358 lines
41 KiB
XML
358 lines
41 KiB
XML
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<mods:title id="0186D42B28584812909DC17125F48165">Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J / K boundary</mods:title>
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<mods:namePart id="DF01DE8744AE0A383FE85B6AD786E013">Allain, Ronan</mods:namePart>
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<mods:namePart id="430A55FE412E62A26E27B18E457E1067">Vullo, Romain</mods:namePart>
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<mods:namePart id="ACE091D3908D8B59BDC0066408C8EAB8">Rozada, Lee</mods:namePart>
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<mods:namePart id="32F4136D0EB630904235F74A16EB4D91">Bourgeais, Renaud</mods:namePart>
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<mods:namePart id="9C1956476C549F8E762B015933266B8F">Fabrègues, Claire Peyre De</mods:namePart>
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<mods:namePart id="A3D70AE1D10C701732B6BE457A207559">Royo-Torres, Rafael</mods:namePart>
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<mods:namePart id="E27DA1D1A4C322FB4A26A66AC07CB65B">Augier, Dominique</mods:namePart>
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<mods:namePart id="2B701B5A63344B644590DF960255F329">Bailly, Gilles</mods:namePart>
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<mods:title id="9304EE4EFEFB33F2D480F7823EE14B89">Geodiversitas</mods:title>
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<mods:part id="E0814D6019AF32ECFDD08824C82AD8EA">
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<mods:date id="D43270EC6EC90F77C67885CF47DF1F75">2022</mods:date>
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<mods:number id="A3BE01284DAA0E67D45C148C539E7AC4">2022-07-21</mods:number>
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<mods:number id="D4C7C7B5C0ACDDCDE6728B0ECC4934B8">44</mods:number>
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<mods:start id="A48C6292CB5E692CFA1C14D7B85F90B4">683</mods:start>
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<mods:identifier id="9EED259590694E817DEE7611525BCEB7" type="CLB-Dataset">107849</mods:identifier>
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<mods:identifier id="E09061C72D352B00A4DE1A27E5E26705" type="DOI">10.5252/geodiversitas2022v44a25</mods:identifier>
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<treatment id="03C887B9FFA6FFF876C9A509FD6AF85D" ID-DOI="http://doi.org/10.5281/zenodo.6928883" ID-GBIF-Taxon="197866714" ID-Zenodo-Dep="6928883" LSID="urn:lsid:plazi:treatment:03C887B9FFA6FFF876C9A509FD6AF85D" httpUri="http://treatment.plazi.org/id/03C887B9FFA6FFF876C9A509FD6AF85D" lastPageId="19" lastPageNumber="700" pageId="16" pageNumber="697">
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<subSubSection id="C37B6524FFA6FFFB76C9A509FB01F8FA" box="[1011,1258,1841,1867]" pageId="16" pageNumber="697" type="nomenclature">
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<paragraph id="8BDE36AFFFA6FFFB76C9A509FB01F8FA" blockId="16.[1011,1258,1841,1900]" box="[1011,1258,1841,1867]" pageId="16" pageNumber="697">
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<taxonomicName id="4C614D2CFFA6FFFB76C9A509FB4FF8FA" authorityName="Nopcsa" authorityYear="1928" box="[1011,1188,1841,1867]" family="Helochelydridae" kingdom="Animalia" pageId="16" pageNumber="697" rank="family">Helochelydridae</taxonomicName>
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indet.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37B6524FFA6FFF8711DA56AFD6AF85D" lastPageId="19" lastPageNumber="700" pageId="16" pageNumber="697" type="description">
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<paragraph id="8BDE36AFFFA6FFFB711DA56AFB5EF8DD" blockId="16.[1011,1258,1841,1900]" box="[1063,1205,1873,1900]" pageId="16" pageNumber="697">
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(
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<figureCitation id="135A2A2AFFA6FFFB710AA569FB46F8DD" box="[1072,1197,1873,1900]" captionStart="FIG" captionStartId="18.[132,143,1940,1957]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-29@18.[698,1267,768,1324]" captionTargetPageId="18" captionText="FIG. 12. — Helochelydrid and thalassochelydian turtle remains from Angeac-Charente: A, peripheral plate of Helochelydridae indet. in dorsal view;B, pleural plate of Helochelydridae indet. in dorsal view; C, D, right hyoplastron of Hylaeochelys belli? (Mantell, 1844) (ANG18-5915) in ventral (C) and dorsal (D) views; E, left humerus of Hylaeochelys? belli? (ANG18-5937) in ventral view; F, right humerus (ANG18-5938) in dorsal view;; G, H, Peripheral of Hylaeochelys? belli? (ANG 18-5993) in dorsal (G) and ventral (H) views. Scale bar: A, 1 cm; E-H, 2 cm; B, 3 cm; C, D, 4 cm." figureDoi="http://doi.org/10.5281/zenodo.6902061" httpUri="https://zenodo.org/record/6902061/files/figure.png" pageId="16" pageNumber="697">Fig. 12A, B</figureCitation>
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)
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</paragraph>
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<paragraph id="8BDE36AFFFA6FFFB7617A5AAFC56F81D" blockId="16.[811,1455,1938,2027]" box="[813,957,1938,1964]" pageId="16" pageNumber="697">DESCRIPTION</paragraph>
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<paragraph id="8BDE36AFFFA6FFF87617A58AFD4CFC9E" blockId="16.[811,1455,1938,2027]" lastBlockId="19.[131,777,215,2028]" lastPageId="19" lastPageNumber="700" pageId="16" pageNumber="697">
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In Angeac-Charente, helochelydrids are mostly represented by incomplete isolated shell elements covered by typical tubercles, pustules and crests (
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<figureCitation id="135A2A2AFFA5FFF874E6A2EFFD8BFF43" box="[476,608,215,242]" captionStart="FIG" captionStartId="18.[132,143,1940,1957]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-29@18.[698,1267,768,1324]" captionTargetPageId="18" captionText="FIG. 12. — Helochelydrid and thalassochelydian turtle remains from Angeac-Charente: A, peripheral plate of Helochelydridae indet. in dorsal view;B, pleural plate of Helochelydridae indet. in dorsal view; C, D, right hyoplastron of Hylaeochelys belli? (Mantell, 1844) (ANG18-5915) in ventral (C) and dorsal (D) views; E, left humerus of Hylaeochelys? belli? (ANG18-5937) in ventral view; F, right humerus (ANG18-5938) in dorsal view;; G, H, Peripheral of Hylaeochelys? belli? (ANG 18-5993) in dorsal (G) and ventral (H) views. Scale bar: A, 1 cm; E-H, 2 cm; B, 3 cm; C, D, 4 cm." figureDoi="http://doi.org/10.5281/zenodo.6902061" httpUri="https://zenodo.org/record/6902061/files/figure.png" pageId="19" pageNumber="700">Fig. 12A, B</figureCitation>
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) that diagnose this clade (
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<bibRefCitation id="EFF04B5EFFA5FFF8743EA2C0FD50FEA0" author="Lapparent de Broin & Murelaga" box="[260,699,247,274]" firstAuthor="Lapparent de Broin" pageId="19" pageNumber="700" pagination="135 - 211" refId="ref42274" refString="LAPPARENT DE BROIN F. DE & MURELAGA X. 1999. - Turtles from the Upper Cretaceous of Lano (Iberian Peninsula). Estudios del Museo de Ciencias Naturales de Alava 14: 135 - 211." type="journal article" year="1999">Lapparent de Broin & Murelaga 1999</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFA5FFF877F6A2C0FF2BFE80" author="Joyce" firstAuthor="Joyce" pageId="19" pageNumber="700" pagination="65 - 113" refId="ref41684" refString="JOYCE W. G. 2017. - A Review of the Fossil Record of Basal Mesozoic Turtles. Bulletin of the Peabody Museum of Natural History 58: 65 - 113. https: // doi. org / 10.3374 / 014.058.0105" type="journal article" year="2017">Joyce 2017</bibRefCitation>
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). Probably based on the previous stratigraphical claim by
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<bibRefCitation id="EFF04B5EFFA5FFF875C8A30FFDE8FEE0" author="Neraudeau" box="[242,515,311,337]" etAl="et al." firstAuthor="Neraudeau" pageId="19" pageNumber="700" pagination="1 - 14" refId="ref44595" refString="NERAUDEAU D., ALLAIN R., BALLEVRE M., BATTEN D. J., BUFFE- TAUT E., COLIN J. P., DABARD M. P., DAVIERO- GOMEZ V., EL ALBANI A., GOMEZ B., GROSHENY D., LE LoeUFF J., LEPRINCE A., MARTIN- CLOSAS C., MASURE E., MAZIN J. - M., PHILIPPE M., POUECH J., TONG H., TOURNEPICHE J. - F. & VULLO R. 2012. - The Hauterivian-Barremian lignitic bone bed of Angeac (Charente, SW France): stratigraphical, palaeobiological and palaeogeographical implications. Cretaceous Research 37: 1 - 14. https: // doi. org / 10.1016 / j. cretres. 2012.01.006" type="journal article" year="2012">
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Néraudeau
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<emphasis id="B915EABDFFA5FFF8744FA300FE46FEE0" box="[373,429,311,337]" italics="true" pageId="19" pageNumber="700">et al.</emphasis>
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(2012)
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</bibRefCitation>
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that Angeac-Charente was Hauterivian-Barremian in age,
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<bibRefCitation id="EFF04B5EFFA5FFF8771EA36FFD53FEC0" author="Joyce" box="[548,696,343,369]" firstAuthor="Joyce" pageId="19" pageNumber="700" pagination="65 - 113" refId="ref41684" refString="JOYCE W. G. 2017. - A Review of the Fossil Record of Basal Mesozoic Turtles. Bulletin of the Peabody Museum of Natural History 58: 65 - 113. https: // doi. org / 10.3374 / 014.058.0105" type="journal article" year="2017">Joyce (2017)</bibRefCitation>
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provisionally referred this material to
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<taxonomicName id="4C614D2CFFA5FFF8773EA34FFD02FE21" authorityName="Lapparent de Broin & Murelaga" authorityYear="1999" box="[516,745,375,401]" class="Reptilia" family="Solemydidae" genus="Helochelydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="700" phylum="Chordata" rank="species" species="nopcsai">
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<emphasis id="B915EABDFFA5FFF8773EA34FFD02FE21" box="[516,745,375,401]" italics="true" pageId="19" pageNumber="700">Helochelydra nopcsai</emphasis>
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</taxonomicName>
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, a species otherwise diagnosed by a shell surface ornamentation consisting of distinct, easily dislocated tubercles. Our observations reveal, however, that the helochelydrid material from Angeac-Charente usually displays distinct tubercles that do not easily dislocate. This ornamentation is more similar to the morphotypes of “
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<taxonomicName id="4C614D2CFFA5FFF874D0A00EFD9FFDE1" authorityName="Nopcsa" authorityYear="1928" box="[490,628,566,592]" class="Reptilia" family="Solemydidae" genus="Helochelydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="700" phylum="Chordata" rank="genus">
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<emphasis id="B915EABDFFA5FFF874D0A00EFD9FFDE1" box="[490,628,566,592]" italics="true" pageId="19" pageNumber="700">Helochelydra</emphasis>
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</taxonomicName>
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”
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<emphasis id="B915EABDFFA5FFF877B2A00EFD3EFDE1" box="[648,725,566,592]" italics="true" pageId="19" pageNumber="700">anglica</emphasis>
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and “
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<taxonomicName id="4C614D2CFFA5FFF875AAA06EFEF0FDC1" authorityName="Nopcsa" authorityYear="1928" box="[144,283,598,624]" class="Reptilia" family="Solemydidae" genus="Helochelydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="700" phylum="Chordata" rank="genus">
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<emphasis id="B915EABDFFA5FFF875AAA06EFEF0FDC1" box="[144,283,598,624]" italics="true" pageId="19" pageNumber="700">Helochelydra</emphasis>
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</taxonomicName>
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”
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<emphasis id="B915EABDFFA5FFF8740BA06EFE7CFDC1" box="[305,407,598,624]" italics="true" pageId="19" pageNumber="700">bakewelli</emphasis>
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(
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<bibRefCitation id="EFF04B5EFFA5FFF87491A06EFDC2FDC1" author="Joyce" box="[427,553,598,624]" firstAuthor="Joyce" pageId="19" pageNumber="700" pagination="65 - 113" refId="ref41684" refString="JOYCE W. G. 2017. - A Review of the Fossil Record of Basal Mesozoic Turtles. Bulletin of the Peabody Museum of Natural History 58: 65 - 113. https: // doi. org / 10.3374 / 014.058.0105" type="journal article" year="2017">Joyce 2017</bibRefCitation>
|
|
), respectively from the Purbeck Group (Berriasian) of
|
|
<collectingRegion id="49A5F84DFFA5FFF8772FA04EFD8AFD21" box="[533,609,630,656]" country="United Kingdom" name="Dorset" pageId="19" pageNumber="700">Dorset</collectingRegion>
|
|
and the Wealden (Valanginian) of Sussex,
|
|
<collectingCountry id="F376763FFFA5FFF874FEA0AEFD69FD01" box="[452,642,662,688]" name="United Kingdom" pageId="19" pageNumber="700">United Kingdom</collectingCountry>
|
|
(
|
|
<bibRefCitation id="EFF04B5EFFA5FFF877A9A0AEFF56FD61" author="Joyce" etAl="et al." firstAuthor="Joyce" pageId="19" pageNumber="700" pagination="75 - 97" refId="ref41777" refString="JOYCE W. G., CHAPMAN S. D., MOODY R. T. J. & WALKER C. A. 2011. - The skull of the solemydid turtle Helochelydra nopcsai from the Early Cretaceous of the Isle of Wight (UK) and a review of Solemydidae. Special Papers in Palaeontology 86: 75 - 97." type="journal article" year="2011">
|
|
Joyce
|
|
<emphasis id="B915EABDFFA5FFF877E9A0AEFCECFD01" box="[723,775,662,688]" italics="true" pageId="19" pageNumber="700">et al.</emphasis>
|
|
2011
|
|
</bibRefCitation>
|
|
). That being said, the systematics of Early Cretaceous helochelydrids remains highly confused. For that reason, the helochelydrid material from Angeac-Charente is herein conservatively referred to
|
|
<taxonomicName id="4C614D2CFFA5FFF8749FA12DFDB6FC9E" authorityName="Nopcsa" authorityYear="1928" box="[421,605,789,815]" family="Helochelydridae" kingdom="Animalia" pageId="19" pageNumber="697" rank="family">Helochelydridae</taxonomicName>
|
|
indet.
|
|
</paragraph>
|
|
<caption id="DF1E6627FFA7FFFA75BEA5FAFE8CF858" ID-DOI="http://doi.org/10.5281/zenodo.6902057" ID-Zenodo-Dep="6902057" httpUri="https://zenodo.org/record/6902057/files/figure.png" pageId="17" pageNumber="698" startId="17.[132,143,1986,2003]" targetBox="[151,1436,215,1904]" targetPageId="17">
|
|
<paragraph id="8BDE36AFFFA7FFFA75BEA5FAFE8CF858" blockId="17.[132,1455,1986,2025]" pageId="17" pageNumber="698">
|
|
FIG. 11. — Vertebra of Caudata indet. from Angeac-Charente:
|
|
<emphasis id="B915EABDFFA7FFFA77B2A5FAFD4CF862" bold="true" box="[648,679,1986,2003]" pageId="17" pageNumber="698">A -F</emphasis>
|
|
, (ANG M-71) in anterior (
|
|
<emphasis id="B915EABDFFA7FFFA764DA5FAFC6FF862" bold="true" box="[887,900,1986,2003]" pageId="17" pageNumber="698">A</emphasis>
|
|
), posterior (
|
|
<emphasis id="B915EABDFFA7FFFA76DDA5FAFC1FF862" bold="true" box="[999,1012,1986,2003]" pageId="17" pageNumber="698">B</emphasis>
|
|
), left lateral (
|
|
<emphasis id="B915EABDFFA7FFFA7164A5FAFB87F862" bold="true" box="[1118,1132,1986,2003]" pageId="17" pageNumber="698">C</emphasis>
|
|
), right lateral (
|
|
<emphasis id="B915EABDFFA7FFFA71D8A5FAFB1BF862" bold="true" box="[1250,1264,1986,2003]" pageId="17" pageNumber="698">D</emphasis>
|
|
), dorsal (
|
|
<emphasis id="B915EABDFFA7FFFA7006A5FAFAA3F862" bold="true" box="[1340,1352,1986,2003]" pageId="17" pageNumber="698">E</emphasis>
|
|
) and ventral (
|
|
<emphasis id="B915EABDFFA7FFFA75B3A5E0FF7FF858" bold="true" box="[137,148,2008,2025]" pageId="17" pageNumber="698">F</emphasis>
|
|
) views. Scale bar: 2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="DF1E6627FFA4FFF975BEA5ACFC49F858" ID-DOI="http://doi.org/10.5281/zenodo.6902061" ID-Zenodo-Dep="6902061" httpUri="https://zenodo.org/record/6902061/files/figure.png" pageId="18" pageNumber="699" startId="18.[132,143,1940,1957]" targetBox="[132,1455,215,1878]" targetPageId="18">
|
|
<paragraph id="8BDE36AFFFA4FFF975BEA5ACFC49F858" blockId="18.[132,1456,1940,2025]" pageId="18" pageNumber="699">
|
|
FIG. 12. — Helochelydrid and thalassochelydian turtle remains from Angeac-Charente:
|
|
<emphasis id="B915EABDFFA4FFF97677A5ACFCB1F814" bold="true" box="[845,858,1940,1957]" pageId="18" pageNumber="699">A</emphasis>
|
|
, peripheral plate of
|
|
<taxonomicName id="4C614D2CFFA4FFF976C6A5ACFB68F814" authorityName="Nopcsa" authorityYear="1928" box="[1020,1155,1940,1957]" family="Helochelydridae" kingdom="Animalia" pageId="18" pageNumber="697" rank="family">Helochelydridae</taxonomicName>
|
|
indet. in dorsal view;
|
|
<emphasis id="B915EABDFFA4FFF97009A5ACFAABF814" bold="true" box="[1331,1344,1940,1957]" pageId="18" pageNumber="699">B</emphasis>
|
|
, pleural plate of
|
|
<taxonomicName id="4C614D2CFFA4FFF975A0A593FECAF80D" authorityName="Nopcsa" authorityYear="1928" box="[154,289,1963,1980]" family="Helochelydridae" kingdom="Animalia" pageId="18" pageNumber="697" rank="family">Helochelydridae</taxonomicName>
|
|
indet. in dorsal view;
|
|
<emphasis id="B915EABDFFA4FFF974E0A593FE03F80D" bold="true" box="[474,488,1963,1980]" pageId="18" pageNumber="699">C</emphasis>
|
|
,
|
|
<emphasis id="B915EABDFFA4FFF974C9A593FDEAF80D" bold="true" box="[499,513,1963,1980]" pageId="18" pageNumber="699">D</emphasis>
|
|
, right hyoplastron of
|
|
<taxonomicName id="4C614D2CFFA4FFF9778FA593FC3EF80D" authority="? (Mantell, 1844)" authorityName="? (Mantell" authorityYear="1844" baseAuthorityName="Mantell" baseAuthorityYear="1844" box="[693,981,1963,1980]" class="Reptilia" genus="Hylaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="18" pageNumber="699" phylum="Chordata" rank="species" species="belli">
|
|
<emphasis id="B915EABDFFA4FFF9778FA593FCA0F80D" box="[693,843,1963,1980]" italics="true" pageId="18" pageNumber="699">Hylaeochelys belli</emphasis>
|
|
? (
|
|
<bibRefCitation id="EFF04B5EFFA4FFF97665A593FC3AF80D" author="Mantell" box="[863,977,1963,1980]" firstAuthor="Mantell" pageId="18" pageNumber="699" refId="ref43015" refString="MANTELL G. A. 1844. - The Medals of Creation; or, First Lessons in Geology, and in the Study of Organic Remains. Vol. 2. Henry G. Bohn, London." type="book" year="1844">Mantell, 1844</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
(ANG18-5915) in ventral (
|
|
<emphasis id="B915EABDFFA4FFF9718AA593FB55F80D" bold="true" box="[1200,1214,1963,1980]" pageId="18" pageNumber="699">C</emphasis>
|
|
) and dorsal (
|
|
<emphasis id="B915EABDFFA4FFF97016A593FAD1F80D" bold="true" box="[1324,1338,1963,1980]" pageId="18" pageNumber="699">D</emphasis>
|
|
) views;
|
|
<emphasis id="B915EABDFFA4FFF97044A593FA61F80D" bold="true" box="[1406,1418,1963,1980]" pageId="18" pageNumber="699">E</emphasis>
|
|
, left humerus of
|
|
<taxonomicName id="4C614D2CFFA4FFF975D3A5FAFEBCF862" authorityName="Lydekker" authorityYear="1889" box="[233,343,1986,2003]" class="Reptilia" genus="Hylaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="18" pageNumber="699" phylum="Chordata" rank="genus">
|
|
<emphasis id="B915EABDFFA4FFF975D3A5FAFEBCF862" box="[233,343,1986,2003]" italics="true" pageId="18" pageNumber="699">Hylaeochelys</emphasis>
|
|
</taxonomicName>
|
|
?
|
|
<taxonomicName id="4C614D2CFFA4FFF9745DA5FAFE62F862" authorityName="? (Mantell" authorityYear="1844" box="[359,393,1986,2003]" class="Reptilia" genus="Hylaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="18" pageNumber="699" phylum="Chordata" rank="species" species="belli">
|
|
<emphasis id="B915EABDFFA4FFF9745DA5FAFE62F862" box="[359,393,1986,2003]" italics="true" pageId="18" pageNumber="699">belli</emphasis>
|
|
</taxonomicName>
|
|
? (ANG18-5937) in ventral view;
|
|
<emphasis id="B915EABDFFA4FFF977A3A5FAFD4FF862" bold="true" box="[665,676,1986,2003]" pageId="18" pageNumber="699">F</emphasis>
|
|
, right humerus (ANG18-5938) in dorsal view;;
|
|
<emphasis id="B915EABDFFA4FFF9710AA5FAFBD5F862" bold="true" box="[1072,1086,1986,2003]" pageId="18" pageNumber="699">G</emphasis>
|
|
,
|
|
<emphasis id="B915EABDFFA4FFF97173A5FAFBBCF862" bold="true" box="[1097,1111,1986,2003]" pageId="18" pageNumber="699">H</emphasis>
|
|
, Peripheral of
|
|
<taxonomicName id="4C614D2CFFA4FFF971E9A5FAFAAAF862" authorityName="Lydekker" authorityYear="1889" box="[1235,1345,1986,2003]" class="Reptilia" genus="Hylaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="18" pageNumber="699" phylum="Chordata" rank="genus">
|
|
<emphasis id="B915EABDFFA4FFF971E9A5FAFAAAF862" box="[1235,1345,1986,2003]" italics="true" pageId="18" pageNumber="699">Hylaeochelys</emphasis>
|
|
</taxonomicName>
|
|
?
|
|
<taxonomicName id="4C614D2CFFA4FFF9706BA5FAFA98F862" authorityName="? (Mantell" authorityYear="1844" box="[1361,1395,1986,2003]" class="Reptilia" genus="Hylaeochelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="18" pageNumber="699" phylum="Chordata" rank="species" species="belli">
|
|
<emphasis id="B915EABDFFA4FFF9706BA5FAFA98F862" box="[1361,1395,1986,2003]" italics="true" pageId="18" pageNumber="699">belli</emphasis>
|
|
</taxonomicName>
|
|
? (ANG 18-5993) in dorsal (
|
|
<emphasis id="B915EABDFFA4FFF9741FA5E0FED8F858" bold="true" box="[293,307,2008,2025]" pageId="18" pageNumber="699">G</emphasis>
|
|
) and ventral (
|
|
<emphasis id="B915EABDFFA4FFF9749EA5E0FE59F858" bold="true" box="[420,434,2008,2025]" pageId="18" pageNumber="699">H</emphasis>
|
|
) views. Scale bar: A, 1 cm; E-H, 2 cm; B, 3 cm; C, D, 4 cm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8BDE36AFFFA5FFF875A1A10DFE61F97D" blockId="19.[131,777,215,2028]" pageId="19" pageNumber="700">
|
|
The posterior part of a large carapace collected in 2018 and 2020 provides new information on the morphology of the helochelydrid from Angeac-Charente. However, the material is not fully prepared yet and will be described in more detail elsewhere. A raised midline keel on the posterior neurals is clearly observable, which is a character shared with other helochelydrids (
|
|
<bibRefCitation id="EFF04B5EFFA5FFF87482A1CCFD89FBBE" author="Lydekker" box="[440,610,1012,1039]" firstAuthor="Lydekker" pageId="19" pageNumber="700" refId="ref42720" refString="LYDEKKER R. 1889. - Catalogue of the Fossil Reptilia and Amphibia in the British Museum (Natural History). Part III, The Order Chelonia. Trustees of the British Museum, London, 239 p." type="book" year="1889">Lydekker 1889</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFF04B5EFFA5FFF87748A1CDFE7EFB9F" author="Lapparent de Broin & Murelaga" firstAuthor="Lapparent de Broin" pageId="19" pageNumber="700" pagination="135 - 211" refId="ref42274" refString="LAPPARENT DE BROIN F. DE & MURELAGA X. 1999. - Turtles from the Upper Cretaceous of Lano (Iberian Peninsula). Estudios del Museo de Ciencias Naturales de Alava 14: 135 - 211." type="journal article" year="1999">Lapparent de Broin & Murelaga 1999</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFF04B5EFFA5FFF8749FA62CFDDFFB9F" author="Milner" box="[421,564,1044,1070]" firstAuthor="Milner" pageId="19" pageNumber="700" pagination="1441 - 1467" refId="ref44294" refString="MILNER A. R. 2004. - The turtles of the Purbeck Limestone Group of Dorset, southern England. Palaeontology 47 (6): 1441 - 1467. https: // doi. org / 10.1111 / j. 0031 - 0239.2004.00418. x" type="journal article" year="2004">Milner 2004</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFF04B5EFFA5FFF87779A62CFCEBFB9F" author="VULLO R. & LAPPARENT DE BROIN F. DE & NERAUDEAU D. & DUR- RIEU N." box="[579,768,1044,1070]" journalOrPublisher="Oryctos" pageId="19" pageNumber="700" pagination="37 - 48" part="9" refId="ref50509" refString="VULLO R., LAPPARENT DE BROIN F. DE, NERAUDEAU D. & DUR- RIEU N. 2010. - Turtles from the Early Cenomanian paralic deposits (Late Cretaceous) of Charentes, France. Oryctos 9: 37 - 48." title="Turtles from the Early Cenomanian paralic deposits (Late Cretaceous) of Charentes, France" type="journal article" year="2010">
|
|
Vullo
|
|
<emphasis id="B915EABDFFA5FFF877B2A62DFD55FB9F" box="[648,702,1044,1070]" italics="true" pageId="19" pageNumber="700">et al.</emphasis>
|
|
2010
|
|
</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFF04B5EFFA5FFF875BEA60DFEAFFBFF" author="Joyce" box="[132,324,1076,1103]" etAl="et al." firstAuthor="Joyce" pageId="19" pageNumber="700" pagination="1257 - 1287" refId="ref41904" refString="JOYCE W. G., STERLI J. & CHAPMAN S. D. 2014. - The skeletal morphology of the solemydid turtle Naomichelys speciosa from the Early Cretaceous of Texas. Journal of Paleontology 88 (6): 1257 - 1287. https: // doi. org / 10.1666 / 14 - 002" type="journal article" year="2014">
|
|
Joyce
|
|
<emphasis id="B915EABDFFA5FFF875F3A60DFEEBFBFF" box="[201,256,1076,1102]" italics="true" pageId="19" pageNumber="700">et al.</emphasis>
|
|
2014
|
|
</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFF04B5EFFA5FFF8746FA60DFE38FBFF" author="Joyce" box="[341,467,1076,1103]" firstAuthor="Joyce" pageId="19" pageNumber="700" pagination="65 - 113" refId="ref41684" refString="JOYCE W. G. 2017. - A Review of the Fossil Record of Basal Mesozoic Turtles. Bulletin of the Peabody Museum of Natural History 58: 65 - 113. https: // doi. org / 10.3374 / 014.058.0105" type="journal article" year="2017">Joyce 2017</bibRefCitation>
|
|
; Pérez-García
|
|
<emphasis id="B915EABDFFA5FFF87741A60DFD58FBFF" box="[635,691,1076,1102]" italics="true" pageId="19" pageNumber="700">et al.</emphasis>
|
|
2020). The general morphology resembles that of other species in which this region is sufficiently known, such as “
|
|
<taxonomicName id="4C614D2CFFA5FFF87793A64CFF52FB1F" authorityName="Nopcsa" authorityYear="1928" class="Reptilia" family="Solemydidae" genus="Helochelydra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="700" phylum="Chordata" rank="genus">
|
|
<emphasis id="B915EABDFFA5FFF87793A64CFF52FB1F" italics="true" pageId="19" pageNumber="700">Helochelydra</emphasis>
|
|
</taxonomicName>
|
|
”
|
|
<emphasis id="B915EABDFFA5FFF875F7A6ACFEF1FB1F" box="[205,282,1172,1198]" italics="true" pageId="19" pageNumber="700">anglica</emphasis>
|
|
,
|
|
<taxonomicName id="4C614D2CFFA5FFF87412A6ACFDEDFB1F" authorityName="Hay" authorityYear="1908" box="[296,518,1172,1198]" class="Reptilia" family="Pleurosternidae" genus="Naomichelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="700" phylum="Chordata" rank="species" species="speciosa">
|
|
<emphasis id="B915EABDFFA5FFF87412A6ACFDEDFB1F" box="[296,518,1172,1198]" italics="true" pageId="19" pageNumber="700">Naomichelys speciosa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C614D2CFFA5FFF8772FA6ACFD14FB1F" authorityName="Pérez-García, Espílez, Mampel & Alcalá" authorityYear="2020" box="[533,767,1172,1198]" class="Reptilia" genus="Aragochersis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="700" phylum="Chordata" rank="species" species="lignitesta">
|
|
<emphasis id="B915EABDFFA5FFF8772FA6ACFD14FB1F" box="[533,767,1172,1198]" italics="true" pageId="19" pageNumber="700">Aragochersis lignitesta</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="4C614D2CFFA5FFF87583A68CFE71FB7F" authorityName="Lapparent de Broin & Murelaga" authorityYear="1996" box="[185,410,1204,1230]" class="Reptilia" family="Solemydidae" genus="Solemys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="700" phylum="Chordata" rank="species" species="vermiculata">
|
|
<emphasis id="B915EABDFFA5FFF87583A68CFE71FB7F" box="[185,410,1204,1230]" italics="true" pageId="19" pageNumber="700">Solemys vermiculata</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<bibRefCitation id="EFF04B5EFFA5FFF87494A68CFDB7FB7F" author="Lydekker" box="[430,604,1204,1230]" firstAuthor="Lydekker" pageId="19" pageNumber="700" refId="ref42720" refString="LYDEKKER R. 1889. - Catalogue of the Fossil Reptilia and Amphibia in the British Museum (Natural History). Part III, The Order Chelonia. Trustees of the British Museum, London, 239 p." type="book" year="1889">Lydekker 1889</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFF04B5EFFA5FFF87754A68CFE7DFB5F" author="Lapparent de Broin & Murelaga" firstAuthor="Lapparent de Broin" pageId="19" pageNumber="700" pagination="135 - 211" refId="ref42274" refString="LAPPARENT DE BROIN F. DE & MURELAGA X. 1999. - Turtles from the Upper Cretaceous of Lano (Iberian Peninsula). Estudios del Museo de Ciencias Naturales de Alava 14: 135 - 211." type="journal article" year="1999">Lapparent de Broin & Murelaga 1999</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFF04B5EFFA5FFF8749CA6ECFDDEFB5F" author="Milner" box="[422,565,1236,1262]" firstAuthor="Milner" pageId="19" pageNumber="700" pagination="1441 - 1467" refId="ref44294" refString="MILNER A. R. 2004. - The turtles of the Purbeck Limestone Group of Dorset, southern England. Palaeontology 47 (6): 1441 - 1467. https: // doi. org / 10.1111 / j. 0031 - 0239.2004.00418. x" type="journal article" year="2004">Milner 2004</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFF04B5EFFA5FFF8777FA6ECFCEBFB5F" author="Joyce" box="[581,768,1236,1262]" etAl="et al." firstAuthor="Joyce" pageId="19" pageNumber="700" pagination="1257 - 1287" refId="ref41904" refString="JOYCE W. G., STERLI J. & CHAPMAN S. D. 2014. - The skeletal morphology of the solemydid turtle Naomichelys speciosa from the Early Cretaceous of Texas. Journal of Paleontology 88 (6): 1257 - 1287. https: // doi. org / 10.1666 / 14 - 002" type="journal article" year="2014">
|
|
Joyce
|
|
<emphasis id="B915EABDFFA5FFF877BDA6ECFD55FB5F" box="[647,702,1236,1262]" italics="true" pageId="19" pageNumber="700">et al.</emphasis>
|
|
2014
|
|
</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFF04B5EFFA5FFF875BEA6CCFEE9FABF" author="Joyce" box="[132,258,1268,1294]" firstAuthor="Joyce" pageId="19" pageNumber="700" pagination="65 - 113" refId="ref41684" refString="JOYCE W. G. 2017. - A Review of the Fossil Record of Basal Mesozoic Turtles. Bulletin of the Peabody Museum of Natural History 58: 65 - 113. https: // doi. org / 10.3374 / 014.058.0105" type="journal article" year="2017">Joyce 2017</bibRefCitation>
|
|
; Pérez-García
|
|
<emphasis id="B915EABDFFA5FFF87493A6CCFE0BFABC" box="[425,480,1267,1293]" italics="true" pageId="19" pageNumber="700">et al.</emphasis>
|
|
2020). Neurals V and VI are elongate and hexagonal. Neurals VII and VIII are probably fused. They are wide and hexagonal anteriorly and posteriorly, and narrower on the midline. There are three suprapygals, which are reminiscent of the condition in FMNH PR273, a nearly complete specimen referred to the North American species
|
|
<taxonomicName id="4C614D2CFFA5FFF8748CA78BFD7BFA7C" authorityName="Hay" authorityYear="1908" box="[438,656,1459,1485]" class="Reptilia" family="Pleurosternidae" genus="Naomichelys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Testudines" pageId="19" pageNumber="700" phylum="Chordata" rank="species" species="speciosa">
|
|
<emphasis id="B915EABDFFA5FFF8748CA78BFD7BFA7C" box="[438,656,1459,1485]" italics="true" pageId="19" pageNumber="700">Naomichelys speciosa</emphasis>
|
|
</taxonomicName>
|
|
. However, the number of suprapygals is known to be rather variable in basal turtles. The morphology of the suprapygals and pygal differs from other known helochelydrids, but detailed comparisons are needed in order to assess the systematic value of these characters. The shell surface sculpturing is less prominent on the neurals and becomes more pronounced on the distal part of the costals and on the peripherals, suprapygals, and pygal.
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</paragraph>
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<paragraph id="8BDE36AFFFA5FFF875A1A4EAFD6AF85D" blockId="19.[131,777,215,2028]" pageId="19" pageNumber="700">
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Helochelydrids are considered by numerous authors to be terrestrial forms (
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<bibRefCitation id="EFF04B5EFFA5FFF8745BA4CAFD14F8BD" author="Lapparent de Broin & Murelaga" box="[353,767,1778,1804]" firstAuthor="Lapparent de Broin" pageId="19" pageNumber="700" pagination="135 - 211" refId="ref42274" refString="LAPPARENT DE BROIN F. DE & MURELAGA X. 1999. - Turtles from the Upper Cretaceous of Lano (Iberian Peninsula). Estudios del Museo de Ciencias Naturales de Alava 14: 135 - 211." type="journal article" year="1999">Lapparent de Broin & Murelaga 1999</bibRefCitation>
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;
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<bibRefCitation id="EFF04B5EFFA5FFF875BEA52AFED7F89D" author="Joyce" box="[132,316,1810,1836]" etAl="et al." firstAuthor="Joyce" pageId="19" pageNumber="700" pagination="75 - 97" refId="ref41777" refString="JOYCE W. G., CHAPMAN S. D., MOODY R. T. J. & WALKER C. A. 2011. - The skull of the solemydid turtle Helochelydra nopcsai from the Early Cretaceous of the Isle of Wight (UK) and a review of Solemydidae. Special Papers in Palaeontology 86: 75 - 97." type="journal article" year="2011">
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Joyce
|
|
<emphasis id="B915EABDFFA5FFF875FCA52AFF10F89D" box="[198,251,1810,1836]" italics="true" pageId="19" pageNumber="700">et al.</emphasis>
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2011
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</bibRefCitation>
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|
;
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|
<bibRefCitation id="EFF04B5EFFA5FFF87471A52AFDF4F89D" author="SCHEYER T. M. & PEREZ- GARCIA A. & MURELAGA X." box="[331,543,1809,1836]" journalOrPublisher="Organisms Diversity & Evolution" pageId="19" pageNumber="700" pagination="199 - 212" part="15" refId="ref48623" refString="SCHEYER T. M., PEREZ- GARCIA A. & MURELAGA X. 2015. - Shell bone histology of solemydid turtles (stem Testudines): palaeoecological implications. Organisms Diversity & Evolution 15: 199 - 212. https: // doi. org / 10.1007 / s 13127 - 014 - 0188 - 0" title="Shell bone histology of solemydid turtles (stem Testudines): palaeoecological implications" type="journal article" year="2015">
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|
Scheyer
|
|
<emphasis id="B915EABDFFA5FFF87492A52AFE35F89D" box="[424,478,1810,1836]" italics="true" pageId="19" pageNumber="700">et al.</emphasis>
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|
2015
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</bibRefCitation>
|
|
). They are known to have limbs and neck covered with osteoderms (
|
|
<bibRefCitation id="EFF04B5EFFA5FFF877B8A50AFF55F8DD" author="Barrett" etAl="et al." firstAuthor="Barrett" pageId="19" pageNumber="700" pagination="279 - 295" refId="ref36054" refString="BARRETT P. M., CLARKE J. B., BRINKMAN D. B., CHAPMAN S. D. & ENSOM P. C. 2002. - Morphology, histology and identification of the ' granicones' from the Purbeck Limestone Formation (Lower Cretaceous: Berriasian) of Dorset, southern England. Cretaceous Research 23: 279 - 295. https: // doi. org / 10.1006 / cres. 2002.1002" type="journal article" year="2002">
|
|
Barrett
|
|
<emphasis id="B915EABDFFA5FFF877EEA50AFCECF8FA" box="[724,775,1841,1867]" italics="true" pageId="19" pageNumber="700">et al.</emphasis>
|
|
2002
|
|
</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFF04B5EFFA5FFF875F7A56AFE6CF8DD" author="Joyce" box="[205,391,1873,1900]" etAl="et al." firstAuthor="Joyce" pageId="19" pageNumber="700" pagination="1257 - 1287" refId="ref41904" refString="JOYCE W. G., STERLI J. & CHAPMAN S. D. 2014. - The skeletal morphology of the solemydid turtle Naomichelys speciosa from the Early Cretaceous of Texas. Journal of Paleontology 88 (6): 1257 - 1287. https: // doi. org / 10.1666 / 14 - 002" type="journal article" year="2014">
|
|
Joyce
|
|
<emphasis id="B915EABDFFA5FFF8742AA56AFEADF8DA" box="[272,326,1873,1899]" italics="true" pageId="19" pageNumber="700">et al.</emphasis>
|
|
2014
|
|
</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="EFF04B5EFFA5FFF874ADA569FD87F8DD" author="SCHEYER T. M. & PEREZ- GARCIA A. & MURELAGA X." box="[407,620,1873,1900]" journalOrPublisher="Organisms Diversity & Evolution" pageId="19" pageNumber="700" pagination="199 - 212" part="15" refId="ref48623" refString="SCHEYER T. M., PEREZ- GARCIA A. & MURELAGA X. 2015. - Shell bone histology of solemydid turtles (stem Testudines): palaeoecological implications. Organisms Diversity & Evolution 15: 199 - 212. https: // doi. org / 10.1007 / s 13127 - 014 - 0188 - 0" title="Shell bone histology of solemydid turtles (stem Testudines): palaeoecological implications" type="journal article" year="2015">
|
|
Scheyer
|
|
<emphasis id="B915EABDFFA5FFF874CFA56AFDC0F8DA" box="[501,555,1873,1899]" italics="true" pageId="19" pageNumber="700">et al.</emphasis>
|
|
2015
|
|
</bibRefCitation>
|
|
). Such osteoderms have also been found at Angeac-Charente. Known cranial material suggests a durophagous diet, although it is still unclear exactly which hard-shelled food elements these turtles fed on (
|
|
<bibRefCitation id="EFF04B5EFFA5FFF87457A5EAFDCCF85A" author="Joyce" box="[365,551,2001,2028]" etAl="et al." firstAuthor="Joyce" pageId="19" pageNumber="700" pagination="75 - 97" refId="ref41777" refString="JOYCE W. G., CHAPMAN S. D., MOODY R. T. J. & WALKER C. A. 2011. - The skull of the solemydid turtle Helochelydra nopcsai from the Early Cretaceous of the Isle of Wight (UK) and a review of Solemydidae. Special Papers in Palaeontology 86: 75 - 97." type="journal article" year="2011">
|
|
Joyce
|
|
<emphasis id="B915EABDFFA5FFF8748AA5EAFE0DF85A" box="[432,486,2001,2027]" italics="true" pageId="19" pageNumber="700">et al.</emphasis>
|
|
2011
|
|
</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="EFF04B5EFFA5FFF8770CA5E9FD9BF85A" author="Joyce" box="[566,624,2001,2027]" firstAuthor="Joyce" pageId="19" pageNumber="700" pagination="1257 - 1287" refId="ref41904" refString="JOYCE W. G., STERLI J. & CHAPMAN S. D. 2014. - The skeletal morphology of the solemydid turtle Naomichelys speciosa from the Early Cretaceous of Texas. Journal of Paleontology 88 (6): 1257 - 1287. https: // doi. org / 10.1666 / 14 - 002" type="journal article" year="2014">2014</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
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</document> |