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<document id="ED56E195833F615E2F20EFC83557CC86" ID-DOI="10.5281/zenodo.206897" ID-GBIF-Dataset="70da07cc-4f7e-4154-a3ed-1a809b5aa039" ID-ISSN="1175-5326" ID-Zenodo-Dep="206897" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460245249099" checkinUser="plazi" docAuthor="Marino, Pablo I., Tóthová, Andrea &amp; Spinelli, Gustavo R" docDate="2011" docId="038C3A09FFFF652FFF39FCA14C26FE58" docLanguage="en" docName="zt02777p068.pdf" docOrigin="Zootaxa 2777" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Atrichopogon (Meloehelea) macrodentatum Marino, Tóthová &amp; Spinelli, 2011, new species" docType="treatment" docVersion="9" lastPageNumber="65" masterDocId="FFB54271FFFE652BFFAEFFCB4879FFDF" masterDocTitle="Two new Patagonian species of Atrichopogon (Meloehelea) (Diptera: Ceratopogonidae)" masterLastPageNumber="68" masterPageNumber="61" pageNumber="62" updateTime="1698241647024" updateUser="plazi">
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<mods:title id="AF8E08818749AC6106AB3DDDF54C6C73">Two new Patagonian species of Atrichopogon (Meloehelea) (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="36ED9837E6585958478B3D8ED043BE59">Marino, Pablo I.</mods:namePart>
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<mods:namePart id="6C356FDCEBF971A6E70C11D0457BE363">Tóthová, Andrea</mods:namePart>
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<mods:namePart id="7E69F09B348CE344516AFA2BCA2F5515">Spinelli, Gustavo R</mods:namePart>
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<mods:title id="E1224F21EA64F0A830E03146A0983B02">Zootaxa</mods:title>
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<mods:date id="475CB1072AB73AC5005BA6B8499EF774">2011</mods:date>
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<treatment id="038C3A09FFFF652FFF39FCA14C26FE58" ID-DOI="http://doi.org/10.5281/zenodo.6188623" ID-GBIF-Taxon="119390351" ID-Zenodo-Dep="6188623" LSID="urn:lsid:plazi:treatment:038C3A09FFFF652FFF39FCA14C26FE58" httpUri="http://treatment.plazi.org/id/038C3A09FFFF652FFF39FCA14C26FE58" lastPageId="4" lastPageNumber="65" pageId="1" pageNumber="62">
<subSubSection id="C33FD894FFFF652AFF39FCA14B39FC5B" box="[151,832,874,900]" pageId="1" pageNumber="62" type="nomenclature">
<paragraph id="8B9A8B1FFFFF652AFF39FCA14B39FC5B" blockId="1.[151,832,874,900]" box="[151,832,874,900]" pageId="1" pageNumber="62">
<heading id="D0D23C73FFFF652AFF39FCA14B39FC5B" bold="true" box="[151,832,874,900]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
<emphasis id="B951570DFFFF652AFF39FCA14B39FC5B" bold="true" box="[151,832,874,900]" pageId="1" pageNumber="62">
<taxonomicName id="4C25F09CFFFF652AFF39FCA14AD0FC5B" ID-CoL="8B5DS" box="[151,681,874,900]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="macrodentatum" status="sp. nov." subGenus="Meloehelea">
<emphasis id="B951570DFFFF652AFF39FCA14944FC5B" bold="true" box="[151,317,874,900]" italics="true" pageId="1" pageNumber="62">Atrichopogon</emphasis>
(
<emphasis id="B951570DFFFF652AFEE0FCA149A3FC5B" bold="true" box="[334,474,874,900]" italics="true" pageId="1" pageNumber="62">Meloehelea</emphasis>
)
<emphasis id="B951570DFFFF652AFE45FCA14AD0FC5B" bold="true" box="[491,681,874,900]" italics="true" pageId="1" pageNumber="62">macrodentatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A262EA76FFFF652AFD1EFCA14B39FC5B" box="[688,832,874,900]" pageId="1" pageNumber="62" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33FD894FFFF652AFF39FC7A4C61FBCE" pageId="1" pageNumber="62" type="diagnosis">
<paragraph id="8B9A8B1FFFFF652AFF39FC7A4C61FBCE" blockId="1.[151,1436,945,2019]" pageId="1" pageNumber="62">
<emphasis id="B951570DFFFF652AFF39FC7A4968FC15" bold="true" box="[151,273,945,970]" pageId="1" pageNumber="62">Diagnosis.</emphasis>
<emphasis id="B951570DFFFF652AFEB6FC7A49E8FC16" box="[280,401,945,969]" italics="true" pageId="1" pageNumber="62">Male adult</emphasis>
: only extant species of
<taxonomicName id="4C25F09CFFFF652AFD34FC7A4BA9FC16" authority="(Meloehelea)" baseAuthorityName="Meloehelea" box="[666,976,945,969]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B951570DFFFF652AFD34FC7A4B4DFC16" box="[666,820,945,969]" italics="true" pageId="1" pageNumber="62">Atrichopogon</emphasis>
(
<emphasis id="B951570DFFFF652AFCEAFC7A4BBEFC16" box="[836,967,945,969]" italics="true" pageId="1" pageNumber="62">Meloehelea</emphasis>
)
</taxonomicName>
from Patagonia with one subapical tooth in the gonostylus and aedeagal-parameral complex with posteromedial portion slightly notched.
<emphasis id="B951570DFFFF652AFB66FC1E4D25FC32" box="[1224,1372,981,1005]" italics="true" pageId="1" pageNumber="62">Female adult</emphasis>
: only extant species of
<taxonomicName id="4C25F09CFFFF652AFEF6FC3249A2FBCE" box="[344,475,1017,1041]" class="Insecta" family="Ceratopogonidae" genus="Meloehelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B951570DFFFF652AFEF6FC3249A2FBCE" box="[344,475,1017,1041]" italics="true" pageId="1" pageNumber="62">Meloehelea</emphasis>
</taxonomicName>
from Patagonia with very strong mandibular teeth.
</paragraph>
</subSubSection>
<subSubSection id="C33FD894FFFF6529FF6BFBD64AC6FE9D" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62" type="description">
<paragraph id="8B9A8B1FFFFF652AFF6BFBD64918F940" blockId="1.[151,1436,945,2019]" pageId="1" pageNumber="62">
<emphasis id="B951570DFFFF652AFF6BFBD6492FFBE9" bold="true" box="[197,342,1053,1078]" pageId="1" pageNumber="62">Description.</emphasis>
<emphasis id="B951570DFFFF652AFECEFBD649A7FBEA" box="[352,478,1053,1077]" italics="true" pageId="1" pageNumber="62">Male adult</emphasis>
:
<emphasis id="B951570DFFFF652AFE5EFBD64A49FBEA" bold="true" box="[496,560,1053,1077]" pageId="1" pageNumber="62">Head</emphasis>
: Dark brown (
<figureCitation id="131E979AFFFF652AFD70FBD64B59FBEA" box="[734,800,1053,1077]" captionStart="FIGURE 1" captionStartId="2.[151,250,1735,1758]" captionTargetBox="[164,1436,367,1701]" captionTargetId="figure@2.[151,1436,367,1713]" captionTargetPageId="2" captionText="FIGURE 1. Male of Atrichopogon (Meloehelea) macrodentatum Marino, Tóthová &amp; Spinelli, new species. A—head (dorsal view), B—mouthparts (dorsal view), C—thorax (dorsal view), D—wing, E—hypopygium (ventral view)." httpUri="https://zenodo.org/record/206898/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
A). Ommatidia without interfacet pubescence, abutting medially for length of 3 ommatidia. Antenna brown; with plume well developed; flagellomeres 58 at least partially fused; flagellomere 910 with plume setae, 1013 elongated; flagellomere 13 with apical projection slightly basally constricted; AR 1.00. Maxillary palpus brown (
<figureCitation id="131E979AFFFF652AFCAAFB424B3DFB7E" box="[772,836,1161,1185]" captionStart="FIGURE 1" captionStartId="2.[151,250,1735,1758]" captionTargetBox="[164,1436,367,1701]" captionTargetId="figure@2.[151,1436,367,1713]" captionTargetPageId="2" captionText="FIGURE 1. Male of Atrichopogon (Meloehelea) macrodentatum Marino, Tóthová &amp; Spinelli, new species. A—head (dorsal view), B—mouthparts (dorsal view), C—thorax (dorsal view), D—wing, E—hypopygium (ventral view)." httpUri="https://zenodo.org/record/206898/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
B); third segment slightly swollen at midlength, with well-developed pit located at midlength; segments 4, 5 separate; PR 3.00. Proboscis short, mouthparts length
<quantity id="4CDD26FAFFFF652AFAC5FB6648BFFB36" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="62" unit="mm" value="0.2">0.20 mm</quantity>
; head width/mouthparts length 2.00.
<emphasis id="B951570DFFFF652AFDD6FB194AA8FB35" bold="true" box="[632,721,1234,1258]" pageId="1" pageNumber="62">Thorax</emphasis>
: dark brown (
<figureCitation id="131E979AFFFF652AFCD5FB1A4BC7FB36" box="[891,958,1233,1257]" captionStart="FIGURE 1" captionStartId="2.[151,250,1735,1758]" captionTargetBox="[164,1436,367,1701]" captionTargetId="figure@2.[151,1436,367,1713]" captionTargetPageId="2" captionText="FIGURE 1. Male of Atrichopogon (Meloehelea) macrodentatum Marino, Tóthová &amp; Spinelli, new species. A—head (dorsal view), B—mouthparts (dorsal view), C—thorax (dorsal view), D—wing, E—hypopygium (ventral view)." httpUri="https://zenodo.org/record/206898/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
C); scutellum slightly paler than scutum; scutum with setae arising directly from surface; paratergite with 1 seta. Anepisternum well developed, sharply bilobed posteriorly. Legs light brown; hindtibia slightly expanded at apex; hindtibial spur length less than width of hindtibia at midlength; hindtibial comb with 8 spines; prothoracic
<collectingCountry id="F332CB8FFFFF652AFC3EFAF54BCDFA8A" box="[912,948,1342,1365]" name="Chile" pageId="1" pageNumber="62">TR</collectingCountry>
3.20; mesothoracic
<collectingCountry id="F332CB8FFFFF652AFB33FAF54CB8FA8A" box="[1181,1217,1342,1365]" name="Chile" pageId="1" pageNumber="62">TR</collectingCountry>
3.00; metathoracic
<collectingCountry id="F332CB8FFFFF652AFF39FAA948C2FAA6" box="[151,187,1378,1401]" name="Chile" pageId="1" pageNumber="62">TR</collectingCountry>
2.50; claws curved, bifid at tip, empodia present.
<emphasis id="B951570DFFFF652AFD48FAAA4B5FFAA5" bold="true" box="[742,806,1377,1402]" pageId="1" pageNumber="62">Wing</emphasis>
(
<figureCitation id="131E979AFFFF652AFC98FAAA4B0AFAA6" box="[822,883,1377,1401]" captionStart="FIGURE 1" captionStartId="2.[151,250,1735,1758]" captionTargetBox="[164,1436,367,1701]" captionTargetId="figure@2.[151,1436,367,1713]" captionTargetPageId="2" captionText="FIGURE 1. Male of Atrichopogon (Meloehelea) macrodentatum Marino, Tóthová &amp; Spinelli, new species. A—head (dorsal view), B—mouthparts (dorsal view), C—thorax (dorsal view), D—wing, E—hypopygium (ventral view)." httpUri="https://zenodo.org/record/206898/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
D) without pattern of pigmented membrane, with macrotrichia apically on r3, wing length
<quantity id="4CDD26FAFFFF652AFDFBFA4E4AC2FA42" box="[597,699,1413,1437]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="62" unit="mm" value="1.4">1.40 mm</quantity>
; breadth
<quantity id="4CDD26FAFFFF652AFC8FFA4E4BFEFA42" box="[801,903,1413,1437]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="62" unit="mm" value="0.46">0.46 mm</quantity>
; CR 0.64. Halter white.
<emphasis id="B951570DFFFF652AFB3AFA4E4D70FA41" bold="true" box="[1172,1289,1413,1438]" pageId="1" pageNumber="62">Abdomen</emphasis>
: tergites uniformly brown. Sternites more or less similarly rectangular shaped.
<emphasis id="B951570DFFFF652AFC3EFA604B87FA1B" bold="true" box="[912,1022,1451,1476]" pageId="1" pageNumber="62">Genitalia</emphasis>
(
<figureCitation id="131E979AFFFF652AFBBEFA674C29FA1B" box="[1040,1104,1452,1476]" captionStart="FIGURE 1" captionStartId="2.[151,250,1735,1758]" captionTargetBox="[164,1436,367,1701]" captionTargetId="figure@2.[151,1436,367,1713]" captionTargetPageId="2" captionText="FIGURE 1. Male of Atrichopogon (Meloehelea) macrodentatum Marino, Tóthová &amp; Spinelli, new species. A—head (dorsal view), B—mouthparts (dorsal view), C—thorax (dorsal view), D—wing, E—hypopygium (ventral view)." httpUri="https://zenodo.org/record/206898/files/figure.png" pageId="1" pageNumber="62">Fig. 1</figureCitation>
E): segment 9 about equal in width to segment 8, tergite 9 short, extending midlength of gonocoxite, posterior margin rounded. Sternite 9 with posterior margin slightly concave, with single row of 10 setae. Gonocoxite without medial lobe, moderately stout, 1.7 times longer than greatest breadth; gonostylus 0.75 as long as gonocoxite, nearly straight, apex gently curved, pointed, subapical stout tooth. Aedeagal-parameral complex broad; posterodorsal projection tapering to cap-like process; posteroventrally broad, posteromedial portion slightly notched. Cercus elongate, barely extending beyond apex of 9th tergite.
</paragraph>
<paragraph id="8B9A8B1FFFFF6529FF69F9604AC6FE9D" blockId="1.[151,1436,945,2019]" lastBlockId="2.[151,1436,151,323]" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62">
<emphasis id="B951570DFFFF652AFF69F9604926F91C" box="[199,351,1707,1731]" italics="true" pageId="1" pageNumber="62">Female adult</emphasis>
: As for male, with following differences.
<emphasis id="B951570DFFFF652AFCE2F9604BF5F91C" bold="true" box="[844,908,1707,1731]" pageId="1" pageNumber="62">Head</emphasis>
: antenna dark brown, with flagellomeres 18 compressed, slightly broader than long; flagellomeres 913 elongate, proportions as shown in
<figureCitation id="131E979AFFFF652AFB49F9044D54F938" box="[1255,1325,1743,1767]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1947]" captionTargetBox="[173,1408,193,1893]" captionTargetId="figure@3.[173,1409,193,1899]" captionTargetPageId="3" captionText="FIGURE 2. Female of Atrichopogon (Meloehelea) macrodentatum Marino, Tóthová &amp; Spinelli, new species. A—head (dorsal view), B—mouthparts (dorsal view), C— wing, D—thorax (lateral view), E— terminal abdominal segments and spermathecae (ventral view)." httpUri="https://zenodo.org/record/206899/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
A, flagellomere 13 with apical nipple, slightly constricted basally; AR 2.47 (2.062.86, n=5). Maxillary palpus with third segment bearing sensory pit slightly beyond midlength; PR 2.95 (2.503.22, n=5); mouthparts length 0.20 (0.16 0.22, n=5) mm, head width/mouthparts length 1.78 (1.631.96, n=5). Mandible well developed (
<figureCitation id="131E979AFFFF652AFB76F8F04D64F88C" box="[1240,1309,1851,1875]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1947]" captionTargetBox="[173,1408,193,1893]" captionTargetId="figure@3.[173,1409,193,1899]" captionTargetPageId="3" captionText="FIGURE 2. Female of Atrichopogon (Meloehelea) macrodentatum Marino, Tóthová &amp; Spinelli, new species. A—head (dorsal view), B—mouthparts (dorsal view), C— wing, D—thorax (lateral view), E— terminal abdominal segments and spermathecae (ventral view)." httpUri="https://zenodo.org/record/206899/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
B), with 18 (1520, n=5) strong teeth (
<figureCitation id="131E979AFFFF652AFE69F8944A74F8A8" box="[455,525,1887,1911]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1947]" captionTargetBox="[173,1408,193,1893]" captionTargetId="figure@3.[173,1409,193,1899]" captionTargetPageId="3" captionText="FIGURE 2. Female of Atrichopogon (Meloehelea) macrodentatum Marino, Tóthová &amp; Spinelli, new species. A—head (dorsal view), B—mouthparts (dorsal view), C— wing, D—thorax (lateral view), E— terminal abdominal segments and spermathecae (ventral view)." httpUri="https://zenodo.org/record/206899/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
B). Laciniae with pale indistinct teeth, without spicules.
<emphasis id="B951570DFFFF652AFB22F8944C9DF8A8" bold="true" box="[1164,1252,1887,1911]" pageId="1" pageNumber="62">Thorax</emphasis>
: Legs (
<figureCitation id="131E979AFFFF652AFA95F8944D07F8A8" box="[1339,1406,1887,1911]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1947]" captionTargetBox="[173,1408,193,1893]" captionTargetId="figure@3.[173,1409,193,1899]" captionTargetPageId="3" captionText="FIGURE 2. Female of Atrichopogon (Meloehelea) macrodentatum Marino, Tóthová &amp; Spinelli, new species. A—head (dorsal view), B—mouthparts (dorsal view), C— wing, D—thorax (lateral view), E— terminal abdominal segments and spermathecae (ventral view)." httpUri="https://zenodo.org/record/206899/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
D) light brown; hind tibia slightly expanded at apex; hindtibial spur length less than width of hind tibia at midlength; hindtibial comb with 8 spines; prothoracic
<collectingCountry id="F332CB8FFFFF652AFD20F8634ACBF860" box="[654,690,1960,1983]" name="Chile" pageId="1" pageNumber="62">TR</collectingCountry>
3.18 (3.003.40, n=5); mesothoracic
<collectingCountry id="F332CB8FFFFF652AFBC4F8634CF7F860" box="[1130,1166,1960,1983]" name="Chile" pageId="1" pageNumber="62">TR</collectingCountry>
2.81 (2.612.92, n=5); metathoracic
<collectingCountry id="F332CB8FFFFF652AFE80F807492BF83C" box="[302,338,1996,2019]" name="Chile" pageId="1" pageNumber="62">TR</collectingCountry>
2.48 (2.392.57, n=5); empodia present.
<emphasis id="B951570DFFFF652AFCB1F8014B26F83C" bold="true" box="[799,863,1994,2019]" pageId="1" pageNumber="62">Wing</emphasis>
(
<figureCitation id="131E979AFFFF652AFCC1F8004BC8F83C" box="[879,945,1995,2019]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1947]" captionTargetBox="[173,1408,193,1893]" captionTargetId="figure@3.[173,1409,193,1899]" captionTargetPageId="3" captionText="FIGURE 2. Female of Atrichopogon (Meloehelea) macrodentatum Marino, Tóthová &amp; Spinelli, new species. A—head (dorsal view), B—mouthparts (dorsal view), C— wing, D—thorax (lateral view), E— terminal abdominal segments and spermathecae (ventral view)." httpUri="https://zenodo.org/record/206899/files/figure.png" pageId="1" pageNumber="62">Fig. 2</figureCitation>
C) without pattern of pigmented membrane, with macrotrichia very abundant in r3 and m1, 15
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m2, 1
<quantity id="4CDD26FAFFFC6529FCE5FF5C4B0FFF70" box="[843,886,151,175]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="2" pageNumber="63" unit="in" value="8.0">8 in</quantity>
cua1, and
<quantity id="4CDD26FAFFFC6529FC46FF5C4C56FF70" box="[1000,1071,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="7.62" metricValueMin="0.0" pageId="2" pageNumber="63" unit="in" value="1.5" valueMax="3.0" valueMin="0.0">03 in</quantity>
anal cell; radial cells with broad lumen, second 4 times longer than first; wing length 1.50 (1.361.64, n=5) mm; breadth 0.63 (0.580.68, n=5) mm; CR 0.71 (0.690.73, n=5). Halter white.
<emphasis id="B951570DFFFC6529FDC5FF294A99FF24" bold="true" box="[619,736,226,251]" pageId="2" pageNumber="63">Abdomen</emphasis>
(
<figureCitation id="131E979AFFFC6529FD5FFF294B4EFF25" box="[753,823,226,250]" captionStart="FIGURE 2" captionStartId="3.[151,250,1924,1947]" captionTargetBox="[173,1408,193,1893]" captionTargetId="figure@3.[173,1409,193,1899]" captionTargetPageId="3" captionText="FIGURE 2. Female of Atrichopogon (Meloehelea) macrodentatum Marino, Tóthová &amp; Spinelli, new species. A—head (dorsal view), B—mouthparts (dorsal view), C— wing, D—thorax (lateral view), E— terminal abdominal segments and spermathecae (ventral view)." httpUri="https://zenodo.org/record/206899/files/figure.png" pageId="2" pageNumber="63">Fig. 2</figureCitation>
E): tergites uniformly brown. Sternite 8 without elongate, curved setae. Two subequal ovoid spermathaecae, each with elongated neck, measuring
<date id="FF9BADDFFFFC6529FB1DFECC4C9CFEC0" box="[1203,1253,263,287]" pageId="2" pageNumber="63" value="1969-10">69 x</date>
49 µm, neck 10 µm, and
<date id="FF9BADDFFFFC6529FF54FEE14952FE9D" box="[250,299,298,322]" pageId="2" pageNumber="63" value="1973-10">73 x</date>
54 µm, neck 11 µm. Cercus brown.
</paragraph>
</subSubSection>
<caption id="DF5ADB97FFFC6529FF39F90C4CBFF923" httpUri="https://zenodo.org/record/206898/files/figure.png" pageId="2" pageNumber="63" targetBox="[164,1436,367,1701]" targetPageId="2">
<paragraph id="8B9A8B1FFFFC6529FF39F90C4CBFF923" blockId="2.[151,1435,1735,1788]" pageId="2" pageNumber="63">
<emphasis id="B951570DFFFC6529FF39F90C496DF903" bold="true" box="[151,276,1735,1758]" pageId="2" pageNumber="63">FIGURE 1.</emphasis>
Male of
<taxonomicName id="4C25F09CFFFC6529FEDAF90C4C24F902" authority="Marino, Tothova &amp; Spinelli" authorityName="Marino, Tothova &amp; Spinelli" box="[372,1117,1735,1757]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="macrodentatum" status="sp. nov." subGenus="Meloehelea">
<emphasis id="B951570DFFFC6529FEDAF90C4986F902" box="[372,511,1735,1757]" italics="true" pageId="2" pageNumber="63">Atrichopogon</emphasis>
(
<emphasis id="B951570DFFFC6529FDBEF90C4AFFF902" box="[528,646,1735,1757]" italics="true" pageId="2" pageNumber="63">Meloehelea</emphasis>
)
<emphasis id="B951570DFFFC6529FD38F90C4B4DF902" box="[662,820,1735,1757]" italics="true" pageId="2" pageNumber="63">macrodentatum</emphasis>
Marino, Tóthová &amp; Spinelli
</taxonomicName>
,
<taxonomicNameLabel id="A262EA76FFFC6529FBC7F9034C9EF902" box="[1129,1255,1735,1757]" pageId="2" pageNumber="63" rank="species">new species</taxonomicNameLabel>
. A—head (dorsal view), B—mouthparts (dorsal view), C—thorax (dorsal view), D—wing, E—hypopygium (ventral view).
</paragraph>
</caption>
<subSubSection id="C33FD894FFFC652FFF6BF8E34C26FE58" lastPageId="4" lastPageNumber="65" pageId="2" pageNumber="63" type="distribution">
<paragraph id="8B9A8B1FFFFC6529FF6BF8E34985F857" blockId="2.[151,1436,1832,2037]" pageId="2" pageNumber="63">
<emphasis id="B951570DFFFC6529FF6BF8E34A6FF89E" bold="true" box="[197,534,1832,1857]" pageId="2" pageNumber="63">Distribution and bionomics.</emphasis>
<taxonomicName id="4C25F09CFFFC6529FDB0F8E24B09F89E" box="[542,880,1833,1857]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="macrodentatum">
<emphasis id="B951570DFFFC6529FDB0F8E24B09F89E" box="[542,880,1833,1857]" italics="true" pageId="2" pageNumber="63">Atrichopogon macrodentatum</emphasis>
</taxonomicName>
inhabits
<taxonomicName id="4C25F09CFFFC6529FC77F8E24C23F89E" box="[985,1114,1833,1857]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="63" phylum="Tracheophyta" rank="genus">
<emphasis id="B951570DFFFC6529FC77F8E24C23F89E" box="[985,1114,1833,1857]" italics="true" pageId="2" pageNumber="63">Nothofagus</emphasis>
</taxonomicName>
forests within the Valdivian forests of
<collectingCountry id="F332CB8FFFFC6529FEA4F8874900F8BB" box="[266,377,1868,1892]" name="Argentina" pageId="2" pageNumber="63">Argentina</collectingCountry>
and
<collectingCountry id="F332CB8FFFFC6529FE1DF887498AF8BB" box="[435,499,1868,1892]" name="Chile" pageId="2" pageNumber="63">Chile</collectingCountry>
, at elevations of
<quantity id="4CDD26FAFFFC6529FD18F8874B40F8BB" box="[694,825,1868,1892]" metricMagnitude="2" metricUnit="m" metricValue="8.705" metricValueMax="9.5" metricValueMin="7.909999999999999" pageId="2" pageNumber="63" unit="m" value="870.5" valueMax="950.0" valueMin="791.0">
791
<specimenCount id="9D234096FFFC6529FD43F8874B40F8BB" box="[749,825,1868,1892]" pageId="2" pageNumber="63" type="generic">950 m</specimenCount>
</quantity>
. The dominant species in these forests are
<taxonomicName id="4C25F09CFFFC6529FA8AF8854DE2F8BA" box="[1316,1435,1869,1893]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="63" phylum="Tracheophyta" rank="species" species="pumilio">
<emphasis id="B951570DFFFC6529FA8AF8854D4EF8BA" box="[1316,1335,1870,1893]" italics="true" pageId="2" pageNumber="63">N</emphasis>
.
<emphasis id="B951570DFFFC6529FAE8F8864DE2F8BA" box="[1350,1435,1869,1893]" italics="true" pageId="2" pageNumber="63">pumilio</emphasis>
</taxonomicName>
(lenga) and
<taxonomicName id="4C25F09CFFFC6529FEB2F8B949D6F856" box="[284,431,1905,1929]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="63" phylum="Tracheophyta" rank="species" species="antarctica">
<emphasis id="B951570DFFFC6529FEB2F8B94956F856" box="[284,303,1906,1929]" italics="true" pageId="2" pageNumber="63">N</emphasis>
.
<emphasis id="B951570DFFFC6529FE93F8B949D6F856" box="[317,431,1906,1929]" italics="true" pageId="2" pageNumber="63">antarctica</emphasis>
</taxonomicName>
(ñire).
</paragraph>
<paragraph id="8B9A8B1FFFFC652FFF6BF85F4A87FF28" blockId="2.[151,1436,1832,2037]" lastBlockId="4.[151,1436,151,392]" lastPageId="4" lastPageNumber="65" pageId="2" pageNumber="63">
<emphasis id="B951570DFFFC6529FF6BF85F49BDF872" bold="true" box="[197,452,1940,1965]" pageId="2" pageNumber="63">Specimens examined.</emphasis>
<typeStatus id="549E35BDFFFC6529FE64F85E4A68F872" box="[458,529,1941,1965]" pageId="2" pageNumber="63">Types</typeStatus>
:
<typeStatus id="549E35BDFFFC6529FDB5F85F4AFAF873" box="[539,643,1940,1964]" pageId="2" pageNumber="63" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F332CB8FFFFC6529FD60F85F4B39F873" box="[718,832,1940,1964]" name="Argentina" pageId="2" pageNumber="63">Argentina</collectingCountry>
, Parque Nacional Nahuel Huapi, arroyo Neuquén-Co,
<geoCoordinate id="EE11EDD8FFFC6529FF39F8734954F80F" box="[151,301,1976,2000]" direction="south" orientation="latitude" pageId="2" pageNumber="63" precision="1" value="-40.480083">40º2848.3”S</geoCoordinate>
<geoCoordinate id="EE11EDD8FFFC6529FE99F87349A1F80F" box="[311,472,1976,2000]" direction="west" orientation="longitude" pageId="2" pageNumber="63" precision="1" value="-71.61225">71º3644.1”W</geoCoordinate>
,
<quantity id="4CDD26FAFFFC6529FE48F8734A49F80F" box="[486,560,1976,2000]" metricMagnitude="2" metricUnit="m" metricValue="8.09" pageId="2" pageNumber="63" unit="m" value="809.0">809 m</quantity>
, 23.i/
<date id="FF9BADDFFFFC6529FDDFF8734AAEF80F" box="[625,727,1976,2000]" pageId="2" pageNumber="63" value="2008-02-04">4.ii.2008</date>
, A. Garré F. Montes de Oca, Malaise trap. Allotype female, same data except CDC light trap (MLP).
<typeStatus id="549E35BDFFFC6529FDCAF8164AADF82A" box="[612,724,2013,2037]" pageId="2" pageNumber="63" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D234096FFFC6529FD71F8174B37F82B" box="[735,846,2012,2036]" pageId="2" pageNumber="63" type="female">4 females</specimenCount>
, as follows: same data as
<typeStatus id="549E35BDFFFC6529FBC1F8174CADF82B" box="[1135,1236,2012,2036]" pageId="2" pageNumber="63" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D234096FFFC6529FB71F8174D39F82B" box="[1247,1344,2012,2036]" pageId="2" pageNumber="63" type="female">1 female</specimenCount>
(MLP);
<collectingCountry id="F332CB8FFFFA652FFF39FF5C4970FF70" box="[151,265,151,175]" name="Argentina" pageId="4" pageNumber="65">Argentina</collectingCountry>
, Parque Nacional Nahuel Huapi, Puerto Blest, Estación Biológica,
<geoCoordinate id="EE11EDD8FFFA652FFBAEFF5C4CE1FF70" box="[1024,1176,151,175]" direction="south" orientation="latitude" pageId="4" pageNumber="65" precision="1" value="-41.026222">41º0134.4”S</geoCoordinate>
<geoCoordinate id="EE11EDD8FFFA652FFB31FF5C4D38FF70" box="[1183,1345,151,175]" direction="west" orientation="longitude" pageId="4" pageNumber="65" precision="1" value="-71.815475">71º4855.7”W</geoCoordinate>
,
<quantity id="4CDD26FAFFFA652FFAE3FF5C4DEEFF70" box="[1357,1431,151,175]" metricMagnitude="2" metricUnit="m" metricValue="7.909999999999999" pageId="4" pageNumber="65" unit="m" value="791.0">791 m</quantity>
, 6/
<date id="FF9BADDFFFFA652FFF01FF774960FF0B" box="[175,281,188,212]" pageId="4" pageNumber="65" value="2007-01-13">13.i.2007</date>
, A. Garré F. Montes de Oca,
<specimenCount id="9D234096FFFA652FFD30FF774B6AFF0B" box="[670,787,188,212]" pageId="4" pageNumber="65" type="female">2 females</specimenCount>
, at light (MLP);
<collectingCountry id="F332CB8FFFFA652FFC4DFF774C5AFF0B" box="[995,1059,188,212]" name="Chile" pageId="4" pageNumber="65">Chile</collectingCountry>
, Cautín Prov., Lago Conguillio,
<date id="FF9BADDFFFFA652FFF39FF144973FF28" box="[151,266,223,247]" pageId="4" pageNumber="65" value="1984-12-06">6.xii.1984</date>
, J. A. Downes,
<specimenCount id="9D234096FFFA652FFE15FF144A64FF28" box="[443,541,223,247]" pageId="4" pageNumber="65" type="female">1 female</specimenCount>
(
<date id="FF9BADDFFFFA652FFD82FF144AE8FF28" box="[556,657,223,247]" pageId="4" pageNumber="65" value="1661-01-08">1661/8/1</date>
) (CNCI).
</paragraph>
<caption id="DF5ADB97FFFD6528FF39F84F4954F807" httpUri="https://zenodo.org/record/206899/files/figure.png" pageId="3" pageNumber="64" targetBox="[173,1408,193,1893]" targetPageId="3">
<paragraph id="8B9A8B1FFFFD6528FF39F84F4954F807" blockId="3.[151,1436,1924,2008]" pageId="3" pageNumber="64">
<emphasis id="B951570DFFFD6528FF39F84F496BF845" bold="true" box="[151,274,1924,1947]" pageId="3" pageNumber="64">FIGURE 2.</emphasis>
Female of
<taxonomicName id="4C25F09CFFFD6528FE2AF84E4C1DF845" authority="Marino, Tothova &amp; Spinelli" authorityName="Marino, Tothova &amp; Spinelli" box="[388,1124,1924,1947]" class="Insecta" family="Ceratopogonidae" genus="Atrichopogon" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="macrodentatum" status="sp. nov." subGenus="Meloehelea">
<emphasis id="B951570DFFFD6528FE2AF84E4A69F844" box="[388,528,1925,1947]" italics="true" pageId="3" pageNumber="64">Atrichopogon</emphasis>
(
<emphasis id="B951570DFFFD6528FDB0F84E4AEDF844" box="[542,660,1925,1947]" italics="true" pageId="3" pageNumber="64">Meloehelea</emphasis>
)
<emphasis id="B951570DFFFD6528FD0DF84E4B38F844" box="[675,833,1925,1947]" italics="true" pageId="3" pageNumber="64">macrodentatum</emphasis>
Marino, Tóthová &amp; Spinelli
</taxonomicName>
,
<taxonomicNameLabel id="A262EA76FFFD6528FBC0F84E4C92F845" box="[1134,1259,1924,1946]" pageId="3" pageNumber="64" rank="species">new species</taxonomicNameLabel>
. A—head (dorsal view), B—mouthparts (dorsal view), C— wing, D—thorax (lateral view), E— terminal abdominal segments and spermathecae (ventral view).
</paragraph>
</caption>
<paragraph id="8B9A8B1FFFFA652FFF6BFEC84CACFEBB" blockId="4.[151,1436,151,392]" pageId="4" pageNumber="65">
<emphasis id="B951570DFFFA652FFF6BFEC849A9FEC3" bold="true" box="[197,464,259,284]" pageId="4" pageNumber="65">Taxonomic discussion.</emphasis>
The female of this new species is easily distinguished from their congeners of the subgenus
<taxonomicName id="4C25F09CFFFA652FFF70FEE34918FE9F" box="[222,353,296,320]" class="Insecta" family="Ceratopogonidae" genus="Meloehelea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B951570DFFFA652FFF70FEE34918FE9F" box="[222,353,296,320]" italics="true" pageId="4" pageNumber="65">Meloehelea</emphasis>
</taxonomicName>
by the very strong mandibular teeth. The male have only one tooth in the gonostylus. Other characters to distinguish this species from the remaining Patagonian species may be found in the key.
</paragraph>
<paragraph id="8B9A8B1FFFFA652FFF6BFEA44C26FE58" blockId="4.[151,1436,151,392]" box="[197,1119,367,392]" pageId="4" pageNumber="65">
<emphasis id="B951570DFFFA652FFF6BFEA44A5AFE57" bold="true" box="[197,547,367,392]" pageId="4" pageNumber="65">Derivation of specific epithet.</emphasis>
A reference to the strong female mandibular teeth.
</paragraph>
</subSubSection>
</treatment>
</document>