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<document id="FAD24F89BB71BDA437593F1BB142A6E3" ID-CLB-Dataset="21673" ID-DOI="10.11646/zootaxa.4789.1.1" ID-GBIF-Dataset="c12aaed8-264c-459c-b1aa-09eaeb686670" ID-ISSN="1175-5326" ID-Zenodo-Dep="3884610" ID-ZooBank="311A27A2-A053-4D20-A43E-EF57DBE55166" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591601966137" checkinUser="plazi" docAuthor="Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit &amp; Katoh, Toru" docDate="2020" docId="03C86C16EB3A5746FF6EDB06FB9EF872" docLanguage="en" docName="zootaxa.4789.1.1.pdf" docOrigin="Zootaxa 4789 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phortica (Ashima) nigrifoliiseta Takada, Momma &amp; Shima" docType="treatment" docVersion="10" lastPageNumber="29" masterDocId="FFF1146EEB26575AFFF9DD08FFEBFFAA" masterDocTitle="Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="29" updateTime="1698842174822" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="233C69B15C58B6F0FCC82BE81A8F91CE">
<mods:title id="A671D133B811AFB9B6A16B6A19410CF2">Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species</mods:title>
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<mods:namePart id="C1EFB21F21C0F7FF40CF0745D881889A">Toda, Masanori J.</mods:namePart>
<mods:affiliation id="A606631F758ED0F04B23C6D834757AD6">Hokkaido University Museum, Hokkaido University, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="D8F43582E82F5F6C42925295F2421F30">Bänziger, Hans</mods:namePart>
<mods:affiliation id="EB85080B5B41AA8A5D019B5E6457853E">Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034</mods:affiliation>
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<mods:namePart id="94DF7BA152D283510E027ED80A83E993">Sati, Pradeep C.</mods:namePart>
<mods:affiliation id="C9071060D969943A18A7346907DC4072">Systematic &amp; Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. &amp; satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575</mods:affiliation>
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<mods:namePart id="3F4E77E26318A4DC1064B27AB86AE56C">Fartyal, Rajendra S.</mods:namePart>
<mods:affiliation id="E5536E15043D57010234DDE81E04F514">Systematic &amp; Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. &amp; rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263</mods:affiliation>
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<mods:namePart id="041EB6923563F3F85C5EA967BDC2D3BA">Suwito, Awit</mods:namePart>
<mods:affiliation id="8F070C4DC972E9B55F34F1D9BD100226">Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com</mods:affiliation>
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<mods:namePart id="507C6FDC2B50080C0DE408E7D4DC5D2B">Katoh, Toru</mods:namePart>
<mods:affiliation id="F0E849EC5B49BD3DE335DA183DEBAC6E">Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo 060 - 0810, Japan t _ katoh @ sci. hokudai. ac. jp; https: // orcid. org / 0000 - 0003 - 3376 - 8545</mods:affiliation>
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<mods:title id="6D0575913BAC1972015B140FFD405D4C">Zootaxa</mods:title>
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<mods:date id="35B501824928BBE5CF30C058C160619D">2020</mods:date>
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<mods:number id="4611EA454FB00E6EAB4425436E894D47">2020-06-08</mods:number>
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<mods:number id="2AA3AF98E9824B930C01D12160D5F2FB">4789</mods:number>
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<mods:identifier id="5636D68480AA7990A548389693BF94B2" type="DOI">10.11646/zootaxa.4789.1.1</mods:identifier>
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<treatment id="03C86C16EB3A5746FF6EDB06FB9EF872" ID-DOI="http://doi.org/10.5281/zenodo.5221323" ID-GBIF-Taxon="164612775" ID-Zenodo-Dep="5221323" LSID="urn:lsid:plazi:treatment:03C86C16EB3A5746FF6EDB06FB9EF872" httpUri="http://treatment.plazi.org/id/03C86C16EB3A5746FF6EDB06FB9EF872" lastPageNumber="29" pageId="28" pageNumber="29">
<subSubSection id="C37B8E8BEB3A5746FF6EDB06FC81F983" box="[151,874,1550,1577]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BDEDD00EB3A5746FF6EDB06FC81F983" blockId="28.[151,874,1550,1577]" box="[151,874,1550,1577]" pageId="28" pageNumber="29">
<heading id="D0966A6CEB3A5746FF6EDB06FC81F983" bold="true" box="[151,874,1550,1577]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<emphasis id="B9150112EB3A5746FF6EDB06FC81F983" bold="true" box="[151,874,1550,1577]" pageId="28" pageNumber="29">
3.1.
<taxonomicName id="4C61A683EB3A5746FF32DB06FDA6F983" ID-CoL="8DBWG" authorityName="Takada, Momma &amp; Shima" box="[203,589,1550,1577]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="nigrifoliiseta" subGenus="Ashima">
<emphasis id="B9150112EB3A5746FF32DB06FED8F983" bold="true" box="[203,307,1550,1577]" italics="true" pageId="28" pageNumber="29">Phortica</emphasis>
(
<emphasis id="B9150112EB3A5746FEBDDB06FE4AF983" bold="true" box="[324,417,1550,1577]" italics="true" pageId="28" pageNumber="29">Ashima</emphasis>
)
<emphasis id="B9150112EB3A5746FE4BDB06FDA6F983" bold="true" box="[434,589,1550,1577]" italics="true" pageId="28" pageNumber="29">nigrifoliiseta</emphasis>
</taxonomicName>
species subgroup, new
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37B8E8BEB3A5746FF6EDB5DFB9EF872" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8BDEDD00EB3A5746FF6EDB5DFB12F93E" blockId="28.[151,1437,1621,2009]" pageId="28" pageNumber="29">
<emphasis id="B9150112EB3A5746FF6EDB5DFEF9F9DA" bold="true" box="[151,274,1621,1648]" pageId="28" pageNumber="29">Diagnosis.</emphasis>
Setae on and/or near ventral margin of epandrium denser than those on other portions (ch.32-1;
<figureCitation id="135AC185EB3A5746FACEDB5EFA65F9DA" box="[1335,1422,1622,1648]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="28" pageNumber="29">Fig. 8G</figureCitation>
); aedeagal, outer membrane ornamented with somewhat irregular, minute spinules (ch.56-3;
<figureCitation id="135AC185EB3A5746FB6ADB72FB07F93E" box="[1171,1260,1658,1684]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="28" pageNumber="29">Fig. 8H</figureCitation>
).
</paragraph>
<paragraph id="8BDEDD00EB3A5746FF3EDB95FD73F81F" blockId="28.[151,1437,1621,2009]" pageId="28" pageNumber="29">
<emphasis id="B9150112EB3A5746FF3EDB95FE46F912" bold="true" box="[199,429,1693,1720]" pageId="28" pageNumber="29">Shared characters.</emphasis>
Dorsomedial, tentorial apodeme &lt;3/5 basal, parallel portion of dorsolateral, tentorial apodeme (ch.1-1). Longest, dorsal branch of arista shorter than longest seta on pedicel; apical, leaf-like expansion heart-shaped, as broad as long, apically notched (
<figureCitation id="135AC185EB3A5746FD34DBEEFCC2F8AA" box="[717,809,1766,1792]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="28" pageNumber="29">Fig. 8A</figureCitation>
). Palpus with hollow-shaped sense organ (ch.9-2). Abdominal sternite III wider than long (ch.22-1;
<figureCitation id="135AC185EB3A5746FD62DA02FD1BF88E" box="[667,752,1802,1828]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="28" pageNumber="29">Fig. 8E</figureCitation>
), without dense setulae on medial surface (ch.26-0;
<figureCitation id="135AC185EB3A5746FAC0DA02FA7BF88E" box="[1337,1424,1802,1828]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="28" pageNumber="29">Fig. 8E</figureCitation>
). Hypandrium symmetric (ch.40-0;
<figureCitation id="135AC185EB3A5746FDE0DA26FD82F8E2" box="[537,617,1838,1864]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="28" pageNumber="29">Fig. 8J</figureCitation>
). Pregonite (
<figureCitation id="135AC185EB3A5746FD03DA26FCACF8E2" box="[762,839,1838,1864]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="28" pageNumber="29">Fig. 8I</figureCitation>
) partly strongly sclerotized (ch.43-0), with basal extension nearly rectangular to main body, apically narrowing (ch.49-1) and sensilla on submedial and basal portions (ch.52,53-1); apical knob with pointed barbule (ch.46,47-0). Sclerotized, posterior postgonites on aedeagal sheath medially separated from each other (ch.66-0).
</paragraph>
<paragraph id="8BDEDD00EB3A5746FF3EDAB6FB9EF872" blockId="28.[151,1437,1621,2009]" box="[199,1141,1981,2009]" pageId="28" pageNumber="29">
<emphasis id="B9150112EB3A5746FF3EDAB6FE79F872" bold="true" box="[199,402,1981,2008]" pageId="28" pageNumber="29">Included species.</emphasis>
<taxonomicName id="4C61A683EB3A5746FE61DAB6FCEBF872" authorityName="Takada, Momma &amp; Shima" box="[408,768,1982,2009]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="nigrifoliiseta" subGenus="Ashima">
<emphasis id="B9150112EB3A5746FE61DAB6FE11F872" box="[408,506,1982,2008]" italics="true" pageId="28" pageNumber="29">Phortica</emphasis>
(
<emphasis id="B9150112EB3A5746FDF2DAB6FDB4F872" box="[523,607,1982,2008]" italics="true" pageId="28" pageNumber="29">Ashima</emphasis>
)
<emphasis id="B9150112EB3A5746FD96DAB6FCEBF872" box="[623,768,1982,2008]" italics="true" pageId="28" pageNumber="29">nigrifoliiseta</emphasis>
</taxonomicName>
; and
<emphasis id="B9150112EB3A5746FCC7DAB7FCA4F872" box="[830,847,1983,2008]" italics="true" pageId="28" pageNumber="29">P</emphasis>
.
<taxonomicName id="4C61A683EB3A5746FCA4DAB6FBFEF872" authority="Toda &amp; Bänziger &amp; Sati &amp; Fartyal &amp; Suwito &amp; Katoh, 2020" authorityName="Toda &amp; Bänziger &amp; Sati &amp; Fartyal &amp; Suwito &amp; Katoh" authorityYear="2020" box="[861,1045,1982,2009]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="kerinciensis" status="sp. nov." subGenus="Ashima">
(
<emphasis id="B9150112EB3A5746FC9FDAB7FC9CF873" box="[870,887,1983,2009]" italics="true" pageId="28" pageNumber="29">A</emphasis>
.)
<emphasis id="B9150112EB3A5746FC76DAB6FBFEF872" box="[911,1045,1982,2008]" italics="true" pageId="28" pageNumber="29">kerinciensis</emphasis>
</taxonomicName>
<emphasis id="B9150112EB3A5746FBE5DAB6FB9EF872" bold="true" box="[1052,1141,1982,2008]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A226BC69EB3A5746FBE5DAB6FB9EF872" box="[1052,1141,1982,2008]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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