168 lines
16 KiB
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168 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.386.6913" ID-GBIF-Dataset="3a2df501-1763-4f11-a2ea-f9abaac99c00" ID-PMC="PMC3970064" ID-Pensoft-Pub="1313-2970-386-29" ID-PubMed="24693214" ID-ZBK="357FE9805295436EB40CFDD307D00D48" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-386-29" ModsDocOrigin="ZooKeys 386" ModsDocTitle="The Megophthalmidia (Diptera, Mycetophilidae) of North America including eight new species" checkinTime="1451246199718" checkinUser="pensoft" docAuthor="Kerr, Peter H." docDate="2014" docId="2FD3F491CAFE5B11BFE9BFA3A6DEC536" docLanguage="en" docName="ZooKeys 386: 29-83" docOrigin="ZooKeys 386" docSource="http://dx.doi.org/10.3897/zookeys.386.6913" docTitle="Megophthalmidia lenimenta Kerr, 2014, sp. n." docType="treatment" docUuid="65C644B7-D627-41F2-96F8-464E57B062E5" docUuidSource="ZooBank" docVersion="4" lastPageNumber="41" masterDocId="FFE4CC07FFFAC50AFFB0FFF4FFD6FFF4" masterDocTitle="The Megophthalmidia (Diptera, Mycetophilidae) of North America including eight new species" masterLastPageNumber="83" masterPageNumber="29" pageNumber="38" updateTime="1668157894205" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Megophthalmidia (Diptera, Mycetophilidae) of North America including eight new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kerr, Peter H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>386</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>29</mods:start>
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<mods:end>83</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.386.6913</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.386.6913</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-386-29</mods:identifier>
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<mods:identifier type="ZBK">357FE9805295436EB40CFDD307D00D48</mods:identifier>
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<mods:identifier type="ZooBank">357FE9805295436EB40CFDD307D00D48</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152051507" LSID="urn:lsid:zoobank.org:act:65C644B7-D627-41F2-96F8-464E57B062E5" httpUri="http://treatment.plazi.org/id/2FD3F491CAFE5B11BFE9BFA3A6DEC536" lastPageId="12" lastPageNumber="41" pageId="9" pageNumber="38">
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<subSubSection pageId="9" pageNumber="38" type="nomenclature">
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<paragraph pageId="9" pageNumber="38">
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<taxonomicName LSID="http://zoobank.org/65C644B7-D627-41F2-96F8-464E57B062E5" class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia lenimenta" order="Diptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="lenimenta">
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<pageBreakToken pageId="9" pageNumber="38" start="start">Megophthalmidia</pageBreakToken>
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lenimenta
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</taxonomicName>
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<taxonomicNameLabel pageId="9" pageNumber="38">sp. n.</taxonomicNameLabel>
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Figs 21-30
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="38" type="type material">
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<paragraph pageId="9" pageNumber="38">Type material.</paragraph>
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<paragraph pageId="9" pageNumber="38">
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Holotype: "USA: CA: Yolo Co., McLaughlin NR, Clover Valley, Oak grassland, MT#1,
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="38.84">38.8400°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-122.3451">122.3451°W</geoCoordinate>
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, 500masl, 24.
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<normalizedToken originalValue="iii–">iii-</normalizedToken>
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29.iv.2010 P. H. Kerr & C. E. Koehler, CSCA10L050" / "HOLOTYPE 13M302,
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia lenimenta" order="Diptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="lenimenta">Megophthalmidia lenimenta</taxonomicName>
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♂, Kerr, 2014" [red label]. Deposited in CSCA, mounted on gray point, complete specimen (Fig. 21). See Fig. 108 for image of type locality.
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</paragraph>
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<paragraph pageId="9" pageNumber="38">
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Paratypes (all bearing a blue paratype label): 3 ♂♂, 2♀♀, same locality as holotype [Locality Fig. 108; SBNM # 13M283 (♂); CSCA 13M317 (dissected ♂; Figs 22-30), 13M337 (♂),13M338 (♀), 13M339 (♀; Fig. 21)]; 3 ♂♂, ♀ "USA: CA: Yolo Co., McLaughlin NR, Clover Valley, Oak grassland, MT#1,
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="38.84">38.8400°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-122.3451">122.3451°W</geoCoordinate>
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, 500masl, 29.
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<normalizedToken originalValue="iv–">iv-</normalizedToken>
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9.vi.2010 P. H. Kerr & C.E. Koehler, 10L187" [LACM # 11H156 (♂); CSCA numbers 13M303 (♂, dissected), 13M340 (♂), and 13M342 (♀)].
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</paragraph>
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<paragraph pageId="9" pageNumber="38">
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Additional material examined: 2 ♂♂, "USA: CALIFORNIA, Stanislaus County, Del Puerto Canyon, Frank Raines Park / ca 1120', 3
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<normalizedToken originalValue="–IV–">-IV-</normalizedToken>
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70, Paul H. Arnaud, Jr., Collector" [CAS; one specimen dissected, #13M588].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="38" type="diagnosis">
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<paragraph pageId="9" pageNumber="38">Diagnosis.</paragraph>
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<paragraph pageId="9" pageNumber="38">
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia lenimenta" order="Diptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="lenimenta">Megophthalmidia lenimenta</taxonomicName>
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sp. n. may be confused with several Nearctic congeners that also have a brown thorax. Among these, it is probably most similar to
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia browni" order="Diptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="browni">Megophthalmidia browni</taxonomicName>
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sp. n. on account of both species having epandria with a medial notch along the posterior margin, a medial depression, and elongated posterior processes. In
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia lenimenta" order="Diptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="lenimenta">Megophthalmidia lenimenta</taxonomicName>
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, one aedeagal tine is very reduced/undeveloped, only slightly longer than wide (Fig. 29; whereas in
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia browni" order="Diptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="browni">Megophthalmidia browni</taxonomicName>
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both tines are many times longer than wide (Fig. 9)). Among other Nearctic congeners with elongate posterior epandrial processes, including
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia browni" order="Diptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="browni">Megophthalmidia browni</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia lenimenta" order="Diptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="lenimenta">Megophthalmidia lenimenta</taxonomicName>
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is also distinguished by having length of dorsomedial epandrial surface at least half the length of epandrium (Fig. 24).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="40" pageId="9" pageNumber="38" type="description">
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<paragraph pageId="9" pageNumber="38">Description.</paragraph>
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<paragraph pageId="9" pageNumber="38">Male. Body length: 2.4-3.1, 2.8 [n/a] mm (n=5). Wing length: 2.6-2.9, 2.8 [2.9] mm (n=7).</paragraph>
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<paragraph pageId="9" pageNumber="38">Coloration (Fig. 21). Male. Head dark brown; antennal scape brown or dark brown, pedicel and flagellomeres brown; face dark brown, clypeus and labrum brown to dark brown; palps and labellum cream-colored to pale yellow (palpomeres 1-3 usually slightly darker than others, palpomere 2 with light patch where sensilla present). Thorax brown to dark brown throughout; scutum setae brown. Coxae lighter in color than thorax, cream-colored to light brown, sometimes with area at base somewhat darker in color, fore coxa slightly lighter in color than mid- and hind coxa; femora cream colored to light brown, becoming gradually darker dorsoapically on mid- and hind femora; tibiae light brown, tarsi light brown to brown; hind tibial comb yellowish, preceded by 0-3 (usually 3) dark brown setae. Wing hyaline without markings, wing veins brown; haltere stem and knob white to cream-colored. Abdominal segments concolorous brown to dark brown. Terminalia brown.</paragraph>
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<caption pageId="9" pageNumber="38">
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<paragraph pageId="9" pageNumber="38">
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Figure 21.
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia lenimenta" order="Diptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="lenimenta">Megophthalmidia lenimenta</taxonomicName>
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sp. n., habitus [holotype male above, # 13M302; female below, # 13M339]. Scale bar = 1 mm.
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</paragraph>
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</caption>
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<paragraph pageId="10" pageNumber="39">
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<pageBreakToken pageId="10" pageNumber="39" start="start">Head</pageBreakToken>
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. Ocelli slightly raised, median ocellus in line with anterior margin of lateral ocelli, median ocellus approx. 0.3
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<normalizedToken originalValue="–0.5×">-0.5x</normalizedToken>
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size of lateral ocelli; lateral ocellus located approx. 2
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<normalizedToken originalValue="×">x</normalizedToken>
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diameter of ocellus from eye margin, separated from median ocellus by approx. 1.9
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<normalizedToken originalValue="–2×">-2x</normalizedToken>
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its own diameter. Eyes with microsetae, which are approximately as long as width of facet. Frons microtrichose, without setae, flattened. Antennal length 1.3-1.7, 1.6 [1.5] mm (n=7). Face clearly longer than wide, setose; clypeus and labrum microtrichose, without setae. Palpus with four palpomeres; palpomere 1 oblong-triangular, without setae; other palpomeres with brown setae; palpomere 2 bearing small pocket of sensilla; palpomere 1 length longer than or subequal in length to palpomere 2; palpomere 3 length subequal to or slightly shorter than combined length of palpomeres 1 and 2; palpomere 4 length 0.7
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<normalizedToken originalValue="–1×">-1x</normalizedToken>
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combined lengths of palpomeres 1-3.
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</paragraph>
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<paragraph pageId="10" pageNumber="39">
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Thorax. Antepronotum, proepisternum, and laterotergite bearing setae; remaining lateral thoracic sclerites bare. Dorsum with evenly-distributed, short, appressed setae, bearing longer setae only along lateral and posterior margins. Costal wing vein extends beyond R5, approx. two-thirds distance between R5 and M1; R1 longer than r-m; cubital fork proximad of r-m base (as in
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia occidentalis" order="Diptera" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="occidentalis">Megophthalmidia occidentalis</taxonomicName>
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, Fig. 52); R1, M1, M2, CuA1, and CuA2 with setae on upper surface (lacking setae on M1 + M2). Wing veins A1 and CuP absent.
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</paragraph>
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<paragraph pageId="11" pageNumber="40">
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<pageBreakToken pageId="11" pageNumber="40" start="start">Male</pageBreakToken>
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genitalia (Figs 22-30). Epandrium dorsal surface with clear medial depression, where setae are lacking; posterior margin narrowly emarginate at center (Fig. 24). Posterior processes of epandrium elongate, approx. 4
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<normalizedToken originalValue="–5×">-5x</normalizedToken>
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longer than narrowest width near base, separated at base by approx. 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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narrowest width of process, length of setae at base of epandrial processes 2
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<normalizedToken originalValue="–3×">-3x</normalizedToken>
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width of process, bare along most of length (Figs 22, 23). Gonocoxites as in Figs 25-27. Adeagal fork uneven; one tine a mere nub, the other, elongate, gently s-curved, and pointed outward (Fig. 29).
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</paragraph>
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<caption pageId="11" pageNumber="40">
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<paragraph pageId="11" pageNumber="40">
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Figures 22-24.
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia lenimenta" order="Diptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="lenimenta">Megophthalmidia lenimenta</taxonomicName>
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sp. n., male epandrium [paratype, # 13M317] 22 Lateral view 23 Posterior view 24 Dorsal view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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<caption pageId="11" pageNumber="40">
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<paragraph pageId="11" pageNumber="40">
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Figures 25-27.
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia lenimenta" order="Diptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="lenimenta">Megophthalmidia lenimenta</taxonomicName>
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sp. n., male hypandrium [paratype, # 13M317] 25 Lateral view 26 Dorsal view 27 Ventral view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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<caption pageId="11" pageNumber="40">
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<paragraph pageId="11" pageNumber="40">
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Figures 28-30.
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Megophthalmidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophthalmidia lenimenta" order="Diptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="lenimenta">Megophthalmidia lenimenta</taxonomicName>
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sp. n., male aedeagus [paratype, # 13M317] 28 Lateral view 29 Dorsal view 30 Posterior view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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<paragraph pageId="11" pageNumber="40">Female. Body length: 2.6-3.1, 2.9 mm (n=3). Antennal length 1.0-1.1, 1.1 mm (n=3). Wing length: 2.6-3.2, 3.0 mm (n=3).</paragraph>
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<paragraph pageId="11" pageNumber="40">Coloration (Fig. 21). Noticeably darker in color throughout body; abdominal segments 8-10 orange-brown to brown, brown along margins; cerci light brown to brown.</paragraph>
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<paragraph pageId="11" pageNumber="40">Head and thorax. Same as male, except palpomere 4 appx. length of palpomeres 1-3.</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="41" type="etymology">
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<paragraph pageId="12" pageNumber="41">
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<pageBreakToken pageId="12" pageNumber="41" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="12" pageNumber="41">
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The species epithet
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<normalizedToken originalValue="“lenimenta”">"lenimenta"</normalizedToken>
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is an adjective, derived from the Latin word for remedy/melioration/reclamation. This name is given in thanks to the preservation efforts of Sylvia Mclaughlin and the University of California Donald and Sylvia Mclaughlin Reserve staff, including Cathy Koehler, and the Homestake Mining Company.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |