidae)</mods:title>
</mods:titleInfo>
<mods:name id="6665F877E1CCCE7B716C2D9B5AB6F250" type="personal">
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<mods:namePart id="2C574F61E35FB86E3ED4A2C9915664A4">Kompier, Tom</mods:namePart>
</mods:name>
<mods:typeOfResource id="8F9DFA2F302C1A0E5529873A8F72CF5F">text</mods:typeOfResource>
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<mods:titleInfo id="AC104DB779F1910A4B24E8E0E76DD7CD">
<mods:title id="446F30456692EE177A11782782622480">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3FF923511BE59473A550246BD4C9F04C">
<mods:date id="5C2EEEBD876454ECB6FCDC6630C9F7B6">2018</mods:date>
<mods:detail id="E03FA7EF095B5494D8C79402ACEF921C" type="pubDate">
<mods:number id="14BF280862B0C210610351CC9F7E0A96">2018-08-28</mods:number>
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<mods:detail id="62D2353D69137CEACB76D25F05B1C95A" type="volume">
<mods:number id="BB52E87870FA41E3BB0E7AF5040CDDA8">4462</mods:number>
</mods:detail>
<mods:detail id="C97A1AA2123A9B69D8B17AB17613A88C" type="issue">
<mods:number id="8024EF3B0924C9B7E2BFA85C3503611B">3</mods:number>
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<mods:classification id="E5F45F7AB5DA21CFFF81D7A38E30C691">journal article</mods:classification>
<mods:identifier id="275DE510E6D5B29B5846AF333287EE71" type="CLB-Dataset">29397</mods:identifier>
<mods:identifier id="CC0FC7809D9D5812D0DA833A338940E0" type="DOI">10.11646/zootaxa.4462.3.1</mods:identifier>
<mods:identifier id="C927980CDC31CC90F542779BE0FCE8D5" type="GBIF-Dataset">ee3428d1-9e3b-495d-99a5-e61ea679b5e9</mods:identifier>
<mods:identifier id="BD25404F377E4B2FAC75979C4701E395" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6300B536C4A6DCB889708613303154B8" type="Zenodo-Dep">1441611</mods:identifier>
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<treatment id="03C887DB60678C06FF67D52B1BA8FDBE" ID-DOI="http://doi.org/10.5281/zenodo.5974765" ID-GBIF-Taxon="148424961" ID-Zenodo-Dep="5974765" LSID="urn:lsid:plazi:treatment:03C887DB60678C06FF67D52B1BA8FDBE" httpUri="http://treatment.plazi.org/id/03C887DB60678C06FF67D52B1BA8FDBE" lastPageId="7" lastPageNumber="308" pageId="2" pageNumber="303">
<subSubSection id="C37B654660678C03FF67D52B1B26FBCF" pageId="2" pageNumber="303" type="nomenclature">
<paragraph id="8BDE36CD60678C03FF67D52B1B31FBA9" blockId="2.[151,624,1041,1101]" box="[151,615,1041,1067]" pageId="2" pageNumber="303">
<heading id="D09681A160678C03FF67D52B1B31FBA9" bold="true" box="[151,615,1041,1067]" fontSize="11" level="1" pageId="2" pageNumber="303" reason="1">
<taxonomicName id="4C614D4E60678C03FF67D52B1B31FBA9" ID-CoL="9X246" authority="(Fraser, 1926)" baseAuthorityName="Fraser" baseAuthorityYear="1926" box="[151,615,1041,1067]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60678C03FF67D52B1B31FBA9" bold="true" box="[151,615,1041,1067]" pageId="2" pageNumber="303">
<emphasis id="B915EADF60678C03FF67D52B18E5FBA9" bold="true" box="[151,435,1041,1067]" italics="true" pageId="2" pageNumber="303">Asiagomphus auricolor</emphasis>
(
<bibRefCitation id="EFF04B3C60678C03FE32D52B1B09FBA9" box="[450,607,1041,1067]" pageId="2" pageNumber="303" refId="ref17247" refString="Fraser, F. C. (1926) Notes on a collection of Dragonflies (Order Odonata) from the Dutch East Indies and descriptions of four new species from the neighbouring continent. Treubia, 8, 467 - 494." type="journal article">Fraser, 1926</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDE36CD60678C03FF67D50E1B26FBCF" blockId="2.[151,624,1041,1101]" box="[151,624,1076,1101]" pageId="2" pageNumber="303">
(
<figureCitation id="135A2A4860678C03FF6FD50E184EFBCF" box="[159,280,1076,1101]" captionStart="FIGURE 4" captionStartId="9.[151,250,1948,1970]" captionTargetBox="[175,1416,193,1930]" captionTargetId="figure@9.[175,1416,193,1931]" captionTargetPageId="9" captionText="FIGURE 4. Anterior and posterior hamuli [a,c,e,g] lateral view, [b,d,f,h] antero-ventral lateral view. (a–b) Asiagomphus auricolor male used for description; (c–d) A. pacificus Cao Bang Prov., 19-V-2015; (e–f) A. monticola sp. nov. holotype; (g–h) A. superciliaris sp. nov. holotype." httpUri="https://zenodo.org/record/1441619/files/figure.png" pageId="2" pageNumber="303">Figs. 4a–b</figureCitation>
,
<figureCitation id="135A2A4860678C03FED3D50E1817FBCF" box="[291,321,1076,1101]" captionStart="FIGURE 5" captionStartId="10.[151,250,1939,1961]" captionTargetBox="[191,1400,193,1919]" captionTargetId="figure@10.[191,1400,193,1920]" captionTargetPageId="10" captionText="FIGURE 5. Male appendages in dorsal view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441621/files/figure.png" pageId="2" pageNumber="303">5d</figureCitation>
,
<figureCitation id="135A2A4860678C03FEBDD50E183DFBCF" box="[333,363,1076,1101]" captionStart="FIGURE 6" captionStartId="11.[151,250,1945,1967]" captionTargetBox="[201,1402,193,1927]" captionTargetId="figure@11.[188,1403,193,1927]" captionTargetPageId="11" captionText="FIGURE 6. Male appendages in lateral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441623/files/figure.png" pageId="2" pageNumber="303">6d</figureCitation>
,
<figureCitation id="135A2A4860678C03FE87D50E18C3FBCF" box="[375,405,1076,1101]" captionStart="FIGURE 7" captionStartId="12.[151,250,1946,1968]" captionTargetBox="[173,1418,193,1934]" captionTargetId="figure@12.[173,1418,193,1934]" captionTargetPageId="12" captionText="FIGURE 7. Male appendages in ventral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. superciliaris paratype Huu Lien NR, 24-V-2014." httpUri="https://zenodo.org/record/1441625/files/figure.png" pageId="2" pageNumber="303">7d</figureCitation>
,
<figureCitation id="135A2A4860678C03FE51D50E18E8FBCE" box="[417,446,1076,1100]" captionStart="FIGURE 8" captionStartId="14.[151,250,1755,1777]" captionTargetBox="[174,1436,447,1721]" captionTargetId="figure@14.[151,1436,447,1733]" captionTargetPageId="14" captionText="FIGURE 8. Right branch of male epiproct in dorso-lateral view. (a) Asiagomphus auricolor male used for description; (b) A. monticola holotype; (c) A. pacificus Cao Bang Prov., 19-V-2015; (d) A. kosterini holotype; (e) A. superciliaris holotype." httpUri="https://zenodo.org/record/1441627/files/figure.png" pageId="2" pageNumber="303">8a</figureCitation>
,
<figureCitation id="135A2A4860678C03FE39D50E18B0FBCE" box="[457,486,1076,1100]" captionStart="FIGURE 9" captionStartId="16.[151,250,1712,1734]" captionTargetBox="[151,1436,337,1691]" captionTargetId="figure@16.[151,1436,337,1691]" captionTargetPageId="16" captionText="FIGURE 9. Vertex in antero-dorsal view. [a–d] male, [e–f] female, (a) Asiagomphus superciliaris holotype; (b) A. superciliaris paratype Huu Lien NR, 24-V-2014; (c) A. superciliaris paratype Bac Kan Prov., 5-IV-2015; (d) A. monticola holotype; (e) A. auricolor female used for description; (f) A. reinhardti female used for description." httpUri="https://zenodo.org/record/1441629/files/figure.png" pageId="2" pageNumber="303">9e</figureCitation>
,
<figureCitation id="135A2A4860678C03FE02D50E1B3EFBCF" box="[498,616,1076,1101]" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [a–d, g] male, [e–f, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="2" pageNumber="303">10a–c, e–f</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37B654660678C03FF67D5461CCAFB10" box="[151,1436,1148,1170]" pageId="2" pageNumber="303" type="reference_group">
<paragraph id="8BDE36CD60678C03FF67D5461CCAFB10" blockId="2.[151,1436,1148,1170]" box="[151,1436,1148,1170]" pageId="2" pageNumber="303">
<taxonomicName id="4C614D4E60678C03FF67D5461B4DFB10" ID-CoL="9X246" authority=": Fraser (1926)" authorityName=": Fraser" authorityYear="1926" box="[151,539,1148,1170]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60678C03FF67D54618D3FB10" box="[151,389,1148,1170]" italics="true" pageId="2" pageNumber="303">Asiagomphus auricolor</emphasis>
:
<bibRefCitation id="EFF04B3C60678C03FE63D5471B4DFB10" box="[403,539,1148,1170]" pageId="2" pageNumber="303" refId="ref17247" refString="Fraser, F. C. (1926) Notes on a collection of Dragonflies (Order Odonata) from the Dutch East Indies and descriptions of four new species from the neighbouring continent. Treubia, 8, 467 - 494." type="journal article">Fraser (1926)</bibRefCitation>
</taxonomicName>
pp. 482–483, fig. 4a [description of female];
<bibRefCitation id="EFF04B3C60678C03FC19D5461D88FB10" box="[1001,1246,1148,1170]" pageId="2" pageNumber="303" refId="ref17651" refString="von Ellenrieder, N., Hauser, M., Gaimari, S. D. &amp; Thai, H. P. (2015) First records of Macromia katae (Macromiidae) and Indothemis carnatica (Libellulidae) from Vietnam (Insecta: Odonata). Check List, 11, 1 - 13. https: // doi. org / 10.15560 / 11.1.1514" type="journal article">
Ellenrieder
<emphasis id="B915EADF60678C03FB90D5471DC5FB10" box="[1120,1171,1148,1170]" italics="true" pageId="2" pageNumber="303">et al.</emphasis>
(2015)
</bibRefCitation>
, p. 5 [species list].
</paragraph>
</subSubSection>
<subSubSection id="C37B654660678C03FF67D5841CDBF971" pageId="2" pageNumber="303" type="materials_examined">
<paragraph id="8BDE36CD60678C03FF67D58418BAFAE5" blockId="2.[151,1437,1214,1959]" pageId="2" pageNumber="303">
<materialsCitation id="3B093C9060678C03FF67D5841D96FB55" ID-GBIF-Occurrence="1913893003" box="[151,1216,1214,1239]" collectingDate="2014-05-01" country="Vietnam" location="Huu Lien" pageId="2" pageNumber="303" specimenCount="1" specimenCount-male="1" stateProvince="Lang Son">
<emphasis id="B915EADF60678C03FF67D58418D4FB55" bold="true" box="[151,386,1214,1239]" pageId="2" pageNumber="303">Material examined:</emphasis>
<specimenCount id="9D67FD4460678C03FE78D58518E2FB54" box="[392,436,1215,1239]" pageId="2" pageNumber="303" type="male">1 ♂</specimenCount>
,
<location id="8EBE601660678C03FE4FD5851B7FFB55" LSID="urn:lsid:plazi:treatment:03C887DB60678C06FF67D52B1BA8FDBE:8EBE601660678C03FE4FD5851B7FFB55" box="[447,553,1214,1239]" country="Vietnam" name="Huu Lien" pageId="2" pageNumber="303" stateProvince="Lang Son">Huu Lien</location>
NR,
<collectingRegion id="49A5F82F60678C03FD94D5851B87FB55" box="[612,721,1215,1239]" country="Vietnam" name="Lang Son" pageId="2" pageNumber="303">Lang Son</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FC73D5841AB3FB55" box="[899,997,1214,1239]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FC00D5851D0AFB55" box="[1008,1116,1215,1239]" pageId="2" pageNumber="303" value="2014-05-01">
<date id="FFDF100D60678C03FC00D5851D0AFB55" box="[1008,1116,1215,1239]" pageId="2" pageNumber="303" value="2014-05-01">1-V-2014</date>
</collectingDate>
, TK leg.
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FB3BD58518E3FB79" ID-GBIF-Occurrence="1913892981" collectingDate="2014-05-24" country="Vietnam" location="Huu Lien" pageId="2" pageNumber="303" specimenCount="1" specimenCount-male="1" stateProvince="Lang Son">
<specimenCount id="9D67FD4460678C03FB3BD5851DA0FB54" box="[1227,1270,1215,1239]" pageId="2" pageNumber="303" type="male">1 ♂</specimenCount>
, same location and collector,
<collectingDate id="EF9BE9E560678C03FEC9D5D918E3FB79" box="[313,437,1251,1275]" pageId="2" pageNumber="303" value="2014-05-24">
<date id="FFDF100D60678C03FEC9D5D918E3FB79" box="[313,437,1251,1275]" pageId="2" pageNumber="303" value="2014-05-24">24-V-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FE32D5D91995FA9D" ID-GBIF-Occurrence="1913893000" collectingDate="2014-06-26" country="Vietnam" location="Bac Kan" pageId="2" pageNumber="303" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Bac Kan">
<specimenCount id="9D67FD4460678C03FE32D5D91B55FB78" box="[450,515,1251,1275]" pageId="2" pageNumber="303" type="male">2 ♂♂</specimenCount>
<specimenCount id="9D67FD4460678C03FDFCD5D91B6EFB78" box="[524,568,1251,1275]" pageId="2" pageNumber="303" type="female">1 ♀</specimenCount>
[
<emphasis id="B915EADF60678C03FDBAD5D81A6AFB79" bold="true" box="[586,828,1250,1275]" pageId="2" pageNumber="303">used for description</emphasis>
],
<collectingRegion id="49A5F82F60678C03FCA6D5D91AEEFB78" box="[854,952,1251,1274]" country="Vietnam" name="Bac Kan" pageId="2" pageNumber="303">Bac Kan</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FB84D5D81D80FB79" box="[1140,1238,1250,1275]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FB12D5D81C3FFB79" box="[1250,1385,1250,1275]" pageId="2" pageNumber="303" value="2014-06-26">
<date id="FFDF100D60678C03FB12D5D81C3FFB79" box="[1250,1385,1250,1275]" pageId="2" pageNumber="303" value="2014-06-26">26-VI-2014</date>
</collectingDate>
, TK leg.
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FF3FD43D1C65FA9D" ID-GBIF-Occurrence="1913892992" box="[207,1331,1286,1311]" collectingDate="2015-04-19" country="Vietnam" location="Huu Lien" pageId="2" pageNumber="303" specimenCount="1" specimenCount-male="1" stateProvince="Lang Son">
<specimenCount id="9D67FD4460678C03FF3FD43D19ACFA9C" box="[207,250,1287,1311]" pageId="2" pageNumber="303" type="male">1 ♂</specimenCount>
<emphasis id="B915EADF60678C03FEF1D43C1B55FA9D" bold="true" box="[257,515,1286,1311]" pageId="2" pageNumber="303">[used for description]</emphasis>
,
<location id="8EBE601660678C03FDE1D43D1B2AFA9D" LSID="urn:lsid:plazi:treatment:03C887DB60678C06FF67D52B1BA8FDBE:8EBE601660678C03FDE1D43D1B2AFA9D" box="[529,636,1286,1311]" country="Vietnam" name="Huu Lien" pageId="2" pageNumber="303" stateProvince="Lang Son">Huu Lien</location>
NR,
<collectingRegion id="49A5F82F60678C03FD49D43D1A71FA9D" box="[697,807,1287,1311]" country="Vietnam" name="Lang Son" pageId="2" pageNumber="303">Lang Son</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FC2DD43C1D69FA9D" box="[989,1087,1286,1311]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FBBAD43C1D98FA9D" box="[1098,1230,1286,1311]" pageId="2" pageNumber="303" value="2015-04-19">
<date id="FFDF100D60678C03FBBAD43C1D98FA9D" box="[1098,1230,1286,1311]" pageId="2" pageNumber="303" value="2015-04-19">19-IV-2015</date>
</collectingDate>
, TK leg.
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FACFD43D1AC9FAC1" ID-GBIF-Occurrence="1913892984" collectingDate="2015-06-17" country="Vietnam" location="Pia Oac" pageId="2" pageNumber="303" specimenCount="1" specimenCount-male="1" stateProvince="Cao Bang">
<specimenCount id="9D67FD4460678C03FACFD43D1C3AFA9C" box="[1343,1388,1287,1311]" pageId="2" pageNumber="303" type="male">1 ♂</specimenCount>
,
<location id="8EBE601660678C03FA88D43C1992FAC1" LSID="urn:lsid:plazi:treatment:03C887DB60678C06FF67D52B1BA8FDBE:8EBE601660678C03FA88D43C1992FAC1" country="Vietnam" name="Pia Oac" pageId="2" pageNumber="303" stateProvince="Cao Bang">Pia Oac</location>
NR,
<collectingRegion id="49A5F82F60678C03FEF8D411182DFAC1" box="[264,379,1323,1347]" country="Vietnam" name="Cao Bang" pageId="2" pageNumber="303">Cao Bang</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FDCCD4101BC8FAC1" box="[572,670,1322,1347]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FD5CD4101A65FAC1" box="[684,819,1322,1347]" pageId="2" pageNumber="303" value="2015-06-17">
<date id="FFDF100D60678C03FD5CD4101A65FAC1" box="[684,819,1322,1347]" pageId="2" pageNumber="303" value="2015-06-17">17-VI-2015</date>
</collectingDate>
, TK leg.
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FC5FD41118BAFAE5" ID-GBIF-Occurrence="1913892996" collectingDate="2015-06-29" country="Vietnam" location="Ba Vi" municipality="Ha Tay" pageId="2" pageNumber="303" specimenCount="1" specimenCount-male="1" stateProvince="Ha Noi">
<specimenCount id="9D67FD4460678C03FC5FD4111A88FAC0" box="[943,990,1323,1347]" pageId="2" pageNumber="303" type="male">1 ♂</specimenCount>
,
<collectingMunicipality id="6BBAACB760678C03FC1FD4111D13FAC1" box="[1007,1093,1323,1347]" pageId="2" pageNumber="303">Ha Tay</collectingMunicipality>
,
<location id="8EBE601660678C03FBA4D4111DCEFAC1" LSID="urn:lsid:plazi:treatment:03C887DB60678C06FF67D52B1BA8FDBE:8EBE601660678C03FBA4D4111DCEFAC1" box="[1108,1176,1322,1347]" country="Vietnam" municipality="Ha Tay" name="Ba Vi" pageId="2" pageNumber="303" stateProvince="Ha Noi">Ba Vi</location>
NP,
<collectingRegion id="49A5F82F60678C03FB26D4111C78FAC1" box="[1238,1326,1322,1347]" country="Vietnam" name="Ha Noi" pageId="2" pageNumber="303">Ha Noi</collectingRegion>
, northern
<collectingCountry id="F376765D60678C03FF67D47419AFFAE5" box="[151,249,1358,1383]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FEF4D47418DDFAE5" box="[260,395,1358,1383]" pageId="2" pageNumber="303" value="2015-06-29">
<date id="FFDF100D60678C03FEF4D47418DDFAE5" box="[260,395,1358,1383]" pageId="2" pageNumber="303" value="2015-06-29">29-VI-2015</date>
</collectingDate>
, TK leg.
</materialsCitation>
</paragraph>
<paragraph id="8BDE36CD60678C03FF37D4481CDBF971" blockId="2.[151,1437,1214,1959]" pageId="2" pageNumber="303">
<materialsCitation id="3B093C9060678C03FF37D44818DCFA2D" ID-GBIF-Occurrence="1913893011" collectingDate="2014-04-21" country="Vietnam" location="Huu Lien" pageId="2" pageNumber="303" specimenCount="1" specimenCount-male="1" stateProvince="Lang Son">
<emphasis id="B915EADF60678C03FF37D4481A26FA09" bold="true" box="[199,880,1394,1419]" pageId="2" pageNumber="303">Additional records based on field observations by TK:</emphasis>
<specimenCount id="9D67FD4460678C03FC8CD4491AFBFA08" box="[892,941,1395,1419]" pageId="2" pageNumber="303" type="male">1 ♂</specimenCount>
,
<location id="8EBE601660678C03FC4ED4491D7BFA09" LSID="urn:lsid:plazi:treatment:03C887DB60678C06FF67D52B1BA8FDBE:8EBE601660678C03FC4ED4491D7BFA09" box="[958,1069,1394,1419]" country="Vietnam" name="Huu Lien" pageId="2" pageNumber="303" stateProvince="Lang Son">Huu Lien</location>
NR,
<collectingRegion id="49A5F82F60678C03FB83D4491DB3FA09" box="[1139,1253,1395,1419]" country="Vietnam" name="Lang Son" pageId="2" pageNumber="303">Lang Son</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FF67D4AC19AFFA2D" box="[151,249,1430,1455]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FEF4D4AD18DCFA2D" box="[260,394,1431,1455]" pageId="2" pageNumber="303" value="2014-04-21">
<date id="FFDF100D60678C03FEF4D4AD18DCFA2D" box="[260,394,1431,1455]" pageId="2" pageNumber="303" value="2014-04-21">21-IV-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FE65D4AC1A44FA2D" ID-GBIF-Occurrence="1913892998" box="[405,786,1430,1455]" collectingDate="2014-04-26" country="Vietnam" location="Huu Lien" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Lang Son">
5 ind., same location,
<collectingDate id="EF9BE9E560678C03FD7DD4AC1A44FA2D" box="[653,786,1430,1455]" pageId="2" pageNumber="303" value="2014-04-26">
<date id="FFDF100D60678C03FD7DD4AC1A44FA2D" box="[653,786,1430,1455]" pageId="2" pageNumber="303" value="2014-04-26">26-IV-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FCEED4AD1DC4FA2D" ID-GBIF-Occurrence="1913892997" box="[798,1170,1430,1455]" collectingDate="2014-05-01" country="Vietnam" location="Huu Lien" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Lang Son">
14 ind., same location,
<collectingDate id="EF9BE9E560678C03FBD4D4AD1DC4FA2D" box="[1060,1170,1431,1455]" pageId="2" pageNumber="303" value="2014-05-01">
<date id="FFDF100D60678C03FBD4D4AD1DC4FA2D" box="[1060,1170,1431,1455]" pageId="2" pageNumber="303" value="2014-05-01">1-V-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FB6DD4AD1845FA51" ID-GBIF-Occurrence="1913892978" collectingDate="2014-05-24" country="Vietnam" location="Huu Lien" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Lang Son">
10 ind., same location,
<collectingDate id="EF9BE9E560678C03FF67D4811845FA51" box="[151,275,1467,1491]" pageId="2" pageNumber="303" value="2014-05-24">
<date id="FFDF100D60678C03FF67D4811845FA51" box="[151,275,1467,1491]" pageId="2" pageNumber="303" value="2014-05-24">24-V-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FED0D4811AD0FA51" ID-GBIF-Occurrence="1913892989" box="[288,902,1466,1491]" collectingDate="2014-06-21" country="Vietnam" location="Bac Kan" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Bac Kan">
1 ind.,
<collectingRegion id="49A5F82F60678C03FE82D4811883FA50" box="[370,469,1467,1490]" country="Vietnam" name="Bac Kan" pageId="2" pageNumber="303">Bac Kan</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FD7FD4801BA7FA51" box="[655,753,1466,1491]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FD0ED4811AD0FA51" box="[766,902,1467,1491]" pageId="2" pageNumber="303" value="2014-06-21">
<date id="FFDF100D60678C03FD0ED4811AD0FA51" box="[766,902,1467,1491]" pageId="2" pageNumber="303" value="2014-06-21">21-VI-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FC63D4811C4FFA51" ID-GBIF-Occurrence="1913892993" box="[915,1305,1466,1491]" collectingDate="2014-06-22" country="Vietnam" location="Bac Kan" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Bac Kan">
1 ind., same location,
<collectingDate id="EF9BE9E560678C03FB62D4811C4FFA51" box="[1170,1305,1467,1491]" pageId="2" pageNumber="303" value="2014-06-22">
<date id="FFDF100D60678C03FB62D4811C4FFA51" box="[1170,1305,1467,1491]" pageId="2" pageNumber="303" value="2014-06-22">22-VI-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FAD7D48118D0FA75" ID-GBIF-Occurrence="1913892974" collectingDate="2014-06-24" country="Vietnam" location="Bac Kan" pageId="2" pageNumber="303" specimenCount="1" specimenCount-male="1" stateProvince="Bac Kan">
<specimenCount id="9D67FD4460678C03FAD7D4811C03FA50" box="[1319,1365,1467,1491]" pageId="2" pageNumber="303" type="male">1 ♂</specimenCount>
, same location,
<collectingDate id="EF9BE9E560678C03FF0FD4E518D0FA75" box="[255,390,1503,1527]" pageId="2" pageNumber="303" value="2014-06-24">
<date id="FFDF100D60678C03FF0FD4E518D0FA75" box="[255,390,1503,1527]" pageId="2" pageNumber="303" value="2014-06-24">24-VI-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FE62D4E51BACFA75" ID-GBIF-Occurrence="1913892988" box="[402,762,1502,1527]" collectingDate="2014-06-25" country="Vietnam" location="Bac Kan" pageId="2" pageNumber="303" specimenCount="1" specimenCount-male="1" stateProvince="Bac Kan">
<specimenCount id="9D67FD4460678C03FE62D4E518E8FA74" box="[402,446,1503,1527]" pageId="2" pageNumber="303" type="male">1 ♂</specimenCount>
, same location,
<collectingDate id="EF9BE9E560678C03FD83D4E41BACFA75" box="[627,762,1502,1527]" pageId="2" pageNumber="303" value="2014-06-25">
<date id="FFDF100D60678C03FD83D4E41BACFA75" box="[627,762,1502,1527]" pageId="2" pageNumber="303" value="2014-06-25">25-VI-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FCF6D4E519AFF999" ID-GBIF-Occurrence="1913892994" collectingDate="2014-06-25" country="Vietnam" location="Pia Oac" pageId="2" pageNumber="303" specimenCount="3" specimenCount-male="3" stateProvince="Cao Bang">
<specimenCount id="9D67FD4460678C03FCF6D4E51A10FA74" box="[774,838,1503,1527]" pageId="2" pageNumber="303" type="male">3 ♂♂</specimenCount>
,
<location id="8EBE601660678C03FCA2D4E41AFCFA75" LSID="urn:lsid:plazi:treatment:03C887DB60678C06FF67D52B1BA8FDBE:8EBE601660678C03FCA2D4E41AFCFA75" box="[850,938,1502,1527]" country="Vietnam" name="Pia Oac" pageId="2" pageNumber="303" stateProvince="Cao Bang">Pia Oac</location>
NR,
<collectingRegion id="49A5F82F60678C03FC16D4E51D03FA75" box="[998,1109,1503,1527]" country="Vietnam" name="Cao Bang" pageId="2" pageNumber="303">Cao Bang</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FAFAD4E41C3AFA75" box="[1290,1388,1502,1527]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FA87D4E419AFF999" pageId="2" pageNumber="303" value="2014-06-25">
<date id="FFDF100D60678C03FA87D4E419AFF999" pageId="2" pageNumber="303" value="2014-06-25">25- VI-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FEF5D7391A34F999" ID-GBIF-Occurrence="1913893012" box="[261,866,1538,1563]" collectingDate="2014-06-26" country="Vietnam" location="Bac Kan" pageId="2" pageNumber="303" specimenCount="3" specimenCount-male="3" stateProvince="Bac Kan">
<specimenCount id="9D67FD4460678C03FEF5D7391811F998" box="[261,327,1539,1563]" pageId="2" pageNumber="303" type="male">3 ♂♂</specimenCount>
,
<collectingRegion id="49A5F82F60678C03FEA4D73918E0F998" box="[340,438,1539,1562]" country="Vietnam" name="Bac Kan" pageId="2" pageNumber="303">Bac Kan</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FD9DD7381B99F999" box="[621,719,1538,1563]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FD2BD7381A34F999" box="[731,866,1538,1563]" pageId="2" pageNumber="303" value="2014-06-26">
<date id="FFDF100D60678C03FD2BD7381A34F999" box="[731,866,1538,1563]" pageId="2" pageNumber="303" value="2014-06-26">26-VI-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FC9FD7391985F9BD" ID-GBIF-Occurrence="1913893001" collectingDate="2014-06-28" country="Vietnam" location="Lang Son" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Lang Son">
8 ind.,
<collectingRegion id="49A5F82F60678C03FC4ED7391D7AF999" box="[958,1068,1539,1563]" country="Vietnam" name="Lang Son" pageId="2" pageNumber="303">Lang Son</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FB12D7381C12F999" box="[1250,1348,1538,1563]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FAA0D7391985F9BD" pageId="2" pageNumber="303" value="2014-06-28">
<date id="FFDF100D60678C03FAA0D7391985F9BD" pageId="2" pageNumber="303" value="2014-06-28">28-VI- 2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FF2DD71D1A7CF9BD" ID-GBIF-Occurrence="1913893004" box="[221,810,1574,1599]" collectingDate="2014-06-29" country="Vietnam" location="Cao Bang" pageId="2" pageNumber="303" specimenCount="1" specimenCount-male="1" stateProvince="Cao Bang">
<specimenCount id="9D67FD4460678C03FF2DD71D185EF9BC" box="[221,264,1575,1599]" pageId="2" pageNumber="303" type="male">1 ♂</specimenCount>
,
<collectingRegion id="49A5F82F60678C03FEE4D71D18D4F9BD" box="[276,386,1575,1599]" country="Vietnam" name="Cao Bang" pageId="2" pageNumber="303">Cao Bang</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FDC6D71C1BCEF9BD" box="[566,664,1574,1599]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FD52D71D1A7CF9BD" box="[674,810,1575,1599]" pageId="2" pageNumber="303" value="2014-06-29">
<date id="FFDF100D60678C03FD52D71D1A7CF9BD" box="[674,810,1575,1599]" pageId="2" pageNumber="303" value="2014-06-29">29-VI-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FCC4D71D1CD2F9BD" ID-GBIF-Occurrence="1913893009" box="[820,1412,1574,1599]" collectingDate="2014-07-06" country="Vietnam" location="Bac Kan" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Bac Kan">
2 ind.,
<collectingRegion id="49A5F82F60678C03FC70D71D1AB7F9BC" box="[896,993,1575,1598]" country="Vietnam" name="Bac Kan" pageId="2" pageNumber="303">Bac Kan</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FB64D71C1DA0F9BD" box="[1172,1270,1574,1599]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FAF1D71C1CD2F9BD" box="[1281,1412,1574,1599]" pageId="2" pageNumber="303" value="2014-07-06">
<date id="FFDF100D60678C03FAF1D71C1CD2F9BD" box="[1281,1412,1574,1599]" pageId="2" pageNumber="303" value="2014-07-06">6-VII-2014</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FA7ED71C1ADBF9E1" ID-GBIF-Occurrence="1913892969" collectingDate="2015-03-22" country="Vietnam" location="Lang Son" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Lang Son">
5 ind., Huu Lien NR,
<collectingRegion id="49A5F82F60678C03FE87D77118B2F9E1" box="[375,484,1611,1635]" country="Vietnam" name="Lang Son" pageId="2" pageNumber="303">Lang Son</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FD69D7701BADF9E1" box="[665,763,1610,1635]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FCF7D7701ADBF9E1" box="[775,909,1610,1635]" pageId="2" pageNumber="303" value="2015-03-22">
<date id="FFDF100D60678C03FCF7D7701ADBF9E1" box="[775,909,1610,1635]" pageId="2" pageNumber="303" value="2015-03-22">22-III-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FC68D7701C43F9E1" ID-GBIF-Occurrence="1913892991" box="[920,1301,1610,1635]" collectingDate="2015-04-19" country="Vietnam" location="Lang Son" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Lang Son">
5 ind., same location,
<collectingDate id="EF9BE9E560678C03FB60D7701C43F9E1" box="[1168,1301,1610,1635]" pageId="2" pageNumber="303" value="2015-04-19">
<date id="FFDF100D60678C03FB60D7701C43F9E1" box="[1168,1301,1610,1635]" pageId="2" pageNumber="303" value="2015-04-19">19-IV-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FAD1D7711B2EF905" ID-GBIF-Occurrence="1913892979" collectingDate="2015-04-30" country="Vietnam" location="Cao Bang" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Cao Bang">
7 ind.,
<collectingRegion id="49A5F82F60678C03FA9FD7711984F905" country="Vietnam" name="Cao Bang" pageId="2" pageNumber="303">Cao Bang</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FE76D75418BEF905" box="[390,488,1646,1671]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FE03D7541B2EF905" box="[499,632,1646,1671]" pageId="2" pageNumber="303" value="2015-04-30">
<date id="FFDF100D60678C03FE03D7541B2EF905" box="[499,632,1646,1671]" pageId="2" pageNumber="303" value="2015-04-30">30-IV-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FD73D7551AA3F905" ID-GBIF-Occurrence="1913893007" box="[643,1013,1646,1671]" collectingDate="2015-05-23" country="Vietnam" location="Cao Bang" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Cao Bang">
2 ind., same location,
<collectingDate id="EF9BE9E560678C03FC89D7541AA3F905" box="[889,1013,1646,1671]" pageId="2" pageNumber="303" value="2015-05-23">
<date id="FFDF100D60678C03FC89D7541AA3F905" box="[889,1013,1646,1671]" pageId="2" pageNumber="303" value="2015-05-23">23-V-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FBF0D75418DCF929" ID-GBIF-Occurrence="1913892985" collectingDate="2015-06-15" country="Vietnam" location="Thai Nguyen" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Thai Nguyen">
15 ind.,
<collectingRegion id="49A5F82F60678C03FBACD7541DB9F905" box="[1116,1263,1646,1671]" country="Vietnam" name="Thai Nguyen" pageId="2" pageNumber="303">Thai Nguyen</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FF67D7A819AFF929" box="[151,249,1682,1707]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FEF3D7A818DCF929" box="[259,394,1682,1707]" pageId="2" pageNumber="303" value="2015-06-15">
<date id="FFDF100D60678C03FEF3D7A818DCF929" box="[259,394,1682,1707]" pageId="2" pageNumber="303" value="2015-06-15">15-VI-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FE66D7A91DF0F929" ID-GBIF-Occurrence="1913893006" box="[406,1190,1682,1707]" collectingDate="2015-06-16" country="Vietnam" location="Cuc Phuong" pageId="2" pageNumber="303" specimenCount="1" specimenCount-male="1" stateProvince="Ninh Binh">
<specimenCount id="9D67FD4460678C03FE66D7A91897F928" box="[406,449,1683,1707]" pageId="2" pageNumber="303" type="male">1 ♂</specimenCount>
,
<location id="8EBE601660678C03FE3CD7A91B03F929" LSID="urn:lsid:plazi:treatment:03C887DB60678C06FF67D52B1BA8FDBE:8EBE601660678C03FE3CD7A91B03F929" box="[460,597,1682,1707]" country="Vietnam" name="Cuc Phuong" pageId="2" pageNumber="303" stateProvince="Ninh Binh">Cuc Phuong</location>
NP,
<collectingRegion id="49A5F82F60678C03FD7AD7A81BA9F929" box="[650,767,1682,1707]" country="Vietnam" name="Ninh Binh" pageId="2" pageNumber="303">Ninh Binh</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FC42D7A81D42F929" box="[946,1044,1682,1707]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FBEFD7A81DF0F929" box="[1055,1190,1682,1707]" pageId="2" pageNumber="303" value="2015-06-16">
<date id="FFDF100D60678C03FBEFD7A81DF0F929" box="[1055,1190,1682,1707]" pageId="2" pageNumber="303" value="2015-06-16">16-VI-2015</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FB41D7A91888F94D" ID-GBIF-Occurrence="1913892963" collectingDate="2016-03-19" collectorName="Huu Lien NR" country="Vietnam" location="Lang Son" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Lang Son">
2 ind.,
<collectorName id="2694531B60678C03FB0ED7A91CCEF928" box="[1278,1432,1682,1707]" pageId="2" pageNumber="303">Huu Lien NR</collectorName>
,
<collectingRegion id="49A5F82F60678C03FF67D78D1853F94D" box="[151,261,1719,1743]" country="Vietnam" name="Lang Son" pageId="2" pageNumber="303">Lang Son</collectingRegion>
Prov.,
<collectingDate id="EF9BE9E560678C03FEA7D78C1888F94D" box="[343,478,1718,1743]" pageId="2" pageNumber="303" value="2016-03-19">
<date id="FFDF100D60678C03FEA7D78C1888F94D" box="[343,478,1718,1743]" pageId="2" pageNumber="303" value="2016-03-19">19-III-2016</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FE1BD78D1C46F94D" ID-GBIF-Occurrence="1913892990" box="[491,1296,1718,1743]" collectingDate="2017-04-16" country="Vietnam" location="Cuc Phuong" pageId="2" pageNumber="303" specimenCount="1" specimenCount-male="1" stateProvince="Ninh Binh">
<specimenCount id="9D67FD4460678C03FE1BD78D1B4EF94C" box="[491,536,1719,1743]" pageId="2" pageNumber="303" type="male">1 ♂</specimenCount>
,
<location id="8EBE601660678C03FDD6D78D1BE7F94D" LSID="urn:lsid:plazi:treatment:03C887DB60678C06FF67D52B1BA8FDBE:8EBE601660678C03FDD6D78D1BE7F94D" box="[550,689,1718,1743]" country="Vietnam" name="Cuc Phuong" pageId="2" pageNumber="303" stateProvince="Ninh Binh">Cuc Phuong</location>
NP,
<collectingRegion id="49A5F82F60678C03FD1BD78C1A34F94D" box="[747,866,1718,1743]" country="Vietnam" name="Ninh Binh" pageId="2" pageNumber="303">Ninh Binh</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FBEDD78C1D29F94D" box="[1053,1151,1718,1743]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FB7BD78C1C46F94D" box="[1163,1296,1718,1743]" pageId="2" pageNumber="303" value="2017-04-16">
<date id="FFDF100D60678C03FB7BD78C1C46F94D" box="[1163,1296,1718,1743]" pageId="2" pageNumber="303" value="2017-04-16">16-IV-2017</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FAEED78D1A16F971" ID-GBIF-Occurrence="1913892975" collectingDate="2017-06-11" country="Vietnam" location="Ninh Binh" pageId="2" pageNumber="303" specimenCount="1" stateProvince="Ninh Binh">
1 ind., Cuc Phuong NP,
<collectingRegion id="49A5F82F60678C03FED2D7E018C1F971" box="[290,407,1754,1779]" country="Vietnam" name="Ninh Binh" pageId="2" pageNumber="303">Ninh Binh</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FDBDD7E01BF9F971" box="[589,687,1754,1779]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FD49D7E11A16F971" box="[697,832,1755,1779]" pageId="2" pageNumber="303" value="2017-06-11">
<date id="FFDF100D60678C03FD49D7E11A16F971" box="[697,832,1755,1779]" pageId="2" pageNumber="303" value="2017-06-11">11-VI-2017</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B093C9060678C03FCBBD7E11CDFF971" ID-GBIF-Occurrence="1913893005" box="[843,1417,1754,1779]" collectingDate="2017-07-08" country="Vietnam" location="Bac Kan" pageId="2" pageNumber="303" specimenCount="1" specimenCount-male="1" stateProvince="Bac Kan">
<specimenCount id="9D67FD4460678C03FCBBD7E11A21F970" box="[843,887,1755,1779]" pageId="2" pageNumber="303" type="male">1 ♂</specimenCount>
,
<collectingRegion id="49A5F82F60678C03FC73D7E11AB3F970" box="[899,997,1755,1778]" country="Vietnam" name="Bac Kan" pageId="2" pageNumber="303">Bac Kan</collectingRegion>
Prov., northern
<collectingCountry id="F376765D60678C03FB69D7E01DADF971" box="[1177,1275,1754,1779]" name="Vietnam" pageId="2" pageNumber="303">Vietnam</collectingCountry>
,
<collectingDate id="EF9BE9E560678C03FAF6D7E11CDFF971" box="[1286,1417,1755,1779]" pageId="2" pageNumber="303" value="2017-07-08">
<date id="FFDF100D60678C03FAF6D7E11CDFF971" box="[1286,1417,1755,1779]" pageId="2" pageNumber="303" value="2017-07-08">8-VII-2017</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B654660678C04FF37D7C41AE8FAF1" lastPageId="5" lastPageNumber="306" pageId="2" pageNumber="303" type="description">
<paragraph id="8BDE36CD60678C03FF37D7C41A48F825" blockId="2.[151,1437,1214,1959]" pageId="2" pageNumber="303">
<emphasis id="B915EADF60678C03FF37D7C418AFF895" bold="true" box="[199,505,1790,1815]" pageId="2" pageNumber="303">First description of male</emphasis>
(
<figureCitation id="135A2A4860678C03FDFCD7C41BDFF895" box="[524,649,1790,1815]" captionStart="FIGURE 4" captionStartId="9.[151,250,1948,1970]" captionTargetBox="[175,1416,193,1930]" captionTargetId="figure@9.[175,1416,193,1931]" captionTargetPageId="9" captionText="FIGURE 4. Anterior and posterior hamuli [a,c,e,g] lateral view, [b,d,f,h] antero-ventral lateral view. (a–b) Asiagomphus auricolor male used for description; (c–d) A. pacificus Cao Bang Prov., 19-V-2015; (e–f) A. monticola sp. nov. holotype; (g–h) A. superciliaris sp. nov. holotype." httpUri="https://zenodo.org/record/1441619/files/figure.png" pageId="2" pageNumber="303">Figs. 4a–b</figureCitation>
,
<figureCitation id="135A2A4860678C03FD67D7C41BE0F895" box="[663,694,1790,1815]" captionStart="FIGURE 5" captionStartId="10.[151,250,1939,1961]" captionTargetBox="[191,1400,193,1919]" captionTargetId="figure@10.[191,1400,193,1920]" captionTargetPageId="10" captionText="FIGURE 5. Male appendages in dorsal view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441621/files/figure.png" pageId="2" pageNumber="303">5d</figureCitation>
,
<figureCitation id="135A2A4860678C03FD35D7C41BB2F895" box="[709,740,1790,1815]" captionStart="FIGURE 6" captionStartId="11.[151,250,1945,1967]" captionTargetBox="[201,1402,193,1927]" captionTargetId="figure@11.[188,1403,193,1927]" captionTargetPageId="11" captionText="FIGURE 6. Male appendages in lateral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441623/files/figure.png" pageId="2" pageNumber="303">6d</figureCitation>
,
<figureCitation id="135A2A4860678C03FD04D7C41A44F895" box="[756,786,1790,1815]" captionStart="FIGURE 7" captionStartId="12.[151,250,1946,1968]" captionTargetBox="[173,1418,193,1934]" captionTargetId="figure@12.[173,1418,193,1934]" captionTargetPageId="12" captionText="FIGURE 7. Male appendages in ventral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. superciliaris paratype Huu Lien NR, 24-V-2014." httpUri="https://zenodo.org/record/1441625/files/figure.png" pageId="2" pageNumber="303">7d</figureCitation>
,
<figureCitation id="135A2A4860678C03FCD2D7C51A69F895" box="[802,831,1791,1815]" captionStart="FIGURE 8" captionStartId="14.[151,250,1755,1777]" captionTargetBox="[174,1436,447,1721]" captionTargetId="figure@14.[151,1436,447,1733]" captionTargetPageId="14" captionText="FIGURE 8. Right branch of male epiproct in dorso-lateral view. (a) Asiagomphus auricolor male used for description; (b) A. monticola holotype; (c) A. pacificus Cao Bang Prov., 19-V-2015; (d) A. kosterini holotype; (e) A. superciliaris holotype." httpUri="https://zenodo.org/record/1441627/files/figure.png" pageId="2" pageNumber="303">8a</figureCitation>
).
<emphasis id="B915EADF60678C03FCA8D7C41AC2F895" box="[856,916,1790,1815]" italics="true" pageId="2" pageNumber="303">Head</emphasis>
. Labium, labrum, postclypeus and antefrons black. Anteclypeus dark brown. Base of mandibles pale yellow. Postfrons with broad yellow stripe, extending anteriorly just down the antefrons, and posterior edge of postfrons black. Cranium black, lateral ocelli framed with a raised ridge posteriorly. Occiput dark brown, wrinkled and slightly raised centrally, occipital ridge with long black hairs. Antennae black, but top of first segment pale.
</paragraph>
<caption id="DF1E664560668C02FF67D6F21A61F85C" box="[151,823,1992,2015]" httpUri="https://zenodo.org/record/1441613/files/figure.png" pageId="3" pageNumber="304" startId="3.[151,250,1992,2014]" targetBox="[204,1391,193,1976]" targetPageId="3">
<paragraph id="8BDE36CD60668C02FF67D6F21A61F85C" blockId="3.[151,823,1992,2015]" box="[151,823,1992,2015]" pageId="3" pageNumber="304">
<emphasis id="B915EADF60668C02FF67D6F21845F85F" bold="true" box="[151,275,1992,2014]" pageId="3" pageNumber="304">FIGURE 1.</emphasis>
<taxonomicName id="4C614D4E60668C02FEE9D6F2188EF85D" box="[281,472,1992,2015]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="acco">
<emphasis id="B915EADF60668C02FEE9D6F2188EF85D" box="[281,472,1992,2015]" italics="true" pageId="3" pageNumber="304">Asiagomphus acco</emphasis>
</taxonomicName>
copula, Xuan Son NP, 30-V-2015.
</paragraph>
</caption>
<paragraph id="8BDE36CD60618C05FF37D1AD1C10FE2F" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
<emphasis id="B915EADF60618C05FF37D1AD184EFF32" box="[199,280,151,176]" italics="true" pageId="4" pageNumber="305">Thorax</emphasis>
. Prothorax black, but anterior lobe with central yellow spot, and median lobe with small yellow central spot and pair of lateral spots. Synthorax black with pale yellow markings as follows: Mesepisternum with dorsal stripe, widening anteriorly and confluent with collar stripe to form two inverted “7” shapes. Small antehumeral spot towards dorsal end of humeral suture. Mesepimeron almost completely covered by broad lateral stripe. Mesokatepisternum blackish brown with large yellow spot. Metepisternum with broad lateral stripe, somewhat constricted just below dorsal end and bordered by clear black stripe over interpleural suture, running through spiracle, margins of yellow stripe less distinct ventrally. Metakatepisternum brown with diffuse yellow center. Metepimeron almost completely yellow, but black stripe over metapleural suture. Coxae and legs all black.
</paragraph>
<paragraph id="8BDE36CD60618C05FF37D08D1BEFFDE2" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
<emphasis id="B915EADF60618C05FF37D08D1867FE52" box="[199,305,439,464]" italics="true" pageId="4" pageNumber="305">Abdomen</emphasis>
. Black with yellow markings as follows: S1 yellow with small black area dorso-laterally covered in long black hairs; S2 yellow with black marking dorsally of yellow auricle, continuing posteriorly over dorsum, but interrupted in middle over dorsal central line by yellow stripe; S3 with antero-ventral spot and short narrow anterodorsal stripe; S4–7 with antero-dorsal half-ring, interrupted dorsally by thin black line on S7; posterior half of S9 with large dorsal mark. S7–9 markedly widened.
</paragraph>
<paragraph id="8BDE36CD60618C05FF37D3561D38FC97" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
<emphasis id="B915EADF60618C05FF37D35618F2FD07" box="[199,420,620,645]" italics="true" pageId="4" pageNumber="305">Accessory genitalia</emphasis>
(
<figureCitation id="135A2A4860618C05FE44D3561B7AFD07" box="[436,556,620,645]" captionStart="FIGURE 4" captionStartId="9.[151,250,1948,1970]" captionTargetBox="[175,1416,193,1930]" captionTargetId="figure@9.[175,1416,193,1931]" captionTargetPageId="9" captionText="FIGURE 4. Anterior and posterior hamuli [a,c,e,g] lateral view, [b,d,f,h] antero-ventral lateral view. (a–b) Asiagomphus auricolor male used for description; (c–d) A. pacificus Cao Bang Prov., 19-V-2015; (e–f) A. monticola sp. nov. holotype; (g–h) A. superciliaris sp. nov. holotype." httpUri="https://zenodo.org/record/1441619/files/figure.png" pageId="4" pageNumber="305">Figs. 4a–b</figureCitation>
). Black, anterior hamulus about
<date id="FFDF100D60618C05FC51D3561AEBFD06" box="[929,957,620,644]" pageId="4" pageNumber="305">4x</date>
longer than wide, apical third constricted, minimally curved inwards and posteriorly at apex, latero-posterior margin of constricted part with distinct small teeth. Posterior hamulus robust, in lateral view
<date id="FFDF100D60618C05FD5CD38E1B9EFD4E" box="[684,712,692,716]" pageId="4" pageNumber="305">3x</date>
length of visible part of anterior hamulus, lateral surface deeply concave in middle part, apex directed inward and anteriorly to form robust hook, lateral surface smooth, without ridge crossing surface distal of concave part and anterior margin smooth in lateral view.
</paragraph>
<paragraph id="8BDE36CD60618C05FF37D21B1DC7FCBA" blockId="4.[151,1437,151,2013]" box="[199,1169,799,824]" pageId="4" pageNumber="305">
<taxonomicName id="4C614D4E60618C05FF37D21B18D4FCBA" box="[199,386,799,824]" class="Insecta" family="Pneumoridae" genus="Vesica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="spermalis">
<emphasis id="B915EADF60618C05FF37D21B18D4FCBA" box="[199,386,799,824]" italics="true" pageId="4" pageNumber="305">Vesica spermalis</emphasis>
</taxonomicName>
. Flagellum extending well beyond median process of apical segment.
</paragraph>
<paragraph id="8BDE36CD60618C05FF37D27F1B3CFC26" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
<emphasis id="B915EADF60618C05FF37D27F1844FCDE" box="[199,274,837,860]" italics="true" pageId="4" pageNumber="305">Wings.</emphasis>
Hyaline with black veins, minimally tinted amber at base. Anal triangle 3-celled, cubito-anal space with one crossvein, all triangles uncrossed, nodal index 12–19:18–
<quantity id="4C999B2860618C05FCAED2521ACCFC02" box="[862,922,871,896]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="4" pageNumber="305" unit="in" value="14.0">14 in</quantity>
FW, 15–12:
<quantity id="4C999B2860618C05FBD9D25D1DC7FC02" box="[1065,1169,871,896]" metricMagnitude="-1" metricUnit="m" metricValue="3.683" metricValueMax="3.81" metricValueMin="3.556" pageId="4" pageNumber="305" unit="in" value="14.5" valueMax="15.0" valueMin="14.0">14–15 in</quantity>
HW. Pt brown,
<quantity id="4C999B2860618C05FAA2D2521CCEFCFD" box="[1362,1432,872,896]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="305" unit="mm" value="4.0">4 mm</quantity>
, covering 5–6 cells in FW and
<quantity id="4C999B2860618C05FE18D2B61B79FC27" box="[488,559,908,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="1.778" metricValueMin="1.524" pageId="4" pageNumber="305" unit="in" value="6.5" valueMax="7.0" valueMin="6.0">6–7 in</quantity>
HW.
</paragraph>
<paragraph id="8BDE36CD60618C05FF37D2951995FB6A" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
<emphasis id="B915EADF60618C05FF37D29518DEFC4A" box="[199,392,943,968]" italics="true" pageId="4" pageNumber="305">Anal appendages</emphasis>
(
<figureCitation id="135A2A4860618C05FE68D29518A0FC4A" box="[408,502,943,968]" captionStart="FIGURE 5" captionStartId="10.[151,250,1939,1961]" captionTargetBox="[191,1400,193,1919]" captionTargetId="figure@10.[191,1400,193,1920]" captionTargetPageId="10" captionText="FIGURE 5. Male appendages in dorsal view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441621/files/figure.png" pageId="4" pageNumber="305">Figs. 5d</figureCitation>
,
<figureCitation id="135A2A4860618C05FDF3D2951B77FC4A" box="[515,545,943,968]" captionStart="FIGURE 6" captionStartId="11.[151,250,1945,1967]" captionTargetBox="[201,1402,193,1927]" captionTargetId="figure@11.[188,1403,193,1927]" captionTargetPageId="11" captionText="FIGURE 6. Male appendages in lateral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441623/files/figure.png" pageId="4" pageNumber="305">6d</figureCitation>
,
<figureCitation id="135A2A4860618C05FDDDD2951B1DFC4A" box="[557,587,943,968]" captionStart="FIGURE 7" captionStartId="12.[151,250,1946,1968]" captionTargetBox="[173,1418,193,1934]" captionTargetId="figure@12.[173,1418,193,1934]" captionTargetPageId="12" captionText="FIGURE 7. Male appendages in ventral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. superciliaris paratype Huu Lien NR, 24-V-2014." httpUri="https://zenodo.org/record/1441625/files/figure.png" pageId="4" pageNumber="305">7d</figureCitation>
,
<figureCitation id="135A2A4860618C05FDA7D28A1B22FC4A" box="[599,628,944,968]" captionStart="FIGURE 8" captionStartId="14.[151,250,1755,1777]" captionTargetBox="[174,1436,447,1721]" captionTargetId="figure@14.[151,1436,447,1733]" captionTargetPageId="14" captionText="FIGURE 8. Right branch of male epiproct in dorso-lateral view. (a) Asiagomphus auricolor male used for description; (b) A. monticola holotype; (c) A. pacificus Cao Bang Prov., 19-V-2015; (d) A. kosterini holotype; (e) A. superciliaris holotype." httpUri="https://zenodo.org/record/1441627/files/figure.png" pageId="4" pageNumber="305">8a</figureCitation>
). Black. Cerci simple, unbranched, widely diverging in dorsal view and apically tapering to a posteriorly directed point (
<figureCitation id="135A2A4860618C05FD43D2EE1A53FC6F" box="[691,773,980,1005]" captionStart="FIGURE 5" captionStartId="10.[151,250,1939,1961]" captionTargetBox="[191,1400,193,1919]" captionTargetId="figure@10.[191,1400,193,1920]" captionTargetPageId="10" captionText="FIGURE 5. Male appendages in dorsal view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441621/files/figure.png" pageId="4" pageNumber="305">Fig. 5d</figureCitation>
), in lateral view minimally curved downward at mid-point, apically inflicted upward to form up-slanting spike, in length cerci approximately 1.5x length of S10 (
<figureCitation id="135A2A4860618C05FAC9D2CD1CD9FB92" box="[1337,1423,1015,1040]" captionStart="FIGURE 6" captionStartId="11.[151,250,1945,1967]" captionTargetBox="[201,1402,193,1927]" captionTargetId="figure@11.[188,1403,193,1927]" captionTargetPageId="11" captionText="FIGURE 6. Male appendages in lateral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441623/files/figure.png" pageId="4" pageNumber="305">Fig. 6d</figureCitation>
). Epiproct deeply bifid, in dorsal view branches rounded, simple (
<figureCitation id="135A2A4860618C05FC9CD5261AE8FBB6" box="[876,958,1052,1077]" captionStart="FIGURE 8" captionStartId="14.[151,250,1755,1777]" captionTargetBox="[174,1436,447,1721]" captionTargetId="figure@14.[151,1436,447,1733]" captionTargetPageId="14" captionText="FIGURE 8. Right branch of male epiproct in dorso-lateral view. (a) Asiagomphus auricolor male used for description; (b) A. monticola holotype; (c) A. pacificus Cao Bang Prov., 19-V-2015; (d) A. kosterini holotype; (e) A. superciliaris holotype." httpUri="https://zenodo.org/record/1441627/files/figure.png" pageId="4" pageNumber="305">Fig. 8a</figureCitation>
), diverging in same measure as cerci (
<figureCitation id="135A2A4860618C05FA9FD52619E0FBDA" captionStart="FIGURE 7" captionStartId="12.[151,250,1946,1968]" captionTargetBox="[173,1418,193,1934]" captionTargetId="figure@12.[173,1418,193,1934]" captionTargetPageId="12" captionText="FIGURE 7. Male appendages in ventral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. superciliaris paratype Huu Lien NR, 24-V-2014." httpUri="https://zenodo.org/record/1441625/files/figure.png" pageId="4" pageNumber="305">Fig. 7d</figureCitation>
). In lateral view relatively straight, upper margin slightly concave, lower margin somewhat rounded, its lowest point one-third from base, its apical two-thirds becoming thinner towards apex. Slightly longer than cerci, apex a blunt point, hooked inward and upward to form small dorsal tooth (
<figureCitation id="135A2A4860618C05FC61D5BD1AB3FB22" box="[913,997,1159,1184]" captionStart="FIGURE 6" captionStartId="11.[151,250,1945,1967]" captionTargetBox="[201,1402,193,1927]" captionTargetId="figure@11.[188,1403,193,1927]" captionTargetPageId="11" captionText="FIGURE 6. Male appendages in lateral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441623/files/figure.png" pageId="4" pageNumber="305">Fig. 6d</figureCitation>
). Posterior margin of epiproct between branches straight, separation between branches about 1.3x wider than length of protruding part of branches (
<figureCitation id="135A2A4860618C05FA9FD59619E0FB6A" captionStart="FIGURE 7" captionStartId="12.[151,250,1946,1968]" captionTargetBox="[173,1418,193,1934]" captionTargetId="figure@12.[173,1418,193,1934]" captionTargetPageId="12" captionText="FIGURE 7. Male appendages in ventral view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. superciliaris paratype Huu Lien NR, 24-V-2014." httpUri="https://zenodo.org/record/1441625/files/figure.png" pageId="4" pageNumber="305">Fig. 7d</figureCitation>
).
</paragraph>
<paragraph id="8BDE36CD60618C05FF37D5CE180BF9D2" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
<emphasis id="B915EADF60618C05FF37D5CE18FDFA8F" bold="true" box="[199,427,1268,1293]" pageId="4" pageNumber="305">Variation in males.</emphasis>
Two of seven specimens without antehumeral spot and with second yellow lateral stripe over metepisternum reduced to postero-dorsal corner. In two more specimens black stripe over interpleural suture broader, including all of spiracle (
<figureCitation id="135A2A4860618C05FDC2D4061BCDFAD7" box="[562,667,1340,1365]" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [a–d, g] male, [e–f, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="4" pageNumber="305">Fig. 10b</figureCitation>
). Yellow spot on mesokatepisternum somewhat variable in size, metakatepisternum varying from all yellowish brown to all dark brown. In one specimen black on S2 reduced, in another more extensive, forming complete dorso-lateral stripe (
<figureCitation id="135A2A4860618C05FCAFD4BE1AE9FA1E" box="[863,959,1412,1437]" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [a–d, g] male, [e–f, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="4" pageNumber="305">Fig. 10c</figureCitation>
), and in one male black connecting across dorsum along posterior margin. Yellow marking on S9 varying between only posterior one-third to three-fifths of segment. In two specimens tiny lateral yellow mark at posterior margin of S8 (
<figureCitation id="135A2A4860618C05FBE0D4F61D24FA67" box="[1040,1138,1484,1509]" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [a–d, g] male, [e–f, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="4" pageNumber="305">Fig. 10b</figureCitation>
). In one male yellow halfring on S4–7 reduced to dorsal spot. In one male cerci as long as epiproct in lateral view. In one specimen a second crossvein in cubito-anal space in one FW and one HW. 12–18 Px in FW, 16–21 Ax in FW, 12–17 Px in HW and 11–14 Ax in HW.
</paragraph>
<paragraph id="8BDE36CD60618C05FF37D7671D45F9F6" blockId="4.[151,1437,151,2013]" box="[199,1043,1628,1653]" pageId="4" pageNumber="305">Measurements (in mm). Total length (incl. appendages) 60–63, HW 38–41.</paragraph>
<paragraph id="8BDE36CD60618C05FF37D74518F2F8F2" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
<emphasis id="B915EADF60618C05FF37D74518B0F91A" bold="true" box="[199,486,1663,1688]" pageId="4" pageNumber="305">Redescription of female</emphasis>
(
<figureCitation id="135A2A4860618C05FE08D7451B00F91A" box="[504,598,1663,1688]" captionStart="FIGURE 9" captionStartId="16.[151,250,1712,1734]" captionTargetBox="[151,1436,337,1691]" captionTargetId="figure@16.[151,1436,337,1691]" captionTargetPageId="16" captionText="FIGURE 9. Vertex in antero-dorsal view. [a–d] male, [e–f] female, (a) Asiagomphus superciliaris holotype; (b) A. superciliaris paratype Huu Lien NR, 24-V-2014; (c) A. superciliaris paratype Bac Kan Prov., 5-IV-2015; (d) A. monticola holotype; (e) A. auricolor female used for description; (f) A. reinhardti female used for description." httpUri="https://zenodo.org/record/1441629/files/figure.png" pageId="4" pageNumber="305">Figs. 9e</figureCitation>
,
<figureCitation id="135A2A4860618C05FD94D7BA1BC6F91A" box="[612,656,1664,1688]" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [a–d, g] male, [e–f, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="4" pageNumber="305">10e</figureCitation>
). As
<typeStatus id="54DA886F60618C05FD3FD7451A67F91A" box="[719,817,1663,1688]" pageId="4" pageNumber="305" type="holotype">holotype</typeStatus>
male, but raised ridges posterior to lateral ocelli with spike at posterior end, and occipital ridge with central spike (
<figureCitation id="135A2A4860618C05FCB8D79E1ACCF93E" box="[840,922,1700,1725]" captionStart="FIGURE 9" captionStartId="16.[151,250,1712,1734]" captionTargetBox="[151,1436,337,1691]" captionTargetId="figure@16.[151,1436,337,1691]" captionTargetPageId="16" captionText="FIGURE 9. Vertex in antero-dorsal view. [a–d] male, [e–f] female, (a) Asiagomphus superciliaris holotype; (b) A. superciliaris paratype Huu Lien NR, 24-V-2014; (c) A. superciliaris paratype Bac Kan Prov., 5-IV-2015; (d) A. monticola holotype; (e) A. auricolor female used for description; (f) A. reinhardti female used for description." httpUri="https://zenodo.org/record/1441629/files/figure.png" pageId="4" pageNumber="305">Fig. 9e</figureCitation>
). Antehumeral spot reduced on one side (
<figureCitation id="135A2A4860618C05FA9FD79E1996F962" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [a–d, g] male, [e–f, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="4" pageNumber="305">Fig. 10e</figureCitation>
) and absent on other side. Second yellow lateral stripe as wide as black stripe over interpleural suture. S2 completely yellow apart from black postero-dorsal marking covering about one-fourth of dorsum of segment. S3 with only small anterolateral spot and no dorsal spot. S8 with minimal yellow line along anterior margin. S9 with postero-dorsal mark covering about two-fifths of segment. Cerci conical, black, as long as S10. Nodal index 15– 18:19–16/17–12:13–16.
</paragraph>
<paragraph id="8BDE36CD60618C05FF37D6461B46F85F" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
<emphasis id="B915EADF60618C05FF37D64618E9F817" bold="true" box="[199,447,1916,1941]" pageId="4" pageNumber="305">Variation in females.</emphasis>
Based on a photo of a female caught on
<date id="FFDF100D60618C05FC7FD6461D47F817" box="[911,1041,1916,1941]" pageId="4" pageNumber="305" value="2014-04-26">26-IV-2014</date>
(
<figureCitation id="135A2A4860618C05FBD0D6461D28F817" box="[1056,1150,1916,1941]" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [a–d, g] male, [e–f, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="4" pageNumber="305">Fig. 10f</figureCitation>
), S1–2 with broad dorsolateral black line completely bisecting the yellow marking. Thorax with distinct antehumeral spot. Cerci whitish. This is a much fresher individual.
</paragraph>
<caption id="DF1E664560608C04FF67D5E01987FAAF" httpUri="https://zenodo.org/record/1441615/files/figure.png" pageId="5" pageNumber="306" startId="5.[151,250,1242,1264]" targetBox="[151,1437,193,1221]" targetPageId="5">
<paragraph id="8BDE36CD60608C04FF67D5E01987FAAF" blockId="5.[151,1436,1242,1325]" pageId="5" pageNumber="306">
<emphasis id="B915EADF60608C04FF67D5E01845FB6D" bold="true" box="[151,275,1242,1264]" pageId="5" pageNumber="306">FIGURE 2.</emphasis>
<taxonomicName id="4C614D4E60608C04FEEAD5E0188FFB72" box="[282,473,1242,1264]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="acco">
<emphasis id="B915EADF60608C04FEEAD5E0188FFB72" box="[282,473,1242,1264]" italics="true" pageId="5" pageNumber="306">Asiagomphus acco</emphasis>
</taxonomicName>
alive in hand. [a–c] male, [e–f] female. (a) Ba Be NP, 21-VII-2013; (b) Quang Nam Prov., 21- VI-2016; (c) Tam Dao NP, 18-V-2014; (d) Yen Bai Prov., 20-VI-2015; (e) Xuan Son NP, 22-V-2015; (f) Lang Son Prov., 2-V- 2015.
</paragraph>
</caption>
<paragraph id="8BDE36CD60608C04FF37D4611AE8FAF1" blockId="5.[151,1437,1370,2007]" box="[199,958,1370,1395]" pageId="5" pageNumber="306">Measurements (in mm). Total length (incl. appendages) 63, HW 41.</paragraph>
</subSubSection>
<subSubSection id="C37B654660608C07FF37D4441C32FE0A" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306" type="discussion">
<paragraph id="8BDE36CD60608C07FF37D4441802FF57" blockId="5.[151,1437,1370,2007]" lastBlockId="6.[151,1437,151,2013]" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306">
<emphasis id="B915EADF60608C04FF37D444181AFA15" bold="true" box="[199,332,1406,1431]" pageId="5" pageNumber="306">Discussion.</emphasis>
When Fraser described
<taxonomicName id="4C614D4E60608C04FD90D4441A6AFA15" authorityName="Fraser" authorityYear="1926" box="[608,828,1406,1431]" class="Insecta" family="Gomphidae" genus="Gomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60608C04FD90D4441A6AFA15" box="[608,828,1406,1431]" italics="true" pageId="5" pageNumber="306">Gomphus auricolor</emphasis>
</taxonomicName>
, he knew only few similar species (notably
<taxonomicName id="4C614D4E60608C04FAC1D4441CCAFA15" authorityName="Leach" authorityYear="1815" box="[1329,1436,1406,1431]" class="Insecta" family="Gomphidae" genus="Gomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B915EADF60608C04FAC1D4441CCAFA15" box="[1329,1436,1406,1431]" italics="true" pageId="5" pageNumber="306">Gomphus</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C614D4E60608C04FF40D4981811FA39" authorityName="Asahina" authorityYear="1985" box="[176,327,1442,1467]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B915EADF60608C04FF40D4981811FA39" box="[176,327,1442,1467]" italics="true" pageId="5" pageNumber="306">Asiagomphus</emphasis>
</taxonomicName>
)
<taxonomicName id="4C614D4E60608C04FEA8D4981BDEFA39" authority="(Laidlaw, 1922))" authorityName="" authorityYear="1922" baseAuthorityName="Laidlaw" baseAuthorityYear="1922" box="[344,648,1442,1467]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="nilgiricus">
<emphasis id="B915EADF60608C04FEA8D4981895FA39" box="[344,451,1442,1467]" italics="true" pageId="5" pageNumber="306">nilgiricus</emphasis>
<bibRefCitation id="EFF04B3C60608C04FE3BD4981BDEFA39" box="[459,648,1442,1467]" pageId="5" pageNumber="306" refId="ref17499" refString="Laidlaw, F. F. (1922) A list of the dragonflies recorded from the Indian Empire. Part V. The subfamily Gomphinae. Records of the Indian Museum, 24, 367 - 426, 1 pl." type="journal article">(Laidlaw, 1922))</bibRefCitation>
</taxonomicName>
and therefore judged the extensively yellow S1–2 of his female to be sufficient to separate his species novum from all others. “The species belongs to the group
<taxonomicName id="4C614D4E60608C04FB61D4FC1DABFA5D" authorityName="" baseAuthorityName="Laidlaw" baseAuthorityYear="1922" box="[1169,1277,1478,1503]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="nilgiricus">
<emphasis id="B915EADF60608C04FB61D4FC1DABFA5D" box="[1169,1277,1478,1503]" italics="true" pageId="5" pageNumber="306">nilgiricus</emphasis>
</taxonomicName>
by its parallel antehumeral [sic] stripes broadly confluent with the short mesothoracic collar, but is easily distinguished from that and all others of the genus [
<taxonomicName id="4C614D4E60608C04FE3FD7341B6CF9A5" authorityName="Leach" authorityYear="1815" box="[463,570,1550,1575]" class="Insecta" family="Gomphidae" genus="Gomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B915EADF60608C04FE3FD7341B6CF9A5" box="[463,570,1550,1575]" italics="true" pageId="5" pageNumber="306">Gomphus</emphasis>
</taxonomicName>
, TK], by the strikingly broad expanse of bright yellow at base of abdomen, a feature which at once meets the eye.” (
<bibRefCitation id="EFF04B3C60608C04FDA5D7091A6CF9C9" box="[597,826,1586,1611]" pageId="5" pageNumber="306" refId="ref17247" refString="Fraser, F. C. (1926) Notes on a collection of Dragonflies (Order Odonata) from the Dutch East Indies and descriptions of four new species from the neighbouring continent. Treubia, 8, 467 - 494." type="journal article">Fraser 1926, p. 483</bibRefCitation>
). Although the pattern of S1–2 of most
<taxonomicName id="4C614D4E60608C04FAF5D7081CCAF9C9" authorityName="Asahina" authorityYear="1985" box="[1285,1436,1586,1611]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
<emphasis id="B915EADF60608C04FAF5D7081CCAF9C9" box="[1285,1436,1586,1611]" italics="true" pageId="5" pageNumber="306">Asiagomphus</emphasis>
</taxonomicName>
species contains large yellow areas, they almost invariably have longitudinal dorso-lateral black stripes on S2. This is also true for all species from northern
<collectingCountry id="F376765D60608C04FD85D7401B81F911" box="[629,727,1658,1683]" name="Vietnam" pageId="5" pageNumber="306">Vietnam</collectingCountry>
, from where
<taxonomicName id="4C614D4E60608C04FC84D7411D56F911" authorityName="Fraser" authorityYear="1926" box="[884,1024,1658,1683]" class="Insecta" family="Gomphidae" genus="Gomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60608C04FC84D7411D56F911" box="[884,1024,1658,1683]" italics="true" pageId="5" pageNumber="306">A. auricolor</emphasis>
</taxonomicName>
was described, apart from
<emphasis id="B915EADF60608C04FACED7411CCDF910" box="[1342,1435,1659,1682]" italics="true" pageId="5" pageNumber="306">
<taxonomicName id="4C614D4E60608C04FACED7411CC1F910" box="[1342,1431,1659,1682]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="acco">A. acco</taxonomicName>
,
</emphasis>
which has identical S1–2 to Fraser’s description, but which has completely black metepisternum and paired postero-dorsal abdominal spots instead of complete half-rings. As is obvious from the female in
<figureCitation id="135A2A4860608C04FB37D7F81C77F959" box="[1223,1313,1730,1755]" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [a–d, g] male, [e–f, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="5" pageNumber="306">Fig. 10f</figureCitation>
, which has a large dorso-lateral black stripe on S1–2 exactly like the male in
<figureCitation id="135A2A4860608C04FC89D7DC1A8EF97D" box="[889,984,1766,1791]" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [a–d, g] male, [e–f, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="5" pageNumber="306">Fig. 10c</figureCitation>
, even in
<taxonomicName id="4C614D4E60608C04FBCDD7DD1D90F97D" authorityName="Fraser" authorityYear="1926" box="[1085,1222,1766,1791]" class="Insecta" family="Gomphidae" genus="Gomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60608C04FBCDD7DD1D90F97D" box="[1085,1222,1766,1791]" italics="true" pageId="5" pageNumber="306">A. auricolor</emphasis>
</taxonomicName>
almost completely yellow S1–2 is not a consistent character. However, females with almost completely yellow S1–2 are not infrequent. The specimen used for description here is one such specimen. It otherwise also fits the description provided by Fraser, apart from the color of the cerci, which appears to be variable (see
<figureCitation id="135A2A4860608C04FB91D6681DE8F8E9" box="[1121,1214,1874,1899]" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [a–d, g] male, [e–f, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="5" pageNumber="306">Fig. 10f</figureCitation>
). It also on one side has a vestigial antehumeral spot, another variable character, as is clear from the variation in males, in some of which the antehumeral spot is absent and in others present. In the male specimens studied the lack of antehumeral spots occurs in combination with reduced second yellow lateral stripe and darkened and worn wings. The same is true of the female used for description, so it appears likely that the antehumeral spot often becomes obscured in older specimens.
</paragraph>
<paragraph id="8BDE36CD60638C07FF37D1E51C32FE0A" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="307">
There are no other
<taxonomicName id="4C614D4E60638C07FE59D1E51B16FF7A" authorityName="Asahina" authorityYear="1985" box="[425,576,223,248]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B915EADF60638C07FE59D1E51B16FF7A" box="[425,576,223,248]" italics="true" pageId="6" pageNumber="307">Asiagomphus</emphasis>
</taxonomicName>
species in northern
<collectingCountry id="F376765D60638C07FCDED1E51ADBFF7A" box="[814,909,223,248]" name="Vietnam" pageId="6" pageNumber="307">Vietnam</collectingCountry>
that so closely fit the original description by Fraser and have almost completely yellow S1–2. The described female was caught in close association with several males of a species of
<taxonomicName id="4C614D4E60638C07FE7BD01D1B74FEC2" authorityName="Asahina" authorityYear="1985" box="[395,546,295,320]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B915EADF60638C07FE7BD01D1B74FEC2" box="[395,546,295,320]" italics="true" pageId="6" pageNumber="307">Asiagomphus</emphasis>
</taxonomicName>
that occurs widely in northern
<collectingCountry id="F376765D60638C07FC75D01D1AB1FEC2" box="[901,999,295,320]" name="Vietnam" pageId="6" pageNumber="307">Vietnam</collectingCountry>
, but does not fit the description of any other known species. These males too have extensively yellow S1–2 and most often a yellow stripe over the metepisternum. These are here described as the male of
<taxonomicName id="4C614D4E60638C07FCFBD04A1AC2FE0A" authorityName="Fraser" authorityYear="1926" box="[779,916,367,392]" class="Insecta" family="Gomphidae" genus="Gomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60638C07FCFBD04A1AC2FE0A" box="[779,916,367,392]" italics="true" pageId="6" pageNumber="307">A. auricolor</emphasis>
</taxonomicName>
and its differential diagnosis is provided.
</paragraph>
</subSubSection>
<subSubSection id="C37B654660638C06FF37D0AE1BA8FDBE" lastPageId="7" lastPageNumber="308" pageId="6" pageNumber="307" type="diagnosis">
<paragraph id="8BDE36CD60638C07FF37D0AE1C2AFAD7" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="307">
<emphasis id="B915EADF60638C07FF37D0AE1899FE2F" bold="true" box="[199,463,404,429]" pageId="6" pageNumber="307">Differential diagnosis.</emphasis>
The male of
<taxonomicName id="4C614D4E60638C07FD99D0AE1A24FE2F" authorityName=": Fraser" authorityYear="1926" box="[617,882,404,429]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60638C07FD99D0AE1A24FE2F" box="[617,882,404,429]" italics="true" pageId="6" pageNumber="307">Asiagomphus auricolor</emphasis>
</taxonomicName>
must be compared with all known
<taxonomicName id="4C614D4E60638C07FAF5D0AE1CCAFE2F" authorityName="Asahina" authorityYear="1985" box="[1285,1436,404,429]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B915EADF60638C07FAF5D0AE1CCAFE2F" box="[1285,1436,404,429]" italics="true" pageId="6" pageNumber="307">Asiagomphus</emphasis>
</taxonomicName>
species, to ascertain it is not a junior or senior synonym of currently known species. Most Chinese, Korean, and Japanese species can easily be eliminated on the basis of combinations of patterning and structural characters. Notably, most of these species have a distinct complete antehumeral stripe in addition to the dorsal stripe, and have lengthwise dorsal stripes on the abdomen.
<bibRefCitation id="EFF04B3C60638C07FD8BD31E1A5FFDBE" box="[635,777,548,573]" pageId="6" pageNumber="307" refId="ref17086" refString="Chao, H. F. (1990) The Gomphid Dragonflies of China (Odonata: Gomphidae). The Science and Technology Publishing House, Fuzhou, Fujian, 14 + 486 pp." type="book">Chao (1990)</bibRefCitation>
separates
<taxonomicName id="4C614D4E60638C07FC71D31E1DA1FDBE" authority="Liu &amp; Chao, 1990" authorityName="Liu &amp; Chao" authorityYear="1990" box="[897,1271,548,573]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="motouensis">
<emphasis id="B915EADF60638C07FC71D31E1D49FDBE" box="[897,1055,548,572]" italics="true" pageId="6" pageNumber="307">A. motouensis</emphasis>
Liu &amp;
<bibRefCitation id="EFF04B3C60638C07FB84D31E1DA1FDBE" box="[1140,1271,548,573]" pageId="6" pageNumber="307" refId="ref17086" refString="Chao, H. F. (1990) The Gomphid Dragonflies of China (Odonata: Gomphidae). The Science and Technology Publishing House, Fuzhou, Fujian, 14 + 486 pp." type="book">Chao, 1990</bibRefCitation>
</taxonomicName>
from all other species in his key as the only species that combines a black occiput with an antehumeral spot, but lacks an antehumeral stripe. However,
<taxonomicName id="4C614D4E60638C07FE1BD3561BDCFD06" authorityName="Liu &amp; Chao" authorityYear="1990" box="[491,650,620,644]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="motouensis">
<emphasis id="B915EADF60638C07FE1BD3561BDCFD06" box="[491,650,620,644]" italics="true" pageId="6" pageNumber="307">A. motouensis</emphasis>
</taxonomicName>
has a completely different thorax pattern, lacking a continuous black stripe over the intrapleural suture and more slender dorsal stripes. The Japanese endemic species all have a complete antehumeral stripe and three out of four have a longitudinal pattern on the abdomen. The fourth,
<taxonomicName id="4C614D4E60638C07FA74D38E1BDDFD72" authority="(Matsumura in Oguma, 1926)" authorityName="" authorityYear="1926" baseAuthorityName="Matsumura in Oguma" baseAuthorityYear="1926" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="yayeyamensis">
<emphasis id="B915EADF60638C07FA74D38E1867FD72" italics="true" pageId="6" pageNumber="307">A. yayeyamensis</emphasis>
(Matsumura in Oguma, 1926)
</taxonomicName>
, has triangular dorsal spots on the abdomen, lacks a black stripe over the interpleural suture, and its female has extensively black S1–2.
<taxonomicName id="4C614D4E60638C07FC65D3C61D3CFC96" box="[917,1130,764,789]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="acco">
<emphasis id="B915EADF60638C07FC65D3C61D3CFC96" box="[917,1130,764,789]" italics="true" pageId="6" pageNumber="307">Asiagomphus acco</emphasis>
</taxonomicName>
has black abdomen, black metepisternum, and different cerci.
<taxonomicName id="4C614D4E60638C07FDC0D2251A16FCBA" authorityName=": Kosterin &amp; Yokoi" authorityYear="2016" box="[560,832,799,824]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="reinhardti">
<emphasis id="B915EADF60638C07FDC0D2251A16FCBA" box="[560,832,799,824]" italics="true" pageId="6" pageNumber="307">Asiagomphus reinhardti</emphasis>
</taxonomicName>
is easily identified by its protrusion of the posterior margin of S10 and different cerci.
<taxonomicName id="4C614D4E60638C07FDE7D27E1AD7FCDF" authority="Wilson, 2005" authorityName="Wilson" authorityYear="2005" box="[535,897,836,861]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="giza">
<emphasis id="B915EADF60638C07FDE7D27E1BB2FCDE" box="[535,740,836,861]" italics="true" pageId="6" pageNumber="307">Asiagomphus giza</emphasis>
<bibRefCitation id="EFF04B3C60638C07FD1BD27E1AD7FCDF" box="[747,897,836,861]" pageId="6" pageNumber="307" refId="ref17824" refString="Wilson, K. D. P. (2005) Odonata of Guangxi Zhuang Autonomous Region, China, part II: Anisoptera. International Journal of Odonatology, 8, 107 - 168. https: // doi. org / 10.1080 / 13887890.2005.9748247" type="journal article">Wilson, 2005</bibRefCitation>
</taxonomicName>
is only known from a female, but has a peculiar triple horn on the occiput, lacking in the female used for description, and has longitudinal markings on the abdomen. The western species
<taxonomicName id="4C614D4E60638C07FDF6D2B61A0FFC26" authority="(Laidlaw, 1922)" authorityName="" authorityYear="1922" baseAuthorityName="Laidlaw" baseAuthorityYear="1922" box="[518,857,908,933]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="nilgiricus">
<emphasis id="B915EADF60638C07FDF6D2B61BC0FC27" box="[518,662,908,933]" italics="true" pageId="6" pageNumber="307">A. nilgiricus</emphasis>
(
<bibRefCitation id="EFF04B3C60638C07FD59D2B61A06FC26" box="[681,848,908,933]" pageId="6" pageNumber="307" refId="ref17499" refString="Laidlaw, F. F. (1922) A list of the dragonflies recorded from the Indian Empire. Part V. The subfamily Gomphinae. Records of the Indian Museum, 24, 367 - 426, 1 pl." type="journal article">Laidlaw, 1922</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C614D4E60638C07FC68D2B61D89FC26" authority="(Selys, 1873)" baseAuthorityName="Selys" baseAuthorityYear="1873" box="[920,1247,908,933]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="personatus">
<emphasis id="B915EADF60638C07FC68D2B61D61FC26" box="[920,1079,908,932]" italics="true" pageId="6" pageNumber="307">A. personatus</emphasis>
(Selys, 1873)
</taxonomicName>
, whether or not synonyms (
<bibRefCitation id="EFF04B3C60638C07FEE8D28A18F5FC4A" box="[280,419,944,968]" pageId="6" pageNumber="307" refId="ref17290" refString="Fraser, F. C. (1934) The Fauna of British India, including Ceylon and Burma. Odonata. Vol. II. Taylor and Francis, London, xxiii + 398 pp., col. pls. I - IV excl." type="book">Fraser 1934</bibRefCitation>
), have longitudinal markings on the abdomen (
<bibRefCitation id="EFF04B3C60638C07FC48D28A1D17FC4A" box="[952,1089,943,968]" pageId="6" pageNumber="307" refId="ref17215" refString="Fraser, F. C. (1925) Indian Dragonflies, Part XXI. Journal of the Bombay Natural History Society, 30, 657 - 663, 1 pl." type="journal article">Fraser 1925</bibRefCitation>
) and
<taxonomicName id="4C614D4E60638C07FB71D28A1CCAFC4A" authority="(Fraser, 1922)" baseAuthorityName="Fraser" baseAuthorityYear="1922" box="[1153,1436,943,968]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="odoneli">
<emphasis id="B915EADF60638C07FB71D28A1DA3FC4A" box="[1153,1269,943,968]" italics="true" pageId="6" pageNumber="307">A. odoneli</emphasis>
(Fraser, 1922)
</taxonomicName>
has differently marked S9, without postero-dorsal yellow spot, but with lateral markings instead.
<taxonomicName id="4C614D4E60638C07FAF5D2EE1B53FB92" authority="(Williamson, 1907)" baseAuthorityName="Williamson" baseAuthorityYear="1907" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="xanthenatus">
<emphasis id="B915EADF60638C07FAF5D2EE1848FB92" italics="true" pageId="6" pageNumber="307">Asiagomphus xanthenatus</emphasis>
(
<bibRefCitation id="EFF04B3C60638C07FEC0D2CD18ABFB92" box="[304,509,1015,1040]" pageId="6" pageNumber="307" refId="ref17719" refString="Williamson, E. B. (1907) The dragonflies (Odonata) of Burma and Lower Siam. II. Subfamilies Cordulegasterinae, Chlorogomphinae, and Gomphinae. Proceedings U. S. National Museum, 33, 267 - 317. https: // doi. org / 10.5479 / si. 00963801.33 - 1571.267" type="journal article">Williamson, 1907</bibRefCitation>
)
</taxonomicName>
is closer in appearance to
<taxonomicName id="4C614D4E60638C07FCB0D2C21A9DFB92" authorityName=": Fraser" authorityYear="1926" box="[832,971,1015,1040]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60638C07FCB0D2C21A9DFB92" box="[832,971,1015,1040]" italics="true" pageId="6" pageNumber="307">A. auricolor</emphasis>
</taxonomicName>
, with half-rings at the base of abdomen segments, but its epiproct has widely diverging branches separated by its semicircular posterior margin (
<figureCitation id="135A2A4860638C07FACBD5261CD8FBB7" box="[1339,1422,1052,1077]" captionStart="FIGURE 5" captionStartId="10.[151,250,1939,1961]" captionTargetBox="[191,1400,193,1919]" captionTargetId="figure@10.[191,1400,193,1920]" captionTargetPageId="10" captionText="FIGURE 5. Male appendages in dorsal view. (a) Asiagomphus monticola holotype; (b) A. pacificus Cao Bang Prov., 19-V- 2015; (c) A. superciliaris holotype; (d) A. auricolor male used for description; (e) A. xanthenatus Quang Nam Prov., 15-V- 2016; (f) A. kosterini holotype; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. acco, Tam Dao NP, 18-V-2014." httpUri="https://zenodo.org/record/1441621/files/figure.png" pageId="6" pageNumber="307">Fig. 5e</figureCitation>
), and the apical hook of the posterior hamulus is directed more anteriorly, not inwardly, whereas the anterior margin of the posterior hamulus is clearly angular with small teeth, not smooth (
<figureCitation id="135A2A4860638C07FC25D55E1D7EFBFE" box="[981,1064,1124,1149]" captionStart="FIGURE 3" captionStartId="8.[151,250,1943,1965]" captionTargetBox="[182,1417,193,1925]" captionTargetId="figure@8.[182,1417,193,1925]" captionTargetPageId="8" captionText="FIGURE 3. Anterior and posterior hamuli [a,c,e,g,h] lateral view, [b,d,f] antero-ventral lateral view. (a–b) Asiagomphus acco, Tam Dao NP, 18-V-2014; (c–d) A. kosterini sp. nov. holotype; (e–f) A. xanthenatus Quang Nam Prov., 15-V-2016; (g) A. reinhardti Lam Dong Prov., 10-V-2017; (h) A. kosterini paratype." httpUri="https://zenodo.org/record/1441617/files/figure.png" pageId="6" pageNumber="307">Fig. 3e</figureCitation>
).
<taxonomicName id="4C614D4E60638C07FBCDD55E1805FB22" authority="(Doi &amp; Okumura, 1937)" baseAuthorityName="Doi &amp; Okumura" baseAuthorityYear="1937" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="coreanus">
<emphasis id="B915EADF60638C07FBCDD55E1C14FBFE" box="[1085,1346,1124,1149]" italics="true" pageId="6" pageNumber="307">Asiagomphus coreanus</emphasis>
(Doi &amp; Okumura, 1937)
</taxonomicName>
, endemic for the Korean Peninsula, has complete antehumeral stripe and interrupted black stripe over interpleural suture, and different appendages, whereas
<taxonomicName id="4C614D4E60638C07FCC3D5961AA1FB47" authorityName="Doi" authorityYear="1943" box="[819,1015,1196,1221]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="melanopsoides">
<emphasis id="B915EADF60638C07FCC3D5961AA1FB47" box="[819,1015,1196,1221]" italics="true" pageId="6" pageNumber="307">A. melanopsoides</emphasis>
</taxonomicName>
from
<collectingCountry id="F376765D60638C07FBCCD5971DD6FB46" box="[1084,1152,1197,1220]" name="South Korea" pageId="6" pageNumber="307">Korea</collectingCountry>
and the Russian Far East also has complete antehumeral stripe and longitudinal stripe over the dorsum of the abdomen.
<taxonomicName id="4C614D4E60638C07FAF5D5F5186AFA8F" authorityName="Yang, Mao &amp; Zhang" authorityYear="2006" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="gongshanensis">
<emphasis id="B915EADF60638C07FAF5D5F5186AFA8F" italics="true" pageId="6" pageNumber="307">Asiagomphus gongshanensis</emphasis>
</taxonomicName>
was described from a female specimen, and is said to resemble
<taxonomicName id="4C614D4E60638C07FBF7D5CE1DF7FA8E" baseAuthorityName="Selys" baseAuthorityYear="1873" box="[1031,1185,1268,1292]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="personatus">
<emphasis id="B915EADF60638C07FBF7D5CE1DF7FA8E" box="[1031,1185,1268,1292]" italics="true" pageId="6" pageNumber="307">A. personatus</emphasis>
</taxonomicName>
. The poorly known
<taxonomicName id="4C614D4E60638C07FA74D5CE18E4FAB2" authority="(Morton, 1928)" baseAuthorityName="Morton" baseAuthorityYear="1928" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="corniger">
<emphasis id="B915EADF60638C07FA74D5CE19A1FAB2" italics="true" pageId="6" pageNumber="307">A. corniger</emphasis>
(Morton, 1928)
</taxonomicName>
has completely different anal appendages. Indeed,
<bibRefCitation id="EFF04B3C60638C07FBF9D42D1DCFFAB2" box="[1033,1177,1303,1328]" pageId="6" pageNumber="307" refId="ref17086" refString="Chao, H. F. (1990) The Gomphid Dragonflies of China (Odonata: Gomphidae). The Science and Technology Publishing House, Fuzhou, Fujian, 14 + 486 pp." type="book">Chao (1990)</bibRefCitation>
chose to include it in
<taxonomicName id="4C614D4E60638C07FF67D4061B4CFAD6" authority="Williamson, 1907" authorityName="Williamson" authorityYear="1907" box="[151,538,1340,1365]" class="Insecta" family="Gomphidae" genus="Burmagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B915EADF60638C07FF67D406181EFAD7" box="[151,328,1340,1365]" italics="true" pageId="6" pageNumber="307">Burmagomphus</emphasis>
<bibRefCitation id="EFF04B3C60638C07FEBFD4061B4CFAD6" box="[335,538,1340,1365]" pageId="6" pageNumber="307" refId="ref17719" refString="Williamson, E. B. (1907) The dragonflies (Odonata) of Burma and Lower Siam. II. Subfamilies Cordulegasterinae, Chlorogomphinae, and Gomphinae. Proceedings U. S. National Museum, 33, 267 - 317. https: // doi. org / 10.5479 / si. 00963801.33 - 1571.267" type="journal article">Williamson, 1907</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C614D4E60638C07FDD4D4061A6FFAD7" baseAuthorityName="Needham" baseAuthorityYear="1930" box="[548,825,1340,1365]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="somnolens">
<emphasis id="B915EADF60638C07FDD4D4061A6FFAD7" box="[548,825,1340,1365]" italics="true" pageId="6" pageNumber="307">Asiagomphus somnolens</emphasis>
</taxonomicName>
(only known from a female) has all yellow labrum.
</paragraph>
<paragraph id="8BDE36CD60638C07FF37D45B1C11F817" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="307">
From the above it is clear that the taxon here described as the male of
<taxonomicName id="4C614D4E60638C07FC26D4651D88FAFA" authorityName=": Fraser" authorityYear="1926" box="[982,1246,1375,1400]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60638C07FC26D4651D88FAFA" box="[982,1246,1375,1400]" italics="true" pageId="6" pageNumber="307">Asiagomphus auricolor</emphasis>
</taxonomicName>
is different from all known
<taxonomicName id="4C614D4E60638C07FEFED4BE18F3FA1F" authorityName="Asahina" authorityYear="1985" box="[270,421,1412,1437]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B915EADF60638C07FEFED4BE18F3FA1F" box="[270,421,1412,1437]" italics="true" pageId="6" pageNumber="307">Asiagomphus</emphasis>
</taxonomicName>
species with known males, and differs in color pattern from the species known only from females as well. It is also easily separated from the three new species described in this paper.
<taxonomicName id="4C614D4E60638C07FAF5D49D1851FA67" authorityName="Kompier" authorityYear="2018" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B915EADF60638C07FAF5D49D1851FA67" italics="true" pageId="6" pageNumber="307">Asiagomphus monticola</emphasis>
</taxonomicName>
and
<taxonomicName id="4C614D4E60638C07FEB2D4F718A7FA67" authorityName="Kompier" authorityYear="2018" box="[322,497,1484,1509]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="superciliaris">
<emphasis id="B915EADF60638C07FEB2D4F718A7FA67" box="[322,497,1484,1509]" italics="true" pageId="6" pageNumber="307">A. superciliaris</emphasis>
</taxonomicName>
are much darker and have different posterior and anterior hamuli, and differently shaped epiproct.
<taxonomicName id="4C614D4E60638C07FEA4D4D51B05F98A" authorityName="Kompier" authorityYear="2018" box="[340,595,1519,1544]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="kosterini">
<emphasis id="B915EADF60638C07FEA4D4D51B05F98A" box="[340,595,1519,1544]" italics="true" pageId="6" pageNumber="307">Asiagomphus kosterini</emphasis>
</taxonomicName>
has completely different abdomen pattern and different anal and secondary appendages. The identification of the female is a challenge. It is not clear how consistent the combination of horns on the cranium and occiput of female
<taxonomicName id="4C614D4E60638C07FDBCD70D1BB5F9D2" authorityName="Asahina" authorityYear="1985" box="[588,739,1591,1616]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="B915EADF60638C07FDBCD70D1BB5F9D2" box="[588,739,1591,1616]" italics="true" pageId="6" pageNumber="307">Asiagomphus</emphasis>
</taxonomicName>
species is, but my experience with for instance
<taxonomicName id="4C614D4E60638C07FAFCD7021CCDF9D2" authorityName=": Kosterin &amp; Yokoi" authorityYear="2016" box="[1292,1435,1591,1616]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="reinhardti">
<emphasis id="B915EADF60638C07FAFCD7021CCDF9D2" box="[1292,1435,1591,1616]" italics="true" pageId="6" pageNumber="307">A. reinhardti</emphasis>
</taxonomicName>
(see also under that species) indicates it may well vary considerably between females of the same species. More specimens must be studied to establish the range of variability. However, from my personal observations it appears that also female specimens only tentatively identified as
<taxonomicName id="4C614D4E60638C07FCECD79E1AF0F93F" authorityName=": Fraser" authorityYear="1926" box="[796,934,1700,1725]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60638C07FCECD79E1AF0F93F" box="[796,934,1700,1725]" italics="true" pageId="6" pageNumber="307">A. auricolor</emphasis>
</taxonomicName>
(some specimens could not be conclusively identified, because of differences with the original description by Fraser and the fact they were not caught in copula with the
<typeStatus id="54DA886F60638C07FF0AD7D7187CF887" box="[250,298,1773,1797]" pageId="6" pageNumber="307">type</typeStatus>
of male described here as male
<taxonomicName id="4C614D4E60638C07FD68D7D61A77F887" authorityName=": Fraser" authorityYear="1926" box="[664,801,1772,1797]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60638C07FD68D7D61A77F887" box="[664,801,1772,1797]" italics="true" pageId="6" pageNumber="307">A. auricolor</emphasis>
</taxonomicName>
) sometimes lack a central spike on the occiput and may have additional spikes adjacent to the lateral ocelli. Cerci may be pale or black (
<figureCitation id="135A2A4860638C07FBC7D6351DEBF8AA" box="[1079,1213,1807,1832]" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [a–d, g] male, [e–f, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="6" pageNumber="307">Figs. 10e–f</figureCitation>
). Antehumeral spot may be present or absent (as in the male). Currently, only females with almost completely yellow S1–2, dorsal halfrings on abdomen segments and at least some pale markings on the metepisternum can be positively identified as
<taxonomicName id="4C614D4E60638C07FF67D6461848F817" authorityName=": Fraser" authorityYear="1926" box="[151,286,1916,1941]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60638C07FF67D6461848F817" box="[151,286,1916,1941]" italics="true" pageId="6" pageNumber="307">A. auricolor</emphasis>
</taxonomicName>
. The female in
<figureCitation id="135A2A4860638C07FE3FD6461B71F817" box="[463,551,1916,1941]" captionStart="FIGURE 10" captionStartId="19.[151,250,1872,1894]" captionTargetBox="[150,1436,481,1851]" captionTargetId="figure@19.[150,1436,481,1851]" captionTargetPageId="19" captionText="FIGURE 10. Asiagomphus spp. alive in hand [a–d, g] male, [e–f, h] female. (a) A. auricolor, Bac Kan Prov., 26-VI-2014; (b) A. auricolor, Huu Lien NR, 1-V-2014; (c) A. auricolor, Huu Lien NR, 24-V-2014; (d) A. xanthenatus, Quang Nam Prov., 15-V- 2016; (e) A. auricolor, used for description; (f) A. auricolor, Huu Lien NR, 26-IV-2014; (g) A. pacificus, Cao Bang Prov., 19-V- 2015; (h) A. pacificus, Bac Kan Prov., 25-VI-2014." httpUri="https://zenodo.org/record/1441631/files/figure.png" pageId="6" pageNumber="307">Fig. 10f</figureCitation>
was also caught in close association with typical males of
<emphasis id="B915EADF60638C07FB4AD6461C11F817" box="[1210,1351,1916,1941]" italics="true" pageId="6" pageNumber="307">
<taxonomicName id="4C614D4E60638C07FB4AD6461C15F817" authorityName=": Fraser" authorityYear="1926" box="[1210,1347,1916,1941]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="auricolor">A. auricolor</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8BDE36CD60638C06FF37D69B1B67FF7A" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1436,151,573]" lastPageId="7" lastPageNumber="308" pageId="6" pageNumber="307">
I am confident that the combination of females from northern
<collectingCountry id="F376765D60638C07FC76D69A1AB3F83B" box="[902,997,1952,1977]" name="Vietnam" pageId="6" pageNumber="307">Vietnam</collectingCountry>
that closely fit the original description of
<taxonomicName id="4C614D4E60638C07FF46D6FE18E9F85F" authorityName=": Fraser" authorityYear="1926" box="[182,447,1988,2013]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60638C07FF46D6FE18E9F85F" box="[182,447,1988,2013]" italics="true" pageId="6" pageNumber="307">Asiagomphus auricolor</emphasis>
</taxonomicName>
from
<collectingCountry id="F376765D60638C07FDF4D6FE1B05F85F" box="[516,595,1988,2013]" name="Vietnam" pageId="6" pageNumber="307">Tonkin</collectingCountry>
(present day northern
<collectingCountry id="F376765D60638C07FCA3D6FE1AE2F85F" box="[851,948,1988,2013]" name="Vietnam" pageId="6" pageNumber="307">Vietnam</collectingCountry>
) and the common occurrence of males that are clearly different from all currently known species and can be considered topotypical with
<taxonomicName id="4C614D4E60628C06FB45D1A21C69FF32" authorityName=": Fraser" authorityYear="1926" box="[1205,1343,151,176]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60628C06FB45D1A21C69FF32" box="[1205,1343,151,176]" italics="true" pageId="7" pageNumber="308">A. auricolor</emphasis>
</taxonomicName>
provide a good enough case to consider this taxon as the elusive
<emphasis id="B915EADF60628C06FCD5D1861AE2FF57" box="[805,948,188,213]" italics="true" pageId="7" pageNumber="308">
<taxonomicName id="4C614D4E60628C06FCD5D1861AE6FF57" authorityName=": Fraser" authorityYear="1926" box="[805,944,188,213]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="auricolor">A. auricolor</taxonomicName>
,
</emphasis>
rather than describing it as a new species, which would be the only alternative.
</paragraph>
<paragraph id="8BDE36CD60628C06FF37D03E1BEBFE2F" blockId="7.[151,1436,151,573]" pageId="7" pageNumber="308">
<emphasis id="B915EADF60628C06FF37D03E18EDFE9F" bold="true" box="[199,443,260,285]" pageId="7" pageNumber="308">Habitat and ecology.</emphasis>
<taxonomicName id="4C614D4E60628C06FE32D03E1B9FFE9F" authorityName=": Fraser" authorityYear="1926" box="[450,713,260,285]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="auricolor">
<emphasis id="B915EADF60628C06FE32D03E1B9FFE9F" box="[450,713,260,285]" italics="true" pageId="7" pageNumber="308">Asiagomphus auricolor</emphasis>
</taxonomicName>
is a rather common species found at a variety of smaller streams in semi-open areas, where they generally perch on vegetation by the side of the streams. It is a difficult species to approach and quickly alights when disturbed, flying up into tree cover. They start emerging from the middle of March as one of the earliest species in northern
<collectingCountry id="F376765D60628C06FD44D0551A40FE0A" box="[692,790,367,392]" name="Vietnam" pageId="7" pageNumber="308">Vietnam</collectingCountry>
, together with for instance
<taxonomicName id="4C614D4E60628C06FBBAD0551985FE2F" authority="Karube, 2015" authorityName="Karube" authorityYear="2015" class="Insecta" family="Gomphidae" genus="Trigomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="kompieri">
<emphasis id="B915EADF60628C06FBBAD0551C6DFE0A" box="[1098,1339,367,392]" italics="true" pageId="7" pageNumber="308">Trigomphus kompieri</emphasis>
Karube, 2015
</taxonomicName>
. By the beginning of July only few survive.
</paragraph>
<paragraph id="8BDE36CD60628C06FF37D08D1BA8FDBE" blockId="7.[151,1436,151,573]" pageId="7" pageNumber="308">
<bibRefCitation id="EFF04B3C60628C06FF37D08D1B26FE52" box="[199,624,439,464]" pageId="7" pageNumber="308" refId="ref17573" refString="Reels, G. &amp; Zhang, H. (2015) A Field Guide to the Dragonflies of Hainan. China Forestry Publishing House, Beijing, 463 pp." type="book">Reels &amp; Zhang (2015, pp. 276–277)</bibRefCitation>
illustrate a similar looking species from
<collectingRegion id="49A5F82F60628C06FBA5D08D1DF0FE52" box="[1109,1190,439,464]" country="China" name="Hainan" pageId="7" pageNumber="308">Hainan</collectingRegion>
as
<taxonomicName id="4C614D4E60628C06FB23D08D1C3CFE52" authorityName="Asahina" authorityYear="1985" box="[1235,1386,439,464]" class="Insecta" family="Gomphidae" genus="Asiagomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B915EADF60628C06FB23D08D1C3CFE52" box="[1235,1386,439,464]" italics="true" pageId="7" pageNumber="308">Asiagomphus</emphasis>
</taxonomicName>
sp., although it has larger lateral markings at the posterior margin of S8 and yellow spots on the labrum. The description mentions (p. 277) that “Males perch on the surface of leaves, close to the stream. Very easily disturbed.” It will be interesting to compare these taxa.
</paragraph>
</subSubSection>
</treatment>
</document>
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