225 lines
30 KiB
XML
225 lines
30 KiB
XML
<document id="6DCA56555FED71C399B074BDA943256B" ID-CLB-Dataset="26356" ID-DOI="10.11646/zootaxa.4624.1.8" ID-GBIF-Dataset="4d45525c-5a3e-4e37-a64a-bbf25ba47313" ID-ISSN="1175-5326" ID-Zenodo-Dep="3259349" ID-ZooBank="7DBFCA0A-C648-4222-837C-D47DADE2232D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1561702388238" checkinUser="plazi" docAuthor="Brooks, Scott E., Cumming, Jeffrey M. & Grimaldi, David A." docDate="2019" docId="8A5E87C987152540FF0D27FAFA7DF809" docLanguage="en" docName="zootaxa.4624.1.8.pdf.imf" docOrigin="Zootaxa 4624 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Schistostoma burmanicum Brooks, Cumming & Grimaldi, 2019, sp. nov." docType="treatment" docVersion="7" lastPageNumber="123" masterDocId="7667FFB187142542FF9A223DFFF1FF9E" masterDocTitle="Remarkable new fossil species of Schistostoma Becker (Diptera: Dolichopodidae Microphorinae) from mid-Cretaceous Burmese amber" masterLastPageNumber="131" masterPageNumber="121" pageNumber="122" updateTime="1698737691181" updateUser="ExternalLinkService">
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<mods:mods id="35A77A3724FC2EE91748BC0906DB5539" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2855F792ADB580FE2446ECC131595491">
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<mods:title id="91A4CA4335C602C10D6D7C50D64C079A">Remarkable new fossil species of Schistostoma Becker (Diptera: Dolichopodidae Microphorinae) from mid-Cretaceous Burmese amber</mods:title>
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<mods:name id="2C106E9399DDB722E6BEDC18C4FE114A" type="personal">
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<mods:namePart id="C98B04C29BADD1B8A968530D754A9D09">Brooks, Scott E.</mods:namePart>
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<mods:namePart id="058980118283DCF0E6EFE173E3744142">Cumming, Jeffrey M.</mods:namePart>
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<mods:name id="3485E788D1B574F07AAB8FA22EB3F121" type="personal">
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<mods:roleTerm id="33AB91387E0A608BB09B25C6501F8130">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="159EAB127B190FEEB757C373AA4F3AA8">Grimaldi, David A.</mods:namePart>
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<mods:typeOfResource id="0C43269B63643B2019F43A4EAC9D72E3">text</mods:typeOfResource>
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<mods:title id="49E1A0E21769D97DA865B5E45CD1A998">Zootaxa</mods:title>
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<mods:part id="2A975E307AB5723DC52CB6FE69B948A9">
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<mods:date id="CB1D9368B50E91A79DE2264973008E96">2019</mods:date>
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<mods:number id="10EBD7FF081254A2BF58D5C0D27D0E75">2019-06-27</mods:number>
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<mods:number id="E2823772029B05FB8CCC7FE7BE34936B">4624</mods:number>
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<mods:detail id="BE0928BF1FBB162A6686CBB7680047DF" type="issue">
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<mods:number id="13440A667ECED28CA0F2A869F78040BF">1</mods:number>
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<mods:identifier id="372AE560EB38AF57C0ED24585E343907" type="DOI">10.11646/zootaxa.4624.1.8</mods:identifier>
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<treatment id="8A5E87C987152540FF0D27FAFA7DF809" ID-GBIF-Taxon="158414992" LSID="urn:lsid:plazi:treatment:8A5E87C987152540FF0D27FAFA7DF809" httpUri="http://treatment.plazi.org/id/8A5E87C987152540FF0D27FAFA7DF809" lastPageId="2" lastPageNumber="123" pageId="1" pageNumber="122">
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<subSubSection id="4AED655487152543FF0D27FAFDB1FA7C" box="[151,576,1479,1506]" pageId="1" pageNumber="122" type="nomenclature">
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<paragraph id="024836DF87152543FF0D27FAFDB1FA7C" blockId="1.[151,576,1479,1542]" box="[151,576,1479,1506]" pageId="1" pageNumber="122">
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<heading id="590081B387152543FF0D27FAFDB1FA7C" bold="true" box="[151,576,1479,1506]" fontSize="11" level="1" pageId="1" pageNumber="122" reason="1">
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<emphasis id="3083EACD87152543FF0D27FAFDB1FA7C" bold="true" box="[151,576,1479,1506]" pageId="1" pageNumber="122">
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<taxonomicName id="C5F74D5C87152543FF0D27FAFE2BFA7C" ID-CoL="9HM6Y" box="[151,474,1479,1506]" class="Insecta" family="Dolichopodidae" genus="Schistostoma" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="species" species="burmanicum" status="sp. nov.">
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<emphasis id="3083EACD87152543FF0D27FAFE2BFA7C" bold="true" box="[151,474,1479,1506]" italics="true" pageId="1" pageNumber="122">Schistostoma burmanicum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2BB057B687152543FE7827F5FDB1FA7C" box="[482,576,1480,1506]" pageId="1" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="4AED655487152543FF0D27D6FEFAF998" box="[151,267,1515,1542]" pageId="1" pageNumber="122" type="description">
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<paragraph id="024836DF87152543FF0D27D6FEFAF998" blockId="1.[151,576,1479,1542]" box="[151,267,1515,1542]" pageId="1" pageNumber="122">
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(
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<figureCitation id="9ACC2A5A87152543FF0527D6FEF3F998" box="[159,258,1515,1542]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–4" captionStart-2="FIGURES 5–8" captionStartId-0="3.[151,264,1531,1556]" captionStartId-1="4.[151,264,1580,1605]" captionStartId-2="6.[151,264,1933,1958]" captionTargetBox-0="[151,1436,181,1493]" captionTargetBox-1="[151,1436,181,1560]" captionTargetBox-2="[224,1362,502,1899]" captionTargetId-0="figure@3.[151,1436,181,1511]" captionTargetId-1="figure@4.[151,1436,181,1560]" captionTargetId-2="figure@6.[212,1376,489,1914]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionText-0="FIGURES 1–2. Schistostoma burmanicum sp. nov. male holotype (AMNH Bu-0029). 1. Habitus photograph, left lateral view. 2. Habitus illustration, left lateral view." captionText-1="FIGURES 3–4. Schistostoma burmanicum sp. nov. male holotype (AMNH Bu-0029). 3. Habitus photograph, right lateral view. 4. Right foreleg and midleg, right lateral view." captionText-2="FIGURES 5–8. Schistostoma burmanicum sp. nov. and Schistostoma foliatum sp. nov. 5. Right antenna of S. burmanicum, left medial view. 6. Left foreleg of S. burmanicum, left lateral view. 7. Hypopygium of S. burmanicum, ventral view. 8. Left foreleg of S. foliatum, left lateral view. Abbreviations: epand—epandrium; hypd—hypandrium; hypd proc—hypandrial process." httpUri-0="https://zenodo.org/record/3259351/files/figure.png" httpUri-1="https://zenodo.org/record/3259353/files/figure.png" httpUri-2="https://zenodo.org/record/3259355/files/figure.png" pageId="1" pageNumber="122">Figs 1–7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="4AED655487152543FF0D2409FB95F924" pageId="1" pageNumber="122" type="materials_examined">
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<paragraph id="024836DF87152543FF0D2409FB95F924" blockId="1.[151,1437,1587,2010]" pageId="1" pageNumber="122">
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<emphasis id="3083EACD87152543FF0D2409FEBBF9D0" bold="true" box="[151,330,1587,1614]" pageId="1" pageNumber="122">Type material.</emphasis>
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<materialsCitation id="B29F3C8287152543FECC240EFB95F924" ID-GBIF-Occurrence="2270855923" collectionCode="AMNH, AMBER" collectorName="Leeward Capitol Corp." country="Myanmar" county="Upper Cretaceous" location="Ledo Rd." municipality="Tanai Village" pageId="1" pageNumber="122" specimenCount="2" specimenCount-male="2" stateProvince="Kachin" typeStatus="holotype">
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<emphasis id="3083EACD87152543FECC240EFE04F9D0" bold="true" box="[342,501,1587,1614]" pageId="1" pageNumber="122">
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<typeStatus id="DD4C887D87152543FECC240EFE04F9D0" box="[342,501,1587,1614]" pageId="1" pageNumber="122" type="holotype">HOLOTYPE</typeStatus>
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</emphasis>
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<specimenCount id="14F1FD5687152543FD9B2409FDE9F9D0" box="[513,536,1588,1614]" pageId="1" pageNumber="122" type="male">♂</specimenCount>
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,
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<collectionCode id="64E6AE1A87152543FDBD2409FD8CF9D0" box="[551,637,1588,1614]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="122">AMNH</collectionCode>
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Bu-0029, in Burmese amber with accompanying labels: “
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<collectionCode id="64E6AE1A87152543FAB42409FA68F9D0" box="[1326,1433,1588,1614]" pageId="1" pageNumber="122">AMBER</collectionCode>
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:
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<collectingCountry id="7AE0764F87152543FF0D2465FED8F9EC" box="[151,297,1624,1650]" name="Myanmar" pageId="1" pageNumber="122">MYANMAR</collectingCountry>
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(BURMA)/
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<collectingCounty id="EB294E5387152543FE262465FD78F9EC" box="[444,649,1624,1650]" pageId="1" pageNumber="122">Upper Cretaceous</collectingCounty>
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?/
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<collectingRegion id="C033F83D87152543FD3E246AFD08F9EF" box="[676,761,1623,1649]" country="Myanmar" name="Kachin" pageId="1" pageNumber="122">Kachin</collectingRegion>
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:
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<collectingMunicipality id="E22CACA587152543FC9F246AFC6CF9EF" box="[773,925,1623,1649]" pageId="1" pageNumber="122">Tanai Village</collectingMunicipality>
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(on
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<location id="0728600487152543FC49246AFBCCF9EF" LSID="urn:lsid:plazi:treatment:8A5E87C987152540FF0D27FAFA7DF809:0728600487152543FC49246AFBCCF9EF" box="[979,1085,1623,1649]" country="Myanmar" county="Upper Cretaceous" municipality="Tanai Village" name="Ledo Rd." pageId="1" pageNumber="122" stateProvince="Kachin">Ledo Rd.</location>
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/
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<locationDeviation id="13FA2FFD87152543FBD4246AFAA3F9EC" box="[1102,1362,1623,1650]" location="Myitkyna" pageId="1" pageNumber="122">
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<quantity id="C50F9B3A87152543FBD4246AFB54F9EF" box="[1102,1189,1623,1650]" metricMagnitude="5" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="122" unit="km" value="105.0">105 km</quantity>
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NW Myitkyna
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</locationDeviation>
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)/ coll.
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<collectorName id="AF02530987152543FF0D2446FE50F908" box="[151,417,1659,1686]" pageId="1" pageNumber="122">Leeward Capitol Corp.</collectorName>
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, 1999/
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<collectionCode id="64E6AE1A87152543FE6D2441FDBCF908" box="[503,589,1660,1686]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="122">AMNH</collectionCode>
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Bu-0029”; “Burmese amber/
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<taxonomicName id="C5F74D5C87152543FC322441FBD3F908" box="[936,1058,1660,1686]" kingdom="Plantae" order="Diptera" pageId="1" pageNumber="122" rank="order">DIPTERA</taxonomicName>
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:/
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<specimenCount id="14F1FD5687152543FBA22441FBBCF908" box="[1080,1101,1660,1686]" pageId="1" pageNumber="122" type="male">♂</specimenCount>
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<taxonomicName id="C5F74D5C87152543FBCD2446FAF0F90B" authorityName="Collin" authorityYear="1960" box="[1111,1281,1659,1685]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="subFamily" subFamily="Microphorinae">Microphorinae</taxonomicName>
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:/ (raptorial)”; “
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<typeStatus id="DD4C887D87152543FF3A249DFEC7F924" box="[160,310,1696,1722]" pageId="1" pageNumber="122" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="C5F74D5C87152543FEDC249DFD9EF924" box="[326,623,1696,1722]" class="Insecta" family="Dolichopodidae" genus="Schistostoma" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="species" species="burmanicum">
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<emphasis id="3083EACD87152543FEDC249DFD9EF924" box="[326,623,1696,1722]" italics="true" pageId="1" pageNumber="122">Schistostoma burmanicum</emphasis>
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</taxonomicName>
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/ Cumming, Brooks & Grimaldi [red label]”.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="4AED655487152543FF5D24FEFC82F89F" pageId="1" pageNumber="122" type="diagnosis">
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<paragraph id="024836DF87152543FF5D24FEFC82F89F" blockId="1.[151,1437,1587,2010]" pageId="1" pageNumber="122">
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<emphasis id="3083EACD87152543FF5D24FEFEB3F940" bold="true" box="[199,322,1731,1758]" pageId="1" pageNumber="122">Diagnosis.</emphasis>
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This species is characterized by its enlarged lamellate apically truncate fore tibia (
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<figureCitation id="9ACC2A5A87152543FB7F24FEFAD8F940" box="[1253,1321,1731,1758]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1933,1958]" captionTargetBox="[224,1362,502,1899]" captionTargetId="figure@6.[212,1376,489,1914]" captionTargetPageId="6" captionText="FIGURES 5–8. Schistostoma burmanicum sp. nov. and Schistostoma foliatum sp. nov. 5. Right antenna of S. burmanicum, left medial view. 6. Left foreleg of S. burmanicum, left lateral view. 7. Hypopygium of S. burmanicum, ventral view. 8. Left foreleg of S. foliatum, left lateral view. Abbreviations: epand—epandrium; hypd—hypandrium; hypd proc—hypandrial process." httpUri="https://zenodo.org/record/3259355/files/figure.png" pageId="1" pageNumber="122">Fig. 6</figureCitation>
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) with distinctive ‘eye spot’ and laminate setae along upper ventral margin.
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</paragraph>
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</subSubSection>
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<subSubSection id="4AED655487152540FF5D2536FB00F8B5" lastPageId="2" lastPageNumber="123" pageId="1" pageNumber="122" type="description">
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<paragraph id="024836DF87152540FF5D2536FB00F8B5" blockId="1.[151,1437,1587,2010]" lastBlockId="2.[151,1437,153,1943]" lastPageId="2" lastPageNumber="123" pageId="1" pageNumber="122">
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<emphasis id="3083EACD87152543FF5D2536FE57F8B8" bold="true" box="[199,422,1803,1830]" pageId="1" pageNumber="122">Description. Male.</emphasis>
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Body length
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<quantity id="C50F9B3A87152543FDDF2531FD51F8B8" box="[581,672,1804,1830]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="122" unit="mm" value="1.8">1.8 mm</quantity>
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. Wing length
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<quantity id="C50F9B3A87152543FCD92531FC6FF8B8" box="[835,926,1804,1830]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="122" unit="mm" value="1.4">1.4 mm</quantity>
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.
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<emphasis id="3083EACD87152543FC312531FC05F8B8" bold="true" box="[939,1012,1804,1830]" pageId="1" pageNumber="122">Head:</emphasis>
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Apparently more or less elliptical in lateral view, but compound eyes, occiput and postgena collapsed. Neck inserted apparently below middle of head, with several short setae adjacent to occipital foramen. Ocellar triangle distinctly protruded with a pair of strong ocellar setae and a pair of weaker postocellar setae. Holoptic with compound eyes contiguous for most of length of frons; compound eye ovoid in lateral view, occupying most of head, bare, dorsal ommatidia distinctly larger than ventral ommatidia, medial edge of eye with emargination slightly below level of antenna giving rise to demarcation line between larger dorsal and smaller ventral ommatidia. Frons small, triangular, mostly obliterated by eyes. Face and clypeus recessed in oral cavity and obscured from view. Occiput with series of 3–4 visible postocular setae. Gena obscured from view but apparently very narrow. Postgena broad. Antenna (
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<figureCitation id="9ACC2A5A87162540FBA622DCFB71FF65" box="[1084,1152,224,251]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1933,1958]" captionTargetBox="[224,1362,502,1899]" captionTargetId="figure@6.[212,1376,489,1914]" captionTargetPageId="6" captionText="FIGURES 5–8. Schistostoma burmanicum sp. nov. and Schistostoma foliatum sp. nov. 5. Right antenna of S. burmanicum, left medial view. 6. Left foreleg of S. burmanicum, left lateral view. 7. Hypopygium of S. burmanicum, ventral view. 8. Left foreleg of S. foliatum, left lateral view. Abbreviations: epand—epandrium; hypd—hypandrium; hypd proc—hypandrial process." httpUri="https://zenodo.org/record/3259355/files/figure.png" pageId="2" pageNumber="123">Fig. 5</figureCitation>
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) inserted near middle of head in profile; scape obscured from view, apparently short; pedicel about
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<date id="7649101F87162540FC412338FC0CFE81" box="[987,1021,261,287]" pageId="2" pageNumber="123">2X</date>
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longer than wide, with several setae; postpedicel elongate subtriangular, about 2.5X longer than wide, clothed in fine hairs; stylus arista-like, terminal, about
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<date id="7649101F87162540FF472370FF0EFEF9" box="[221,255,333,359]" pageId="2" pageNumber="123">2X</date>
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longer than postpedicel, 2-articled, basal article short. Palpus obscured from view. Proboscis narrow, directed anteroventrally.
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<emphasis id="3083EACD87162540FE0D234CFE08FE15" bold="true" box="[407,505,369,395]" pageId="2" pageNumber="123">Thorax:</emphasis>
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Slightly longer than wide in dorsal view. Mesoscutum strongly arched, prescutellar depression present. Proepisternum bare. Postpronotal lobe with 1 strong seta. Acrostichal setae present, apparently biserial; other bristles of thorax well-differentiated, each side with: approximately 9 dorsocentrals, anterior dorsocentrals short and closely spaced, 2 posteriormost dorsocentrals strong and widely separated; 3 apparent supra-alar setae; 2 strong notopleural setae; 1 strong postalar seta. Scutellum with posterior margin rounded, with 2 pairs of setae, outer pair weak, inner pair strong. Mesopleuron apparently bare.
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<emphasis id="3083EACD87162540FC0B2018FC23FDA1" bold="true" box="[913,978,549,575]" pageId="2" pageNumber="123">Legs:</emphasis>
|
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Fairly long; foreleg and midleg with specialized morphology and chaetotaxy; tarsal claws, pulvilli and empodium normally developed on all legs.
|
||
<emphasis id="3083EACD87162540FADF2074FA6CFDFD" box="[1349,1437,585,611]" italics="true" pageId="2" pageNumber="123">Foreleg</emphasis>
|
||
(
|
||
<figureCitation id="9ACC2A5A87162540FF052050FF16FD19" box="[159,231,621,647]" captionStart="FIGURES 3–4" captionStartId="4.[151,264,1580,1605]" captionTargetBox="[151,1436,181,1560]" captionTargetId="figure@4.[151,1436,181,1560]" captionTargetPageId="4" captionText="FIGURES 3–4. Schistostoma burmanicum sp. nov. male holotype (AMNH Bu-0029). 3. Habitus photograph, right lateral view. 4. Right foreleg and midleg, right lateral view." httpUri="https://zenodo.org/record/3259353/files/figure.png" pageId="2" pageNumber="123">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="9ACC2A5A87162540FF682050FEF2FD16" box="[242,259,621,648]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1933,1958]" captionTargetBox="[224,1362,502,1899]" captionTargetId="figure@6.[212,1376,489,1914]" captionTargetPageId="6" captionText="FIGURES 5–8. Schistostoma burmanicum sp. nov. and Schistostoma foliatum sp. nov. 5. Right antenna of S. burmanicum, left medial view. 6. Left foreleg of S. burmanicum, left lateral view. 7. Hypopygium of S. burmanicum, ventral view. 8. Left foreleg of S. foliatum, left lateral view. Abbreviations: epand—epandrium; hypd—hypandrium; hypd proc—hypandrial process." httpUri="https://zenodo.org/record/3259355/files/figure.png" pageId="2" pageNumber="123">6</figureCitation>
|
||
): Coxa without visible setae; femur with series of 4 visible strong dorsolateral setae, with large basiventral tubercle bearing 4 apical setae, ventral surface immediately distal to basiventral tubercle with series of 7 short setae, each borne on a tiny tubercle; tibia greatly enlarged and lamellate with apical margin truncate, flattened laterally and slightly concave medially, distinctly patterned with ‘eye spot’ on apical third, lateral surface with short setae along dorsal margin as well as two longitudinal series of short setae near middle and adjacent to ventral margin, ventral margin adorned with modified laminate, thickened and curved setae, medial surface with series of short setae along apical margin, otherwise bare; tarsus with short setae, tarsomere 1 as long as combined length of tarsomeres 2–4.
|
||
<emphasis id="3083EACD87162540FF0D2154FF17FC1D" box="[151,230,873,899]" italics="true" pageId="2" pageNumber="123">Midleg</emphasis>
|
||
: Coxa with strong elongate anterior seta at extreme base and smaller seta immediately below, two weak anterior setae near middle; trochanter with several weak setae; femur with strong basidorsal seta, series of 4 similarly strong anterodorsal setae along basal half, series of short dorsal setae running towards apex, prominent basiventral tubercle bearing thick curved seta and 2 ventral setae near middle (distal seta weaker); tibia with flange on basal half bearing close-set series of short modified setae (some hook-shaped), 1 long erect anterior seta near mid-length; tarsomere 1 as long as combined length of tarsomeres 2–5, with series of erect setae dorsally, otherwise with short setae similar to other tarsomeres.
|
||
<emphasis id="3083EACD87162540FD8E267CFD9FFBC5" box="[532,622,1089,1115]" italics="true" pageId="2" pageNumber="123">Hindleg</emphasis>
|
||
: Coxa with 1 strong lateral seta at basal third, 2 gradually smaller setae apically, several weak apical setae along anterior margin; femur with series of strong dorsal and ventral setae, ventral setae longer; tibia with series of strong dorsal setae, series of weaker ventral setae along basal half; tarsomere 1 as long as combined length of tarsomeres 2–5, with series of stronger setae dorsally, otherwise with short setae similar to other tarsomeres.
|
||
<emphasis id="3083EACD87162540FE3A26EDFE1BFB75" bold="true" box="[416,490,1232,1259]" pageId="2" pageNumber="123">Wing:</emphasis>
|
||
Pterostigma absent, membrane entirely covered with minute microtrichia. Anal lobe obscured from view. Costa circumambient. Extreme anterior base of costa with 2 strong setae. Anterior costal section bearing row of short spine-like setae intermixed with fine setae from humeral crossvein to R
|
||
<subScript id="9E73349A87162540FB472717FB07FAA6" attach="left" box="[1245,1270,1322,1336]" fontSize="6" pageId="2" pageNumber="123">2+3</subScript>
|
||
, posterior part of costa beyond R
|
||
<subScript id="9E73349A87162540FEF92773FE8DFAC2" attach="left" box="[355,380,1358,1372]" fontSize="6" pageId="2" pageNumber="123">2+3</subScript>
|
||
with only fine setae. Longitudinal veins (except CuA+CuP) complete, reaching wing margin. Sc faint apically. R
|
||
<subScript id="9E73349A87162540FEF1274FFE82FA1E" attach="left" box="[363,371,1394,1408]" fontSize="6" pageId="2" pageNumber="123">1</subScript>
|
||
reaching costa beyond middle of wing (or beyond base of M
|
||
<subScript id="9E73349A87162540FB80274FFBD3FA1E" attach="left" box="[1050,1058,1394,1408]" fontSize="6" pageId="2" pageNumber="123">2</subScript>
|
||
). Base of Rs obscured from view. R
|
||
<subScript id="9E73349A87162540FF3027ABFF32FA3A" attach="left" box="[170,195,1430,1444]" fontSize="6" pageId="2" pageNumber="123">2+3</subScript>
|
||
diverging from R
|
||
<subScript id="9E73349A87162540FE1627ABFE54FA3A" attach="left" box="[396,421,1430,1444]" fontSize="6" pageId="2" pageNumber="123">4+5</subScript>
|
||
apically. R
|
||
<subScript id="9E73349A87162540FDB927ABFDCCFA3A" attach="left" box="[547,573,1430,1444]" fontSize="6" pageId="2" pageNumber="123">4+5</subScript>
|
||
weakly sinuous. M
|
||
<subScript id="9E73349A87162540FC8D27ABFCEEFA3A" attach="left" box="[791,799,1430,1444]" fontSize="6" pageId="2" pageNumber="123">1</subScript>
|
||
diverging from R
|
||
<subScript id="9E73349A87162540FC7D27ABFBF0FA3A" attach="left" box="[999,1025,1430,1444]" fontSize="6" pageId="2" pageNumber="123">4+5</subScript>
|
||
beyond cell dm. M
|
||
<subScript id="9E73349A87162540FB4127ABFB12FA3A" attach="left" box="[1243,1251,1430,1444]" fontSize="6" pageId="2" pageNumber="123">1</subScript>
|
||
and M
|
||
<subScript id="9E73349A87162540FAA827ABFACBFA3A" attach="left" box="[1330,1338,1430,1444]" fontSize="6" pageId="2" pageNumber="123">2</subScript>
|
||
strongly diverging beyond cell dm. M
|
||
<subScript id="9E73349A87162540FE7A2787FE19FA56" attach="left" box="[480,488,1466,1480]" fontSize="6" pageId="2" pageNumber="123">2</subScript>
|
||
and M
|
||
<subScript id="9E73349A87162540FDA32787FDB0FA56" attach="left" box="[569,577,1466,1480]" fontSize="6" pageId="2" pageNumber="123">4</subScript>
|
||
weakly diverging beyond cell dm. Short r-m crossvein present in basal portion of wing, distal to base of R
|
||
<subScript id="9E73349A87162540FE6927E3FDFCFA72" attach="left" box="[499,525,1502,1516]" fontSize="6" pageId="2" pageNumber="123">4+5</subScript>
|
||
. Crossvein bm-m apparently complete. Cell dm present, closed by base of M
|
||
<subScript id="9E73349A87162540FAFF27E3FA9CFA72" attach="left" box="[1381,1389,1502,1516]" fontSize="6" pageId="2" pageNumber="123">2</subScript>
|
||
and crossvein dm-m, cell extended to middle of wing. Cells br, bm and cua in basal fourth of wing, partially obscured from view. Cell cua apparently closed, rounded apically with CuA curved. Vein CuA+CuP apparently present and short. Calypter obscured from view.
|
||
<emphasis id="3083EACD87162540FDB42405FD5DF9CD" bold="true" box="[558,684,1592,1619]" pageId="2" pageNumber="123">Abdomen:</emphasis>
|
||
Tergites and sternites 1–6 with short setae. Segment 7 tubular, with 1 weak seta visible on sternite. Sternite 8 subrectangular, forming dome-like cap over dorsal region of hypopygium, with several strong marginal setae.
|
||
<emphasis id="3083EACD87162540FDBB24BFFD41F905" box="[545,688,1666,1691]" italics="true" pageId="2" pageNumber="123">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="9ACC2A5A87162540FD2524BCFCF5F905" box="[703,772,1665,1691]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1933,1958]" captionTargetBox="[224,1362,502,1899]" captionTargetId="figure@6.[212,1376,489,1914]" captionTargetPageId="6" captionText="FIGURES 5–8. Schistostoma burmanicum sp. nov. and Schistostoma foliatum sp. nov. 5. Right antenna of S. burmanicum, left medial view. 6. Left foreleg of S. burmanicum, left lateral view. 7. Hypopygium of S. burmanicum, ventral view. 8. Left foreleg of S. foliatum, left lateral view. Abbreviations: epand—epandrium; hypd—hypandrium; hypd proc—hypandrial process." httpUri="https://zenodo.org/record/3259355/files/figure.png" pageId="2" pageNumber="123">Fig. 7</figureCitation>
|
||
): Lateroflexed to right; inverted with posterior end directed anteriorly; small, about 1/5 length of abdomen, apparently symmetrical. Right and left epandrial lamellae rounded basally and oblong in posterior view, each with pair of narrow subequal apical epandrial lobes, cluster of short setae dorsally near base of epandrial lobes, otherwise bare. Hypandrium rounded basally with narrow elongate medial hypandrial process flanked by pair of strong setae. Cercus obscured from view.
|
||
<emphasis id="3083EACD87162540FB8E252CFB85F8B5" bold="true" box="[1044,1140,1809,1835]" pageId="2" pageNumber="123">Female:</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4AED655487162540FF5D2508FC2BF8D1" box="[199,986,1845,1871]" pageId="2" pageNumber="123" type="etymology">
|
||
<paragraph id="024836DF87162540FF5D2508FC2BF8D1" blockId="2.[151,1437,153,1943]" box="[199,986,1845,1871]" pageId="2" pageNumber="123">
|
||
<emphasis id="3083EACD87162540FF5D2508FEBFF8D1" bold="true" box="[199,334,1845,1871]" pageId="2" pageNumber="123">Etymology.</emphasis>
|
||
The specific epithet refers to
|
||
<collectingCountry id="7AE0764F87162540FD062508FD1AF8D1" box="[668,747,1845,1871]" name="Myanmar" pageId="2" pageNumber="123">Burma</collectingCountry>
|
||
, source of the amber.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4AED655487162540FF5D2564FA7DF809" pageId="2" pageNumber="123" type="discussion">
|
||
<paragraph id="024836DF87162540FF5D2564FA7DF809" blockId="2.[151,1437,153,1943]" pageId="2" pageNumber="123">
|
||
<emphasis id="3083EACD87162540FF5D2564FECAF8ED" bold="true" box="[199,315,1881,1907]" pageId="2" pageNumber="123">Remarks.</emphasis>
|
||
The specimen has clusters of small spheres near the legs (
|
||
<figureCitation id="9ACC2A5A87162540FC462564FBD7F8ED" box="[988,1062,1881,1907]" captionStart="FIGURES 1–2" captionStartId="3.[151,264,1531,1556]" captionTargetBox="[151,1436,181,1493]" captionTargetId="figure@3.[151,1436,181,1511]" captionTargetPageId="3" captionText="FIGURES 1–2. Schistostoma burmanicum sp. nov. male holotype (AMNH Bu-0029). 1. Habitus photograph, left lateral view. 2. Habitus illustration, left lateral view." httpUri="https://zenodo.org/record/3259351/files/figure.png" pageId="2" pageNumber="123">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="9ACC2A5A87162540FBAE2564FBB5F8EA" box="[1076,1092,1881,1908]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1933,1958]" captionTargetBox="[224,1362,502,1899]" captionTargetId="figure@6.[212,1376,489,1914]" captionTargetPageId="6" captionText="FIGURES 5–8. Schistostoma burmanicum sp. nov. and Schistostoma foliatum sp. nov. 5. Right antenna of S. burmanicum, left medial view. 6. Left foreleg of S. burmanicum, left lateral view. 7. Hypopygium of S. burmanicum, ventral view. 8. Left foreleg of S. foliatum, left lateral view. Abbreviations: epand—epandrium; hypd—hypandrium; hypd proc—hypandrial process." httpUri="https://zenodo.org/record/3259355/files/figure.png" pageId="2" pageNumber="123">6</figureCitation>
|
||
), which could be spores, very small pollen grains, or brochosomes (see
|
||
<bibRefCitation id="66664B2E87162540FDFD2540FCBAF809" author="Suludere, Z. & Lavigne, R. & Acikgoz, F. B." box="[615,843,1917,1943]" pageId="2" pageNumber="123" pagination="413 - 416" refId="ref5609" refString="Suludere, Z., Lavigne, R. & Acikgoz, F. B. (2018) First record of accidental contamination of Asilidae (Insecta: Diptera) with brochosomes. Entomological News, 127, 413 - 416. https: // doi. org / 10.3157 / 021.127.0504" type="journal article" year="2018">
|
||
Suludere
|
||
<emphasis id="3083EACD87162540FD482543FCFBF809" box="[722,778,1917,1943]" italics="true" pageId="2" pageNumber="123">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
) that will require higher magnification to ascertain.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |