123 lines
18 KiB
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123 lines
18 KiB
XML
<document id="6190467D0139247603600061FB15F55A" ID-DOI="10.11646/zootaxa.3847.3.6" ID-GBIF-Dataset="733d91cd-37eb-436a-abeb-a66007ed52e5" ID-ISSN="1175-5326" ID-Zenodo-Dep="224831" ID-ZooBank="FE775E06-1C07-4792-BF95-27484FE77957" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460879163011" checkinUser="plazi" docAuthor="Burke, Alan F. & Zolnerowich, Gregory" docDate="2014" docId="D76087FEFFE78E01FF71E26FFE36B6C4" docLanguage="en" docName="zt03847p430.pdf" docOrigin="Zootaxa 3847 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cymatodera ochlera" docType="treatment" docVersion="8" lastPageNumber="428" masterDocId="2B59FF86FFE38E04FFE6E47CFFD8B42B" masterDocTitle="Cymatodera ochlera Barr, a junior synonym of Cymatodera wolcotti Barr, with a comparison to similar species (Coleoptera: Cleridae)" masterLastPageNumber="430" masterPageNumber="423" pageNumber="427" updateTime="1698358244104" updateUser="plazi">
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<mods:mods id="8AAC2DC1356270F65F8534C22FCA7F6F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9E008299EC0AB3FFDD336A094280CD42">
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<mods:title id="17F962D5AB1A3B4C0928940228AC2CC1">Cymatodera ochlera Barr, a junior synonym of Cymatodera wolcotti Barr, with a comparison to similar species (Coleoptera: Cleridae)</mods:title>
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<mods:name id="53FBE19713FF90F0252C98B0C465DE25" type="personal">
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<mods:namePart id="0DF91CC160495CE8ABCBB908545C9928">Burke, Alan F.</mods:namePart>
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<mods:name id="659A0FA50FF8781A7DC5446599F041DD" type="personal">
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<mods:role id="A109F90E24FD791024CC2C9B5E9BA8DE">
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<mods:roleTerm id="279DED8688FD1DF7ECD0EBB51A101A86">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="20FB7AA406716BB05C4402A1BEB3F59C">Zolnerowich, Gregory</mods:namePart>
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<mods:typeOfResource id="BDB5DD59A5ECAFFC01BBCE0E8E0D2AA7">text</mods:typeOfResource>
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<mods:titleInfo id="1675FCAAFA69608657427FC216D9FF76">
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<mods:title id="D027683CB2086D0A57477DD125C448D1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3E7217855B1BAE1EFF8CFA34721C6E59">
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<mods:date id="06525947F19B7155CDE1492C4BBBA136">2014</mods:date>
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<mods:detail id="402D9CC3EB846079A501230CEB41ED75" type="volume">
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<mods:number id="DA55247909BCDD698202F7FAB0E4BBD1">3847</mods:number>
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</mods:detail>
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<mods:detail id="16907350928AF80ED2FDC0AAEA77651E" type="issue">
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<mods:number id="3D5EF1512FD24BF83B5D2A5F75AA3AE4">3</mods:number>
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<mods:end id="B81FB7640347FCDD4D7C96E2BF539919">430</mods:end>
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<mods:identifier id="79CF8D188696CC6FD50F05A3438E2E55" type="DOI">10.11646/zootaxa.3847.3.6</mods:identifier>
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<mods:identifier id="B43C66271BB9D7A5536F4B3EEAB2EF09" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="39DD1C2DAEEF83A9FFBB67E671016359" type="Zenodo-Dep">224831</mods:identifier>
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<treatment id="D76087FEFFE78E01FF71E26FFE36B6C4" ID-DOI="http://doi.org/10.5281/zenodo.6144690" ID-GBIF-Taxon="125818440" ID-Zenodo-Dep="6144690" LSID="urn:lsid:plazi:treatment:D76087FEFFE78E01FF71E26FFE36B6C4" httpUri="http://treatment.plazi.org/id/D76087FEFFE78E01FF71E26FFE36B6C4" lastPageId="5" lastPageNumber="428" pageId="4" pageNumber="427">
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<subSubSection id="17D36563FFE78E01FF71E26FFE36B6C4" lastPageId="5" lastPageNumber="428" pageId="4" pageNumber="427" type="nomenclature">
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<paragraph id="5F7636E8FFE78E00FF71E26FFD92B206" blockId="4.[151,586,1555,1582]" box="[151,586,1555,1582]" pageId="4" pageNumber="427">
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<heading id="043E8184FFE78E00FF71E26FFD92B206" bold="true" box="[151,586,1555,1582]" fontSize="11" level="1" pageId="4" pageNumber="427" reason="1">
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<emphasis id="6DBDEAFAFFE78E00FF71E26FFD92B206" bold="true" box="[151,586,1555,1582]" pageId="4" pageNumber="427">
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<taxonomicName id="98C94D6BFFE78E00FF71E26FFE54B206" box="[151,396,1555,1581]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="ochlera">
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<emphasis id="6DBDEAFAFFE78E00FF71E26FFE54B206" bold="true" box="[151,396,1555,1581]" italics="true" pageId="4" pageNumber="427">Cymatodera ochlera</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="98C94D6BFFE78E00FE2CE26FFD92B206" box="[458,586,1555,1581]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="wolcotti">
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<emphasis id="6DBDEAFAFFE78E00FE2CE26FFD92B206" bold="true" box="[458,586,1555,1581]" italics="true" pageId="4" pageNumber="427">C. wolcotti</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="5F7636E8FFE78E01FF71E226FDF3B5A3" blockId="4.[151,1437,1626,2011]" lastBlockId="5.[151,1437,151,752]" lastPageId="5" lastPageNumber="428" pageId="4" pageNumber="427">
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The diagnostic characters cited by Barr indicate that the sixth visible ventrite of male
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<taxonomicName id="98C94D6BFFE78E00FB9FE227FB29B259" box="[1145,1265,1626,1650]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="ochlera">
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<emphasis id="6DBDEAFAFFE78E00FB9FE227FB29B259" box="[1145,1265,1626,1650]" italics="true" pageId="4" pageNumber="427">C. ochlera</emphasis>
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</taxonomicName>
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has the lateral margins moderately oblique and the posterolateral angles moderately curved inward, while male specimens of
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<taxonomicName id="98C94D6BFFE78E00FA64E203FF2AB291" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="wolcotti">
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<emphasis id="6DBDEAFAFFE78E00FA64E203FF2AB291" italics="true" pageId="4" pageNumber="427">C. wolcotti</emphasis>
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</taxonomicName>
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have the lateral margins of the sixth ventrite straight and the posterolateral angles extended posteriorly. A number of male specimens of both species were examined, and the sixth tergite of these specimens was very consistent and only slight variation in the posterior portion of the lateral margins was observed. Specifically, this portion of the segment can be moderately curved inwardly (
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<figureCitation id="C7F22A6DFFE78E00FCACE372FC44B30C" box="[842,924,1806,1831]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1827,1850]" captionTargetBox="[159,1424,207,1768]" captionTargetId="figure@3.[151,1436,193,1775]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Sixth abdominal segment of: 9 Cymatodera balteata (male, dorsal view); 10 C. ochlera (male, dorsal view); 11 C. wolcotti (male, dorsal view); 12 C. undulata (male, dorsal view); 13 C. balteata (male, ventral view); 14 C. ochlera (male, ventral view); 15 C. wolcotti (male, ventral view); 16 C. wolcotti paratype (male, ventral view); 17 C. wolcotti (male, ventral view); 18 C. undulata (male, ventral view). For Figs. 10 – 11, arrows indicate curvature of posterolateral margins. For Figs. 14 – 17, arrows indicate direction of hind angles." httpUri="https://zenodo.org/record/224834/files/figure.png" pageId="4" pageNumber="427">Fig. 11</figureCitation>
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) to slightly curved outwardly (
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<figureCitation id="C7F22A6DFFE78E00FAE2E372FA82B30C" box="[1284,1370,1806,1831]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1827,1850]" captionTargetBox="[159,1424,207,1768]" captionTargetId="figure@3.[151,1436,193,1775]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Sixth abdominal segment of: 9 Cymatodera balteata (male, dorsal view); 10 C. ochlera (male, dorsal view); 11 C. wolcotti (male, dorsal view); 12 C. undulata (male, dorsal view); 13 C. balteata (male, ventral view); 14 C. ochlera (male, ventral view); 15 C. wolcotti (male, ventral view); 16 C. wolcotti paratype (male, ventral view); 17 C. wolcotti (male, ventral view); 18 C. undulata (male, ventral view). For Figs. 10 – 11, arrows indicate curvature of posterolateral margins. For Figs. 14 – 17, arrows indicate direction of hind angles." httpUri="https://zenodo.org/record/224834/files/figure.png" pageId="4" pageNumber="427">Fig. 10</figureCitation>
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). The lateral margins of the sixth visible ventrite were very consistent among these individuals, with no noticeable variation. Similarly, male specimens identified as
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<taxonomicName id="98C94D6BFFE78E00FD23E32BFC98B345" box="[709,832,1878,1902]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="wolcotti">
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<emphasis id="6DBDEAFAFFE78E00FD23E32BFC98B345" box="[709,832,1878,1902]" italics="true" pageId="4" pageNumber="427">C. wolcotti</emphasis>
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</taxonomicName>
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exhibited the posterolateral angles of the sixth visible ventrite from strongly curved inwardly (
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<figureCitation id="C7F22A6DFFE78E00FD85E306FD60B3B8" box="[611,696,1914,1939]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1827,1850]" captionTargetBox="[159,1424,207,1768]" captionTargetId="figure@3.[151,1436,193,1775]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Sixth abdominal segment of: 9 Cymatodera balteata (male, dorsal view); 10 C. ochlera (male, dorsal view); 11 C. wolcotti (male, dorsal view); 12 C. undulata (male, dorsal view); 13 C. balteata (male, ventral view); 14 C. ochlera (male, ventral view); 15 C. wolcotti (male, ventral view); 16 C. wolcotti paratype (male, ventral view); 17 C. wolcotti (male, ventral view); 18 C. undulata (male, ventral view). For Figs. 10 – 11, arrows indicate curvature of posterolateral margins. For Figs. 14 – 17, arrows indicate direction of hind angles." httpUri="https://zenodo.org/record/224834/files/figure.png" pageId="4" pageNumber="427">Fig. 15</figureCitation>
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), to posteriorly extended (
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<figureCitation id="C7F22A6DFFE78E00FC00E306FBE2B3B8" box="[998,1082,1914,1939]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1827,1850]" captionTargetBox="[159,1424,207,1768]" captionTargetId="figure@3.[151,1436,193,1775]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Sixth abdominal segment of: 9 Cymatodera balteata (male, dorsal view); 10 C. ochlera (male, dorsal view); 11 C. wolcotti (male, dorsal view); 12 C. undulata (male, dorsal view); 13 C. balteata (male, ventral view); 14 C. ochlera (male, ventral view); 15 C. wolcotti (male, ventral view); 16 C. wolcotti paratype (male, ventral view); 17 C. wolcotti (male, ventral view); 18 C. undulata (male, ventral view). For Figs. 10 – 11, arrows indicate curvature of posterolateral margins. For Figs. 14 – 17, arrows indicate direction of hind angles." httpUri="https://zenodo.org/record/224834/files/figure.png" pageId="4" pageNumber="427">Fig. 17</figureCitation>
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), with a number of specimens, identified either as
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<taxonomicName id="98C94D6BFFE78E00FE93E3E3FE2BB39D" box="[373,499,1950,1974]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="wolcotti">
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<emphasis id="6DBDEAFAFFE78E00FE93E3E3FE2BB39D" box="[373,499,1950,1974]" italics="true" pageId="4" pageNumber="427">C. wolcotti</emphasis>
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</taxonomicName>
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or as
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<taxonomicName id="98C94D6BFFE78E00FDD8E3E3FD6EB39D" box="[574,694,1950,1974]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="427" phylum="Arthropoda" rank="species" species="ochlera">
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<emphasis id="6DBDEAFAFFE78E00FDD8E3E3FD6EB39D" box="[574,694,1950,1974]" italics="true" pageId="4" pageNumber="427">C. ochlera</emphasis>
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</taxonomicName>
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, with these angles moderately extended inwardly (
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<figureCitation id="C7F22A6DFFE78E00FAE7E3E2FA57B39C" box="[1281,1423,1950,1975]" captionStart="FIGURES 9 – 18" captionStartId="3.[151,264,1827,1850]" captionTargetBox="[159,1424,207,1768]" captionTargetId="figure@3.[151,1436,193,1775]" captionTargetPageId="3" captionText="FIGURES 9 – 18. Sixth abdominal segment of: 9 Cymatodera balteata (male, dorsal view); 10 C. ochlera (male, dorsal view); 11 C. wolcotti (male, dorsal view); 12 C. undulata (male, dorsal view); 13 C. balteata (male, ventral view); 14 C. ochlera (male, ventral view); 15 C. wolcotti (male, ventral view); 16 C. wolcotti paratype (male, ventral view); 17 C. wolcotti (male, ventral view); 18 C. undulata (male, ventral view). For Figs. 10 – 11, arrows indicate curvature of posterolateral margins. For Figs. 14 – 17, arrows indicate direction of hind angles." httpUri="https://zenodo.org/record/224834/files/figure.png" pageId="4" pageNumber="427">Figs. 14, 16</figureCitation>
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). Moreover, the two species share very similar male genitalia. Male genitalia were extracted and examined from a number of specimens from both species, and minor variation was observed, with the general structure of the aedeagus as follows: shape rather robust; parameres developed, broad, subtriangular in shape, lateral margins sinuate, distally pointed; phallobase narrowing toward base; phallus with copulatory piece acuminate at apex and partially covered by tegmen; phallic plate armed with two rows of denticles that gradually become reduced in size toward apex; phallobasic apodeme robust distally, endophallic struts distally slender (
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<figureCitation id="C7F22A6DFFE68E01FBBFE55BFB1CB56B" box="[1113,1220,295,320]" captionStart="FIGURES 5 – 8" captionStartId="2.[151,264,1802,1825]" captionTargetBox="[454,1124,234,1740]" captionTargetId="figure@2.[443,1136,220,1748]" captionTargetPageId="2" captionText="FIGURES 5 – 8. Male genitalia of: 5 Cymatodera balteata; 6 C. ochlera; 7 C. wolcotti; 8 C. undulata." httpUri="https://zenodo.org/record/224833/files/figure.png" pageId="5" pageNumber="428">Figs. 6–7</figureCitation>
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). Variation in male genitalia observed among those revised specimens are: length of each row of denticles somewhat variable, and phallus feebly to moderately robust.
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</paragraph>
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<paragraph id="5F7636E8FFE68E01FF21E5E9FE36B6C4" blockId="5.[151,1437,151,752]" pageId="5" pageNumber="428">
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Barr indicated females of these species are remarkably similar, but they could be separated based on the distance between the eyes. For the females of
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<taxonomicName id="98C94D6BFFE68E01FD42E5C4FCC2B5FB" box="[676,794,440,464]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="428" phylum="Arthropoda" rank="species" species="ochlera">
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<emphasis id="6DBDEAFAFFE68E01FD42E5C4FCC2B5FB" box="[676,794,440,464]" italics="true" pageId="5" pageNumber="428">C. ochlera</emphasis>
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</taxonomicName>
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, the distance between the eyes is twice the length of the eleventh antennomere, while for
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<taxonomicName id="98C94D6BFFE68E01FDEAE5A0FD50B5DF" box="[524,648,476,500]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="428" phylum="Arthropoda" rank="species" species="wolcotti">
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<emphasis id="6DBDEAFAFFE68E01FDEAE5A0FD50B5DF" box="[524,648,476,500]" italics="true" pageId="5" pageNumber="428">C. wolcotti</emphasis>
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</taxonomicName>
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females, the eyes are separated by a distance that is at least 2½ times the length of the eleventh antennomere. The female specimens examined here showed considerable variability for this character, thus, it is considered unreliable for distinguishing these species. Additionally, the female pygidium of these species is very similar, and noticeable variation can be observed. These observations show that the characters given by Barr do not indicate a clear discontinuity between both species, but instead, specimens of
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<taxonomicName id="98C94D6BFFE68E01FA64E610FF34B683" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="428" phylum="Arthropoda" rank="species" species="ochlera">
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<emphasis id="6DBDEAFAFFE68E01FA64E610FF34B683" italics="true" pageId="5" pageNumber="428">C. ochlera</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="98C94D6BFFE68E01FEC4E6ECFE44B683" box="[290,412,656,680]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="428" phylum="Arthropoda" rank="species" species="wolcotti">
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<emphasis id="6DBDEAFAFFE68E01FEC4E6ECFE44B683" box="[290,412,656,680]" italics="true" pageId="5" pageNumber="428">C. wolcotti</emphasis>
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</taxonomicName>
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have a continuous array of variation. Whether these differences are caused by geographical distribution, population variability, or altitudinal clines, is unknown. Thus,
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<taxonomicName id="98C94D6BFFE68E01FBEEE6C8FB5BB6E7" box="[1032,1155,692,716]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="428" phylum="Arthropoda" rank="species" species="ochlera">
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<emphasis id="6DBDEAFAFFE68E01FBEEE6C8FB5BB6E7" box="[1032,1155,692,716]" italics="true" pageId="5" pageNumber="428">C. ochlera</emphasis>
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</taxonomicName>
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should be considered a junior synonym of
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<taxonomicName id="98C94D6BFFE68E01FE8AE6A4FE3FB6DB" box="[364,487,728,752]" class="Insecta" family="Cleridae" genus="Cymatodera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="428" phylum="Arthropoda" rank="species" species="wolcotti">
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<emphasis id="6DBDEAFAFFE68E01FE8AE6A4FE3FB6DB" box="[364,487,728,752]" italics="true" pageId="5" pageNumber="428">C. wolcotti</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |