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<document id="15345DC23D93C6BB2502064FAC695595" ID="10.11646/zootaxa.4175.3.4" ID-DOI="10.11646/zootaxa.4175.3.4" ID-GBIF-Dataset="bdf7360a-7135-4dd0-8aef-4ed6cf96d748" ID-ISSN="1175-5326" ID-Zenodo-Dep="160654" ID-ZooBank="27F01A0C-47D4-4B0B-BE96-B0D66F2DC381" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476428291595" checkinUser="plazi" docAuthor="Viana, Jéssica Herzog" docDate="2016" docId="C421D67B7E17FFB45ACAF9EDBF1AFE57" docLanguage="en" docName="zootaxa.4175.3.4.pdf" docOrigin="Zootaxa 4175 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acanthoscelides infractus Johnson &amp; Kingsolver 1973, comb n." docType="treatment" docVersion="14" lastPageNumber="250" masterDocId="3818AE037E16FFB65A5DFF91BE26FFDF" masterDocTitle="Nomenclatural changes and lectotype designations in the seed-beetle genus Sennius Bridwell: with the synonymization of Megasennius Whitehead &amp; Kingsolver (Coleoptera: Chrysomelidae: Bruchinae)" masterLastPageNumber="260" masterPageNumber="249" pageNumber="250" updateTime="1698389827584" updateUser="plazi">
<mods:mods id="044211B7FC4F40DE2F5410034FD0F5FF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E2DFF74F396BF8EA4D10DD1C59CF7929">
<mods:title id="CBD115D00313DBD0DD69E6BAC651DCB4">Nomenclatural changes and lectotype designations in the seed-beetle genus Sennius Bridwell: with the synonymization of Megasennius Whitehead &amp; Kingsolver (Coleoptera: Chrysomelidae: Bruchinae)</mods:title>
</mods:titleInfo>
<mods:name id="43B2BA4B4697AAC6AA391172A362168B" type="personal">
<mods:role id="2ACE3C9C2D1984D5AB15BADDFA56F4CA">
<mods:roleTerm id="EE473AB2ECBF861F9947E58002B2AC5E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="62266A44AFA7A5576022344DCABB04F2">Viana, Jéssica Herzog</mods:namePart>
</mods:name>
<mods:typeOfResource id="6B75356F513DC644C63F7F6754D3831B">text</mods:typeOfResource>
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<mods:title id="C23A64E105FC79379021631CB15ACE64">Zootaxa</mods:title>
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<mods:date id="3E61819BD43FC69E9FD70E4EE17E60C5">2016</mods:date>
<mods:detail id="5D54307A5D3BDDF126B3EC32294A7470" type="volume">
<mods:number id="1D8C4DE67C6B07E26DA2EAE888232A52">4175</mods:number>
</mods:detail>
<mods:detail id="2CACEA13AE46CC876B5FC4F41F8FFE06" type="issue">
<mods:number id="B7A7E2D092EF288807F8DA4A9D50E63C">3</mods:number>
</mods:detail>
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<mods:start id="8286E805D1E5462539253A66E1BD6176">249</mods:start>
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<mods:classification id="548FE0AF3C760882350375326C95770D">journal article</mods:classification>
<mods:identifier id="8EAA02D191EF14B01545686F6007FDA5" type="DOI">10.11646/zootaxa.4175.3.4</mods:identifier>
<mods:identifier id="EE7AB3D9317E8AE56B0A7FD267AC0BDF" type="GBIF-Dataset">bdf7360a-7135-4dd0-8aef-4ed6cf96d748</mods:identifier>
<mods:identifier id="F01DC73048CFABB57F499611AC1D8590" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B92A595EA25D677939B1C28C7F22AAB7" type="Zenodo-Dep">160654</mods:identifier>
<mods:identifier id="93EA2DF1E1E8FB1F5E281D7601E0A555" type="ZooBank">27F01A0C-47D4-4B0B-BE96-B0D66F2DC381</mods:identifier>
</mods:mods>
<treatment id="C421D67B7E17FFB45ACAF9EDBF1AFE57" ID-DOI="http://doi.org/10.5281/zenodo.5465545" ID-GBIF-Taxon="124996822" ID-Zenodo-Dep="5465545" LSID="urn:lsid:plazi:treatment:C421D67B7E17FFB45ACAF9EDBF1AFE57" httpUri="http://treatment.plazi.org/id/C421D67B7E17FFB45ACAF9EDBF1AFE57" lastPageId="2" lastPageNumber="250" pageId="1" pageNumber="250">
<subSubSection id="049234E67E17FFB75ACAF9EDBD9CF949" box="[151,954,1660,1686]" pageId="1" pageNumber="250" type="nomenclature">
<paragraph id="4C37676D7E17FFB75ACAF9EDBD9CF949" blockId="1.[151,954,1660,1686]" box="[151,954,1660,1686]" pageId="1" pageNumber="250">
<heading id="177FD0017E17FFB75ACAF9EDBD9CF949" bold="true" box="[151,954,1660,1686]" fontSize="11" level="1" pageId="1" pageNumber="250" reason="1">
<emphasis id="7EFCBB7F7E17FFB75ACAF9EDBD9CF949" bold="true" box="[151,954,1660,1686]" pageId="1" pageNumber="250">
<taxonomicName id="8B881CEE7E17FFB75ACAF9EDBD6FF949" authority="Johnson &amp; Kingsolver, 1973" authorityName="Johnson &amp; Kingsolver" authorityYear="1973" box="[151,841,1660,1686]" class="Insecta" family="Chrysomelidae" genus="Acanthoscelides" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="infractus" status="comb. nov.">
<emphasis id="7EFCBB7F7E17FFB75ACAF9EDBFE9F949" bold="true" box="[151,463,1660,1686]" italics="true" pageId="1" pageNumber="250">Acanthoscelides infractus</emphasis>
(
<bibRefCitation id="28191A9C7E17FFB75B83F9EDBD67F949" author="Johnson" box="[478,833,1660,1686]" pageId="1" pageNumber="250" refString="Johnson, C. D. &amp; Kingsolver, J. M. (1973) A Revision of the genus Sennius of North and Central America (Coleoptera: Bruchidae). United States Department of Agriculture, Technical Bulletin, 1462, 1 - 135." type="journal article" year="1973">Johnson &amp; Kingsolver, 1973</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="65CF06047E17FFB7590AF9EDBD9CF949" box="[855,954,1660,1686]" pageId="1" pageNumber="250">comb n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="049234E67E17FFB75ACAF952BDB8F8C9" pageId="1" pageNumber="250" type="reference_group">
<paragraph id="4C37676D7E17FFB75ACAF952BDB8F8C9" blockId="1.[151,1436,1730,1814]" pageId="1" pageNumber="250">
<treatmentCitationGroup id="6C9840437E17FFB75ACAF952BDB8F8C9" pageId="1" pageNumber="250">
<taxonomicName id="8B881CEE7E17FFB75ACAF952BC89F907" authority="Johnson &amp; Kingsolver, 1973: 67" authorityName="Johnson &amp; Kingsolver" authorityPageNumber="67" authorityYear="1973" box="[151,687,1730,1752]" class="Insecta" family="Chrysomelidae" genus="Sennius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="infractus">
<emphasis id="7EFCBB7F7E17FFB75ACAF952BF6CF907" box="[151,330,1730,1752]" italics="true" pageId="1" pageNumber="250">Sennius infractus</emphasis>
<treatmentCitation id="CD29417C7E17FFB75B08F953BC89F907" author="Johnson" box="[341,687,1730,1752]" page="67" pageId="1" pageNumber="250" year="1973">
<bibRefCitation id="28191A9C7E17FFB75B08F953BCA1F907" author="Johnson" box="[341,647,1730,1752]" pageId="1" pageNumber="250" refString="Johnson, C. D. &amp; Kingsolver, J. M. (1973) A Revision of the genus Sennius of North and Central America (Coleoptera: Bruchidae). United States Department of Agriculture, Technical Bulletin, 1462, 1 - 135." type="journal article" year="1973">Johnson &amp; Kingsolver, 1973</bibRefCitation>
: 67
</treatmentCitation>
</taxonomicName>
–68, 125 (description, key, figures,
<typeStatus id="9333D9CF7E17FFB75E76F952BA7FF907" box="[1067,1113,1731,1752]" pageId="1" pageNumber="250">type</typeStatus>
, taxonomy, distribution, host);
<treatmentCitation id="CD29417C7E17FFB75A9AF970BC0CF928" author="Johnson" box="[199,554,1761,1783]" page="419" pageId="1" pageNumber="250" year="1981">
<bibRefCitation id="28191A9C7E17FFB75A9AF970BFD0F928" author="Johnson" box="[199,502,1761,1783]" pageId="1" pageNumber="250" refString="Johnson, C. D. &amp; Kingsolver, J. M. (1981) Checklist of the Bruchidae (Coleoptera) of Canada, United States, Mexico, Central America and West Indies. The Coleopterists Bulletin, 35 (4), 409 - 422." type="journal article" year="1981">Johnson &amp; Kingsolver, 1981</bibRefCitation>
: 419
</treatmentCitation>
(catalog);
<treatmentCitation id="CD29417C7E17FFB758C3F970BD7BF928" author="Johnson" box="[670,861,1761,1783]" page="56" pageId="1" pageNumber="250" year="1984">
<bibRefCitation id="28191A9C7E17FFB758C3F970BD11F928" author="Johnson" box="[670,823,1761,1783]" pageId="1" pageNumber="250" refString="Johnson, C. D. (1984) Sennius yucatan, n. sp., redescription of S. infractus, and new host records for other Sennius (Coleoptera: Bruchidae). Annals of the Entomological Society of America, 77 (1), 56 - 64. http: // dx. doi. org / 10.1093 / aesa / 77.1.56" type="journal article" year="1984">Johnson, 1984</bibRefCitation>
: 56
</treatmentCitation>
–59 (redescription, male genitalia description, figures);
<treatmentCitation id="CD29417C7E17FFB75A9AF891BFBAF8C9" author="Borowiec" box="[199,412,1792,1814]" page="181" pageId="1" pageNumber="250" year="1987">
<bibRefCitation id="28191A9C7E17FFB75A9AF891BF4AF8C9" author="Borowiec" box="[199,364,1792,1814]" pageId="1" pageNumber="250" refString="Borowiec, L. (1987) The genera of seed beetles (Coleoptera, Bruchidae). Polskie Pismo Entomologiczne, 57, 3 - 207." type="journal article" year="1987">Borowiec, 1987</bibRefCitation>
: 181
</treatmentCitation>
(figure);
<treatmentCitation id="CD29417C7E17FFB75BA0F891BD10F8C9" author="Udayagiri" box="[509,822,1792,1814]" page="104" pageId="1" pageNumber="250" year="1989">
<bibRefCitation id="28191A9C7E17FFB75BA0F891BD21F8C9" author="Udayagiri" box="[509,775,1792,1814]" pageId="1" pageNumber="250" refString="Udayagiri, S. &amp; Wadhi, S. R. (1989) Catalog of Bruchidae. Memoirs of the American Entomological Institute, 45, 1 - 301." type="journal article" year="1989">Udayagiri &amp; Wadhi, 1989</bibRefCitation>
: 104
</treatmentCitation>
(catalog).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="049234E67E17FFB75ACAF8D3BA45F87C" pageId="1" pageNumber="250" type="materials_examined">
<paragraph id="4C37676D7E17FFB75ACAF8D3BA45F87C" blockId="1.[151,1436,1858,2027]" pageId="1" pageNumber="250">
<emphasis id="7EFCBB7F7E17FFB75ACAF8D3BFE7F884" bold="true" box="[151,449,1858,1883]" pageId="1" pageNumber="250">
<typeStatus id="9333D9CF7E17FFB75ACAF8D3BEF4F884" box="[151,210,1858,1883]" pageId="1" pageNumber="250">Type</typeStatus>
material (1,
<collectionCode id="2A99FFA87E17FFB75B30F8D3BF95F884" box="[365,435,1858,1883]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="1" pageNumber="250">MCZ</collectionCode>
):
</emphasis>
<typeStatus id="9333D9CF7E17FFB75B94F8D3BC17F884" box="[457,561,1858,1883]" pageId="1" pageNumber="250" type="holotype">Holotype</typeStatus>
female studied by photo (
<collectionCode id="2A99FFA87E17FFB75907F8D2BDBFF884" box="[858,921,1859,1883]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="1" pageNumber="250">MCZ</collectionCode>
Online Database), with labels: “M-tee Dist. / Brit. Hond. / Jan-Feb. ’06” “
<typeStatus id="9333D9CF7E17FFB75BB6F8F6BCA4F8A0" box="[491,642,1895,1919]" pageId="1" pageNumber="250" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="8B881CEE7E17FFB758FCF8F6BD41F8A0" box="[673,871,1895,1919]" class="Insecta" family="Chrysomelidae" genus="Sennius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="infractus">
<emphasis id="7EFCBB7F7E17FFB758FCF8F6BD41F8A0" box="[673,871,1895,1919]" italics="true" pageId="1" pageNumber="250">Sennius infractus</emphasis>
</taxonomicName>
/ Johnson &amp; Kingsolver” “M.C.Z. /
<typeStatus id="9333D9CF7E17FFB75F7DF8F7BBAEF8A0" box="[1312,1416,1894,1919]" pageId="1" pageNumber="250" type="holotype">Holotype</typeStatus>
/ 32916” “
<taxonomicName id="8B881CEE7E17FFB75AA1F81ABC38F87C" box="[252,542,1931,1955]" class="Insecta" family="Chrysomelidae" genus="Acanthoscelides" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="infractus">
<emphasis id="7EFCBB7F7E17FFB75AA1F81ABC38F87C" box="[252,542,1931,1955]" italics="true" pageId="1" pageNumber="250">Acanthoscelides infractus</emphasis>
</taxonomicName>
/ (
<bibRefCitation id="28191A9C7E17FFB75867F81BBD58F87C" author="Johnson" box="[570,894,1930,1955]" pageId="1" pageNumber="250" refString="Johnson, C. D. &amp; Kingsolver, J. M. (1973) A Revision of the genus Sennius of North and Central America (Coleoptera: Bruchidae). United States Department of Agriculture, Technical Bulletin, 1462, 1 - 135." type="journal article" year="1973">Johnson &amp; Kingsolver, 1973</bibRefCitation>
) / Det. J. H. Viana”.
</paragraph>
</subSubSection>
<subSubSection id="049234E67E17FFB45A9AF83FBF1AFE57" lastPageId="2" lastPageNumber="251" pageId="1" pageNumber="250" type="discussion">
<paragraph id="4C37676D7E17FFB45A9AF83FBF1AFE57" blockId="1.[151,1436,1858,2027]" lastBlockId="2.[151,1436,151,392]" lastPageId="2" lastPageNumber="251" pageId="1" pageNumber="250">
<materialsCitation id="FCE06D307E17FFB45A9AF83FBF1AFE57" ID-GBIF-Occurrence="1322993004" collectionCode="MCZ, USNM, NEW, COMBINATION" country="Honduras" county="Kingsolver" lastPageId="2" lastPageNumber="251" location="There" municipality="Johnson" pageId="1" pageNumber="250" specimenCount="2" typeStatus="holotype">
<emphasis id="7EFCBB7F7E17FFB75A9AF83FBF1DF818" bold="true" box="[199,315,1966,1991]" pageId="1" pageNumber="250">Remarks.</emphasis>
<taxonomicName id="8B881CEE7E17FFB75B18F83EBD79F818" authority="Johnson &amp; Kingsolver, 1973" authorityName="Johnson &amp; Kingsolver" authorityYear="1973" box="[325,863,1966,1991]" class="Insecta" family="Chrysomelidae" genus="Sennius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="250" phylum="Arthropoda" rank="species" species="infractus">
<emphasis id="7EFCBB7F7E17FFB75B18F83EBC2CF818" box="[325,522,1967,1991]" italics="true" pageId="1" pageNumber="250">Sennius infractus</emphasis>
<bibRefCitation id="28191A9C7E17FFB75849F83FBD79F818" author="Johnson" box="[532,863,1966,1991]" pageId="1" pageNumber="250" refString="Johnson, C. D. &amp; Kingsolver, J. M. (1973) A Revision of the genus Sennius of North and Central America (Coleoptera: Bruchidae). United States Department of Agriculture, Technical Bulletin, 1462, 1 - 135." type="journal article" year="1973">Johnson &amp; Kingsolver, 1973</bibRefCitation>
</taxonomicName>
was originally described based only on a female collected in
<collectingCountry id="349F27FD7E17FFB75B75F843BFBFF834" box="[296,409,2002,2027]" name="Honduras" pageId="1" pageNumber="250">Honduras</collectingCountry>
.
<location id="495731B67E17FFB75BFAF843BFCEF834" LSID="urn:lsid:plazi:treatment:C421D67B7E17FFB45ACAF9EDBF1AFE57:495731B67E17FFB75BFAF843BFCEF834" box="[423,488,2002,2027]" country="Honduras" county="Kingsolver" municipality="Johnson" name="There" pageId="1" pageNumber="250">There</location>
is an error in the
<collectionCode id="2A99FFA87E17FFB7589AF842BD22F834" box="[711,772,2003,2027]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="1" pageNumber="250">MCZ</collectionCode>
catalog number for this specimen, 31966, in
<bibRefCitation id="28191A9C7E17FFB45F42F843BF40FF6F" author="Johnson" lastPageId="2" lastPageNumber="251" pageId="1" pageNumber="250" refString="Johnson, C. D. &amp; Kingsolver, J. M. (1973) A Revision of the genus Sennius of North and Central America (Coleoptera: Bruchidae). United States Department of Agriculture, Technical Bulletin, 1462, 1 - 135." type="journal article" year="1973">
<collectingMunicipality id="AC53FD177E17FFB75F42F843BB5DF834" box="[1311,1403,2002,2027]" pageId="1" pageNumber="250">Johnson</collectingMunicipality>
&amp;
<collectingCounty id="A5561FE17E14FFB45ACAFF06BF32FF6F" box="[151,276,151,176]" pageId="2" pageNumber="251">Kingsolver</collectingCounty>
(1973)
</bibRefCitation>
and the correct catalog number is
<collectionCode id="2A99FFA87E14FFB458ADFF09BD0BFF6F" box="[752,813,152,176]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="2" pageNumber="251">MCZ</collectionCode>
32916 (
<collectionCode id="2A99FFA87E14FFB459D6FF09BDECFF6F" box="[907,970,152,176]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="2" pageNumber="251">MCZ</collectionCode>
Online Database). Later,
<bibRefCitation id="28191A9C7E14FFB45EB3FF06BBBAFF6F" author="Johnson" box="[1262,1436,151,176]" pageId="2" pageNumber="251" refString="Johnson, C. D. (1984) Sennius yucatan, n. sp., redescription of S. infractus, and new host records for other Sennius (Coleoptera: Bruchidae). Annals of the Entomological Society of America, 77 (1), 56 - 64. http: // dx. doi. org / 10.1093 / aesa / 77.1.56" type="journal article" year="1984">Johnson (1984)</bibRefCitation>
described the male genitalia of specimens from the same data collection as the female
<typeStatus id="9333D9CF7E14FFB45E3CFF2DBAE0FF0A" box="[1121,1222,188,213]" pageId="2" pageNumber="251" type="holotype">holotype</typeStatus>
, and indicated it as not belonging to
<taxonomicName id="8B881CEE7E14FFB45B00FF71BF92FF28" box="[349,436,224,247]" class="Insecta" family="Chrysomelidae" genus="Sennius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="genus">
<emphasis id="7EFCBB7F7E14FFB45B00FF71BF92FF28" box="[349,436,224,247]" italics="true" pageId="2" pageNumber="251">Sennius</emphasis>
</taxonomicName>
, but did not indicate its proximity with species of any other genus. In this study male specimens from
<collectionCode id="2A99FFA87E14FFB45B0CFE95BF84FEC3" box="[337,418,260,284]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="251">USNM</collectionCode>
described by
<bibRefCitation id="28191A9C7E14FFB4581DFE95BCC8FEC3" author="Johnson" box="[576,750,260,285]" pageId="2" pageNumber="251" refString="Johnson, C. D. (1984) Sennius yucatan, n. sp., redescription of S. infractus, and new host records for other Sennius (Coleoptera: Bruchidae). Annals of the Entomological Society of America, 77 (1), 56 - 64. http: // dx. doi. org / 10.1093 / aesa / 77.1.56" type="journal article" year="1984">Johnson (1984)</bibRefCitation>
with the same data as the
<typeStatus id="9333D9CF7E14FFB45E44FE95BA5DFEC2" box="[1049,1147,260,285]" pageId="2" pageNumber="251" type="holotype">holotype</typeStatus>
were examined, and here this species is here transferred to
<taxonomicName id="8B881CEE7E14FFB45853FEB9BCE2FE9F" box="[526,708,296,320]" class="Insecta" family="Chrysomelidae" genus="Acanthoscelides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="genus">
<emphasis id="7EFCBB7F7E14FFB45853FEB9BCE2FE9F" box="[526,708,296,320]" italics="true" pageId="2" pageNumber="251">Acanthoscelides</emphasis>
</taxonomicName>
,
<emphasis id="7EFCBB7F7E14FFB4588EFEB8BA32FE9F" box="[723,1044,296,320]" italics="true" pageId="2" pageNumber="251">
as
<taxonomicName id="8B881CEE7E14FFB458AEFEB9BA32FE9F" box="[755,1044,296,320]" class="Insecta" family="Chrysomelidae" genus="Acanthoscelides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="251" phylum="Arthropoda" rank="species" species="infractus">Acanthoscelides infractus</taxonomicName>
</emphasis>
<emphasis id="7EFCBB7F7E14FFB45E46FEB6BB68FE9F" bold="true" box="[1051,1358,295,320]" pageId="2" pageNumber="251">
(
<collectionCode id="2A99FFA87E14FFB45E7EFEB6BA4EFE9F" box="[1059,1128,295,320]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="2" pageNumber="251">NEW</collectionCode>
<collectionCode id="2A99FFA87E14FFB45E33FEB6BB64FE9F" box="[1134,1346,295,320]" pageId="2" pageNumber="251">COMBINATION</collectionCode>
),
</emphasis>
due to proximity with species of
<emphasis id="7EFCBB7F7E14FFB45BE2FEDCBC64FEBB" box="[447,578,333,356]" italics="true" pageId="2" pageNumber="251">megacornis</emphasis>
group (Johnson 1990b) and the lack of hinge sclerites in the internal sac of male genitalia.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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