treatments-xml/data/03/AA/F7/03AAF74DFF90996F1DF5FE7E6054F88A.xml
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<document id="8958D5A5835F9D66C438E90A3CF45BD5" ID-CLB-Dataset="55502" ID-DOI="10.11646/zootaxa.5138.2.4" ID-GBIF-Dataset="48c6ef6a-bd20-4f98-8d6b-7a1b79ab2937" ID-ISSN="1175-5326" ID-Zenodo-Dep="6557093" ID-ZooBank="0B0E6A73-9FC2-44BD-9F30-57AD9B9FE4F0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652772555109" checkinUser="plazi" docAuthor="Leavengood, John M., Pinkerton, Morgan G. &amp; Rifkind, Jacques" docDate="2022" docId="03AAF74DFF90996F1DF5FE7E6054F88A" docLanguage="en" docName="zootaxa.5138.2.4.pdf" docOrigin="Zootaxa 5138 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tarsobaenus piper Leavengood, Pinkerton and Rifkind 2022, new species" docType="treatment" docVersion="7" lastPageNumber="173" masterDocId="FF938F35FF9599691D62FFC86558FFA2" masterDocTitle="Description of the new genus Tarsobaenus and three new species from Costa Rica (Coleoptera: Cleridae: Clerinae: Hydnocerini)" masterLastPageNumber="176" masterPageNumber="167" pageNumber="172" updateTime="1699337758796" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CED7443D884F7474248F0E88AEB3B55C">Description of the new genus Tarsobaenus and three new species from Costa Rica (Coleoptera: Cleridae: Clerinae: Hydnocerini)</mods:title>
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<mods:namePart id="0681FB1A3569B6AC8FBCA4C774100DAE">Leavengood, John M.</mods:namePart>
<mods:nameIdentifier id="92C75BC89AA63A1B8831B3C5AEDBE057" type="ORCID">0000-0003-3223-455X</mods:nameIdentifier>
<mods:affiliation id="A727A42D00C1C11BEB667D9CDA20AF9F">United States Department of Agriculture, APHIS, PPQ, 9325 Bay Plaza Blvd, Suite 206, Tampa, FL 33619 USA John. M. Leavengood @ usda. gov; https: // orcid. org / 0000 - 0003 - 3223 - 455 X</mods:affiliation>
<mods:nameIdentifier id="D2329181DD4722BCE71E3352EC877474" type="email">eavengood@usda.gov</mods:nameIdentifier>
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<mods:namePart id="0A334011FCB649AF1DE8D71B09563CD2">Pinkerton, Morgan G.</mods:namePart>
<mods:affiliation id="93142E9F7AFE4FF8AC8CC155DEB59704">UF / IFAS Extension Seminole County, 250 West County Home Road, Sanford, FL 32773 USA</mods:affiliation>
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<mods:namePart id="C5F6656DC8A77090C539B4105FD944FB">Rifkind, Jacques</mods:namePart>
<mods:nameIdentifier id="CD0768145E57C47646DA3E9B83708C49" type="ORCID">0000-0002-7738-7800</mods:nameIdentifier>
<mods:affiliation id="0A8DE6A4B49AA7373764BE8846E7F8D2">Research Associate, California State Collection of Arthropods, 3294 Meadowview Road Sacramento, CA 95832 USA clerid @ aol. com; https: // orcid. org / 0000 - 0002 - 7738 - 7800</mods:affiliation>
<mods:nameIdentifier id="DFAC69E33D3E300864EFFC4DAD0ED028" type="email">clerid@aol.com</mods:nameIdentifier>
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<mods:date id="B1CCB79160751E4FE6B9D02CE8D723CA">2022</mods:date>
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<subSubSection id="C31915D0FF90996C1DF5FE7E61DCFE57" pageId="5" pageNumber="172" type="nomenclature">
<paragraph id="8BBC465BFF90996C1DF5FE7E6686FE72" blockId="5.[151,1156,438,537]" box="[151,990,438,465]" pageId="5" pageNumber="172">
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<taxonomicName id="4C033DD8FF90996C1DF5FE7E6619FE72" authority="Leavengood, Pinkerton and Rifkind" authorityName="Leavengood, Pinkerton and Rifkind" authorityYear="2022" box="[151,833,438,465]" class="Insecta" family="Cleridae" genus="Tarsobaenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="172" phylum="Arthropoda" rank="species" species="piper" status="sp. nov.">
<emphasis id="B9779A49FF90996C1DF5FE7E642EFE73" bold="true" box="[151,374,438,465]" italics="true" pageId="5" pageNumber="172">Tarsobaenus piper</emphasis>
Leavengood, Pinkerton and Rifkind
</taxonomicName>
,
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="C31915D0FF90996C1DF5FE376079F87F" pageId="5" pageNumber="172" type="description">
<paragraph id="8BBC465BFF90996C1DF5FE376447FDBB" blockId="5.[151,1156,438,537]" box="[151,287,511,537]" pageId="5" pageNumber="172">
(
<figureCitation id="13385ADEFF90996C1DFDFE37644FFDBB" box="[159,279,511,537]" captionStart="FIGURES 711" captionStartId="6.[151,264,1411,1435]" captionTargetBox="[151,1436,181,1386]" captionTargetId="figure-26@6.[151,1436,181,1387]" captionTargetPageId="6" captionText="FIGURES 711. Tarsobaenus piper, n. sp. (holotype, paratype): 7—holotype, dorsal habitus; 8—head; 9—paratype, dorsal habitus; 10—elytral plica; 11—lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.6557097" httpUri="https://zenodo.org/record/6557097/files/figure.png" pageId="5" pageNumber="172">Figs. 711</figureCitation>
)
</paragraph>
<paragraph id="8BBC465BFF90996C1DF5FD8F67B7FD08" blockId="5.[151,1437,582,2013]" pageId="5" pageNumber="172">
<materialsCitation id="3B6B4C06FF90996C1DF5FD8F64BFFD24" ID-GBIF-Occurrence="3771643309" collectingDate="1989-12" collectionCode="INBio" country="Costa Rica" elevation="500" location="San Jose" pageId="5" pageNumber="172" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
<typeStatus id="54B8F8F9FF90996C1DF5FD8F65A7FDC3" box="[151,255,583,609]" pageId="5" pageNumber="172" type="holotype">Holotype</typeStatus>
: Female.
<collectingCountry id="F31406CBFF90996C1C13FD8F6741FDC3" box="[369,537,583,609]" name="Costa Rica" pageId="5" pageNumber="172">COSTA RICA</collectingCountry>
: Prov.
<collectingRegion id="49C788B9FF90996C1F05FD8F6796FDC3" box="[615,718,583,609]" country="Costa Rica" name="San Jose" pageId="5" pageNumber="172">San Jose</collectingRegion>
: Est. Bijagual,
<quantity id="4CFBEBBEFF90996C1E1EFD8E66E6FDC3" box="[892,958,582,609]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="172" unit="m" value="500.0">
<elevation id="002EA168FF90996C1E1EFD8E66E6FDC3" box="[892,958,582,609]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="172" unit="m" value="500.0">500m</elevation>
</quantity>
, Res. Biol. Carara,
<date id="FFBD609BFF90996C19FCFD8F6051FDC3" box="[1182,1289,583,609]" pageId="5" pageNumber="172" value="1989-12">
<collectingDate id="EFF99973FF90996C19FCFD8F6051FDC3" box="[1182,1289,583,609]" pageId="5" pageNumber="172" value="1989-12">XII-1989</collectingDate>
</date>
, Tp Malaise, L-N 192250, 474760 (
<collectionCode id="ED12DE9EFF90996C1CF1FDA36487FD24" box="[403,479,619,646]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="5" pageNumber="172">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B6B4C06FF90996C1C91FDA367B3FD08" ID-GBIF-Occurrence="3771643312" collectingDate="1990-06" collectionCode="INBio" collectorName="E. Bello" country="Costa Rica" elevation="50" location="Puntarenas" pageId="5" pageNumber="172" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
<typeStatus id="54B8F8F9FF90996C1C91FDA3670DFD27" box="[499,597,619,645]" pageId="5" pageNumber="172" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D058DD2FF90996C1F06FDA367BEFD27" box="[612,742,619,645]" pageId="5" pageNumber="172" type="generic">1 specimen</specimenCount>
. Prov.
<collectingRegion id="49C788B9FF90996C1E51FDA366EAFD27" box="[819,946,619,645]" country="Costa Rica" name="Puntarenas" pageId="5" pageNumber="172">Puntarenas</collectingRegion>
: Est. Queb. Bonita,
<quantity id="4CFBEBBEFF90996C19F1FDA2619FFD27" box="[1171,1223,618,645]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="172" unit="m" value="50.0">
<elevation id="002EA168FF90996C19F1FDA2619FFD27" box="[1171,1223,618,645]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="172" unit="m" value="50.0">50m</elevation>
</quantity>
, Res. Biol. Carara,
<date id="FFBD609BFF90996C1DF5FD4765A0FD0B" box="[151,248,655,681]" pageId="5" pageNumber="172" value="1990-06">
<collectingDate id="EFF99973FF90996C1DF5FD4765A0FD0B" box="[151,248,655,681]" pageId="5" pageNumber="172" value="1990-06">VI-1990</collectingDate>
</date>
,
<collectorName id="26F6238DFF90996C1C66FD47643AFD0B" box="[260,354,655,681]" pageId="5" pageNumber="172">E. Bello</collectorName>
, L-N-194500, 469850 (
<collectionCode id="ED12DE9EFF90996C1F09FD4767EDFD0B" box="[619,693,655,681]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="5" pageNumber="172">INBio</collectionCode>
, 1 f)
</materialsCitation>
.
</paragraph>
<paragraph id="8BBC465BFF90996C1DA5FD7A6079F87F" blockId="5.[151,1437,582,2013]" pageId="5" pageNumber="172">
<emphasis id="B9779A49FF90996C1DA5FD7A6403FD6E" bold="true" box="[199,347,690,716]" pageId="5" pageNumber="172">Description:</emphasis>
<typeStatus id="54B8F8F9FF90996C1C06FD7B6494FD6F" box="[356,460,691,717]" pageId="5" pageNumber="172" type="holotype">Holotype</typeStatus>
(female). Body length:
<quantity id="4CFBEBBEFF90996C1FBDFD7A661FFD6F" box="[735,839,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" pageId="5" pageNumber="172" unit="mm" value="5.05">5.05 mm</quantity>
. Head, basal antennal segments, mouthparts, thorax, legs, basal tarsomeres, scutellum, elytra, abdomen orangish-testaceous (paler on femora and abdomen); apical tarsomeres, antennomeres III-VI darkening apically), outer margins of pro- and metatibiae (hardly on metatibiae), and mandibles weakly infuscate basally (palest basally, darkening toward apices); antennomeres VII-X, large metathoracic macula, metepisternum, lateral margins of pronotum, distinct band on apical 1/3 of femora (only on outer margin of profemora), and lateral margins of elytra black.
<emphasis id="B9779A49FF90996C1E2BFCAE66C8FC23" bold="true" box="[841,912,870,897]" italics="true" pageId="5" pageNumber="172">Head:</emphasis>
Slightly wider than humeri, surface smooth, impunctate, moderately covered in fine, long, pale, erect, semirecumbent setae; setae on eyes and posterolateral margin of head (behind eyes) longer; eyes prominent, width across eyes distinctly wider than at pronotal tubercles, sparsely covered with fine, long, pale, erect setae; antennae 10-segmented with distinct single-segment club; frons with two dorsally convergent elongate depressions; mouthparts with clypeus longitudinally subdivided; mandibles bifid, right mandible dorsally excavated to receive the left; apical maxillary palpomeres securiform; apical labial palpomeres digitiform.
<emphasis id="B9779A49FF90996C1CC0FBF66759FBFB" bold="true" box="[418,513,1086,1113]" italics="true" pageId="5" pageNumber="172">Thorax:</emphasis>
Pronotum widest at lateral angles but still narrower than width of head, covered in long, pale and testaceous, erect setae interspersed with short, pale, semirecumbent setae; pronotal disc shiny, sparsely, finely punctate; punctures obsolete on anterior and posterior pronotal collars, anterior collar finely rugose; disc very convex, anterior collar strongly constricted, upturned at head, transverse impression complete, anterior fringe of short, pale, erect setae projecting over head; posterior collar abruptly curved upward at scutellar margin with posterior fringe of long, pale, erect setae projecting over scutellum; scutellum with long, pale, erect setae projecting over base of elytral suture. Ventral prothorax sparsely covered with long, pale, suberect setae which become shorter and more recumbent towards middle; procoxal cavities open; mesothorax more densely covered with pale, recumbent setae which continue onto the metepisternum, becoming less dense; metathorax sparsely covered with long, pale, erect setae. Metathorax large, moderately inflated; surface smooth, minutely granulate, shiny.
<emphasis id="B9779A49FF90996C1845FA4B603EFA3F" bold="true" box="[1319,1382,1411,1437]" pageId="5" pageNumber="172">
<emphasis id="B9779A49FF90996C1845FA4B6005FA3F" bold="true" box="[1319,1373,1411,1437]" italics="true" pageId="5" pageNumber="172">Legs</emphasis>
:
</emphasis>
Surface shiny, covered with long, fine, testaceous, erect setae of varying lengths; femora expanded distally (metafemora most strongly),weakly laterally compressed, nearly twice the width of tibia; tibiae narrow, elongate, covered with long, fine, testaceous, erect setae of varying lengths, vestiture on distal 1/3 short, fine, testaceous, semirecumbent; hind legs slightly longer, metafemora extended well past apex of elytra when laid alongside; tibial spur formula 1- 2(?)-1(?); tarsal claw bifid (ungues present).
<emphasis id="B9779A49FF90996C1FF2F9FF67B9F9F3" bold="true" box="[656,737,1591,1617]" pageId="5" pageNumber="172">
<emphasis id="B9779A49FF90996C1FF2F9FF6780F9F3" bold="true" box="[656,728,1591,1617]" italics="true" pageId="5" pageNumber="172">Wings</emphasis>
:
</emphasis>
Elytra shiny, parallel to subparallel (widest at humeri), apices not covering abdomen, disc subflattened dorsoventrally, distinctly punctate with large, widely spaced punctures, surface covered in long, fine, pale, erect, semirecumbent setae of varying lengths; internal facing margin of plica forming elongate excavation with distinct margin, excavation testaceous with ventral margin projecting outwards more so than dorsal margin; apical 1/3 strongly dehiscent; lateral margins weakly serrulate, each serrulate denticle with single seta, serrulations strongest apically, weakest at anterior 1/3; elytral apices serrate, independently rounded-subtruncate (serration reduced at apical truncation). Hindwing dusky-testaceous radiating from venation, apical 1/4 entirely cloudy black visible from dorsal view (obscures tip of abdomen from visibility).
<emphasis id="B9779A49FF90996C19AAF8FA606EF8EF" bold="true" box="[1224,1334,1842,1869]" italics="true" pageId="5" pageNumber="172">Abdomen</emphasis>
: Weakly shiny, sparsely covered with long, pale, erect setae;
<emphasis id="B9779A49FF90996C1FB5F89F666EF8D3" box="[727,822,1879,1905]" italics="true" pageId="5" pageNumber="172">Females</emphasis>
with broad, gradual, smooth emargination on posterior margin of fifth sternite, sixth sternite divided longitudinally into two sclerites with widely-spaced, long, testaceous, subapical setae and dense apical fringe of shorter setae on each sclerite, apical tergite unmodified, but with pale setae of varying lengths, some being apical and subapical setae quite long and curved. Male not known.
</paragraph>
</subSubSection>
<caption id="DF7C16D3FF93996F1DF5FA4B6736FA1D" ID-DOI="http://doi.org/10.5281/zenodo.6557097" ID-Zenodo-Dep="6557097" httpUri="https://zenodo.org/record/6557097/files/figure.png" pageId="6" pageNumber="173" startId="6.[151,264,1411,1435]" targetBox="[151,1436,181,1386]" targetPageId="6">
<paragraph id="8BBC465BFF93996F1DF5FA4B6736FA1D" blockId="6.[151,1436,1411,1471]" pageId="6" pageNumber="173">
<emphasis id="B9779A49FF93996F1DF5FA4B6410FA39" bold="true" box="[151,328,1411,1436]" pageId="6" pageNumber="173">FIGURES 711.</emphasis>
<taxonomicName id="4C033DD8FF93996F1C32FA4B6756FA39" authority="Leavengood &amp; Pinkerton &amp; Rifkind, 2022" authorityName="Leavengood &amp; Pinkerton &amp; Rifkind" authorityYear="2022" box="[336,526,1411,1435]" class="Insecta" family="Cleridae" genus="Tarsobaenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="173" phylum="Arthropoda" rank="species" species="piper" status="sp. nov.">
<emphasis id="B9779A49FF93996F1C32FA4B6756FA39" box="[336,526,1411,1435]" italics="true" pageId="6" pageNumber="173">Tarsobaenus piper</emphasis>
</taxonomicName>
,
<emphasis id="B9779A49FF93996F1F7CFA4C6701FA3E" bold="true" box="[542,601,1412,1436]" pageId="6" pageNumber="173">
<taxonomicNameLabel id="A2442732FF93996F1F7CFA4C6701FA3E" box="[542,601,1412,1436]" pageId="6" pageNumber="173" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(holotype, paratype): 7—holotype, dorsal habitus; 8—head; 9—paratype, dorsal habitus; 10—elytral plica; 11—lateral habitus.
</paragraph>
</caption>
<subSubSection id="C31915D0FF93996F1DA5FA256677F98E" pageId="6" pageNumber="173" type="diagnosis">
<paragraph id="8BBC465BFF93996F1DA5FA256677F98E" blockId="6.[151,1437,1517,1832]" pageId="6" pageNumber="173">
<emphasis id="B9779A49FF93996F1DA5FA256462F9A5" bold="true" box="[199,314,1517,1543]" pageId="6" pageNumber="173">Diagnosis</emphasis>
: This species is the only member of
<taxonomicName id="4C033DD8FF93996F1F80FA266628F9AA" authorityName="Leavengood, Pinkerton and Rifkind" authorityYear="2022" box="[738,880,1518,1544]" class="Insecta" family="Cleridae" genus="Tarsobaenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="B9779A49FF93996F1F80FA266628F9AA" box="[738,880,1518,1544]" italics="true" pageId="6" pageNumber="173">Tarsobaenus</emphasis>
</taxonomicName>
lacking a black elytral fascia. Except for the extreme epipleural margin, the elytra are concolorous orange.
</paragraph>
</subSubSection>
<subSubSection id="C31915D0FF93996F1DA5F9FD6055F9D7" pageId="6" pageNumber="173" type="description">
<paragraph id="8BBC465BFF93996F1DA5F9FD6055F9D7" blockId="6.[151,1437,1517,1832]" pageId="6" pageNumber="173">
<emphasis id="B9779A49FF93996F1DA5F9FD6460F9ED" bold="true" box="[199,312,1589,1615]" pageId="6" pageNumber="173">Variation</emphasis>
: Black markings on the lateral margins of the pronotum and elytra are absent in the more brightly colored
<typeStatus id="54B8F8F9FF93996F1D90F992640AF9D6" box="[242,338,1626,1652]" pageId="6" pageNumber="173" type="paratype">paratype</typeStatus>
(approx..
<quantity id="4CFBEBBEFF93996F1CA4F9916775F9D6" box="[454,557,1625,1652]" metricMagnitude="-3" metricUnit="m" metricValue="5.94" pageId="6" pageNumber="173" unit="mm" value="5.94">5.94 mm</quantity>
; specimen with head projecting and abdomen perhaps distended).
</paragraph>
</subSubSection>
<subSubSection id="C31915D0FF93996F1DA5F9B567FEF942" pageId="6" pageNumber="173" type="biology_ecology">
<paragraph id="8BBC465BFF93996F1DA5F9B567FEF942" blockId="6.[151,1437,1517,1832]" pageId="6" pageNumber="173">
<emphasis id="B9779A49FF93996F1DA5F9B56479F935" bold="true" box="[199,289,1661,1687]" pageId="6" pageNumber="173">Biology</emphasis>
:
<bibRefCitation id="EF923BAAFF93996F1C52F9B66759F93A" author="Letourneau, D. K." box="[304,513,1662,1688]" pageId="6" pageNumber="173" pagination="215 - 217" refId="ref7504" refString="Letourneau, D. K. (1990) Code of ant-plant mutualism broken by parasite. Science, 248, 215 - 217. https: // doi. org / 10.1126 / science. 248.4952.215" type="journal article" year="1990">Letourneau (1990)</bibRefCitation>
observed
<taxonomicName id="4C033DD8FF93996F1F11F9B66791F935" authorityName="Leavengood &amp; Pinkerton &amp; Rifkind" authorityYear="2022" box="[627,713,1662,1687]" class="Insecta" family="Cleridae" genus="Tarsobaenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="173" phylum="Arthropoda" rank="species" species="piper">
<emphasis id="B9779A49FF93996F1F11F9B66791F935" box="[627,713,1662,1687]" italics="true" pageId="6" pageNumber="173">T. piper</emphasis>
</taxonomicName>
in the petiole chambers of
<emphasis id="B9779A49FF93996F1E9AF9B6619CF93A" box="[1016,1220,1662,1688]" italics="true" pageId="6" pageNumber="173">Piper cenocladum</emphasis>
(
<taxonomicName id="4C033DD8FF93996F19B0F9B66017F93B" box="[1234,1359,1662,1689]" class="Magnoliopsida" family="Piperaceae" kingdom="Plantae" order="Piperales" pageId="6" pageNumber="173" phylum="Tracheophyta" rank="family">Piperaceae</taxonomicName>
) in the presence of plant-produced food bodies.
<bibRefCitation id="EF923BAAFF93996F1F07F96A666FF91E" author="Letourneau, D. K." box="[613,823,1698,1724]" pageId="6" pageNumber="173" pagination="4 - 8" refId="ref7541" refString="Letourneau, D. K. (1993) Ants that pay the Piper. Natural History, 102 (10), 4 - 8." type="journal article" year="1993">Letourneau (1993)</bibRefCitation>
suggested that both
<taxonomicName id="4C033DD8FF93996F1943F96A6121F919" authorityName="Leavengood &amp; Pinkerton &amp; Rifkind" authorityYear="2022" box="[1057,1145,1698,1723]" class="Insecta" family="Cleridae" genus="Tarsobaenus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="173" phylum="Arthropoda" rank="species" species="piper">
<emphasis id="B9779A49FF93996F1943F96A6121F919" box="[1057,1145,1698,1723]" italics="true" pageId="6" pageNumber="173">T. piper</emphasis>
</taxonomicName>
and
<taxonomicName id="4C033DD8FF93996F19D3F96A604DF91E" authorityName="Westwood" authorityYear="1839" box="[1201,1301,1698,1724]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="B9779A49FF93996F19D3F96A604DF91E" box="[1201,1301,1698,1724]" italics="true" pageId="6" pageNumber="173">Pheidole</emphasis>
</taxonomicName>
ants induce food body production by chemical stimulation.
</paragraph>
</subSubSection>
<subSubSection id="C31915D0FF93996F1DA5F92166CFF8A6" box="[199,919,1769,1796]" pageId="6" pageNumber="173" type="distribution">
<paragraph id="8BBC465BFF93996F1DA5F92166CFF8A6" blockId="6.[151,1437,1517,1832]" box="[199,919,1769,1796]" pageId="6" pageNumber="173">
<emphasis id="B9779A49FF93996F1DA5F9216401F8A1" bold="true" box="[199,345,1769,1795]" pageId="6" pageNumber="173">Distribution</emphasis>
:
<collectingRegion id="49C788B9FF93996F1C0AF9226491F8A6" box="[360,457,1770,1796]" country="Costa Rica" name="San Jose" pageId="6" pageNumber="173">San Jose</collectingRegion>
and
<collectingRegion id="49C788B9FF93996F1F62F9226723F8A6" box="[512,635,1770,1796]" country="Costa Rica" name="Puntarenas" pageId="6" pageNumber="173">Puntarenas</collectingRegion>
Provinces of
<collectingCountry id="F31406CBFF93996F1E77F92266CBF8A6" box="[789,915,1770,1796]" name="Costa Rica" pageId="6" pageNumber="173">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31915D0FF93996F1DA5F8C66054F88A" box="[199,1292,1806,1832]" pageId="6" pageNumber="173" type="etymology">
<paragraph id="8BBC465BFF93996F1DA5F8C66054F88A" blockId="6.[151,1437,1517,1832]" box="[199,1292,1806,1832]" pageId="6" pageNumber="173">
<emphasis id="B9779A49FF93996F1DA5F8C66410F88A" bold="true" box="[199,328,1806,1832]" pageId="6" pageNumber="173">Etymology</emphasis>
: This species is named after the genus (
<emphasis id="B9779A49FF93996F1E6AF8C6661EF885" box="[776,838,1806,1831]" italics="true" pageId="6" pageNumber="173">Piper</emphasis>
) of the plant with which it is associated.
</paragraph>
</subSubSection>
</treatment>
</document>