232 lines
31 KiB
XML
232 lines
31 KiB
XML
<document id="79135CF2F6439B86B9723D06111984F8" ID-CLB-Dataset="21673" ID-DOI="10.11646/zootaxa.4789.1.1" ID-GBIF-Dataset="c12aaed8-264c-459c-b1aa-09eaeb686670" ID-ISSN="1175-5326" ID-Zenodo-Dep="3884610" ID-ZooBank="311A27A2-A053-4D20-A43E-EF57DBE55166" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591601966137" checkinUser="plazi" docAuthor="Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit & Katoh, Toru" docDate="2020" docId="03C86C16EB2D5751FF6EDC13FB8AFA69" docLanguage="en" docName="zootaxa.4789.1.1.pdf" docOrigin="Zootaxa 4789 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phortica (Ashima) efragmentata Toda & Bänziger & Sati & Fartyal & Suwito & Katoh 2020, sp. nov." docType="treatment" docVersion="10" lastPageNumber="12" masterDocId="FFF1146EEB26575AFFF9DD08FFEBFFAA" masterDocTitle="Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="12" updateTime="1698842174822" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="3099AD9583552D3E9068625AA3845098" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="D063877AEEB2E90DACCD0AEB1D1F3177">
|
||
<mods:title id="C3940FB13A74CFC584AE4AE578C22877">Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="708526B301B19A6938B8B58CA6DAF7F4" type="personal">
|
||
<mods:role id="D42C3E5742444914BAFE9BAFEFEB4696">
|
||
<mods:roleTerm id="8097D242322C1FBF4C9E4FA70A4ACF58">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="26EFC198250C6288397C97D157433758">Toda, Masanori J.</mods:namePart>
|
||
<mods:affiliation id="9CE11EE3BFB60338B47EF2A04928C67E">Hokkaido University Museum, Hokkaido University, Sapporo 060 - 0810, Japan</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="BC6FE982D8CC1A29D9E2D1A1E9022C16" type="personal">
|
||
<mods:role id="94748A1A6EDF74DB28AE7B25CFDCACDE">
|
||
<mods:roleTerm id="1F62D9C30D9ABCC23756D460ACED9431">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1D09F208D4DFB92A0305F02F9F058643">Bänziger, Hans</mods:namePart>
|
||
<mods:affiliation id="BB7101E915DDF73BFC36E0D63D0E5DA8">Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="720BEA632396B378DE82F56E26929285" type="personal">
|
||
<mods:role id="D1C873528550383B74D33DCB5A75C634">
|
||
<mods:roleTerm id="13DF7F9232DF0514A660EA165F75F23E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E04CC5BAE3E1FB0E8C21DD667F12E5E1">Sati, Pradeep C.</mods:namePart>
|
||
<mods:affiliation id="1C5538EAC4357A2DD9A3950EF0E73193">Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="90917B681161D4CA97615871D9A75C35" type="personal">
|
||
<mods:role id="DD556B173DADBD7B08187F876E3D41ED">
|
||
<mods:roleTerm id="83771608370D5CC095B4807C60C5C8CE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="BEBD2BA259498BACCDC75EF5FA56ABC9">Fartyal, Rajendra S.</mods:namePart>
|
||
<mods:affiliation id="8E894D2763E04929C8AF09F1E9E3F894">Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="9074769B8597DF9ED1DB2F68DD99017F" type="personal">
|
||
<mods:role id="4006174930C4591CCAD70EF9718AF061">
|
||
<mods:roleTerm id="C5AE576CF4ABD04D8DFEDF56DB8CDC14">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="BEA476B70993E4C789186C5501923E34">Suwito, Awit</mods:namePart>
|
||
<mods:affiliation id="4E6925CC96641992E326ADB654F8EB0C">Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="59FF3AD8F7EA2CEBC42F3914E27239E9" type="personal">
|
||
<mods:role id="041827D67B1388A2C0DA9B9083A7E042">
|
||
<mods:roleTerm id="60C727460CD3C29CFF09DA2D04A61E6C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DE892A485B68C54E63E991E75CB340C3">Katoh, Toru</mods:namePart>
|
||
<mods:affiliation id="119B3176C641A046FF0F7300CF2B247F">Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo 060 - 0810, Japan t _ katoh @ sci. hokudai. ac. jp; https: // orcid. org / 0000 - 0003 - 3376 - 8545</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="65FADFE042B9310CF1981D0BE66C63AE">text</mods:typeOfResource>
|
||
<mods:relatedItem id="BB94C60B0D62CCF64371B3B5C00D07F6" type="host">
|
||
<mods:titleInfo id="176318DF98A0D4BE743ADE4E0ECD281B">
|
||
<mods:title id="639346CEAAC2A2E7185B468A8C05A07D">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="AFF323B806E796477E4B151B4F882AFA">
|
||
<mods:date id="833132474D4712502B255CF6AA3376E4">2020</mods:date>
|
||
<mods:detail id="94C4E7D2F8CC2ED7F8F5E077ACA78608" type="pubDate">
|
||
<mods:number id="B66EA5F980FA797DC4731DBE78E47FEB">2020-06-08</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="CBC28B83B0999B04E30919E0DAC060B9" type="volume">
|
||
<mods:number id="6D142C4732AAEB220D4DF4E568746593">4789</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="3AC2B619308BFDA6056FEA9299BCDC2D" type="issue">
|
||
<mods:number id="F4C506FA9D5FFBFD04240686D93D07F4">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="21D7BF1B74152E72E20E7C2F91332327" unit="page">
|
||
<mods:start id="70CB6768BEA770B206D0458B999D1E98">1</mods:start>
|
||
<mods:end id="05CA5BA0B6706D63621C65CD3B5DA70A">54</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="948E4545AEAD64F31686EF4F31102357">journal article</mods:classification>
|
||
<mods:identifier id="7FDB9AD2C385F3D808EFB669AB92D2D8" type="CLB-Dataset">21673</mods:identifier>
|
||
<mods:identifier id="35C9C5147930C4E73AB96D216560EFB9" type="DOI">10.11646/zootaxa.4789.1.1</mods:identifier>
|
||
<mods:identifier id="A7A4428F90225C86B038BDBA3D67BD34" type="GBIF-Dataset">c12aaed8-264c-459c-b1aa-09eaeb686670</mods:identifier>
|
||
<mods:identifier id="0B6B5F3E544131D7A7F3DCA9A6E3B5DA" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="0C3BDD3396901B2212BAF4697957A26B" type="Zenodo-Dep">3884610</mods:identifier>
|
||
<mods:identifier id="2E3038CB51E3AB1D8FA5C03650D43DD8" type="ZooBank">311A27A2-A053-4D20-A43E-EF57DBE55166</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03C86C16EB2D5751FF6EDC13FB8AFA69" ID-DOI="http://doi.org/10.5281/zenodo.5221274" ID-GBIF-Taxon="164612799" ID-Zenodo-Dep="5221274" LSID="urn:lsid:plazi:treatment:03C86C16EB2D5751FF6EDC13FB8AFA69" httpUri="http://treatment.plazi.org/id/03C86C16EB2D5751FF6EDC13FB8AFA69" lastPageNumber="12" pageId="11" pageNumber="12">
|
||
<subSubSection id="C37B8E8BEB2D5751FF6EDC13FD19FE9C" box="[151,754,283,310]" pageId="11" pageNumber="12" type="nomenclature">
|
||
<paragraph id="8BDEDD00EB2D5751FF6EDC13FD19FE9C" blockId="11.[151,754,283,346]" box="[151,754,283,310]" pageId="11" pageNumber="12">
|
||
<heading id="D0966A6CEB2D5751FF6EDC13FD19FE9C" bold="true" box="[151,754,283,310]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
|
||
<emphasis id="B9150112EB2D5751FF6EDC13FD19FE9C" bold="true" box="[151,754,283,310]" pageId="11" pageNumber="12">
|
||
4)
|
||
<taxonomicName id="4C61A683EB2D5751FF4EDC13FD6CFE9C" ID-CoL="8DBTY" authority="Toda" authorityName="Toda & Bänziger & Sati & Fartyal & Suwito & Katoh" authorityYear="2020" box="[183,647,283,310]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="efragmentata" status="sp. nov." subGenus="Ashima">
|
||
<emphasis id="B9150112EB2D5751FF4EDC13FEF4FE9C" bold="true" box="[183,287,283,310]" italics="true" pageId="11" pageNumber="12">Phortica</emphasis>
|
||
(
|
||
<emphasis id="B9150112EB2D5751FEC9DC13FE66FE9C" bold="true" box="[304,397,283,310]" italics="true" pageId="11" pageNumber="12">Ashima</emphasis>
|
||
)
|
||
<emphasis id="B9150112EB2D5751FE67DC13FDD4FE9F" bold="true" box="[414,575,283,309]" italics="true" pageId="11" pageNumber="12">efragmentata</emphasis>
|
||
Toda
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A226BC69EB2D5751FD6DDC14FD19FE9C" box="[660,754,284,310]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B8E8BEB2D5751FF6EDC37FEA9FEF0" box="[151,322,319,346]" pageId="11" pageNumber="12" type="description">
|
||
<paragraph id="8BDEDD00EB2D5751FF6EDC37FEA9FEF0" blockId="11.[151,754,283,346]" box="[151,322,319,346]" pageId="11" pageNumber="12">
|
||
(
|
||
<figureCitation id="135AC185EB2D5751FF66DC37FF03FEF0" box="[159,232,319,346]" captionStart="FIGURE 2" captionStartId="12.[151,250,1856,1881]" captionTargetBox="[151,1435,181,1830]" captionTargetId="figure@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURE 2. Phortica (Ashima) efragmentata Toda, sp. nov. (♂ holotype).A, Antenna; B, palpus; C, cibarium (dorsal view); D, proboscis (lateral view); E, fore tarsus; F, abdomen (dorsal view); G, legs; H, abdominal sternites. Abbreviations: a s, anterior sensilla; m s, medial sensilla; p s, posterior sensilla; s c, sensilla campaniformia; s-I–IV, sternites I–IV; t-I–IV, tergites I–IV. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm in A–D; 0.5 mm in E–H." figureDoi="http://doi.org/10.5281/zenodo.3884616" httpUri="https://zenodo.org/record/3884616/files/figure.png" pageId="11" pageNumber="12">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2D5751FF0DDC48FEEBFEF0" box="[244,256,320,346]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="11" pageNumber="12">3</figureCitation>
|
||
D–G)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B8E8BEB2D5751FF6EDC88FEFEFE14" pageId="11" pageNumber="12" type="diagnosis">
|
||
<paragraph id="8BDEDD00EB2D5751FF6EDC88FEFEFE14" blockId="11.[151,1437,384,1475]" pageId="11" pageNumber="12">
|
||
<emphasis id="B9150112EB2D5751FF6EDC88FEF9FE31" bold="true" box="[151,274,384,411]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
|
||
Posterior piece of aedeagal median rod widely bifurcated (
|
||
<figureCitation id="135AC185EB2D5751FC5ADC88FC13FE30" box="[931,1016,384,410]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="11" pageNumber="12">Fig. 3F</figureCitation>
|
||
); aedeagal free sclerite absent (ch.59- 0;
|
||
<figureCitation id="135AC185EB2D5751FF4DDCACFEE3FE14" box="[180,264,420,446]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="11" pageNumber="12">Fig. 3F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B8E8BEB2D5751FF3EDCCEFB96FABB" pageId="11" pageNumber="12" type="description">
|
||
<paragraph id="8BDEDD00EB2D5751FF3EDCCEFC41FD39" blockId="11.[151,1437,384,1475]" pageId="11" pageNumber="12">
|
||
<emphasis id="B9150112EB2D5751FF3EDCCEFEB2FE48" bold="true" box="[199,345,454,482]" pageId="11" pageNumber="12">Description.</emphasis>
|
||
Supracervical setae 18–19. Dorsomedial, tentorial apodeme 3/4 as long as basal, parallel portion of dorsolateral, tentorial apodeme.Arista with arb = 8/4 (
|
||
<figureCitation id="135AC185EB2D5751FCF2DCE3FC8FFDAF" box="[779,868,491,517]" captionStart="FIGURE 2" captionStartId="12.[151,250,1856,1881]" captionTargetBox="[151,1435,181,1830]" captionTargetId="figure@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURE 2. Phortica (Ashima) efragmentata Toda, sp. nov. (♂ holotype).A, Antenna; B, palpus; C, cibarium (dorsal view); D, proboscis (lateral view); E, fore tarsus; F, abdomen (dorsal view); G, legs; H, abdominal sternites. Abbreviations: a s, anterior sensilla; m s, medial sensilla; p s, posterior sensilla; s c, sensilla campaniformia; s-I–IV, sternites I–IV; t-I–IV, tergites I–IV. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm in A–D; 0.5 mm in E–H." figureDoi="http://doi.org/10.5281/zenodo.3884616" httpUri="https://zenodo.org/record/3884616/files/figure.png" pageId="11" pageNumber="12">Fig. 2A</figureCitation>
|
||
). Palpus with hollow-shaped sense organ (
|
||
<figureCitation id="135AC185EB2D5751FAC1DCE3FA64FDAF" box="[1336,1423,491,517]" captionStart="FIGURE 2" captionStartId="12.[151,250,1856,1881]" captionTargetBox="[151,1435,181,1830]" captionTargetId="figure@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURE 2. Phortica (Ashima) efragmentata Toda, sp. nov. (♂ holotype).A, Antenna; B, palpus; C, cibarium (dorsal view); D, proboscis (lateral view); E, fore tarsus; F, abdomen (dorsal view); G, legs; H, abdominal sternites. Abbreviations: a s, anterior sensilla; m s, medial sensilla; p s, posterior sensilla; s c, sensilla campaniformia; s-I–IV, sternites I–IV; t-I–IV, tergites I–IV. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm in A–D; 0.5 mm in E–H." figureDoi="http://doi.org/10.5281/zenodo.3884616" httpUri="https://zenodo.org/record/3884616/files/figure.png" pageId="11" pageNumber="12">Fig. 2B</figureCitation>
|
||
). Cibarial, medial sensilla 12–17 per side; posterior sensilla 6–7 per side (
|
||
<figureCitation id="135AC185EB2D5751FC4FDF06FBE5FD82" box="[950,1038,526,552]" captionStart="FIGURE 2" captionStartId="12.[151,250,1856,1881]" captionTargetBox="[151,1435,181,1830]" captionTargetId="figure@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURE 2. Phortica (Ashima) efragmentata Toda, sp. nov. (♂ holotype).A, Antenna; B, palpus; C, cibarium (dorsal view); D, proboscis (lateral view); E, fore tarsus; F, abdomen (dorsal view); G, legs; H, abdominal sternites. Abbreviations: a s, anterior sensilla; m s, medial sensilla; p s, posterior sensilla; s c, sensilla campaniformia; s-I–IV, sternites I–IV; t-I–IV, tergites I–IV. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm in A–D; 0.5 mm in E–H." figureDoi="http://doi.org/10.5281/zenodo.3884616" httpUri="https://zenodo.org/record/3884616/files/figure.png" pageId="11" pageNumber="12">Fig. 2C</figureCitation>
|
||
). All tarsi entirely yellow (
|
||
<figureCitation id="135AC185EB2D5751FACCDF06FA64FD82" box="[1333,1423,526,552]" captionStart="FIGURE 2" captionStartId="12.[151,250,1856,1881]" captionTargetBox="[151,1435,181,1830]" captionTargetId="figure@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURE 2. Phortica (Ashima) efragmentata Toda, sp. nov. (♂ holotype).A, Antenna; B, palpus; C, cibarium (dorsal view); D, proboscis (lateral view); E, fore tarsus; F, abdomen (dorsal view); G, legs; H, abdominal sternites. Abbreviations: a s, anterior sensilla; m s, medial sensilla; p s, posterior sensilla; s c, sensilla campaniformia; s-I–IV, sternites I–IV; t-I–IV, tergites I–IV. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm in A–D; 0.5 mm in E–H." figureDoi="http://doi.org/10.5281/zenodo.3884616" httpUri="https://zenodo.org/record/3884616/files/figure.png" pageId="11" pageNumber="12">Fig. 2G</figureCitation>
|
||
); fore tarsomere I as long as 3 succeeding tarsomeres together; mid one slightly longer than rest tarsomeres together; hind one as long as rest together. Aedeagal sheath ventro-apically with a pair of strongly sclerotized postgonites much smaller than bifurcated pieces of aedeagal median rod (
|
||
<figureCitation id="135AC185EB2D5751FCB1DF71FC76FD39" box="[840,925,633,659]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="11" pageNumber="12">Fig. 3F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDEDD00EB2D5751FF3EDF95FCE7FD57" blockId="11.[151,1437,384,1475]" pageId="11" pageNumber="12">
|
||
Measurements (
|
||
<typeStatus id="54DA63A2EB2D5751FE81DF94FE35FD1C" box="[376,478,668,694]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
, in mm): BL (straight distance from anterior edge of pedicel to tip of abdomen) = 3.65, ThL (distance from anterior notal margin to apex of scutellum) = 1.56, WL (distance from humeral cross vein to wing apex) = 2.46, WW (maximum wing width) = 1.08.
|
||
</paragraph>
|
||
<paragraph id="8BDEDD00EB2D5751FF3EDE0EFB96FABB" blockId="11.[151,1437,384,1475]" pageId="11" pageNumber="12">
|
||
Indices (
|
||
<typeStatus id="54DA63A2EB2D5751FED1DE0EFE65FC8A" box="[296,398,774,800]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
, in ratio): FW/HW (frontal width/head width) = 0.44, ch/o (maximum width of gena/maximum diameter of eye) = 0.08, prorb (proclinate orbital seta length/posterior reclinate orbital seta length) = 1.78, rcorb (anterior reclinate orbital seta length /posterior reclinate orbital seta length) = 0.91, vb (subvibrissal seta length /vibrissa length) = 0.30, dcl (anterior dorsocentral seta length/posterior dorsocentral seta length) = (n/a), presctl (prescutellar seta length /posterior dorsocentral seta length) = 0.52, sctl (basal scutellar seta length /apical scutellar seta length) = (n/a), sterno (anterior katepisternal seta length/posterior katepisternal seta length) = (n/a), orbito (distance between proclinate and posterior reclinate orbital setae/distance between inner vertical and posterior reclinate orbital setae) = 2.08, dcp (distance between ipsilateral dorsocentral setae/distance between anterior dorsocentral setae) = 0.24, sctlp (distance between ipsilateral scutellar setae/distance between apical scutellar setae) = 1.20, C (2nd costal section between subcostal break and R
|
||
<subScript id="17E5DF45EB2D5751FD77D95FFD43FBCF" attach="left" box="[654,680,1111,1125]" fontSize="6" pageId="11" pageNumber="12">2+3</subScript>
|
||
/3rd costal section between R
|
||
<subScript id="17E5DF45EB2D5751FC09D95FFBE2FBCF" attach="left" box="[1008,1033,1111,1125]" fontSize="6" pageId="11" pageNumber="12">2+3</subScript>
|
||
and R
|
||
<subScript id="17E5DF45EB2D5751FBABD95FFB87FBCF" attach="left" box="[1106,1132,1111,1125]" fontSize="6" pageId="11" pageNumber="12">4+5</subScript>
|
||
) = 2.35, 4c (3rd costal section between R
|
||
<subScript id="17E5DF45EB2D5751FEC6D973FEB3FB23" attach="left" box="[319,344,1147,1161]" fontSize="6" pageId="11" pageNumber="12">2+3</subScript>
|
||
and R
|
||
<subScript id="17E5DF45EB2D5751FE66D973FE52FB23" attach="left" box="[415,441,1147,1161]" fontSize="6" pageId="11" pageNumber="12">4+5</subScript>
|
||
/M
|
||
<subScript id="17E5DF45EB2D5751FE20D973FE0AFB23" attach="left" box="[473,481,1147,1161]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
|
||
between r-m and dm-cu) = 1.50, 4v (M
|
||
<subScript id="17E5DF45EB2D5751FC6FD973FC75FB23" attach="left" box="[918,926,1147,1161]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
|
||
between dm-cu and wing margin/M
|
||
<subScript id="17E5DF45EB2D5751FAC8D973FAD2FB23" attach="left" box="[1329,1337,1147,1161]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
|
||
between r-m and dm-cu) = 3.03, 5x (CuA
|
||
<subScript id="17E5DF45EB2D5751FDF2D996FDF8FB06" attach="left" box="[523,531,1182,1196]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
|
||
between dm-cu and wing margin/dm-cu between M
|
||
<subScript id="17E5DF45EB2D5751FB9CD996FB86FB06" attach="left" box="[1125,1133,1182,1196]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
|
||
and CuA
|
||
<subScript id="17E5DF45EB2D5751FB22D996FB08FB06" attach="left" box="[1243,1251,1182,1196]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
|
||
) = 1.00, ac (3rd costal section between R
|
||
<subScript id="17E5DF45EB2D5751FE57D9CAFE2CFB7A" attach="left" box="[430,455,1218,1232]" fontSize="6" pageId="11" pageNumber="12">2+3</subScript>
|
||
and R
|
||
<subScript id="17E5DF45EB2D5751FDE8D9C9FDC0FB65" attach="left" box="[529,555,1217,1231]" fontSize="6" pageId="11" pageNumber="12">4+5</subScript>
|
||
/distance between distal ends of R
|
||
<subScript id="17E5DF45EB2D5751FC53D9C9FC2FFB65" attach="left" box="[938,964,1217,1231]" fontSize="6" pageId="11" pageNumber="12">4+5</subScript>
|
||
and M
|
||
<subScript id="17E5DF45EB2D5751FBEAD9CAFBF0FB7A" attach="left" box="[1043,1051,1218,1232]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
|
||
) = 3.88, M (CuA
|
||
<subScript id="17E5DF45EB2D5751FB18D9CAFB02FB7A" attach="left" box="[1249,1257,1218,1232]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
|
||
between dm-cu and wing margin/M
|
||
<subScript id="17E5DF45EB2D5751FE8CD9EDFE96FB59" attach="left" box="[373,381,1253,1267]" fontSize="6" pageId="11" pageNumber="12">1</subScript>
|
||
between r-m and dm-cu) = 0.66, C3F [length of heavy setation in 3rd costal section/(length of heavy setation in 3rd costal section+length of light setation in 3rd costal section)] = 0.71.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B8E8BEB2D5751FF3ED813FBB7FA9E" box="[199,1116,1306,1333]" pageId="11" pageNumber="12" type="materials_examined">
|
||
<paragraph id="8BDEDD00EB2D5751FF3ED813FBB7FA9E" blockId="11.[151,1437,384,1475]" box="[199,1116,1306,1333]" pageId="11" pageNumber="12">
|
||
<materialsCitation id="3B09D75DEB2D5751FF3ED813FBB7FA9E" ID-GBIF-Occurrence="2636268353" box="[199,1116,1306,1333]" collectingDate="2013-12-08" collectionCode="SEHU" collectorName="M. J. Toda" country="Malaysia" location="Ulu Gombak" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" stateProvince="Selangor" typeStatus="holotype">
|
||
<emphasis id="B9150112EB2D5751FF3ED813FED0FA9F" bold="true" box="[199,315,1307,1333]" pageId="11" pageNumber="12">
|
||
<typeStatus id="54DA63A2EB2D5751FF3ED813FEDDFA9F" box="[199,310,1307,1333]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="9D671689EB2D5751FEBBD813FEBCFA9F" box="[322,343,1307,1333]" pageId="11" pageNumber="12" type="male">♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED7045C5EB2D5751FE9FD813FE5EFA9F" box="[358,437,1307,1333]" pageId="11" pageNumber="12">SEHU</collectionCode>
|
||
), “
|
||
<location id="8EBE8BDBEB2D5751FE2BD812FD83FA9E" LSID="urn:lsid:plazi:treatment:03C86C16EB2D5751FF6EDC13FB8AFA69:8EBE8BDBEB2D5751FE2BD812FD83FA9E" box="[466,616,1306,1332]" country="Malaysia" name="Ulu Gombak" pageId="11" pageNumber="12" stateProvince="Selangor">Ulu Gombak</location>
|
||
,
|
||
<collectingRegion id="49A513E2EB2D5751FD8DD812FD32FA9E" box="[628,729,1306,1332]" country="Malaysia" name="Selangor" pageId="11" pageNumber="12">Selangor</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3769D90EB2D5751FD1DD812FCA6FA9E" box="[740,845,1306,1332]" name="Malaysia" pageId="11" pageNumber="12">Malaysia</collectingCountry>
|
||
,
|
||
<date id="FFDFFBC0EB2D5751FCA0D812FC26FA9E" box="[857,973,1306,1332]" pageId="11" pageNumber="12" value="2013-12-08">
|
||
<collectingDate id="EF9B0228EB2D5751FCA0D812FC26FA9E" box="[857,973,1306,1332]" pageId="11" pageNumber="12" value="2013-12-08">8.xii.2013</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="2694B8D6EB2D5751FC21D813FBA4FA9E" box="[984,1103,1306,1333]" pageId="11" pageNumber="12">M.J. Toda</collectorName>
|
||
”.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B8E8BEB2D5751FF3ED835FDB8FAF2" box="[199,595,1341,1368]" pageId="11" pageNumber="12" type="distribution">
|
||
<paragraph id="8BDEDD00EB2D5751FF3ED835FDB8FAF2" blockId="11.[151,1437,384,1475]" box="[199,595,1341,1368]" pageId="11" pageNumber="12">
|
||
<emphasis id="B9150112EB2D5751FF3ED835FE8AFAF2" bold="true" box="[199,353,1341,1368]" pageId="11" pageNumber="12">Distribution.</emphasis>
|
||
Peninsular
|
||
<collectingCountry id="F3769D90EB2D5751FE1CD836FDA4FAF2" box="[485,591,1342,1368]" name="Malaysia" pageId="11" pageNumber="12">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B8E8BEB2D5751FF3ED86AFD62FA35" pageId="11" pageNumber="12" type="discussion">
|
||
<paragraph id="8BDEDD00EB2D5751FF3ED86AFD62FA35" blockId="11.[151,1437,384,1475]" pageId="11" pageNumber="12">
|
||
<emphasis id="B9150112EB2D5751FF3ED86AFED0FAD6" bold="true" box="[199,315,1378,1404]" pageId="11" pageNumber="12">Remarks.</emphasis>
|
||
This species resembles
|
||
<taxonomicName id="4C61A683EB2D5751FDB2D86AFD33FAD1" box="[587,728,1377,1404]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="afoliolata">
|
||
<emphasis id="B9150112EB2D5751FDB2D86AFDB7FAD1" box="[587,604,1378,1403]" italics="true" pageId="11" pageNumber="12">P</emphasis>
|
||
.
|
||
<emphasis id="B9150112EB2D5751FD92D869FD33FAD1" box="[619,728,1377,1403]" italics="true" pageId="11" pageNumber="12">afoliolata</emphasis>
|
||
</taxonomicName>
|
||
in having the small postgonites (
|
||
<figureCitation id="135AC185EB2D5751FBB4D869FB65FAD6" box="[1101,1166,1377,1404]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="11" pageNumber="12">Fig. 3</figureCitation>
|
||
F–H), but can be distinguished from it by the diagnostic characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B8E8BEB2D5751FF3ED8A1FB8AFA69" box="[199,1121,1449,1475]" pageId="11" pageNumber="12" type="etymology">
|
||
<paragraph id="8BDEDD00EB2D5751FF3ED8A1FB8AFA69" blockId="11.[151,1437,384,1475]" box="[199,1121,1449,1475]" pageId="11" pageNumber="12">
|
||
<emphasis id="B9150112EB2D5751FF3ED8A1FEA5FA69" bold="true" box="[199,334,1449,1475]" pageId="11" pageNumber="12">Etymology.</emphasis>
|
||
Referring to the lack of free sclerite within aedeagal outer membrane.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |