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<mods:title id="3830C1E566334F3152B9CF185E09CAFA">A revision of the genus Pachygrapsus Randall, 1840 (Crustacea: Decapoda: Brachyura, Grapsidae), with special reference to the Southwest Pacific species</mods:title>
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<mods:namePart id="2CE3DDCD4126B16F25FAA0D462DEA115">Cexus, Jean-Christophe</mods:namePart>
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<taxonomicName id="23944D01ED49FFF66C1CFA88FCD4FAF8" ID-CoL="4BSC3" authority="(de Saussure, 1858)" baseAuthorityName="de Saussure" baseAuthorityYear="1858" box="[264,779,1364,1390]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="D6E0EA90ED49FFF66C1CFA88FDCDFAF8" bold="true" box="[264,530,1364,1390]" italics="true" pageId="14" pageNumber="15">Pachygrapsus gracilis</emphasis>
(de
<bibRefCitation id="80054B73ED49FFF66F5CFA88FCDCFAF8" author="Saussure" box="[584,771,1364,1390]" pageId="14" pageNumber="15" pagination="417 - 496" refId="ref33810" refString="Saussure de, H. (1858) Memoire sur divers crustaces nouveaux du Mexique et des Antilles. Memoires de la Societe de Physique et d'Histoire naturelle de Geneve, 14 (2), 417 - 496 [1 - 82], pl. 1 - 6." type="journal article" year="1858">Saussure, 1858</bibRefCitation>
)
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(
<figureCitation id="7CAF2A07ED49FFF66E0FFA88FC63FAF8" box="[795,956,1364,1390]" captionStart="FIGURE 4" captionStartId="16.[264,367,1652,1676]" captionTargetBox="[276,1317,294,1638]" captionTargetId="figure-77@16.[266,1322,281,1638]" captionTargetPageId="16" captionText="FIGURE 4. —Pachygrapsus gracilis (de Saussure, 1858). a) dorsal view; b) front; c) left chela, outer face; d) right P5; e) male abdomen. a, ce, male 14.9 x 19.5 mm, Florida (USNM 170178); b) ov. female 8.5 x 11 mm, Angola (MNHN B12903)." figureDoi="http://doi.org/10.5281/zenodo.5049240" httpUri="https://zenodo.org/record/5049240/files/figure.png" pageId="14" pageNumber="15">Figures 4ae</figureCitation>
,
<figureCitation id="7CAF2A07ED49FFF66EDEFA88FC26FAF8" box="[970,1017,1364,1390]" captionStart="FIGURE 14" captionStartId="47.[264,368,1403,1427]" captionTargetBox="[274,1309,325,1344]" captionTargetId="figure-211@47.[265,1322,294,1379]" captionTargetPageId="47" captionText="FIGURE 14. —Infraorbital margin: a) Pachygrapsus fakaravensis Rathbun (1907), ov. female 15.0 x 16.5 mm, Taiaro, French Polynesia (MNHN B25782); b) Pachygrapsus laevimanus Stimpson, 1858, neotype male 18.0 x 21.7 mm, Sydney, Australia (AM P7478); c) Pachygrapsus minutus A. Milne­Edwards, 1873, male 5.2 x 7.0 mm, Moruroa, French Polynesia (MNHN B29273); d) Pachygrapsus planifrons de Man, 1888, male 7.0 x 9.0 mm, Rangiroa, French Polynesia (UF 1490); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), male 12.1 x 15.3 mm, Taiaro, French Polynesia (MNHN B29279); f) Pachygrapsus corrugatus (von Martens, 1872) ov. female 13.6 x 15.2 mm, Virgin Is. (USNM 72340); g) Pachygrapsus crassipes Randall, 1840, male 31.9 x 36.6 mm, Sagami Bay, Japan (NSMT Cr7516); h) Pachygrapsus gracilis (de Saussure, 1858), ov. female 8.5 x 11 mm, Angola (MNHN B12903); i) Pachygrapsus loveridgei Chace, 1966, male 8.1 x 10 mm, St. Helena I. (MNHN B17836); j) Pachygrapsus marmoratus (Fabricius, 1787), female 12.7 x 14.3 mm, Préfailles, France (MNHN B22654); k) Pachygrapsus maurus (Lucas, 1846), female 12.7 x 14.3 mm, Banyuls, France (MNHN B12933); l) Pachygrapsus transversus (Gibbes, 1850), male 17.0 x 22.5 mm, French Guiana (MNHN B16013). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5049262" httpUri="https://zenodo.org/record/5049262/files/figure.png" pageId="14" pageNumber="15">14h</figureCitation>
,
<figureCitation id="7CAF2A07ED49FFF66912FA88FBEAFAF8" box="[1030,1077,1364,1390]" captionStart="FIGURE 15" captionStartId="48.[264,368,1262,1286]" captionTargetBox="[381,1213,284,1247]" captionTargetId="figure-298@48.[340,1254,282,1260]" captionTargetPageId="48" captionText="FIGURE 15. —Male gonopods: a) Pachygrapsus fakaravensis Rathbun (1907), holotype 18.0 x 19.4 mm, Fakarava, French Polynesia (USNM 32844), right G1, abdominal face (left) and detail of terminal portion, sternal face (right); b) Pachygrapsus laevimanus Stimpson, 1858, neotype 18.0 x 21.7 mm, Sydney, Australia (AM P7478), left G1, abdominal (left) and sternal face (right), setae omitted; c) Pachygrapsus minutus A. Milne­Edwards, 1873, cl 8.2 mm, Caroline Is., lateral (left) and abdominal face (right), setae omitted; d) Pachygrapsus planifrons de Man, 1888, 9.5 x 11.8 mm, Hawaiian Is. (USNM 29350), right G1, abdominal (left) and sternal face (right); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), cl 11.7 mm, Hawaiian Is., lateral (left) and abdominal face (right); f) Pachygrapsus corrugatus (von Martens, 1872), cl 8.5 mm, Ascension I., left G1 abdominal face (left) and detail of terminal portion, setae omitted (right); g) Pachygrapsus crassipes Randall, 1840, cl 35.6 mm, California, lateral (left) and abdominal face (right) setae omitted; h) Pachygrapsus gracilis (de Saussure, 1858), cl 8.7 mm, Virgin Is., lateral (left) and abdominal face (right), setae omitted; i) Pachygrapsus loveridgei Chace, 1966, holotype 10.3 x 13.1 mm, St. Helena I. (USNM 112457), left G1, abdominal (left) and sternal face (right); j) Pachygrapsus marmoratus (Fabricius, 1787), cl 30.5 mm, Gibraltar, lateral (left) and abdominal face (right), setae omitted; k) Pachygrapsus maurus (Lucas, 1846), cl 8.5 mm, Canary Is., lateral (left) and abdominal face (right) setae omitted; l) Pachygrapsus transversus (Gibbes, 1850), cl 11.1 mm, Key West, Florida, lateral (left) and abdominal face (right), setae omitted. (c, e, gl, from Chace, 1966, fig. 1011; f, from Manning &amp; Chace, 1990, fig. 43). Approximate magnification x 6 (x 12 for detail of terminal portion)." figureDoi="http://doi.org/10.5281/zenodo.5049264" httpUri="https://zenodo.org/record/5049264/files/figure.png" pageId="14" pageNumber="15">15h</figureCitation>
)
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<emphasis id="D6E0EA90ED49FFF66C1CFA43FDDAFA20" box="[264,517,1439,1462]" italics="true" pageId="14" pageNumber="15">Metopograpsus gracilis</emphasis>
de
<bibRefCitation id="80054B73ED49FFF66F3FFA42FCDEFA23" author="Saussure" box="[555,769,1438,1461]" pageId="14" pageNumber="15" pagination="417 - 496" refId="ref33810" refString="Saussure de, H. (1858) Memoire sur divers crustaces nouveaux du Mexique et des Antilles. Memoires de la Societe de Physique et d'Histoire naturelle de Geneve, 14 (2), 417 - 496 [1 - 82], pl. 1 - 6." type="journal article" year="1858">Saussure, 1858: 443</bibRefCitation>
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[27], pl. 2, fig. 15 [
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locality: St. Thomas, Virgin Is.].
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<emphasis id="D6E0EA90ED49FFF66C1CFA03FDA3FA60" box="[264,636,1503,1526]" italics="true" pageId="14" pageNumber="15">Grapsus (Leptograpsus) rugulosus</emphasis>
von
<bibRefCitation id="80054B73ED49FFF66FA5FA02FC5DFA63" author="Martens von, E. &amp; Dr. J. Gundlach's" box="[689,898,1502,1525]" pageId="14" pageNumber="15" pagination="77 - 147" refId="ref31783" refString="Martens von, E. (1872) Uber kubanische Crustaceen nach den Sammlungen Dr. J. Gundlach's. Archiv fur Naturgeschichte, 38 (1), 77 - 147, 257, 258, pl. 4 - 5." type="journal article" year="1872">Martens, 1872: 108</bibRefCitation>
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(
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)
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, not
<taxonomicName id="23944D01ED49FFF6691DFA03FDFEF983" ID-CoL="6PGLY" authority="H. Milne Edwards, 1863" authorityName="H. Milne Edwards" authorityYear="1863" class="Malacostraca" family="Grapsidae" genus="Leptograpsus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="rugulosus">
<emphasis id="D6E0EA90ED49FFF6691DFA03FAD5FA60" box="[1033,1290,1503,1526]" italics="true" pageId="14" pageNumber="15">Leptograpsus rugulosus</emphasis>
H. Milne Edwards, 1863
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<emphasis id="D6E0EA90ED49FFF66C1CF9C3FE27F9A0" box="[264,504,1567,1590]" italics="true" pageId="14" pageNumber="15">Pachygrapsus gracilis</emphasis>
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<bibRefCitation id="80054B73ED49FFF66F08F9C2FD01F9A3" author="Verrill, A. E." box="[540,734,1566,1589]" pageId="14" pageNumber="15" pagination="299 - 474" refId="ref34598" refString="Verrill, A. E. (1908) Decapod Crustacea of Bermuda. I. Brachyura and Anomura. Their distribution, variations, and habits. Transactions of the Connecticut Academy of Arts &amp; Sciences, 13 (6), 299 - 474, pl. 9 - 28." type="journal article" year="1908">Verrill, 1908: 324</bibRefCitation>
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, fig. 6a, b, pl. 12, fig. 2 (
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). —
<treatmentCitation id="65351093ED49FFF66991F9C2FEBDF9C3" author="Rathbun, M. J." page="249" pageId="14" pageNumber="15" year="1918">
<bibRefCitation id="80054B73ED49FFF66991F9C2FEBDF9C3" author="Rathbun, M. J." pageId="14" pageNumber="15" pagination="1 - 461" refId="ref33490" refString="Rathbun, M. J. (1918) The grapsoid crabs of America. Bulletin of the United States National Museum, 97, i - xxii, 1 - 461, pl. 1 - 161." type="journal article" year="1918">Rathbun, 1918: 249</bibRefCitation>
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, pl. 60, fig. 3, pl. 61, fig. 1 (
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,
<collectingCountry id="9C837612ED49FFF66E1CF9E2FCBEF9C3" box="[776,865,1598,1621]" name="Jamaica" pageId="14" pageNumber="15">Jamaica</collectingCountry>
,
<collectingCountry id="9C837612ED49FFF66E78F9E2FC31F9C3" box="[876,1006,1598,1621]" name="Puerto Rico" pageId="14" pageNumber="15">Puerto Rico</collectingCountry>
,
<collectingCountry id="9C837612ED49FFF66EEEF9E2FBB9F9C3" box="[1018,1126,1598,1621]" name="Colombia" pageId="14" pageNumber="15">Colombia</collectingCountry>
,
<collectingCountry id="9C837612ED49FFF66966F9E2FB69F9C3" box="[1138,1206,1598,1621]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
,
<collectingCountry id="9C837612ED49FFF669D5F9E2FAF9F9C3" box="[1217,1318,1598,1621]" name="Bermuda" pageId="14" pageNumber="15">Bermuda</collectingCountry>
; synonymy); 1921: 445, pl. 40, fig. 1 (
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); 1933: 89 (
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). —
<treatmentCitation id="65351093ED49FFF66974F982FAFAF9E3" author="Capart, A." box="[1120,1317,1630,1653]" page="187" pageId="14" pageNumber="15" year="1951">
<bibRefCitation id="80054B73ED49FFF66974F982FAFAF9E3" author="Capart, A." box="[1120,1317,1630,1653]" pageId="14" pageNumber="15" pagination="11 - 205" refId="ref29189" refString="Capart, A. (1951) Crustaces Decapodes Brachyures. Expedition oceanographique Belge dans les eaux cotieres africaines de l'Atlantique Sud (1948 - 1949). Resultats Scientifiques, 3 (1), 11 - 205, pl. 1 - 3." type="journal article" year="1951">Capart, 1951: 187</bibRefCitation>
</treatmentCitation>
, fig. 74, pl. 3, fig. 19 (West Africa). —
<treatmentCitation id="65351093ED49FFF66FCDF9A2FC7DF903" author="Monod, T." box="[729,930,1662,1685]" page="419" pageId="14" pageNumber="15" year="1956">
<bibRefCitation id="80054B73ED49FFF66FCDF9A2FC7DF903" author="Monod, T." box="[729,930,1662,1685]" pageId="14" pageNumber="15" pagination="1 - 674" refId="ref32147" refString="Monod, T. (1956) Hippidea et Brachyura ouest-africains. Memoires de l'Institut Francais d'Afrique Noire, 45, 1 - 674." type="journal article" year="1956">Monod, 1956: 419</bibRefCitation>
</treatmentCitation>
, fig. 569 (West Africa). —
<treatmentCitation id="65351093ED49FFF669DDF9A2FE78F923" author="Holthuis, L. B." page="239" pageId="14" pageNumber="15" year="1959">
<bibRefCitation id="80054B73ED49FFF669DDF9A2FE78F923" author="Holthuis, L. B." pageId="14" pageNumber="15" pagination="1 - 296" refId="ref30959" refString="Holthuis, L. B. (1959) The Crustacea Decapoda of Suriname (Dutch Guiana). Zoologische Verhandelingen, 44, 1 - 296, pl. 1 - 16." type="journal article" year="1959">Holthuis, 1959: 239</bibRefCitation>
</treatmentCitation>
, pl. 10, fig. 3 (
<collectingCountry id="9C837612ED49FFF66F5EF942FD68F923" box="[586,695,1694,1717]" name="Suriname" pageId="14" pageNumber="15">Suriname</collectingCountry>
). —
<treatmentCitation id="65351093ED49FFF66FF0F942FBD1F923" author="Chace, F. A. &amp; Hobbs, H. H." box="[740,1038,1694,1717]" page="167" pageId="14" pageNumber="15" year="1969">
<bibRefCitation id="80054B73ED49FFF66FF0F942FBD1F923" author="Chace, F. A. &amp; Hobbs, H. H." box="[740,1038,1694,1717]" pageId="14" pageNumber="15" pagination="1 - 5" refId="ref29302" refString="Chace, F. A. &amp; Hobbs, H. H. (1969) The freshwater and terrestrial decapod crustaceans of the West Indies with special reference to Dominica. Bredin-Archbold-Smithsonian Biological Survey of Dominica. Bulletin United States National Museum, 292, v + 1 - 258, pl. 1 - 5." type="journal article" year="1969">Chace &amp; Hobbs, 1969: 167</bibRefCitation>
</treatmentCitation>
, fig. 51, 52j (West Indies; references). —
<treatmentCitation id="65351093ED49FFF66CCEF962FD17F943" author="Rodriguez, G." box="[474,712,1726,1749]" page="380" pageId="14" pageNumber="15" year="1980">
<bibRefCitation id="80054B73ED49FFF66CCEF962FD17F943" author="Rodriguez, G." box="[474,712,1726,1749]" pageId="14" pageNumber="15" refId="ref33692" refString="Rodriguez, G. (1980) Crustaceos Decapodos de Venezuela. Instituto Venezolano de Investigaciones Cientificas, Caracas, 494 pp." type="book" year="1980">Rodríguez, 1980: 380</bibRefCitation>
</treatmentCitation>
, pl. 59 (
<collectingCountry id="9C837612ED49FFF66E32F962FC44F943" box="[806,923,1726,1749]" name="Venezuela" pageId="14" pageNumber="15">Venezuela</collectingCountry>
). —
<treatmentCitation id="65351093ED49FFF66EDEF962FAF5F943" author="Manning, R. B. &amp; Holthuis, L. B." box="[970,1322,1726,1749]" page="233" pageId="14" pageNumber="15" year="1981">
<bibRefCitation id="80054B73ED49FFF66EDEF962FAF5F943" author="Manning, R. B. &amp; Holthuis, L. B." box="[970,1322,1726,1749]" pageId="14" pageNumber="15" pagination="1 - 379" refId="ref31742" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African Brachyuran Crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306, i - xii, 1 - 379." type="journal article" year="1981">Manning &amp; Holthuis, 1981: 233</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="9C837612ED49FFF66C2BF902FE4AF963" box="[319,405,1758,1781]" name="Senegal" pageId="14" pageNumber="15">Senegal</collectingCountry>
to
<collectingCountry id="9C837612ED49FFF66CACF902FDD5F963" box="[440,522,1758,1781]" name="Angola" pageId="14" pageNumber="15">Angola</collectingCountry>
; references). —
<treatmentCitation id="65351093ED49FFF66FA1F902FB96F963" author="Rodrigues, M. D. &amp; Brossi-Garcia, A. L." box="[693,1097,1758,1781]" page="63" pageId="14" pageNumber="15" year="1989">
<bibRefCitation id="80054B73ED49FFF66FA1F902FB96F963" author="Rodrigues, M. D. &amp; Brossi-Garcia, A. L." box="[693,1097,1758,1781]" pageId="14" pageNumber="15" pagination="63 - 66" refId="ref33715" refString="Rodrigues, M. D. &amp; Brossi-Garcia, A. L. (1989) Notas sobre novas ocorrencias de Pachygrapsus gracilis (Saussure, 1858) (Crustacea: Brachyura: Grapsidae). Ciencia e Cultura, 41 (1), 63 - 66." type="journal article" year="1989">Rodrigues &amp; Brossi­Garcia, 1989: 63</bibRefCitation>
</treatmentCitation>
, fig. 15 (
<collectingCountry id="9C837612ED49FFF669A3F902FB21F963" box="[1207,1278,1758,1781]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
). —
<treatmentCitation id="65351093ED49FFF66C2CF922FE34F883" author="Melo &amp; G. A" box="[312,491,1790,1813]" page="453" pageId="14" pageNumber="15" year="1996">
<bibRefCitation id="80054B73ED49FFF66C2CF922FE34F883" author="Melo &amp; G. A" box="[312,491,1790,1813]" pageId="14" pageNumber="15" refId="ref31826" refString="Melo de, G. A. S. (1996) Manual de identificacao dos Brachyura (Caranguejos e Siris) do Litoral Brasileiro. Sao Paulo: Pleiade / Fundacao de Amparo a Pesquisa do Estado de Sao Paulo (FAPESP), 604 pp." type="book" year="1996">Melo, 1996: 453</bibRefCitation>
</treatmentCitation>
, fig. 1 (
<collectingCountry id="9C837612ED49FFF66F29F922FD5BF883" box="[573,644,1790,1813]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
); 1998: 499 (
<collectingCountry id="9C837612ED49FFF66E04F922FC88F883" box="[784,855,1790,1813]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="AC8E6509ED48FFF76C1CFEC0FBF1FD8F" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="E42B3682ED48FFF76C1CFEC0FE76FEA3" blockId="15.[264,1321,284,390]" box="[264,425,284,309]" pageId="15" pageNumber="16">
<emphasis id="D6E0EA90ED48FFF76C1CFEC0FE76FEA3" box="[264,425,284,309]" italics="true" pageId="15" pageNumber="16">
<typeStatus id="3B2F8820ED48FFF76C1CFEC0FEE0FEA3" box="[264,319,284,309]" pageId="15" pageNumber="16">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="E42B3682ED48FFF76C2CFE98FE76FE10" blockId="15.[264,1321,284,390]" pageId="15" pageNumber="16">
<materialsCitation id="54FC3CDFED48FFF76C2CFE98FE79FE10" collectionCode="MHNG" collectorName="P. J. Schwendinger" pageId="15" pageNumber="16" specimenCount="1" typeStatus="holotype">
One dry specimen (
<collectionCode id="8285AE47ED48FFF76F3CFE98FD5CFEC8" box="[552,643,324,350]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="16" type="Museum">MHNG</collectionCode>
, frame n° 190, no record number) (
<collectorName id="49615354ED48FFF76929FE98FAFAFEC8" box="[1085,1317,324,350]" pageId="15" pageNumber="16">P. J. Schwendinger</collectorName>
, pers. comm.)
</materialsCitation>
.
</paragraph>
<paragraph id="E42B3682ED48FFF76C1CFE73FE34FE5E" blockId="15.[264,1323,431,737]" box="[264,491,431,456]" pageId="15" pageNumber="16">
<emphasis id="D6E0EA90ED48FFF76C1CFE73FE34FE5E" box="[264,491,431,456]" italics="true" pageId="15" pageNumber="16">Material Examined</emphasis>
</paragraph>
<paragraph id="E42B3682ED48FFF76C2CFE0AFBF1FD8F" blockId="15.[264,1323,431,737]" pageId="15" pageNumber="16">
<emphasis id="D6E0EA90ED48FFF76C2CFE0AFE3CFE66" bold="true" box="[312,483,470,496]" pageId="15" pageNumber="16">East Atlantic.</emphasis>
<materialsCitation id="54FC3CDFED48FFF76CFFFE0AFBF5FD8F" accessionNumber="B12903" collectionCode="MNHN" country="Angola" location="Lobito" pageId="15" pageNumber="16" specimenCount="1">
<emphasis id="D6E0EA90ED48FFF76CFFFE0AFD9BFE66" bold="true" box="[491,580,470,496]" pageId="15" pageNumber="16">
<collectingCountry id="9C837612ED48FFF76CFFFE0AFD9BFE66" box="[491,580,470,496]" name="Angola" pageId="15" pageNumber="16">Angola</collectingCountry>
</emphasis>
:
<location id="E14B6059ED48FFF76F40FE0BFD7AFE67" LSID="urn:lsid:plazi:treatment:6C3D8794ED49FFE96C1CFA88FAF4FD08:E14B6059ED48FFF76F40FE0BFD7AFE67" box="[596,677,471,497]" country="Angola" name="Lobito" pageId="15" pageNumber="16">Lobito</location>
, intertidal in oysters, 2 M 7.8 x 10.0, 9.4 x 12.0 mm, 1 F 9.2 x 12.0 mm, 4 F ov. 6.4 x 7.98.5 x
<quantity id="236C9B67ED48FFF76FF3FE23FC92FD8F" box="[743,845,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="15" pageNumber="16" unit="mm" value="11.1">11.1 mm</quantity>
(
<collectionCode id="8285AE47ED48FFF76E49FE23FC67FD8F" box="[861,952,511,537]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="16">MNHN</collectionCode>
B12903)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AC8E6509ED48FFE96C2CFDFAFEB1FEA3" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="description">
<paragraph id="E42B3682ED48FFF76C2CFDFAFD70FD77" blockId="15.[264,1323,431,737]" pageId="15" pageNumber="16">
<emphasis id="D6E0EA90ED48FFF76C2CFDFAFD91FDD6" bold="true" box="[312,590,550,576]" pageId="15" pageNumber="16">West Atlantic. Florida</emphasis>
:
<materialsCitation id="54FC3CDFED48FFF76F49FDFBFBE8FD07" collectingDate="1972-08-10" collectionCode="USNM" collectorName="R. H. Gore" elevation="2" location="Banana river" municipality="Cap Canaveral" pageId="15" pageNumber="16" specimenCode="USNM 170178" specimenCount="1">
<location id="E14B6059ED48FFF76F49FDFBFD29FDD7" LSID="urn:lsid:plazi:treatment:6C3D8794ED49FFE96C1CFA88FAF4FD08:E14B6059ED48FFF76F49FDFBFD29FDD7" box="[605,758,551,577]" municipality="Cap Canaveral" name="Banana river" pageId="15" pageNumber="16">Banana river</location>
,
<collectingMunicipality id="044FACF8ED48FFF76E16FDFBFC6DFDD7" box="[770,946,551,577]" pageId="15" pageNumber="16">Cap Canaveral</collectingMunicipality>
, on rock rubble and oysters,
<quantity id="236C9B67ED48FFF76810FDFBFEFEFDFF" metricMagnitude="0" metricUnit="m" metricValue="1.8" pageId="15" pageNumber="16" unit="m" value="1.8">
<elevation id="6FB9D1B1ED48FFF76810FDFBFEFEFDFF" metricMagnitude="0" metricUnit="m" metricValue="1.8" pageId="15" pageNumber="16" unit="m" value="1.8">1.8 m</elevation>
</quantity>
, coll.
<collectorName id="49615354ED48FFF76C7CFD93FE33FDFF" box="[360,492,591,617]" pageId="15" pageNumber="16">R. H. Gore</collectorName>
,
<date id="902A1042ED48FFF76CECFD93FD69FDFF" box="[504,694,591,617]" pageId="15" pageNumber="16" value="1972-08-10">
<collectingDate id="806EE9AAED48FFF76CECFD93FD69FDFF" box="[504,694,591,617]" pageId="15" pageNumber="16" value="1972-08-10">10 August 1972</collectingDate>
</date>
, 3 M 11.3 x 14.314.9 x
<quantity id="236C9B67ED48FFF76EC9FD93FB96FDFF" box="[989,1097,591,617]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="15" pageNumber="16" unit="mm" value="19.5">19.5 mm</quantity>
, 2 ov. F 7.2 x 10.0, 12.4 x
<quantity id="236C9B67ED48FFF76C49FDABFE15FD07" box="[349,458,631,657]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="15" pageNumber="16" unit="mm" value="16.7">16.7 mm</quantity>
, 3 F 10.7 x 14.113.5 x
<quantity id="236C9B67ED48FFF76FE3FDABFCBEFD07" box="[759,865,631,657]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="15" pageNumber="16" unit="mm" value="14.3">14.3 mm</quantity>
(
<specimenCode id="B4329EF9ED48FFF76E67FDABFBEFFD07" box="[883,1072,631,657]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="16" type="Museum">USNM 170178</specimenCode>
)
</materialsCitation>
. —
<materialsCitation id="54FC3CDFED48FFF76977FDAAFD73FD77" accessionNumber="B29570, B16013" collectionCode="MNHN" collectorName="F. Geay" country="French Guiana" location="French Guiana" pageId="15" pageNumber="16" specimenCount="1">
<emphasis id="D6E0EA90ED48FFF76977FDAAFAFDFD06" bold="true" box="[1123,1314,630,656]" pageId="15" pageNumber="16">
<collectingCountry id="9C837612ED48FFF76977FDAAFAFDFD06" box="[1123,1314,630,656]" name="French Guiana" pageId="15" pageNumber="16">French Guiana</collectingCountry>
</emphasis>
: coll.
<collectorName id="49615354ED48FFF76C52FD43FE7BFD2F" box="[326,420,671,697]" pageId="15" pageNumber="16">F. Geay</collectorName>
1906, 1 F 11.5 x 16.0 mm (
<collectionCode id="8285AE47ED48FFF76E10FD43FC80FD2F" box="[772,863,671,697]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="16">MNHN</collectionCode>
B29570; mixed with specimens of
<taxonomicName id="23944D01ED48FFF76801FD43FE50FD76" baseAuthorityName="Gibbes" baseAuthorityYear="1850" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="D6E0EA90ED48FFF76801FD43FE50FD76" italics="true" pageId="15" pageNumber="16">P. transversus</emphasis>
</taxonomicName>
under
<collectionCode id="8285AE47ED48FFF76CF5FD1BFDE4FD77" box="[481,571,711,737]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="16">MNHN</collectionCode>
B16013)
</materialsCitation>
.
</paragraph>
<paragraph id="E42B3682ED48FFF76C1CFCD5FEA0FCB4" blockId="15.[264,1325,777,1763]" box="[264,383,777,802]" pageId="15" pageNumber="16">
<emphasis id="D6E0EA90ED48FFF76C1CFCD5FEA0FCB4" box="[264,383,777,802]" italics="true" pageId="15" pageNumber="16">Diagnosis</emphasis>
</paragraph>
<paragraph id="E42B3682ED48FFF76C2CFCEDFDE2FC7D" blockId="15.[264,1325,777,1763]" pageId="15" pageNumber="16">
Carapace markedly striated on gastric, hepatic, branchial regions; cardiac, intestinal regions smooth or only with short striae; lateral margins strongly convergent posteriorly, with tooth behind exorbital angle (
<figureCitation id="7CAF2A07ED48FFF76FB6FC5DFD25FC0D" box="[674,762,897,923]" captionStart="FIGURE 4" captionStartId="16.[264,367,1652,1676]" captionTargetBox="[276,1317,294,1638]" captionTargetId="figure-77@16.[266,1322,281,1638]" captionTargetPageId="16" captionText="FIGURE 4. —Pachygrapsus gracilis (de Saussure, 1858). a) dorsal view; b) front; c) left chela, outer face; d) right P5; e) male abdomen. a, ce, male 14.9 x 19.5 mm, Florida (USNM 170178); b) ov. female 8.5 x 11 mm, Angola (MNHN B12903)." figureDoi="http://doi.org/10.5281/zenodo.5049240" httpUri="https://zenodo.org/record/5049240/files/figure.png" pageId="15" pageNumber="16">Fig. 4a</figureCitation>
). Front 0.6 times exorbital width; anterior margin straight (
<figureCitation id="7CAF2A07ED48FFF76C8AFC75FE28FC55" box="[414,503,937,963]" captionStart="FIGURE 4" captionStartId="16.[264,367,1652,1676]" captionTargetBox="[276,1317,294,1638]" captionTargetId="figure-77@16.[266,1322,281,1638]" captionTargetPageId="16" captionText="FIGURE 4. —Pachygrapsus gracilis (de Saussure, 1858). a) dorsal view; b) front; c) left chela, outer face; d) right P5; e) male abdomen. a, ce, male 14.9 x 19.5 mm, Florida (USNM 170178); b) ov. female 8.5 x 11 mm, Angola (MNHN B12903)." figureDoi="http://doi.org/10.5281/zenodo.5049240" httpUri="https://zenodo.org/record/5049240/files/figure.png" pageId="15" pageNumber="16">Fig. 4b</figureCitation>
). Infraorbital margin almost smooth, at most slightly granulate; without outer notch (
<figureCitation id="7CAF2A07ED48FFF76CDDFC0DFDEFFC7D" box="[457,560,977,1003]" captionStart="FIGURE 14" captionStartId="47.[264,368,1403,1427]" captionTargetBox="[274,1309,325,1344]" captionTargetId="figure-211@47.[265,1322,294,1379]" captionTargetPageId="47" captionText="FIGURE 14. —Infraorbital margin: a) Pachygrapsus fakaravensis Rathbun (1907), ov. female 15.0 x 16.5 mm, Taiaro, French Polynesia (MNHN B25782); b) Pachygrapsus laevimanus Stimpson, 1858, neotype male 18.0 x 21.7 mm, Sydney, Australia (AM P7478); c) Pachygrapsus minutus A. Milne­Edwards, 1873, male 5.2 x 7.0 mm, Moruroa, French Polynesia (MNHN B29273); d) Pachygrapsus planifrons de Man, 1888, male 7.0 x 9.0 mm, Rangiroa, French Polynesia (UF 1490); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), male 12.1 x 15.3 mm, Taiaro, French Polynesia (MNHN B29279); f) Pachygrapsus corrugatus (von Martens, 1872) ov. female 13.6 x 15.2 mm, Virgin Is. (USNM 72340); g) Pachygrapsus crassipes Randall, 1840, male 31.9 x 36.6 mm, Sagami Bay, Japan (NSMT Cr7516); h) Pachygrapsus gracilis (de Saussure, 1858), ov. female 8.5 x 11 mm, Angola (MNHN B12903); i) Pachygrapsus loveridgei Chace, 1966, male 8.1 x 10 mm, St. Helena I. (MNHN B17836); j) Pachygrapsus marmoratus (Fabricius, 1787), female 12.7 x 14.3 mm, Préfailles, France (MNHN B22654); k) Pachygrapsus maurus (Lucas, 1846), female 12.7 x 14.3 mm, Banyuls, France (MNHN B12933); l) Pachygrapsus transversus (Gibbes, 1850), male 17.0 x 22.5 mm, French Guiana (MNHN B16013). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5049262" httpUri="https://zenodo.org/record/5049262/files/figure.png" pageId="15" pageNumber="16">Fig. 14h</figureCitation>
).
</paragraph>
<paragraph id="E42B3682ED48FFF76C2CFC25FD51FB25" blockId="15.[264,1325,777,1763]" pageId="15" pageNumber="16">
Outer face of chelae smooth and regularly convex, sometimes with a faint stria near lower margin, becoming more obvious on outer face of fixed finger; dorsal margin of palm with a weak carina and several oblique striae on inner side; dorsal margin of dactyl tuberculated. Cutting edges of fingers widely gaping, with small triangular teeth; tip of fingers spoon shaped, glabrous (
<figureCitation id="7CAF2A07ED48FFF76F3FFB45FD5EFB25" box="[555,641,1177,1203]" captionStart="FIGURE 4" captionStartId="16.[264,367,1652,1676]" captionTargetBox="[276,1317,294,1638]" captionTargetId="figure-77@16.[266,1322,281,1638]" captionTargetPageId="16" captionText="FIGURE 4. —Pachygrapsus gracilis (de Saussure, 1858). a) dorsal view; b) front; c) left chela, outer face; d) right P5; e) male abdomen. a, ce, male 14.9 x 19.5 mm, Florida (USNM 170178); b) ov. female 8.5 x 11 mm, Angola (MNHN B12903)." figureDoi="http://doi.org/10.5281/zenodo.5049240" httpUri="https://zenodo.org/record/5049240/files/figure.png" pageId="15" pageNumber="16">Fig. 4c</figureCitation>
).
</paragraph>
<paragraph id="E42B3682ED48FFF76C2CFB1DFEA9FAC5" blockId="15.[264,1325,777,1763]" pageId="15" pageNumber="16">
Ambulatory legs with few scattered long bristles and longitudinal rows of short setae on carpi, propodi; propodi without brush of setae. P5 merus with lower margin marked by several oblique striae on outer face; with a somewhat more noticeable submedian tubercle (
<figureCitation id="7CAF2A07ED48FFF76C05FAE5FEB6FAC5" box="[273,361,1337,1363]" captionStart="FIGURE 4" captionStartId="16.[264,367,1652,1676]" captionTargetBox="[276,1317,294,1638]" captionTargetId="figure-77@16.[266,1322,281,1638]" captionTargetPageId="16" captionText="FIGURE 4. —Pachygrapsus gracilis (de Saussure, 1858). a) dorsal view; b) front; c) left chela, outer face; d) right P5; e) male abdomen. a, ce, male 14.9 x 19.5 mm, Florida (USNM 170178); b) ov. female 8.5 x 11 mm, Angola (MNHN B12903)." figureDoi="http://doi.org/10.5281/zenodo.5049240" httpUri="https://zenodo.org/record/5049240/files/figure.png" pageId="15" pageNumber="16">Fig. 4d</figureCitation>
).
</paragraph>
<paragraph id="E42B3682ED48FFF76C2CFABDFDEFFA5D" blockId="15.[264,1325,777,1763]" pageId="15" pageNumber="16">
Abdominal tergites smooth. Male abdomen with sixth somite broader than telson, sixth somite and telson not forming triangle. G1 bluntly rounded at tip, with short corneous process (
<figureCitation id="7CAF2A07ED48FFF76CA8FA6DFDFDFA5D" box="[444,546,1457,1483]" captionStart="FIGURE 15" captionStartId="48.[264,368,1262,1286]" captionTargetBox="[381,1213,284,1247]" captionTargetId="figure-298@48.[340,1254,282,1260]" captionTargetPageId="48" captionText="FIGURE 15. —Male gonopods: a) Pachygrapsus fakaravensis Rathbun (1907), holotype 18.0 x 19.4 mm, Fakarava, French Polynesia (USNM 32844), right G1, abdominal face (left) and detail of terminal portion, sternal face (right); b) Pachygrapsus laevimanus Stimpson, 1858, neotype 18.0 x 21.7 mm, Sydney, Australia (AM P7478), left G1, abdominal (left) and sternal face (right), setae omitted; c) Pachygrapsus minutus A. Milne­Edwards, 1873, cl 8.2 mm, Caroline Is., lateral (left) and abdominal face (right), setae omitted; d) Pachygrapsus planifrons de Man, 1888, 9.5 x 11.8 mm, Hawaiian Is. (USNM 29350), right G1, abdominal (left) and sternal face (right); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), cl 11.7 mm, Hawaiian Is., lateral (left) and abdominal face (right); f) Pachygrapsus corrugatus (von Martens, 1872), cl 8.5 mm, Ascension I., left G1 abdominal face (left) and detail of terminal portion, setae omitted (right); g) Pachygrapsus crassipes Randall, 1840, cl 35.6 mm, California, lateral (left) and abdominal face (right) setae omitted; h) Pachygrapsus gracilis (de Saussure, 1858), cl 8.7 mm, Virgin Is., lateral (left) and abdominal face (right), setae omitted; i) Pachygrapsus loveridgei Chace, 1966, holotype 10.3 x 13.1 mm, St. Helena I. (USNM 112457), left G1, abdominal (left) and sternal face (right); j) Pachygrapsus marmoratus (Fabricius, 1787), cl 30.5 mm, Gibraltar, lateral (left) and abdominal face (right), setae omitted; k) Pachygrapsus maurus (Lucas, 1846), cl 8.5 mm, Canary Is., lateral (left) and abdominal face (right) setae omitted; l) Pachygrapsus transversus (Gibbes, 1850), cl 11.1 mm, Key West, Florida, lateral (left) and abdominal face (right), setae omitted. (c, e, gl, from Chace, 1966, fig. 1011; f, from Manning &amp; Chace, 1990, fig. 43). Approximate magnification x 6 (x 12 for detail of terminal portion)." figureDoi="http://doi.org/10.5281/zenodo.5049264" httpUri="https://zenodo.org/record/5049264/files/figure.png" pageId="15" pageNumber="16">Fig. 15h</figureCitation>
).
</paragraph>
<paragraph id="E42B3682ED48FFF76C2CFA05FC68F9FD" blockId="15.[264,1325,777,1763]" pageId="15" pageNumber="16">
Colour (from photograph): Carapace light green to cream with transverse lines composed of small violet spots. Chelipeds and ambulatory legs pale violet with numerous darker violet spots. The spots retained on the carapace even after several years in preservative (
<figureCitation id="7CAF2A07ED48FFF76C50F98DFE48F9FD" box="[324,407,1617,1643]" captionStart="FIGURE 4" captionStartId="16.[264,367,1652,1676]" captionTargetBox="[276,1317,294,1638]" captionTargetId="figure-77@16.[266,1322,281,1638]" captionTargetPageId="16" captionText="FIGURE 4. —Pachygrapsus gracilis (de Saussure, 1858). a) dorsal view; b) front; c) left chela, outer face; d) right P5; e) male abdomen. a, ce, male 14.9 x 19.5 mm, Florida (USNM 170178); b) ov. female 8.5 x 11 mm, Angola (MNHN B12903)." figureDoi="http://doi.org/10.5281/zenodo.5049240" httpUri="https://zenodo.org/record/5049240/files/figure.png" pageId="15" pageNumber="16">Fig. 4a</figureCitation>
) is a useful character to recognize the species.
</paragraph>
<paragraph id="E42B3682ED48FFF76C2CF9A5FE74F975" blockId="15.[264,1325,777,1763]" pageId="15" pageNumber="16">
Measurements: medium size; carapace of specimens examined ranging from: males 7.8 x 10.014.9 x
<quantity id="236C9B67ED48FFF76CCAF97DFD95F92D" box="[478,586,1697,1723]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="15" pageNumber="16" unit="mm" value="19.5">19.5 mm</quantity>
; females 6.4 x 7.913.5 x
<quantity id="236C9B67ED48FFF76E95F97DFC32F92D" box="[897,1005,1697,1723]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="15" pageNumber="16" unit="mm" value="14.3">14.3 mm</quantity>
; smallest ovigerous female 6.4 x
<quantity id="236C9B67ED48FFF76C5EF915FE78F975" box="[330,423,1737,1763]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="15" pageNumber="16" unit="mm" value="7.9">7.9 mm</quantity>
.
</paragraph>
<paragraph id="E42B3682ED48FFF76C1CF8DBFE48F8B6" blockId="15.[264,1322,1799,1905]" box="[264,407,1799,1824]" pageId="15" pageNumber="16">
<emphasis id="D6E0EA90ED48FFF76C1CF8DBFE48F8B6" box="[264,407,1799,1824]" italics="true" pageId="15" pageNumber="16">Distribution</emphasis>
</paragraph>
<paragraph id="E42B3682ED48FFF76C2CF8F2FE37F8E7" blockId="15.[264,1322,1799,1905]" pageId="15" pageNumber="16">
<emphasis id="D6E0EA90ED48FFF76C2CF8F2FE3AF8DE" bold="true" box="[312,485,1838,1864]" pageId="15" pageNumber="16">West Atlantic</emphasis>
.
<collectingCountry id="9C837612ED48FFF76CE3F8F3FDB0F8DF" box="[503,623,1839,1865]" name="Argentina" pageId="15" pageNumber="16">Argentine</collectingCountry>
,
<collectingCountry id="9C837612ED48FFF76F6BF8F3FD16F8DF" box="[639,713,1839,1865]" name="Brazil" pageId="15" pageNumber="16">Brazil</collectingCountry>
,
<collectingCountry id="9C837612ED48FFF76FCDF8F3FC52F8DF" box="[729,909,1839,1865]" name="French Guiana" pageId="15" pageNumber="16">French Guiana</collectingCountry>
, Caribbean, Texas.
<emphasis id="D6E0EA90ED48FFF76968F8F2FAFCF8DE" bold="true" box="[1148,1315,1838,1864]" pageId="15" pageNumber="16">East Atlantic</emphasis>
.
<collectingCountry id="9C837612ED48FFF76C1CF88BFEBBF8E7" box="[264,356,1879,1905]" name="Senegal" pageId="15" pageNumber="16">Senegal</collectingCountry>
to
<collectingCountry id="9C837612ED48FFF76C9EF88BFE3BF8E7" box="[394,484,1879,1905]" name="Angola" pageId="15" pageNumber="16">Angola</collectingCountry>
.
</paragraph>
<caption id="B0EB660AED57FFE86C1CF9A8FCF4F947" ID-DOI="http://doi.org/10.5281/zenodo.5049240" ID-Zenodo-Dep="5049240" httpUri="https://zenodo.org/record/5049240/files/figure.png" pageId="16" pageNumber="17" startId="16.[264,367,1652,1676]" targetBox="[276,1317,294,1638]" targetPageId="16">
<paragraph id="E42B3682ED57FFE86C1CF9A8FCF4F947" blockId="16.[264,1323,1652,1745]" pageId="16" pageNumber="17">
<emphasis id="D6E0EA90ED57FFE86C1CF9A8FE53F91A" bold="true" box="[264,396,1652,1676]" pageId="16" pageNumber="17">FIGURE 4.</emphasis>
<taxonomicName id="23944D01ED57FFE86CA4F9A9FC5AF91D" authority="(de Saussure, 1858)" baseAuthorityName="de Saussure" baseAuthorityYear="1858" box="[432,901,1652,1676]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="D6E0EA90ED57FFE86CA4F9A9FD7EF91A" box="[432,673,1653,1676]" italics="true" pageId="16" pageNumber="17">Pachygrapsus gracilis</emphasis>
(de
<bibRefCitation id="80054B73ED57FFE86FC2F9A8FCA2F91D" author="Saussure" box="[726,893,1652,1675]" pageId="16" pageNumber="17" pagination="417 - 496" refId="ref33810" refString="Saussure de, H. (1858) Memoire sur divers crustaces nouveaux du Mexique et des Antilles. Memoires de la Societe de Physique et d'Histoire naturelle de Geneve, 14 (2), 417 - 496 [1 - 82], pl. 1 - 6." type="journal article" year="1858">Saussure, 1858</bibRefCitation>
)
</taxonomicName>
. a) dorsal view; b) front; c) left chela, outer face; d) right P5; e) male abdomen. a, ce, male 14.9 x 19.5 mm, Florida (USNM 170178); b) ov. female 8.5 x 11 mm, Angola (MNHN B12903).
</paragraph>
</caption>
<paragraph id="E42B3682ED57FFE86C1CF8D9FEBEF888" blockId="16.[264,1320,1797,1903]" box="[264,353,1797,1822]" pageId="16" pageNumber="17">
<emphasis id="D6E0EA90ED57FFE86C1CF8D9FEBEF888" box="[264,353,1797,1822]" italics="true" pageId="16" pageNumber="17">Habitat</emphasis>
</paragraph>
<paragraph id="E42B3682ED57FFE86C2CF8F1FE8BF8F9" blockId="16.[264,1320,1797,1903]" pageId="16" pageNumber="17">
Intertidal. Among mangrove roots and on river banks near the sea (
<bibRefCitation id="80054B73ED57FFE86974F8F1FE98F8F9" author="Chace, F. A. &amp; Hobbs, H. H." pageId="16" pageNumber="17" pagination="1 - 5" refId="ref29302" refString="Chace, F. A. &amp; Hobbs, H. H. (1969) The freshwater and terrestrial decapod crustaceans of the West Indies with special reference to Dominica. Bredin-Archbold-Smithsonian Biological Survey of Dominica. Bulletin United States National Museum, 292, v + 1 - 258, pl. 1 - 5." type="journal article" year="1969">Chace &amp; Hobbs, 1969</bibRefCitation>
).
</paragraph>
<paragraph id="E42B3682ED56FFE96C1CFEC0FEB1FEA3" blockId="17.[264,1323,284,670]" box="[264,366,284,309]" pageId="17" pageNumber="18">
<emphasis id="D6E0EA90ED56FFE96C1CFEC0FEB1FEA3" box="[264,366,284,309]" italics="true" pageId="17" pageNumber="18">Remarks</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="AC8E6509ED56FFE96C2CFE98FAF4FD08" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="E42B3682ED56FFE96C2CFE98FAF4FD08" blockId="17.[264,1323,284,670]" pageId="17" pageNumber="18">
<taxonomicName id="23944D01ED56FFE96C2CFE98FD9DFECB" ID-CoL="4BSC3" baseAuthorityName="de Saussure" baseAuthorityYear="1858" box="[312,578,324,349]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="D6E0EA90ED56FFE96C2CFE98FD9DFECB" box="[312,578,324,349]" italics="true" pageId="17" pageNumber="18">Pachygrapsus gracilis</emphasis>
</taxonomicName>
is morphologically close to
<taxonomicName id="23944D01ED56FFE96E8BFE98FB0AFEC8" ID-CoL="4BSC9" authority="Chace, 1966" authorityName="Chace" authorityYear="1966" box="[927,1237,324,350]" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="loveridgei">
<emphasis id="D6E0EA90ED56FFE96E8BFE98FBE9FECB" box="[927,1078,324,349]" italics="true" pageId="17" pageNumber="18">P. loveridgei</emphasis>
<bibRefCitation id="80054B73ED56FFE96954FE98FB0AFEC8" author="Chace, F. A." box="[1088,1237,324,350]" pageId="17" pageNumber="18" pagination="623 - 660" refId="ref29268" refString="Chace, F. A. (1966) Decapod crustaceans from St. Helena Island, South Atlantic. Proceedings of the United States National Museum, 118, 623 - 660." type="journal article" year="1966">Chace, 1966</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="23944D01ED56FFE96801FE98FD94FE10" ID-CoL="4BSCP" authority="(Gibbes, 1850)" baseAuthorityName="Gibbes" baseAuthorityYear="1850" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="transversus">
<emphasis id="D6E0EA90ED56FFE96801FE98FE50FE13" italics="true" pageId="17" pageNumber="18">P. transversus</emphasis>
(
<bibRefCitation id="80054B73ED56FFE96C8BFEB0FD9DFE10" author="Gibbes, L. R." box="[415,578,364,390]" pageId="17" pageNumber="18" pagination="165 - 201" refId="ref30581" refString="Gibbes, L. R. (1850) On the carcinological Collections of the Cabinets of Natural History in the United States. Proceedings of the Third Meeting of American Association for Advancement of Science, 3, 165 - 201." type="journal article" year="1850">Gibbes, 1850</bibRefCitation>
)
</taxonomicName>
, both also Atlantic species of similar size, with only one tooth behind the exorbital angle and with the male abdomen not triangular at the tip. The three species have very distinct G1 that should not be confused (
<figureCitation id="7CAF2A07ED56FFE96EC6FE60FBB3FE40" box="[978,1132,444,470]" captionStart="FIGURE 15" captionStartId="48.[264,368,1262,1286]" captionTargetBox="[381,1213,284,1247]" captionTargetId="figure-298@48.[340,1254,282,1260]" captionTargetPageId="48" captionText="FIGURE 15. —Male gonopods: a) Pachygrapsus fakaravensis Rathbun (1907), holotype 18.0 x 19.4 mm, Fakarava, French Polynesia (USNM 32844), right G1, abdominal face (left) and detail of terminal portion, sternal face (right); b) Pachygrapsus laevimanus Stimpson, 1858, neotype 18.0 x 21.7 mm, Sydney, Australia (AM P7478), left G1, abdominal (left) and sternal face (right), setae omitted; c) Pachygrapsus minutus A. Milne­Edwards, 1873, cl 8.2 mm, Caroline Is., lateral (left) and abdominal face (right), setae omitted; d) Pachygrapsus planifrons de Man, 1888, 9.5 x 11.8 mm, Hawaiian Is. (USNM 29350), right G1, abdominal (left) and sternal face (right); e) Pachygrapsus plicatus (H. Milne Edwards, 1837), cl 11.7 mm, Hawaiian Is., lateral (left) and abdominal face (right); f) Pachygrapsus corrugatus (von Martens, 1872), cl 8.5 mm, Ascension I., left G1 abdominal face (left) and detail of terminal portion, setae omitted (right); g) Pachygrapsus crassipes Randall, 1840, cl 35.6 mm, California, lateral (left) and abdominal face (right) setae omitted; h) Pachygrapsus gracilis (de Saussure, 1858), cl 8.7 mm, Virgin Is., lateral (left) and abdominal face (right), setae omitted; i) Pachygrapsus loveridgei Chace, 1966, holotype 10.3 x 13.1 mm, St. Helena I. (USNM 112457), left G1, abdominal (left) and sternal face (right); j) Pachygrapsus marmoratus (Fabricius, 1787), cl 30.5 mm, Gibraltar, lateral (left) and abdominal face (right), setae omitted; k) Pachygrapsus maurus (Lucas, 1846), cl 8.5 mm, Canary Is., lateral (left) and abdominal face (right) setae omitted; l) Pachygrapsus transversus (Gibbes, 1850), cl 11.1 mm, Key West, Florida, lateral (left) and abdominal face (right), setae omitted. (c, e, gl, from Chace, 1966, fig. 1011; f, from Manning &amp; Chace, 1990, fig. 43). Approximate magnification x 6 (x 12 for detail of terminal portion)." figureDoi="http://doi.org/10.5281/zenodo.5049264" httpUri="https://zenodo.org/record/5049264/files/figure.png" pageId="17" pageNumber="18">Fig. 15h, i, l</figureCitation>
).
<taxonomicName id="23944D01ED56FFE96990FE60FEBEFE6B" ID-CoL="4BSC3" baseAuthorityName="de Saussure" baseAuthorityYear="1858" class="Malacostraca" family="Grapsidae" genus="Pachygrapsus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="D6E0EA90ED56FFE96990FE60FEBEFE6B" italics="true" pageId="17" pageNumber="18">Pachygrapsus gracilis</emphasis>
</taxonomicName>
is also distinct by the lateral carapace margins being strongly convergent posteriorly, the straight anterior margin of the front, the infraorbital margin being almost smooth and without an outer notch, the upper margin of the dactyl of the chela being tuberculated, the absence of a brush of setae on the propodi of the ambulatory legs, and by its colour pattern, having spots that are retained on the carapace even after several years of preservation.
</paragraph>
</subSubSection>
</treatment>
</document>