291 lines
28 KiB
XML
291 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/alpento.4.50408" ID-GBIF-Dataset="586342a0-2920-4cda-a127-0174e38900ac" ID-GBIF-Taxon="163641915" ID-Pensoft-Pub="2535-0889-4-29" ID-Pensoft-UUID="CE022010D00B55C688AB9F10AC804858" ID-ZooBank="7C1EAAF32DD34307ABA1D9AAD6F795F2" LSID="urn:lsid:zoobank.org:act:49BA78BD-7423-410B-A311-9BD18A1C134D" ModsDocID="2535-0889-4-29" checkinTime="1587051829217" checkinUser="pensoft" docAuthor="Huber, Charles & Schnitter, Peer Hajo" docDate="2020" docId="42E5E0CA916250D49AAAFD179099D0D4" docLanguage="en" docName="AlpEntomol 4: 29-38" docOrigin="Alpine Entomology 4" docSource="http://dx.doi.org/10.3897/alpento.4.50408" docTitle="Nebria (Pseudonebriola) tsambagarav Huber & Schnitter 2020, sp. nov." docType="treatment" docUuid="49BA78BD-7423-410B-A311-9BD18A1C134D" docUuidSource="ZooBank" docVersion="3" id="CE022010D00B55C688AB9F10AC804858" lastPageNumber="29" masterDocId="CE022010D00B55C688AB9F10AC804858" masterDocTitle="Nebria (Pseudonebriola) tsambagarav sp. nov., a new alpine species from the Mongolian Altai (Coleoptera, Carabidae)" masterLastPageNumber="38" masterPageNumber="29" pageId="0" pageNumber="29" updateTime="1643444276011" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Nebria (Pseudonebriola) tsambagarav sp. nov., a new alpine species from the Mongolian Altai (Coleoptera, Carabidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Huber, Charles</mods:namePart>
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<mods:affiliation>Natural History Museum, Bernastrasse 15, CH- 3005 Bern, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0064-145X</mods:nameIdentifier>
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<mods:nameIdentifier type="email">charles.huber@nmbe.ch</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schnitter, Peer Hajo</mods:namePart>
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<mods:affiliation>Gartenstadtstrasse 8, D- 06126 Halle (Saale), Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Alpine Entomology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>29</mods:start>
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<mods:end>38</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/alpento.4.50408</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/alpento.4.50408</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2535-0889-4-29</mods:identifier>
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<mods:identifier type="ZooBank">7C1EAAF32DD34307ABA1D9AAD6F795F2</mods:identifier>
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<mods:identifier type="Pensoft-UUID">CE022010D00B55C688AB9F10AC804858</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="163641915" LSID="urn:lsid:zoobank.org:act:49BA78BD-7423-410B-A311-9BD18A1C134D" httpUri="http://treatment.plazi.org/id/42E5E0CA916250D49AAAFD179099D0D4" lastPageNumber="29" pageId="0" pageNumber="29">
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<subSubSection pageId="0" pageNumber="29" type="nomenclature">
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<paragraph pageId="0" pageNumber="29">
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<taxonomicName LSID="42E5E0CA-9162-50D4-9AAA-FD179099D0D4" authority="Huber & Schnitter, 2020" authorityName="Huber & Schnitter" authorityYear="2020" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nebria (Pseudonebriola) tsambagarav" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="tsambagarav" status="sp. nov." subGenus="Pseudonebriola">Nebria (Pseudonebriola) tsambagarav</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="29">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 3" captionStartId="F4" captionText="Figure 3. Nebria (Pseudonebriola) tsambagarav sp. nov., habitus, holotype. Scale bar: 5 mm." figureDoi="10.3897/alpento.4.50408.figure3" httpUri="https://binary.pensoft.net/fig/400474" pageId="0" pageNumber="29">Figs 3</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Second antennomere of right antenna of Nebria (Pseudonebriola) tsambagarav sp. nov., paratype, with four apical setae. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.50408.figure5" httpUri="https://binary.pensoft.net/fig/400476" pageId="0" pageNumber="29">, 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronotum: A, Nebria (Pseudonebriola) kerzhneri Shilenkov, 1982, holotype; B, N. (P.) medvedevi Shilenkov, 1982, paratype; C, N. (P.) tsambagarav sp. nov., paratype. Scale bar: 1 mm." figureDoi="10.3897/alpento.4.50408.figure6" httpUri="https://binary.pensoft.net/fig/400477" pageId="0" pageNumber="29">, 6</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Median lobes; A-B dried lobes, C-D embedded lobes, A-C with partially everted endophallus: A, Nebria (Pseudonebriola) kerzhneri Shilenkov, 1982, holotype; B, N. (P.) medvedevi Shilenkov, 1982, paratype; C, N. (P.) tsambagarav sp. nov., holotype; D, N. (P.) kaszabi Shilenkov, 1982, Urunkhaika, Marakol Lake, Kazakhstan. Scale bar: 1 mm." figureDoi="10.3897/alpento.4.50408.figure7" httpUri="https://binary.pensoft.net/fig/400478" pageId="0" pageNumber="29">, 7</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Right gonocoxa (in ventral view): A, Nebria (Pseudonebriola) tsambagarav sp. nov., paratype; B, N. (P.) kaszabi Shilenkov, 1982; Urunkhaika, Marakol Lake KAZ. Scale bar: 0.5 mm." figureDoi="10.3897/alpento.4.50408.figure8" httpUri="https://binary.pensoft.net/fig/400479" pageId="0" pageNumber="29">, 8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="type material">
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<paragraph pageId="0" pageNumber="29">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" pageId="0" pageNumber="29">Holotype</emphasis>
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♂:
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<geoCoordinate degrees="48" direction="north" minutes="41" orientation="latitude" precision="1" seconds="27.1" value="48.69086">
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48°41
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<normalizedToken originalValue="’27.1”">'27.1"</normalizedToken>
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N
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</geoCoordinate>
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,
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<geoCoordinate degrees="90" direction="east" minutes="40" orientation="longitude" precision="1" seconds="33.4" value="90.67594">
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90°40
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<normalizedToken originalValue="’33.4”">'33.4"</normalizedToken>
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E
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</geoCoordinate>
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; Mongolei, Bajan Ulgij Aimak, Tsambagarav uul; alpine Zone: Bachufer; 3168
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<normalizedToken originalValue="üNN">ueNN</normalizedToken>
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; 1. 7. 2016; 20 MG 2016 [internal abbreviation of the collector: location No. 20, Mongolia expedition 2016]; leg. Schnitter (NMBE).
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</paragraph>
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" pageId="0" pageNumber="29">Paratypes</emphasis>
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: 31 ♂, 22 ♀, same data as holotype (NMBE, ZIN, cBK, cSCHN).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="29" type="description">
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<paragraph pageId="0" pageNumber="29">Description.</paragraph>
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<paragraph pageId="0" pageNumber="29">Body size: 7.5-9 mm.</paragraph>
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<paragraph pageId="0" pageNumber="29">Colour: Dorsal surface black, head black, without lightened spots on the vertex; appendages of the head brownish lightened, always lighter than the head. Antennal scape black, antennomeres 2-11 brown with darkened apices. Femora black, tibiae and tarsomeres brown, apices of tarsomeres usually black.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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Head: Mandibles short; stipes flat, smooth, with 4-6 setae. Anterior margin of the labrum straight, bearing six setae. Anterior margin of the clypeus straight or slightly concave; clypeus laterally unisetose. Frontoclypeal suture concave. Frons with transverse wrinkles, which roundly merge into the lateral and longitudinal supraorbital wrinkles. Eyes protruding, temples present, oblique. Head supraorbitally unisetose. Disc impunctate with an isodiametric microreticulation. Antennae long and thin, extending to the end of the basal third of the elytra. Antennal scape as long as the
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<normalizedToken originalValue="eye’s">eye's</normalizedToken>
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diameter (ratio antennal scape length/eye length = 0.98
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.03 (0.93-1.04), slightly conical to suboval, distinctly narrowed basally and slightly apically, with one dorsoapical seta (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F3" captionText="Figure 4. Antennal scape: A, Nebria (Pseudonebriola) tsambagarav sp. nov. paratype; B, N. (P.) medvedevi Shilenkov, 1982, paratype. Scale bar: 0.5 mm." figureDoi="10.3897/alpento.4.50408.figure4" httpUri="https://binary.pensoft.net/fig/400475" pageId="0" pageNumber="29">4</figureCitation>
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). Ratio antennal scape length/scape width = 2.16
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.07 (2.05-2.26). 2nd antennomere with one long ventroapical and one long dorsoapical seta, and generally (in 89% of the examined specimens) with 1-2 additional, often hardly visible short setae on the apex; therefore the 2nd antennomere trisetose (in 47%) or even quadrisetose (in 42%) (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Second antennomere of right antenna of Nebria (Pseudonebriola) tsambagarav sp. nov., paratype, with four apical setae. Scale bar: 0.1 mm." figureDoi="10.3897/alpento.4.50408.figure5" httpUri="https://binary.pensoft.net/fig/400476" pageId="0" pageNumber="29">5</figureCitation>
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). Ligula short, triangular, broad at base, with two apical setae. Penultimate labial palpomere trisetose. Mentum bidentate, medially hardly incised. Median teeth with one seta at each side. Lobus lateralis of the mentum wide, apically faintly rounded or even rectilinearly narrowed, spina apicalis short, triangular, incisio lateralis distinct. Submentum bilaterally with 3-4 setae.
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</paragraph>
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<caption doi="10.3897/alpento.4.50408.figure3" httpUri="https://binary.pensoft.net/fig/400474" pageId="0" pageNumber="29" start="Figure 3" startId="F4">
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" pageId="0" pageNumber="29">Figure 3.</emphasis>
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<taxonomicName authorityName="Huber & Schnitter" authorityYear="2020" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nebria (Pseudonebriola) tsambagarav" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="tsambagarav" subGenus="Pseudonebriola">Nebria (Pseudonebriola) tsambagarav</taxonomicName>
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sp. nov., habitus, holotype. Scale bar: 5 mm.
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</paragraph>
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</caption>
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<caption doi="10.3897/alpento.4.50408.figure4" httpUri="https://binary.pensoft.net/fig/400475" pageId="0" pageNumber="29" start="Figure 4" startId="F3">
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" pageId="0" pageNumber="29">Figure 4.</emphasis>
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Antennal scape:
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<emphasis bold="true" pageId="0" pageNumber="29">A</emphasis>
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,
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<taxonomicName authorityName="Huber & Schnitter" authorityYear="2020" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nebria (Pseudonebriola) tsambagarav" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="tsambagarav" subGenus="Pseudonebriola">Nebria (Pseudonebriola) tsambagarav</taxonomicName>
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sp. nov. paratype;
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<emphasis bold="true" pageId="0" pageNumber="29">B</emphasis>
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,
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<taxonomicName genus="N." lsidName="N. (P.) medvedevi" pageId="0" pageNumber="29" rank="species" species="medvedevi" subGenus="P.">N. (P.) medvedevi</taxonomicName>
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Shilenkov, 1982, paratype. Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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<caption doi="10.3897/alpento.4.50408.figure5" httpUri="https://binary.pensoft.net/fig/400476" pageId="0" pageNumber="29" start="Figure 5" startId="F5">
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" pageId="0" pageNumber="29">Figure 5.</emphasis>
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Second antennomere of right antenna of
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<taxonomicName authorityName="Huber & Schnitter" authorityYear="2020" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nebria (Pseudonebriola) tsambagarav" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="tsambagarav" subGenus="Pseudonebriola">Nebria (Pseudonebriola) tsambagarav</taxonomicName>
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sp. nov., paratype, with four apical setae. Scale bar: 0.1 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="29">
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Pronotum (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronotum: A, Nebria (Pseudonebriola) kerzhneri Shilenkov, 1982, holotype; B, N. (P.) medvedevi Shilenkov, 1982, paratype; C, N. (P.) tsambagarav sp. nov., paratype. Scale bar: 1 mm." figureDoi="10.3897/alpento.4.50408.figure6" httpUri="https://binary.pensoft.net/fig/400477" pageId="0" pageNumber="29">6</figureCitation>
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): Cordate, widest at apical third or even forth; ratio maximum width/length = 1.43
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.03 (1.39-1.49). Anterior angles widely rounded, faintly protruding. Lateral margin evenly rounded to the anterior angles, faintly rounded to the posterior angles with a distinct sinuation in front of the posterior angle. Lateral margin of the posterior angles long. Posterior angles rectangular or even acutely turned outwards. Ratio apical pronotal width/maximum pronotal width = 0.80
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.02 (0.78-0.84). Apical margination of the pronotum restricted to lateral one-third. Basal margin straight, occasionally laterally faintly curved, the posterior angles not protruding backwards. Basal width of the pronotum 0.92 (0.88-0.97) times the anterior width. Lateral groove distinct, narrow, flat, impunctate or with isolated dents, laterally narrowly edged, slightly broadened to the anterior angle, posteriorly joining the deep basal fovea. One midlateral seta present near widest point of the pronotum; occasionally bilateral-bisetose (2%) or asymmetrically uni-/bisetose (2%). Basolateral seta present. Apical and basal transverse impression coarsely but sparsely punctate. Median longitudinal impression distinct, reaching the posterior border of the pronotum. Disc with isodiametric microreticulation. Prosternal process wide, triangular, laterally slightly margined, medially flat, apically bulging and immarginate. Proepisternum smooth and impunctate.
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</paragraph>
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<caption doi="10.3897/alpento.4.50408.figure6" httpUri="https://binary.pensoft.net/fig/400477" pageId="0" pageNumber="29" start="Figure 6" startId="F6">
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" pageId="0" pageNumber="29">Figure 6.</emphasis>
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Pronotum:
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<emphasis bold="true" pageId="0" pageNumber="29">A</emphasis>
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,
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<taxonomicName authorityName="Shilenkov" authorityYear="1982" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nebria (Pseudonebriola) kerzhneri" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="kerzhneri" subGenus="Pseudonebriola">Nebria (Pseudonebriola) kerzhneri</taxonomicName>
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Shilenkov, 1982, holotype;
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<emphasis bold="true" pageId="0" pageNumber="29">B</emphasis>
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,
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<taxonomicName genus="N." lsidName="N. (P.) medvedevi" pageId="0" pageNumber="29" rank="species" species="medvedevi" subGenus="P.">N. (P.) medvedevi</taxonomicName>
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Shilenkov, 1982, paratype;
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<emphasis bold="true" pageId="0" pageNumber="29">C</emphasis>
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,
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<taxonomicName genus="N." lsidName="N. (P.) tsambagarav" pageId="0" pageNumber="29" rank="species" species="tsambagarav" subGenus="P.">N. (P.) tsambagarav</taxonomicName>
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sp. nov., paratype. Scale bar: 1 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="29">
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Elytra: Silhouette suboval, apically faintly expanded, maximum width at two thirds. Ratio length/width of the elytra = 1.53
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.02 (1.51-1.57). Elytral width = 1.59
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.04 (1.51-1.66) times the pronotal width. Lateral margin faintly rounded, apex acutely rounded. Basal margination slightly curved, joined at an obtuse angle with the lateral margination. Shoulder flatly curved. Hindwings vestigial. Humeral and apical carinae little pronounced. Striae 1-7 distinct, punctate, stria 8 as a row of punctures. Striae 1-3 reaching the apex, the other striae obliterate slightly in front of the apex. Intervals on disc flat, interval 3 generally asetose (76%) on disc, disregarding the always present apical seta, occasionally with one seta on the disc (24%). Scutellar seta absent. Microsculpture isodiametric.
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</paragraph>
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<paragraph pageId="0" pageNumber="29">Ventral surface: Mes- and Metepisterna smooth and impunctate. Metacoxa basally and apically unisetose. Sternum II laterally faintly and widely dispersed punctate. Sternum III medially asetose. Sterna IV-VI each with one posterior paramedial seta. Anal sternum paralaterally unisetose in the male, bisetose in the female.</paragraph>
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<paragraph pageId="0" pageNumber="29">Legs: All tarsomeres dorsally glabrous; metatarsomere 4 ventrally with a short projecting tooth. Metafemur dorsoapically with a few short setae.</paragraph>
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<paragraph pageId="0" pageNumber="29">
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Male genitalia: Edeagus (Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Median lobes; A-B dried lobes, C-D embedded lobes, A-C with partially everted endophallus: A, Nebria (Pseudonebriola) kerzhneri Shilenkov, 1982, holotype; B, N. (P.) medvedevi Shilenkov, 1982, paratype; C, N. (P.) tsambagarav sp. nov., holotype; D, N. (P.) kaszabi Shilenkov, 1982, Urunkhaika, Marakol Lake, Kazakhstan. Scale bar: 1 mm." figureDoi="10.3897/alpento.4.50408.figure7" httpUri="https://binary.pensoft.net/fig/400478" pageId="0" pageNumber="29">7C</figureCitation>
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): Base of the median lobe small; the base decreasing abruptly to the thin mid-shaft. Mid-shaft strongly curved at base on inner side, moderately curved to the apex. Apex straight, faintly deflected to the left, in front of the long tip ventrally faintly convex. Mid-shaft of the endophallus with short setae.
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</paragraph>
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<paragraph pageId="0" pageNumber="29">
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Female genitalia: Gonocoxa (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Right gonocoxa (in ventral view): A, Nebria (Pseudonebriola) tsambagarav sp. nov., paratype; B, N. (P.) kaszabi Shilenkov, 1982; Urunkhaika, Marakol Lake KAZ. Scale bar: 0.5 mm." figureDoi="10.3897/alpento.4.50408.figure8" httpUri="https://binary.pensoft.net/fig/400479" pageId="0" pageNumber="29">8A</figureCitation>
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): Gonocoxite 2 slender, dagger-like, curved, two fifth the length of gonocoxite 1, broad at base, distinctly narrowing near base, narrowing towards apex which is faintly deflected. Apex rounded, hardly arcuate and grooved dorsally. Ventral preapical insertion furrow short-oval, with two nematiform setae. Gonocoxites ventrally unjointed and continuously sclerotized, dorsally separated by a membranous area.
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</paragraph>
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<caption doi="10.3897/alpento.4.50408.figure7" httpUri="https://binary.pensoft.net/fig/400478" pageId="0" pageNumber="29" start="Figure 7" startId="F7">
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<paragraph pageId="0" pageNumber="29">
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<emphasis bold="true" pageId="0" pageNumber="29">Figure 7.</emphasis>
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Median lobes; A-B dried lobes, C-D embedded lobes, A-C with partially everted endophallus:
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<emphasis bold="true" pageId="0" pageNumber="29">A</emphasis>
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,
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<taxonomicName authorityName="Shilenkov" authorityYear="1982" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nebria (Pseudonebriola) kerzhneri" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="kerzhneri" subGenus="Pseudonebriola">Nebria (Pseudonebriola) kerzhneri</taxonomicName>
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Shilenkov, 1982, holotype;
|
||
<emphasis bold="true" pageId="0" pageNumber="29">B</emphasis>
|
||
,
|
||
<taxonomicName genus="N." lsidName="N. (P.) medvedevi" pageId="0" pageNumber="29" rank="species" species="medvedevi" subGenus="P.">N. (P.) medvedevi</taxonomicName>
|
||
Shilenkov, 1982, paratype;
|
||
<emphasis bold="true" pageId="0" pageNumber="29">C</emphasis>
|
||
,
|
||
<taxonomicName genus="N." lsidName="N. (P.) tsambagarav" pageId="0" pageNumber="29" rank="species" species="tsambagarav" subGenus="P.">N. (P.) tsambagarav</taxonomicName>
|
||
sp. nov., holotype;
|
||
<emphasis bold="true" pageId="0" pageNumber="29">D</emphasis>
|
||
,
|
||
<taxonomicName genus="N." lsidName="N. (P.) kaszabi" pageId="0" pageNumber="29" rank="species" species="kaszabi" subGenus="P.">N. (P.) kaszabi</taxonomicName>
|
||
Shilenkov, 1982, Urunkhaika, Marakol Lake, Kazakhstan. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/alpento.4.50408.figure8" httpUri="https://binary.pensoft.net/fig/400479" pageId="0" pageNumber="29" start="Figure 8" startId="F8">
|
||
<paragraph pageId="0" pageNumber="29">
|
||
<emphasis bold="true" pageId="0" pageNumber="29">Figure 8.</emphasis>
|
||
Right gonocoxa (in ventral view):
|
||
<emphasis bold="true" pageId="0" pageNumber="29">A</emphasis>
|
||
,
|
||
<taxonomicName authorityName="Huber & Schnitter" authorityYear="2020" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nebria (Pseudonebriola) tsambagarav" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="tsambagarav" subGenus="Pseudonebriola">Nebria (Pseudonebriola) tsambagarav</taxonomicName>
|
||
sp. nov., paratype;
|
||
<emphasis bold="true" pageId="0" pageNumber="29">B</emphasis>
|
||
,
|
||
<taxonomicName genus="N." lsidName="N. (P.) kaszabi" pageId="0" pageNumber="29" rank="species" species="kaszabi" subGenus="P.">N. (P.) kaszabi</taxonomicName>
|
||
Shilenkov, 1982; Urunkhaika, Marakol Lake KAZ. Scale bar: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="29" type="habitat">
|
||
<paragraph pageId="0" pageNumber="29">Habitat</paragraph>
|
||
<paragraph pageId="0" pageNumber="29">
|
||
(Figs
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Tsambagarav uul, Bajan Oelgii province, Western Mongolia. Photo: Peer Schnitter." figureDoi="10.3897/alpento.4.50408.figure9" httpUri="https://binary.pensoft.net/fig/400480" pageId="0" pageNumber="29">9</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitat of Nebria (Pseudonebriola) tsambagarav sp. nov. on Tsambagarav uul; rivulet at 3168 m a. s. l. Photo: Peer Schnitter." figureDoi="10.3897/alpento.4.50408.figure10" httpUri="https://binary.pensoft.net/fig/400481" pageId="0" pageNumber="29">10</figureCitation>
|
||
). The new species was found in the alpine region of the Tsambagarav uul exclusively under bigger stones along the riverbanks of small rivulets. The water ran off higher snowfields. Directly at the snow fields not any specimen was found.
|
||
</paragraph>
|
||
<caption doi="10.3897/alpento.4.50408.figure9" httpUri="https://binary.pensoft.net/fig/400480" pageId="0" pageNumber="29" start="Figure 9" startId="F9">
|
||
<paragraph pageId="0" pageNumber="29">
|
||
<emphasis bold="true" pageId="0" pageNumber="29">Figure 9.</emphasis>
|
||
Tsambagarav uul, Bajan
|
||
<normalizedToken originalValue="Ölgii">Oelgii</normalizedToken>
|
||
province, Western Mongolia. Photo: Peer Schnitter.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/alpento.4.50408.figure10" httpUri="https://binary.pensoft.net/fig/400481" pageId="0" pageNumber="29" start="Figure 10" startId="F10">
|
||
<paragraph pageId="0" pageNumber="29">
|
||
<emphasis bold="true" pageId="0" pageNumber="29">Figure 10.</emphasis>
|
||
Habitat of
|
||
<taxonomicName authorityName="Huber & Schnitter" authorityYear="2020" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nebria (Pseudonebriola) tsambagarav" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="tsambagarav" subGenus="Pseudonebriola">Nebria (Pseudonebriola) tsambagarav</taxonomicName>
|
||
sp. nov. on Tsambagarav uul; rivulet at 3168 m a.s.l. Photo: Peer Schnitter.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="29" type="ecology">
|
||
<paragraph pageId="0" pageNumber="29">Ecology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="29">Hydrophilic, ripicol, cold-preferring, alpine level.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="29" type="etymology">
|
||
<paragraph pageId="0" pageNumber="29">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="29">The specific epithet refers to the type locality, the Tsambagarav uul; noun in apposition.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="29" type="distribution">
|
||
<paragraph pageId="0" pageNumber="29">Distribution</paragraph>
|
||
<paragraph pageId="0" pageNumber="29">
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Map of the Altai Mountain range displaying the distribution of Pseudonebriola species." figureDoi="10.3897/alpento.4.50408.figure11" httpUri="https://binary.pensoft.net/fig/400482" pageId="0" pageNumber="29">11</figureCitation>
|
||
).
|
||
<taxonomicName genus="N." lsidName="N. tsambagarav" pageId="0" pageNumber="29" rank="species" species="tsambagarav">
|
||
<emphasis italics="true" pageId="0" pageNumber="29">N. tsambagarav</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is known only from the type locality at an altitude of 3168 m a.s.l. at the Tsambagarav uul, a mountain (with the summit at 4165 m a.s.l.) of the Mongolian Altai in the westernmost province of Bayan
|
||
<normalizedToken originalValue="Ölgii">Oelgii</normalizedToken>
|
||
of Mongolia.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="29">
|
||
The Mongolian
|
||
<taxonomicName authority="Ledoux & Roux, 1989" authorityName="Ledoux & Roux" authorityYear="1989" class="Insecta" family="Carabidae" genus="Pseudonebriola" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudonebriola" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="29">Pseudonebriola</emphasis>
|
||
</taxonomicName>
|
||
species are restricted to the main ridge of the Mongolian Altai (
|
||
<taxonomicName genus="N." lsidName="N. medvedevi" pageId="0" pageNumber="29" rank="species" species="medvedevi">
|
||
<emphasis italics="true" pageId="0" pageNumber="29">N. medvedevi</emphasis>
|
||
</taxonomicName>
|
||
), to the Tsambagarav mountain, the easternmost ridge bordering the Great Lakes Depression (
|
||
<taxonomicName genus="N." lsidName="N. tsambagarav" pageId="0" pageNumber="29" rank="species" species="tsambagarav">
|
||
<emphasis italics="true" pageId="0" pageNumber="29">N. tsambagarav</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.), and to the Northwestern part of the Gobi Altai (
|
||
<taxonomicName genus="N." lsidName="N. kerzhneri" pageId="0" pageNumber="29" rank="species" species="kerzhneri">
|
||
<emphasis italics="true" pageId="0" pageNumber="29">N. kerzhneri</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/alpento.4.50408.figure11" httpUri="https://binary.pensoft.net/fig/400482" pageId="0" pageNumber="29" start="Figure 11" startId="F11">
|
||
<paragraph pageId="0" pageNumber="29">
|
||
<emphasis bold="true" pageId="0" pageNumber="29">Figure 11.</emphasis>
|
||
Map of the Altai Mountain range displaying the distribution of
|
||
<taxonomicName authority="Ledoux & Roux, 1989" authorityName="Ledoux & Roux" authorityYear="1989" class="Insecta" family="Carabidae" genus="Pseudonebriola" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudonebriola" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="29">Pseudonebriola</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |