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<document id="48651E2E4B9E5D81EAD2685DA4FA8CAA" ID-GBIF-Dataset="a60d5d32-bb18-492f-8385-b14df30e93b2" checkinTime="1447190589017" checkinUser="donat" docAuthor="Dlussky, G. M., Rasnitsyn, A. P. &amp; Perfilieva, K. S." docDate="2015" docId="0B04695EFFF36644FC6CFB3BFF36FE5F" docLanguage="en" docName="29875.pdf.imf" docOrigin="Caucasian Entomological Bulletin 11 (1)" docTitle="Gesomyrmex Mayr 1868" docType="treatment" docVersion="11" lastPageNumber="138" masterDocId="F73D1126FFF56643FFC7FFCDFFCDFFBB" masterDocTitle="The Ants (Hymenoptera: Formicidae) of Bolshaya Svetlovodnaya (Late Eocene of Sikhote-Alin, Russian Far East)" masterLastPageNumber="152" masterPageNumber="131" pageNumber="137" updateTime="1694997511864" updateUser="admin">
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<mods:title id="076D53B05E19E5259BB1339FE1844EF3">The Ants (Hymenoptera: Formicidae) of Bolshaya Svetlovodnaya (Late Eocene of Sikhote-Alin, Russian Far East)</mods:title>
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<mods:namePart id="F9B60AC3C24CBE0E633CD7C0662D6953">Dlussky, G. M.</mods:namePart>
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<treatment id="0B04695EFFF36644FC6CFB3BFF36FE5F" ID-DOI="http://doi.org/10.5281/zenodo.6105367" ID-GBIF-Taxon="114926168" ID-Zenodo-Dep="6105367" LSID="urn:lsid:plazi:treatment:0B04695EFFF36644FC6CFB3BFF36FE5F" httpUri="http://treatment.plazi.org/id/0B04695EFFF36644FC6CFB3BFF36FE5F" lastPageId="7" lastPageNumber="138" pageId="6" pageNumber="137">
<subSubSection id="CBB78BC3FFF36645FC6CFB3BFC55FAD9" pageId="6" pageNumber="137" type="nomenclature">
<paragraph id="8312D848FFF36645FC6CFB3BFB35FAB0" box="[939,1272,1270,1291]" pageId="6" pageNumber="137">
<heading id="D85A6F24FFF36645FC6CFB3BFB35FAB0" bold="true" box="[939,1272,1270,1291]" centered="true" fontSize="9" level="3" pageId="6" pageNumber="137" reason="0">
<emphasis id="B1D9045AFFF36645FC6CFB3BFB35FAB0" bold="true" box="[939,1272,1270,1291]" pageId="6" pageNumber="137">
Genus
<taxonomicName id="44ADA3CBFFF36645FC33FB3BFB35FAB0" ID-CoL="4N9D" ID-ENA="46648" authority="Mayr, 1868" authorityName="Mayr" authorityYear="1868" box="[1012,1272,1270,1291]" class="Insecta" family="Formicidae" genus="Gesomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="B1D9045AFFF36645FC33FB3BFBB0FAB0" bold="true" box="[1012,1149,1270,1291]" italics="true" pageId="6" pageNumber="137">Gesomyrmex</emphasis>
<bibRefCitation id="CB300153D9FDE468A3A2827DAF86960E" author="Mayr G." journalOrPublisher="Beitrage zur Naturkunde Preussens" pagination="1 - 102" part="1" refId="ref21040" refString="Mayr G. 1868. Die Ameisen des baltischen Bernsteins. Beiträge zur Naturkunde Preussens. 1: 1 102." title="Die Ameisen des baltischen Bernsteins" type="journal article" year="1868">Mayr, 1868</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8312D848FFF36645FC9BFAFDFC55FAD9" pageId="6" pageNumber="137">
Type species
<taxonomicName id="44ADA3CBFFF36645FC2AFAE2FA82FAFE" authority="Mayr, 1868" authorityName="Mayr" authorityYear="1868" box="[1005,1359,1327,1349]" class="Insecta" family="Formicidae" genus="Gesomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="hoernesi">
<emphasis id="B1D9045AFFF36645FC2AFAE2FB00FAFE" bold="true" box="[1005,1229,1327,1349]" italics="true" pageId="6" pageNumber="137">Gesomyrmex hoernesi</emphasis>
<bibRefCitation id="00FDB6B714213807008F09CB84BA6D49" author="Mayr G." journalOrPublisher="Beitrage zur Naturkunde Preussens" pagination="1 - 102" part="1" refId="ref21040" refString="Mayr G. 1868. Die Ameisen des baltischen Bernsteins. Beiträge zur Naturkunde Preussens. 1: 1 102." title="Die Ameisen des baltischen Bernsteins" type="journal article" year="1868">Mayr, 1868</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="CBB78BC3FFF36644FC9BFAA6FEA8FE2F" lastPageId="7" lastPageNumber="138" pageId="6" pageNumber="137" type="diagnosis">
<paragraph id="8312D848FFF36645FC9BFAA6FC63F979" pageId="6" pageNumber="137">
<emphasis id="B1D9045AFFF36645FC9BFAA6FC0BFA3B" bold="true" box="[860,966,1387,1408]" pageId="6" pageNumber="137">Diagnosis</emphasis>
(for fossil imprints). Gyne. Antennae geniculate, 10-segmented, with very short scape. Clypeus extends back between antennal sockets. Antennal insertions widely separated, situated close to the posterior clypeal margin. Frontal lobes close to antennal insertions. Mandibles massive, triangular, with 410 teeth. Palp formula 6, 4. Posterolateral corners of the head and propodeum without spines. Petiole with thick scale. Fore wings with closed cells 1+2r, 3r and mcu. Veins RS and M after 2r-rs usually partly united and form short section RS+M. Posterior margin of cell 3r (formed by RS) convex or S-formed.
</paragraph>
<paragraph id="8312D848FFF36644FC9BF906FEA8FE2F" lastPageId="7" lastPageNumber="138" pageId="6" pageNumber="137">
<emphasis id="B1D9045AFFF36645FC9BF906FBD2F95B" bold="true" box="[860,1055,1739,1760]" pageId="6" pageNumber="137">Species included.</emphasis>
Six extant species are known which are distributed in the Oriental tropics [
<bibRefCitation id="BE2A788B0A786E9B3017A9DFD755B8DB" author="Mayr G." journalOrPublisher="Beitrage zur Naturkunde Preussens" pagination="1 - 102" part="1" refId="ref21040" refString="Mayr G. 1868. Die Ameisen des baltischen Bernsteins. Beiträge zur Naturkunde Preussens. 1: 1 102." title="Die Ameisen des baltischen Bernsteins" type="journal article" year="1868">Mayr, 1868</bibRefCitation>
]. Six fossil species are known:
<taxonomicName id="44ADA3CBFFF36645FB9FF8C8FC90F88C" authority="Mayr, 1868" authorityName="Mayr" authorityYear="1868" class="Insecta" family="Formicidae" genus="Gesomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="hoernesi">
<emphasis id="B1D9045AFFF36645FB9FF8C8FAF8F8A1" bold="true" box="[1112,1333,1797,1818]" italics="true" pageId="6" pageNumber="137">Gesomyrmex hoernesi</emphasis>
<bibRefCitation id="49DFE5B9C3877A4FA04DFD3ECC40756C" author="Mayr G." journalOrPublisher="Beitrage zur Naturkunde Preussens" pagination="1 - 102" part="1" refId="ref21040" refString="Mayr G. 1868. Die Ameisen des baltischen Bernsteins. Beiträge zur Naturkunde Preussens. 1: 1 102." title="Die Ameisen des baltischen Bernsteins" type="journal article" year="1868">Mayr, 1868</bibRefCitation>
</taxonomicName>
(Baltic amber, Late Eocene),
<taxonomicName id="44ADA3CBFFF36645FB4CF8EFFC4FF8EE" authority="Dlussky, Wappler" authorityName="Dlussky, Wappler" class="Insecta" family="Formicidae" genus="Gesomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="germanicus">
<emphasis id="B1D9045AFFF36645FB4CF8EFFAD6F88C" bold="true" box="[1163,1307,1826,1847]" italics="true" pageId="6" pageNumber="137">G.germanicus</emphasis>
<bibRefCitation id="D78D8FB766BAABC3BA4C381734AD080A" author="Dlussky G. M. &amp; Wappler T. &amp; Wedmann S." journalOrPublisher="Zootaxa" pagination="1 - 20" part="2031" refId="ref20353" refString="Dlussky G. M., Wappler T., Wedmann S. 2009. Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities. Zootaxa. 2031: 1 20." title="Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities" type="journal article" year="2009">Dlussky, Wappler et Wedmann, 2009</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="44ADA3CBFFF36645FB42F8F2FC4FF8C9" authority="Dlussky, Wappler" authorityName="Dlussky, Wappler" class="Insecta" family="Formicidae" genus="Gesomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="flavescens">
<emphasis id="B1D9045AFFF36645FB42F8F2FADDF8EE" bold="true" box="[1157,1296,1855,1877]" italics="true" pageId="6" pageNumber="137">G. flavescens</emphasis>
<bibRefCitation id="F55DFAE78263FF78DB56B3DBA85DCF9A" author="Dlussky G. M. &amp; Wappler T. &amp; Wedmann S." journalOrPublisher="Zootaxa" pagination="1 - 20" part="2031" refId="ref20353" refString="Dlussky G. M., Wappler T., Wedmann S. 2009. Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities. Zootaxa. 2031: 1 20." title="Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities" type="journal article" year="2009">Dlussky, Wappler et Wedmann, 2009</bibRefCitation>
</taxonomicName>
(Eckfeld, Germany, Middle Eocene),
<taxonomicName id="44ADA3CBFFF36645FC53F8B7FB1FF834" authority="Dlussky, Wappler" authorityName="Dlussky, Wappler" box="[916,1234,1914,1935]" class="Insecta" family="Formicidae" genus="Gesomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="B1D9045AFFF36645FC53F8B7FBCBF834" bold="true" box="[916,1030,1914,1935]" italics="true" pageId="6" pageNumber="137">G. pulcher</emphasis>
<bibRefCitation id="0313E9FA78C404CF68283D8F7759622C" author="Dlussky G. M. &amp; Wappler T. &amp; Wedmann S." journalOrPublisher="Zootaxa" pagination="1 - 20" part="2031" refId="ref20353" refString="Dlussky G. M., Wappler T., Wedmann S. 2009. Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities. Zootaxa. 2031: 1 20." title="Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities" type="journal article" year="2009">Dlussky, Wappler et Wedmann, 2009</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="44ADA3CBFFF36645FCABF85AFB6EF816" authority="Dlussky, Wappler" authorityName="Dlussky, Wappler" box="[876,1187,1943,1965]" class="Insecta" family="Formicidae" genus="Gesomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="breviceps">
<emphasis id="B1D9045AFFF36645FCABF85AFC2AF816" bold="true" box="[876,999,1943,1965]" italics="true" pageId="6" pageNumber="137">G. breviceps</emphasis>
<bibRefCitation id="36992278B809334F8CD2515A981BAA38" author="Dlussky G. M. &amp; Wappler T. &amp; Wedmann S." journalOrPublisher="Zootaxa" pagination="1 - 20" part="2031" refId="ref20353" refString="Dlussky G. M., Wappler T., Wedmann S. 2009. Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities. Zootaxa. 2031: 1 20." title="Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities" type="journal article" year="2009">Dlussky, Wappler et Wedmann, 2009</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="44ADA3CBFFF36645FCEAF878FB94F871" authority="Dlussky, Wappler" authorityName="Dlussky, Wappler" box="[813,1113,1972,1994]" class="Insecta" family="Formicidae" genus="Gesomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="137" phylum="Arthropoda" rank="species" species="curiosus">
<emphasis id="B1D9045AFFF36645FCEAF878FC53F871" bold="true" box="[813,926,1973,1994]" italics="true" pageId="6" pageNumber="137">G. curiosus</emphasis>
<bibRefCitation id="F6526A5E0555E4E5B0C2FA3559093C3E" author="Dlussky G. M. &amp; Wappler T. &amp; Wedmann S." journalOrPublisher="Zootaxa" pagination="1 - 20" part="2031" refId="ref20353" refString="Dlussky G. M., Wappler T., Wedmann S. 2009. Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities. Zootaxa. 2031: 1 20." title="Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities" type="journal article" year="2009">Dlussky, Wappler et Wedmann, 2009</bibRefCitation>
</taxonomicName>
(Grube Messel, Germany, Middle Eocene) [
<bibRefCitation id="E819C3F239E43CCEA406F601DFD169F6" author="Mayr G." journalOrPublisher="Beitrage zur Naturkunde Preussens" pagination="1 - 102" part="1" refId="ref21040" refString="Mayr G. 1868. Die Ameisen des baltischen Bernsteins. Beiträge zur Naturkunde Preussens. 1: 1 102." title="Die Ameisen des baltischen Bernsteins" type="journal article" year="1868">Mayr, 1868</bibRefCitation>
;
<bibRefCitation id="BFF1E07D535E84DB0FB35D503FAAE080" author="Dlussky G. M." journalOrPublisher="Paleontological Journal" pagination="1043 - 1086" part="43" refId="ref20031" refString="Dlussky G. M. 2009. The ant subfamilies Ponerinae, Cerapachyinae, and Pseudomyrmecinae (Hymenoptera, Formicidae) in the Late Eocene ambers of Europe. Paleontological Journal. 43: 1043 1086." title="The ant subfamilies Ponerinae, Cerapachyinae, and Pseudomyrmecinae (Hymenoptera, Formicidae) in the Late Eocene ambers of Europe" type="journal article" year="2009">Dlussky et al., 2009</bibRefCitation>
].
<taxonomicName id="44ADA3CBFFF26644FE9AFED9FCCAFE92" authority="Théobald, 1937" authorityName="Theobald" authorityYear="1937" box="[349,775,272,297]" class="Insecta" family="Formicidae" genus="Gesomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="expectans">
<emphasis id="B1D9045AFFF26644FE9AFED9FD9FFE92" bold="true" box="[349,594,276,297]" italics="true" pageId="7" pageNumber="138">Gesomyrmex expectans</emphasis>
Théobald, 1937
</taxonomicName>
(Kleinkembs, France, Oligocene) and
<taxonomicName id="44ADA3CBFFF26644FD84FEE2FF20FEE4" authority="Théobald, 1937" authorityName="Theobald" authorityYear="1937" class="Insecta" family="Formicidae" genus="Gesomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="miegi">
<emphasis id="B1D9045AFFF26644FD84FEE2FD57FEFF" bold="true" box="[579,666,303,324]" italics="true" pageId="7" pageNumber="138">G. miegi</emphasis>
Théobald, 1937
</taxonomicName>
(Haut-Rhin, France, Oligocene) excluded from
<taxonomicName id="44ADA3CBFFF26644FF7AFEA9FEF2FEC2" authorityName="Mayr" authorityYear="1868" box="[189,319,356,377]" class="Insecta" family="Formicidae" genus="Gesomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="B1D9045AFFF26644FF7AFEA9FEF2FEC2" bold="true" box="[189,319,356,377]" italics="true" pageId="7" pageNumber="138">Gesomyrmex</emphasis>
</taxonomicName>
[
<bibRefCitation id="A941FA9539316F94958F4A9401D1D686" author="Dlussky G. M." journalOrPublisher="Paleontological Journal" pagination="1043 - 1086" part="43" refId="ref20031" refString="Dlussky G. M. 2009. The ant subfamilies Ponerinae, Cerapachyinae, and Pseudomyrmecinae (Hymenoptera, Formicidae) in the Late Eocene ambers of Europe. Paleontological Journal. 43: 1043 1086." title="The ant subfamilies Ponerinae, Cerapachyinae, and Pseudomyrmecinae (Hymenoptera, Formicidae) in the Late Eocene ambers of Europe" type="journal article" year="2009">Dlussky et al., 2009</bibRefCitation>
]. Three new species are described below.
</paragraph>
</subSubSection>
<subSubSection id="CBB78BC3FFF26644FF2AFE57FF36FE5F" pageId="7" pageNumber="138" type="discussion">
<paragraph id="8312D848FFF26644FF2AFE57FF36FE5F" pageId="7" pageNumber="138">
<emphasis id="B1D9045AFFF26644FF2AFE57FEFCFE14" bold="true" box="[237,305,410,431]" pageId="7" pageNumber="138">Notes.</emphasis>
Living species of
<taxonomicName id="44ADA3CBFFF26644FE3CFE54FDB0FE14" authorityName="Mayr" authorityYear="1868" box="[507,637,409,431]" class="Insecta" family="Formicidae" genus="Gesomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="B1D9045AFFF26644FE3CFE54FDB0FE14" bold="true" box="[507,637,409,431]" italics="true" pageId="7" pageNumber="138">Gesomyrmex</emphasis>
</taxonomicName>
are arboreal ants constructing nests in tree twigs [
<bibRefCitation id="98E9DD16D1BF4793AC4EEC66151F4640" author="Wheeler W. M." box="[189,746,404,421]" journalOrPublisher="Psyche" pagination="35 - 40" part="37" refId="ref21983" refString="Wheeler W. M. 1930. A second note on Gesomyrmex. Psyche. 37: 35 40." title="A second note on Gesomyrmex" type="journal article" year="1930">Wheeler, 1930</bibRefCitation>
;
<bibRefCitation id="AF9D0096A014EBF55276CBCEF5730BB0" author="Cole A. C., Jr." journalOrPublisher="Annals of the Entomological Society of America" pagination="71 - 76" part="42" refId="ref19570" refString="Cole A. C., Jr. 1949. A study of the genus Gesomyrmex Mayr, and a description of a species new to the genus (Hymenoptera: Formicidae). Annals of the Entomological Society of America. 42: 71 76." title="A study of the genus Gesomyrmex Mayr, and a description of a species new to the genus (Hymenoptera: Formicidae)" type="journal article" year="1949">Cole, 1949</bibRefCitation>
].
</paragraph>
</subSubSection>
</treatment>
</document>