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<document ID-CLB-Dataset="279481" ID-DOI="http://dx.doi.org/10.3897/zookeys.1185.112099" ID-GBIF-Dataset="2f7bf0ee-f894-4872-a09f-6bfd7985e2fa" ID-PMC="PMC10696598" ID-Pensoft-Pub="1313-2970-1185-1" ID-Pensoft-UUID="99994BD90C195466A249943D2E939FB4" ID-PubMed="38058424" ID-ZooBank="5B12796B210249948D3353FAD1A1B763" ModsDocID="1313-2970-1185-1" checkinTime="1701152605290" checkinUser="pensoft" docAuthor="Lee, Chi-Feng" docDate="2023" docId="F967FCF857665534A1D0DA5F559F0B5D" docLanguage="en" docName="ZooKeys 1185: 1-19" docOrigin="ZooKeys 1185" docPubDate="2023-11-27" docSource="http://dx.doi.org/10.3897/zookeys.1185.112099" docTitle="Sphaeroderma changi Lee 2023, sp. nov." docType="treatment" docUuid="880B2056-1C8E-4C62-AFBB-434F07506557" docUuidSource="ZooBank" docVersion="4" id="99994BD90C195466A249943D2E939FB4" lastPageNumber="1" masterDocId="99994BD90C195466A249943D2E939FB4" masterDocTitle="Four new species of Sphaeroderma Stephens (Coleoptera, Chrysomelidae, Galerucinae, Alticini) from Taiwan, with discussion on genus boundaries based on S. flavonotatum Chujo and S. jungchani sp. nov." masterLastPageNumber="19" masterPageNumber="1" pageNumber="1" updateTime="1732692427602" updateUser="ExternalLinkService">
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<mods:title id="6FB370B889B231681E7AF1811877A221">Four new species of Sphaeroderma Stephens (Coleoptera, Chrysomelidae, Galerucinae, Alticini) from Taiwan, with discussion on genus boundaries based on S. flavonotatum Chujo and S. jungchani sp. nov.</mods:title>
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<mods:name id="EEC0DFDA97D0C022447D3BB10EF39309" type="personal">
<mods:role id="1778C2763B5FBAB2FF736714A4246022">
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<mods:namePart id="4B7F731892C935B61EF5844B76A82732">Lee, Chi-Feng</mods:namePart>
<mods:nameIdentifier id="AADFACC2C1459F4ECD33F3FB5D9C717E" type="ORCID">https://orcid.org/0000-0003-1996-0557</mods:nameIdentifier>
<mods:affiliation id="8B85F0B0B69A6E19A8F6641C15277E64">Applied Zoology Division, Taiwan Agricultural Research Institute, Taichung 413, Taiwan</mods:affiliation>
<mods:nameIdentifier id="A7E5F148FEF96B60D3712C6FFC459008" type="email">chifeng@tari.gov.tw</mods:nameIdentifier>
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<mods:typeOfResource id="986CFCAF1836C56E71354F6C325C2DE9">text</mods:typeOfResource>
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<mods:title id="3AE8022F2F0CBBA9B7EDD3EB2B8813E4">ZooKeys</mods:title>
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<mods:part id="D8E7F28E28DCBF3C1CD7A07C5C016F61">
<mods:date id="B325457BB41A4743BFB2016B1C2A0930">2023</mods:date>
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<mods:number id="3E73D38516A920B57AB046941F608751">2023-11-27</mods:number>
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<mods:number id="C4E4967C4190455FFC5A23166973EF3C">1185</mods:number>
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<mods:classification id="897BEE4AF58FA6B02FCADA04D921628F">journal article</mods:classification>
<mods:identifier id="EAEF488E243D42A12665319322DE54B2" type="DOI">http://dx.doi.org/10.3897/zookeys.1185.112099</mods:identifier>
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<treatment id="F967FCF857665534A1D0DA5F559F0B5D" ID-GBIF-Taxon="219268494" LSID="urn:lsid:zoobank.org:act:880B2056-1C8E-4C62-AFBB-434F07506557" httpUri="http://treatment.plazi.org/id/F967FCF857665534A1D0DA5F559F0B5D" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Chrysomelidae" scope_order="Coleoptera" scope_subFamily="Galerucinae" scope_tribe="Alticini">
<subSubSection id="0FE2DCBF8131C3C7FD036363F269A23B" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="BCFF2BB6686826CA06B4647C05A87844" pageId="0" pageNumber="1">
<taxonomicName id="7233A8EBE325B92AD15159E5ADD38583" LSID="https://zoobank.org/880B2056-1C8E-4C62-AFBB-434F07506557" authority="Lee, 2023" authorityName="Lee" authorityYear="2023" class="Insecta" family="Chrysomelidae" genus="Sphaeroderma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeroderma changi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="changi" status="sp. nov.">Sphaeroderma changi</taxonomicName>
<taxonomicNameLabel id="31C180247DEF99DD3247F08941BF3F77" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="5B95775F88E10682DCA57E00B0540A38" pageId="0" pageNumber="1" type="description">
<paragraph id="F3D711C9CA8ADCB25880BD02F9532AE4" pageId="0" pageNumber="1">
<figureCitation id="CB2CA7D6C9AFCAC72F78E9982F19D01A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Sphaeroderma species in frontal view A S. hsui sp. nov. B S. changi sp. nov. C S. sheipaensis sp. nov. D S. flavonotatum Chujo E S. jungchani sp. nov." figureDoi="10.3897/zookeys.1185.112099.figure2" httpUri="https://binary.pensoft.net/fig/942952" pageId="0" pageNumber="1">Figs 2B</figureCitation>
<figureCitation id="13A4228107807E1531DADDAAFE02ABDF" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus of Sphaeroderma changi sp. nov. and S. sheipaensis sp. nov. A S. changi sp. nov., female, dorsal view B ditto, ventral view C ditto, lateral view D S. sheipaensis sp. nov., female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1185.112099.figure5" httpUri="https://binary.pensoft.net/fig/942955" pageId="0" pageNumber="1">, 5A-C</figureCitation>
<figureCitation id="C0430D9BFD1157829EB2CBEF9CB146BF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Sphaeroderma changi sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1">, 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="ABE06BCBBE0F278172FB42F09518CC82" pageId="0" pageNumber="1" type="types">
<paragraph id="77DD3BC0E5B91760BDAE9606358A86E9" pageId="0" pageNumber="1">Types.</paragraph>
<paragraph id="621CC58E0EDBE1374C3D705E8EC11B3C" pageId="0" pageNumber="1">
<emphasis id="8A6F836D6833253373DBC7BD51993AFB" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
♂ (TARI). Taiwan: Miaoli, Hsishihshan trail (西勢山林道), 26.IX.2021, leg. Y.-F. Hsu.
<emphasis id="1BFCC95CA234E2FBEC49BBCB38C4C9CE" bold="true" pageId="0" pageNumber="1">
<emphasis id="CF16E3E048E13C18DF6EAE528F66DF71" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
.
</emphasis>
1♀ (TARI), same but with
<normalizedToken id="B5245182BCBB55A99F7600AF60B3B666" originalValue="“28.VIII.2021”">&quot;28.VIII.2021&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="CAD528D05FD1877DBDA477B174241064" pageId="0" pageNumber="1" type="description">
<paragraph id="C228E22115F7C6E273C63211C8DA8C5F" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="102E85D288A476B14D479B2C227030BE" pageId="0" pageNumber="1">
Length 3.2 mm, width 2.2 mm. Body color (Fig.
<figureCitation id="D449DEF6589AC6601676BD93FFEE183B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus of Sphaeroderma changi sp. nov. and S. sheipaensis sp. nov. A S. changi sp. nov., female, dorsal view B ditto, ventral view C ditto, lateral view D S. sheipaensis sp. nov., female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1185.112099.figure5" httpUri="https://binary.pensoft.net/fig/942955" pageId="0" pageNumber="1">5A-C</figureCitation>
) yellowish or reddish-brown; head, pronotum, and legs blackish-brown; antennae blackish-brown except four basal antennomeres yellowish-brown. Antennae filiform in males (Fig.
<figureCitation id="2F8667B269EEE6285A635830849D06D7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Sphaeroderma changi sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1">6A</figureCitation>
), length ratios of antennomeres I-XI 1.0:0.7:0.4:0.5:0.7:0.7:0.8:0.8:0.8:0.8:1.2, length to width ratios of antennomeres I-XI 2.8:2.2:1.9:2.0:2.7:2.5:2.5:2.3:2.3:2.4:3.5; similar in females (Fig.
<figureCitation id="D738C3ED9710FB4B13DBDF085A42F330" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Sphaeroderma changi sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1">6B</figureCitation>
), length ratios of antennomeres I-XI 1.0:0.6:0.5:0.5:0.7:0.6:0.7:0.7:0.7:0.7:0.9, length to width ratios of antennomeres I-XI 2.9:2.2:2.1:2.1:2.7:2.4:2.7:2.6:2.5:2.4:3.2. Antennal calli transverse, with supracallinal sulci well developed. Pronotum 1.7
<normalizedToken id="06728A2713434FFD0094DFC849D8523C" originalValue="×">x</normalizedToken>
wider than long, disc with fine, scattered punctures the same size as those on elytra; lateral margins almost straight; anterolateral callosity protruding forward; posterolateral callosity poorly developed. Elytra 1.1-1.2 times longer than wide, sides widely rounded; disc with punctures entirely confused; humeral calli well developed. Abdominal ventrite V without internal median ridge in both sexes. Male genitalia: aedeagus (Fig.
<figureCitation id="11E6B4CB02A58E41DE379C6837B914CA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Sphaeroderma changi sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1">6C, D</figureCitation>
) wide in dorsal view, 3.1
<normalizedToken id="6490FC9E0E60BE4B83AF23A33282B9FA" originalValue="×">x</normalizedToken>
longer than wide; parallel-sided, apex narrowly rounded, strongly widened at apical 1/8; moderately curved at middle in lateral view; ventral surface with one pair of oblique ridges from apical 1/5 of lateral margins; ostium membranous and with Y-shaped sclerotized area. Female genitalia: ventrite VIII (Fig.
<figureCitation id="FCD80D4BBF2B9072CFF66DF1077E8D11" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Sphaeroderma changi sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1">6E</figureCitation>
) with apical part triangular, but sides slightly narrowed near apex, apical margin smooth and with seven pairs of setae medially, disc with medial part membranous, spiculum short; gonocoxae (Fig.
<figureCitation id="A9EE288BB67B6956AE60D533D0F1F70F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Sphaeroderma changi sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1">6G</figureCitation>
) separated, longitudinally triangular, with dense, long setae along apical margins, with one slender and transverse basal sclerite; receptacle of spermatheca (Fig.
<figureCitation id="3E8C229616A9D53116A397C95654E814" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Sphaeroderma changi sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1">6F</figureCitation>
) moderately swollen; pump short and strongly curved, transverse wrinkles present on entire pump and extending onto half of receptacle; sclerotized proximal spermathecal duct long, with ramus oblong.
</paragraph>
</subSubSection>
<subSubSection id="BDBE9CC00CD8C1B88E36F72DD0D5B1E7" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="90EA26E74201BF4B29649CAC53348525" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="33AD7F2B60C30FA59C4AB23822C168E9" pageId="0" pageNumber="1">
Adults of
<taxonomicName id="3ED40C51A6CA680F48E8FB089D8E51AE" authorityName="Lee" authorityYear="2023" class="Insecta" family="Chrysomelidae" genus="Sphaeroderma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeroderma changi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="changi">
<emphasis id="A0B9B9F6E6B2D52517C24500C5C15AEB" italics="true" pageId="0" pageNumber="1">Sphaeroderma changi</emphasis>
</taxonomicName>
sp. nov. have a characteristic and diagnostic color pattern (Fig.
<figureCitation id="9C209E2BACFE11068FE7A43941C3B432" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus of Sphaeroderma changi sp. nov. and S. sheipaensis sp. nov. A S. changi sp. nov., female, dorsal view B ditto, ventral view C ditto, lateral view D S. sheipaensis sp. nov., female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1185.112099.figure5" httpUri="https://binary.pensoft.net/fig/942955" pageId="0" pageNumber="1">5A-C</figureCitation>
). In addition, this new species differs from the other two Taiwanese species (
<taxonomicName id="25F7C65177845CA532584AA8FD07497C" lsidName="S. hsui" pageId="0" pageNumber="1" rank="species" species="hsui">
<emphasis id="B31DF08F7FE172CE067178DBF6CFFF1D" italics="true" pageId="0" pageNumber="1">S. hsui</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="16CB231DB7E2F52241F2D6EB1E42F8D7" lsidName="S. sheipaensis" pageId="0" pageNumber="1" rank="species" species="sheipaensis">
<emphasis id="C4C97C596145C2B2606FCD4ABD2B435A" italics="true" pageId="0" pageNumber="1">S. sheipaensis</emphasis>
</taxonomicName>
sp. nov.) based on the following combination of characters: yellowish-brown antennomeres I-IV and black antennomeres V-XI (Fig.
<figureCitation id="F89460A4D4C875492BF70C3144666B75" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus of Sphaeroderma changi sp. nov. and S. sheipaensis sp. nov. A S. changi sp. nov., female, dorsal view B ditto, ventral view C ditto, lateral view D S. sheipaensis sp. nov., female, dorsal view E ditto, ventral view F ditto, lateral view." figureDoi="10.3897/zookeys.1185.112099.figure5" httpUri="https://binary.pensoft.net/fig/942955" pageId="0" pageNumber="1">5A-C</figureCitation>
) [entire yellowish-brown antennae in
<taxonomicName id="07708F44553CCB0C543DB4D0A07BA240" lsidName="S. hsui" pageId="0" pageNumber="1" rank="species" species="hsui">
<emphasis id="FD0B494F07386D204B1789F5827240A5" italics="true" pageId="0" pageNumber="1">S. hsui</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="7F02E848B3C93DE95CC23EB2A953A9DA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Sphaeroderma hsui sp. nov. A typical form, male, dorsal view B ditto, ventral view C ditto, lateral view D color variation, male, dorsal view E ditto, ventral view F ditto, lateral view G color variation, female, dorsal view H ditto, ventral view I ditto, lateral view." figureDoi="10.3897/zookeys.1185.112099.figure3" httpUri="https://binary.pensoft.net/fig/942953" pageId="0" pageNumber="1">3</figureCitation>
)]; transverse antennal calli with well-developed supracallinal sulci (Fig.
<figureCitation id="071008F603977AA3BBFF080A43D0881F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Sphaeroderma species in frontal view A S. hsui sp. nov. B S. changi sp. nov. C S. sheipaensis sp. nov. D S. flavonotatum Chujo E S. jungchani sp. nov." figureDoi="10.3897/zookeys.1185.112099.figure2" httpUri="https://binary.pensoft.net/fig/942952" pageId="0" pageNumber="1">2B</figureCitation>
) [longitudinal antennal calli with poorly delimited supracallinal sulci in
<taxonomicName id="E7633604F14F49B7541A188904CFD0A5" lsidName="S. hsui" pageId="0" pageNumber="1" rank="species" species="hsui">
<emphasis id="52773BF4B2E5789FF974F54F2E27695C" italics="true" pageId="0" pageNumber="1">S. hsui</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="6DF0E74272BEA0E7A4380DB64B9D8F2F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Sphaeroderma species in frontal view A S. hsui sp. nov. B S. changi sp. nov. C S. sheipaensis sp. nov. D S. flavonotatum Chujo E S. jungchani sp. nov." figureDoi="10.3897/zookeys.1185.112099.figure2" httpUri="https://binary.pensoft.net/fig/942952" pageId="0" pageNumber="1">2A</figureCitation>
)]; wide aedeagus, 3.1
<normalizedToken id="24602D36D5C97454FC5EA834658C5A5F" originalValue="×">x</normalizedToken>
longer than wide, ventral disc with one pair of oblique ridges from apical 1/5 in lateral margin (Fig.
<figureCitation id="97DAE869A58E2F7F33EDF6C223616D3D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Sphaeroderma changi sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1">6C, D</figureCitation>
) [slender aedeagus, 4.0
<normalizedToken id="3A8060D6F56D06C4F09342232D74E39D" originalValue="×">x</normalizedToken>
longer than wide and ventral disc without oblique ridges in
<taxonomicName id="D6808FEF855E9F1580C6E344997E9A57" lsidName="S. hsui" pageId="0" pageNumber="1" rank="species" species="hsui">
<emphasis id="9810CB84C21266681BBEDEC406649B25" italics="true" pageId="0" pageNumber="1">S. hsui</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="755367B4553E21C2FCDB06D6FEACDA0B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Sphaeroderma hsui sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure4" httpUri="https://binary.pensoft.net/fig/942954" pageId="0" pageNumber="1">4C, D</figureCitation>
)]; triangular abdominal ventrite VIII in females with apical margin with seven pairs of setae at medial part, subapically and slightly narrowed sides (Fig.
<figureCitation id="87240FB0501196FC25DF5BB2FCE6D5F8" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Sphaeroderma changi sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1">6E</figureCitation>
) [apical margin lacking setae; subapically and moderately narrowed sides in
<taxonomicName id="EEBB5CCB5E29DECA136D4789C19460AA" lsidName="S. hsui" pageId="0" pageNumber="1" rank="species" species="hsui">
<emphasis id="0DDDDAB530C60590E10222C8463BFFF8" italics="true" pageId="0" pageNumber="1">S. hsui</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="B89A07EBEB8B71D482802C073C1539A0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Sphaeroderma hsui sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure4" httpUri="https://binary.pensoft.net/fig/942954" pageId="0" pageNumber="1">4E</figureCitation>
)]; longitudinally triangular gonocoxae (Fig.
<figureCitation id="5E73E25BE574EE4AB96E98299A593EF5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Diagnostic characters of Sphaeroderma changi sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure6" httpUri="https://binary.pensoft.net/fig/942956" pageId="0" pageNumber="1">6G</figureCitation>
) [transversely triangular gonocoxae in
<taxonomicName id="B31B4E3756B474290641BD19AEB36401" lsidName="S. hsui" pageId="0" pageNumber="1" rank="species" species="hsui">
<emphasis id="31235F58772AA79678F4BCA165C6A8D3" italics="true" pageId="0" pageNumber="1">S. hsui</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="3F4B5691465F24AD669BB869EB6C6455" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Sphaeroderma hsui sp. nov. A antenna, male B antenna, female C aedeagus, dorsal view D aedeagus, lateral view E abdominal ventrite VIII, female F spermatheca G gonocoxae." figureDoi="10.3897/zookeys.1185.112099.figure4" httpUri="https://binary.pensoft.net/fig/942954" pageId="0" pageNumber="1">4G</figureCitation>
) and
<taxonomicName id="FC8D61C564180E0B391F93CC2A8BC552" lsidName="S. sheipaensis" pageId="0" pageNumber="1" rank="species" species="sheipaensis">
<emphasis id="1B088C5A14970EF83943368C2A80F334" italics="true" pageId="0" pageNumber="1">S. sheipaensis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="04CA236FB06A7BAA6133E64DD61221EE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Diagnostic characters of Sphaeroderma sheipaensis sp. nov. and S. jungchani Lee, sp. nov. A antenna, male, S. jungchani sp. nov. B antenna, female, Sphaeroderma sheipaensis sp. nov. C aedeagus, dorsal view, S. jungchani sp. nov. D ditto, lateral view E abdominal ventrite VIII, female, S. sheipaensis sp. nov. F spermatheca, S. sheipaensis sp. nov. G gonocoxae, S. sheipaensis sp. nov." figureDoi="10.3897/zookeys.1185.112099.figure7" httpUri="https://binary.pensoft.net/fig/942957" pageId="0" pageNumber="1">7G</figureCitation>
)].
</paragraph>
</subSubSection>
<subSubSection id="0DA6CD344AAF8A8DFD60823A476E31B2" pageId="0" pageNumber="1" type="etymology">
<paragraph id="90FCE82B265A6C84D4978C58CC9CA476" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="27865D68ABF997F5008318C8DD83D349" pageId="0" pageNumber="1">This species is named for Mr Li-Jen Chang (張勵仁) for his assistance in conducting the project.</paragraph>
</subSubSection>
<subSubSection id="412CC08CA4A4CA46C2815BC17B86854A" pageId="0" pageNumber="1" type="distribution">
<paragraph id="8B0A84037C048BF4E373D8DADA0EDC26" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="415463C5388C2F0C967093F5E04B558E" pageId="0" pageNumber="1">
<taxonomicName id="EA17E8A02FE4782E03B1F60F7A330A20" authorityName="Lee" authorityYear="2023" class="Insecta" family="Chrysomelidae" genus="Sphaeroderma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeroderma changi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="changi">
<emphasis id="E6778DE773C6FCA1535FAE4442B8FE4E" italics="true" pageId="0" pageNumber="1">Sphaeroderma changi</emphasis>
</taxonomicName>
sp. nov. is a rarely collected species known from only one locality in Shei-Pa National Park.
</paragraph>
</subSubSection>
</treatment>
</document>