treatments-xml/data/69/3D/02/693D028CC37E57D9896790CBCA0F24F7.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.952.52695" ID-GBIF-Dataset="ae804d69-5527-45cc-b501-d8203d096d1d" ID-PMC="PMC7394776" ID-Pensoft-Pub="1313-2970-952-65" ID-Pensoft-UUID="2BA2D209D7AA5CFA8272664366B9D9D5" ID-PubMed="32774112" ID-ZooBank="9752C98C0C8F4BCC974BEEF6A47F0F59" ModsDocID="1313-2970-952-65" checkinTime="1595568097234" checkinUser="pensoft" docAuthor="Sutcharit, Chirasak, Inkhavilay, Khamla &amp; Panha, Somsak" docDate="2020" docId="693D028CC37E57D9896790CBCA0F24F7" docLanguage="en" docName="ZooKeys 952: 65-93" docOrigin="ZooKeys 952" docSource="http://dx.doi.org/10.3897/zookeys.952.52695" docTitle="Trichelix Ancey 1887" docType="treatment" docVersion="4" id="2BA2D209D7AA5CFA8272664366B9D9D5" lastPageNumber="65" masterDocId="2BA2D209D7AA5CFA8272664366B9D9D5" masterDocTitle="Taxonomic note on Trichelix horrida (Pfeiffer, 1863) from Laos, with a type catalogue of Moellendorffia, Trichelix, and Moellendorffiella (Heterobranchia, Camaenidae)" masterLastPageNumber="93" masterPageNumber="65" pageNumber="65" updateTime="1668169153165" updateUser="ExternalLinkService">
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<mods:title>Taxonomic note on Trichelix horrida (Pfeiffer, 1863) from Laos, with a type catalogue of Moellendorffia, Trichelix, and Moellendorffiella (Heterobranchia, Camaenidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Inkhavilay, Khamla</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Panha, Somsak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>952</mods:number>
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<mods:start>65</mods:start>
<mods:end>93</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.952.52695</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.952.52695</mods:identifier>
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<treatment ID-GBIF-Taxon="165940712" LSID="urn:lsid:plazi:treatment:693D028CC37E57D9896790CBCA0F24F7" httpUri="http://treatment.plazi.org/id/693D028CC37E57D9896790CBCA0F24F7" lastPageNumber="65" pageId="0" pageNumber="65">
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Genus 
<taxonomicName LSID="693D028C-C37E-57D9-8967-90CBCA0F24F7" authority="Ancey, 1887" authorityName="Ancey" authorityYear="1887" class="Gastropoda" family="Camaenidae" genus="Trichelix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichelix" order="Stylommatophora" pageId="0" pageNumber="65" phylum="Mollusca" rank="genus">Trichelix Ancey, 1887</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="65" type="reference_group">
<paragraph pageId="0" pageNumber="65">
<taxonomicName authorityName="Ancey" authorityYear="1887" class="Gastropoda" family="Camaenidae" genus="Trichelix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichelix" order="Stylommatophora" pageId="0" pageNumber="65" phylum="Mollusca" rank="genus">Trichelix</taxonomicName>
<bibRefCitation author="Ancey, C-F" pageId="0" pageNumber="65" publicationUrl="https://www.biodiversitylibrary.org/page/28528930" refId="B1" refString="Ancey, C-F, 1887. Description of new genera or subgenera of Helicidae. The Conchologists Exchange 1: 64. https://www.biodiversitylibrary.org/page/28528930" title="Description of new genera or subgenera of Helicidae. The Conchologists Exchange 1: 64." url="https://www.biodiversitylibrary.org/page/28528930" year="1887">Ancey 1887</bibRefCitation>
: 64. 
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica, Supplement" pageId="0" pageNumber="65" pagination="1467 - 1626" refId="B52" refString="Schileyko, AA, 2003. Treatise on recent terrestrial pulmonate mollusks. Part 11: Trigonochlamydidae, Papillodesmidae, Vitrinidae, Limacidae, Bielziidae, Agriolimacidae, Boettgerillidae, Camaenidae. Ruthenica, Supplement 2: 1467 - 1626" title="Treatise on recent terrestrial pulmonate mollusks. Part 11: Trigonochlamydidae, Papillodesmidae, Vitrinidae, Limacidae, Bielziidae, Agriolimacidae, Boettgerillidae, Camaenidae." volume="2" year="2003">Schileyko 2003</bibRefCitation>
: 1513.
</paragraph>
<paragraph pageId="0" pageNumber="65">
<taxonomicName authority="" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Trihelix)" order="Stylommatophora" pageId="0" pageNumber="65" phylum="Mollusca" rank="subGenus" subGenus="Trihelix">Helix (Trihelix)</taxonomicName>
: 
<bibRefCitation author="Pilsbry, HA" journalOrPublisher="Academy of Natural Science Philadelphia, PA" pageId="0" pageNumber="65" publicationUrl="https://www.biodiversitylibrary.org/page/23626780" refId="B45" refString="Pilsbry, HA, 1890. [1890-1891] Manual of Conchology, Structure and Systematic, with Illustrations of the Species, Second Series, Volume 6 (Helicidae, Vol. IV). Academy of Natural Science Philadelphia, PA, https://www.biodiversitylibrary.org/page/23626780" title="[1890 - 1891] Manual of Conchology, Structure and Systematic, with Illustrations of the Species, Second Series, Volume 6 (Helicidae, Vol. IV)." url="https://www.biodiversitylibrary.org/page/23626780" year="1890">Pilsbry 1890</bibRefCitation>
: 9 (incorrect subsequent spelling). 
<bibRefCitation author="Pilsbry, HA" journalOrPublisher="Academy of Natural Science Philadelphia, PA" pageId="0" pageNumber="65" publicationUrl="https://www.biodiversitylibrary.org/page/1103099" refId="B46" refString="Pilsbry, HA, 1895. [1893-1895] Manual of Conchology, Structure and Systematic, with Illustrations of the Species, Second Series, Volume 9 (Helicidae, Vol. 7. Guide to the Study of Helices). Academy of Natural Science Philadelphia, PA, https://www.biodiversitylibrary.org/page/1103099" title="[1893 - 1895] Manual of Conchology, Structure and Systematic, with Illustrations of the Species, Second Series, Volume 9 (Helicidae, Vol. 7. Guide to the Study of Helices)." url="https://www.biodiversitylibrary.org/page/1103099" year="1895">Pilsbry 1895</bibRefCitation>
: 289.
</paragraph>
<paragraph pageId="0" pageNumber="65">
<taxonomicName authorityName="Ancey" authorityYear="1887" class="Gastropoda" family="Camaenidae" genus="Moellendorffia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Moellendorffia" order="Stylommatophora" pageId="0" pageNumber="65" phylum="Mollusca" rank="genus">Moellendorffia</taxonomicName>
(
<taxonomicName authorityName="Ancey" authorityYear="1898" class="Gastropoda" family="Camaenidae" genus="Trihelix" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trihelix" order="Stylommatophora" pageId="0" pageNumber="65" phylum="Mollusca" rank="genus">Trihelix</taxonomicName>
[sic]): 
<bibRefCitation author="Pilsbry, HA" journalOrPublisher="The Nautilus" pageId="0" pageNumber="65" pagination="63 - 67" publicationUrl="https://www.biodiversitylibrary.org/page/1755561" refId="B48" refString="Pilsbry, HA, 1905. Notes on Moellendorffia and Stegodera. The Nautilus 19: 63 - 67, https://www.biodiversitylibrary.org/page/1755561" title="Notes on Moellendorffia and Stegodera." url="https://www.biodiversitylibrary.org/page/1755561" volume="19" year="1905">Pilsbry 1905</bibRefCitation>
: 65.
</paragraph>
<paragraph pageId="0" pageNumber="65">
<taxonomicName authority="" class="Gastropoda" family="Camaenidae" genus="Moellendorffia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Moellendorffia (Trichelix)" order="Stylommatophora" pageId="0" pageNumber="65" phylum="Mollusca" rank="subGenus" subGenus="Trichelix">Moellendorffia (Trichelix)</taxonomicName>
: 
<bibRefCitation author="Zilch, A" journalOrPublisher="Gebrueder Borntraeger, Berlin" pageId="0" pageNumber="65" refId="B65" refString="Zilch, A, 1960 [1959-1960]. Gastropoda, Euthyneura. In: Schindewolf OH (Ed.) Handbuch der Palaeozoologie, Band 6, Gastropoda. Gebrueder Borntraeger, Berlin" title="Gastropoda, Euthyneura. In: Schindewolf OH (Ed.) Handbuch der Palaeozoologie, Band 6, Gastropoda." year="1960 [1959 - 1960]">Zilch 1960</bibRefCitation>
: 612.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="type species">
<paragraph pageId="0" pageNumber="65">Type species.</paragraph>
<paragraph pageId="0" pageNumber="65">
<taxonomicName authorityName="L.Pfeiffer" authorityYear="1863" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix horrida" order="Stylommatophora" pageId="0" pageNumber="65" phylum="Mollusca" rank="species" species="horrida">
<emphasis italics="true" pageId="0" pageNumber="65">Helix horrida</emphasis>
</taxonomicName>
Pfeiffer, 1863; by original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="description">
<paragraph pageId="0" pageNumber="65">Description.</paragraph>
<paragraph pageId="0" pageNumber="65">Shell small to medium-sized, flattened to concave, rather thin, umbilicate, and corneous to brownish. Spire shrunken; embryonic shell nearly smooth; following whorls granulated and with short to long periostracal hairs arranged in oblique rows along the lines of growth. Last whorl rounded and descending anteriorly. Aperture ventral or subvertical; trigonal or subcircular; with strong or weak barriers inside the aperture at upper periphery and below periphery, and externally marked with strong to weak longitudinal furrows. Peristome expanded and continuous or discontinuous; parietal callus thin or thickened and little elevated.</paragraph>
<paragraph pageId="0" pageNumber="65">Genitalia typical of camaenids, without either dart apparatus or accessory glands. Penis and epiphallus long, penial verge present, and flagellum short. Internal wall of penis and vagina with longitudinal pilasters.</paragraph>
<paragraph pageId="0" pageNumber="65">Radular teeth arranged in V-shaped rows; central and lateral teeth triangular.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="65" type="remarks">
<paragraph pageId="0" pageNumber="65">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="65">
The genus is currently comprised of six nominal species (
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica, Supplement" pageId="0" pageNumber="65" pagination="1467 - 1626" refId="B52" refString="Schileyko, AA, 2003. Treatise on recent terrestrial pulmonate mollusks. Part 11: Trigonochlamydidae, Papillodesmidae, Vitrinidae, Limacidae, Bielziidae, Agriolimacidae, Boettgerillidae, Camaenidae. Ruthenica, Supplement 2: 1467 - 1626" title="Treatise on recent terrestrial pulmonate mollusks. Part 11: Trigonochlamydidae, Papillodesmidae, Vitrinidae, Limacidae, Bielziidae, Agriolimacidae, Boettgerillidae, Camaenidae." volume="2" year="2003">Schileyko 2003</bibRefCitation>
; 
<bibRefCitation author="Minato, H" journalOrPublisher="Chiribotan" pageId="0" pageNumber="65" pagination="20 - 34" refId="B30" refString="Minato, H, 2011. A memorandum of the studies of Japanese land snails (8), the genus Moellendorffia (Pulmonata: Camaenidae) from Amami Islands. Chiribotan 45: 20 - 34" title="A memorandum of the studies of Japanese land snails (8), the genus Moellendorffia (Pulmonata: Camaenidae) from Amami Islands." volume="45" year="2011">Minato 2011</bibRefCitation>
). Two species occur in northern Laos and southern China (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Approximate geographical position of the type locality of all nominal species of the genera Moellendorffia (circle), Trichelix (square), and Moellendorffiella (triangle). The numbers indicated correspond to the nominal species listed in the catalogue of the type specimen." figureDoi="10.3897/zookeys.952.52695.figure3" httpUri="https://binary.pensoft.net/fig/435099" pageId="0" pageNumber="65">3</figureCitation>
), viz. 
<taxonomicName lsidName="T. horrida" pageId="0" pageNumber="65" rank="species" species="horrida">
<emphasis italics="true" pageId="0" pageNumber="65">T. horrida</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="T. biscalpta" pageId="0" pageNumber="65" rank="species" species="biscalpta">
<emphasis italics="true" pageId="0" pageNumber="65">T. biscalpta</emphasis>
</taxonomicName>
(Heude, 1885), and one has been recorded from Taiwan, viz. 
<taxonomicName lsidName="T. hiraseana" pageId="0" pageNumber="65" rank="species" species="hiraseana">
<emphasis italics="true" pageId="0" pageNumber="65">T. hiraseana</emphasis>
</taxonomicName>
(Pilsbry, 1905). Three species occur on the Amami Islands, Central Ryukyu Islands, Japan, viz. 
<taxonomicName lsidName="T. eucharista" pageId="0" pageNumber="65" rank="species" species="eucharista">
<emphasis italics="true" pageId="0" pageNumber="65">T. eucharista</emphasis>
</taxonomicName>
(Pilsbry, 1901), 
<taxonomicName lsidName="T. diminuta" pageId="0" pageNumber="65" rank="species" species="diminuta">
<emphasis italics="true" pageId="0" pageNumber="65">T. diminuta</emphasis>
</taxonomicName>
(Pilsbry &amp; Hirase, 1905) and 
<taxonomicName lsidName="T. tokunoensis" pageId="0" pageNumber="65" rank="species" species="tokunoensis">
<emphasis italics="true" pageId="0" pageNumber="65">T. tokunoensis</emphasis>
</taxonomicName>
(Pilsbry &amp; Hirase, 1905).
</paragraph>
</subSubSection>
</treatment>
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