228 lines
27 KiB
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228 lines
27 KiB
XML
<document ID-CLB-Dataset="53461" ID-DOI="http://dx.doi.org/10.3897/phytokeys.219.101779" ID-GBIF-Dataset="6cda71cf-ef59-4fbd-860d-96dfb7021bc1" ID-PMC="PMC10194777" ID-Pensoft-Pub="1314-2003-219-145" ID-Pensoft-UUID="A2ECEB672C4D5027A90252D384FB1795" ID-PubMed="37215999" ModsDocID="1314-2003-219-145" checkinTime="1677023374729" checkinUser="pensoft" docAuthor="Moonlight, Peter W. & Cardoso, Domingos Benicio Oliveira Silva" docDate="2023" docId="F2F096E986A450F9843106C6C6B21042" docLanguage="en" docName="PhytoKeys 219: 145-170" docOrigin="PhytoKeys 219" docPubDate="2023-02-20" docSource="http://dx.doi.org/10.3897/phytokeys.219.101779" docTitle="Keraunea Moonlight & Cardoso, 2023, nom. cons." docType="treatment" docVersion="5" id="A2ECEB672C4D5027A90252D384FB1795" lastPageNumber="145" masterDocId="A2ECEB672C4D5027A90252D384FB1795" masterDocTitle="A taxonomic revision of Keraunea, including three new species and its phylogenetic realignment with Ehretiaceae (Boraginales)" masterLastPageNumber="170" masterPageNumber="145" pageNumber="145" updateTime="1732751842401" updateUser="ExternalLinkService">
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<mods:mods id="80228DB231B54089B13A67A1A1D1B2F2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BBE433DEB3DBFBFDF03473618B868E95">
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<mods:title id="D94CA70C022B0975D2047F65D126489F">A taxonomic revision of Keraunea, including three new species and its phylogenetic realignment with Ehretiaceae (Boraginales)</mods:title>
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</mods:titleInfo>
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<mods:name id="A215E0E0F8359ED26FCC730A648BA639" type="personal">
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<mods:role id="EBDF9F91FEF262C9E265EAB71ECA6B4A">
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<mods:roleTerm id="89E8902B22F31061BFB1A903FB0FF0AA">Author</mods:roleTerm>
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<mods:namePart id="639AA097796A973BBDCEE9E0BA40FC2F">Moonlight, Peter W.</mods:namePart>
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<mods:nameIdentifier id="121834C80E6D6DEC8F7F7DC210054B70" type="ORCID">https://orcid.org/0000-0003-4342-2089</mods:nameIdentifier>
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<mods:affiliation id="BB475986A5555D24655DA5E32C10A78E">Botany Department, School of Natural Sciences, Trinity College Dublin, Dublin 2, Ireland</mods:affiliation>
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<mods:nameIdentifier id="7CF716AB2A2083579F9DE61B2CCF2572" type="email">p.moonlight@rbge.ac.uk</mods:nameIdentifier>
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<mods:roleTerm id="25AC32BB8E07176A67F4B7A5F6A6376D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CAAA2016AE1E305A84F3DA12E37B5E3D">Cardoso, Domingos Benicio Oliveira Silva</mods:namePart>
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<mods:nameIdentifier id="66927D90F8400C455FBE59B65A2D5642" type="ORCID">https://orcid.org/0000-0001-7072-2656</mods:nameIdentifier>
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<mods:affiliation id="ACF2C9036EE0F2372D8C8626C69C2BAA">Instituto de Pesquisas Jardim Botanico do Rio de Janeiro, Rio de Janeiro, Brazil & Universidade Federal da Bahia, Salvador, Bahia, Brazil</mods:affiliation>
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<mods:typeOfResource id="3ACE63E891A2E2F4C5D086ECB54287D5">text</mods:typeOfResource>
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<mods:titleInfo id="2CE476605E4E4DB761A40D8CEF36FF02">
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<mods:title id="6D381F1A2318BC3BABBC00B434C1C74C">PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="E3851F1C35384087223A6D857374AC71">
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<mods:date id="A0D9A1062FFE18C6303D3582F6779CA1">2023</mods:date>
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<mods:detail id="8F219CFE755004AAE754B09B18FD84A5" type="pubDate">
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<mods:number id="D58C7ADAD1A92D989ED1E8222B8FE7BF">2023-02-20</mods:number>
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</mods:detail>
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<mods:detail id="237D7489904D9B2352F5DF8015C1AE22" type="volume">
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<mods:number id="38D08E8B82A7C7C593DA13142570E18C">219</mods:number>
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</mods:detail>
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<mods:start id="6D961732E9BD567257EB5A32EB76CF3E">145</mods:start>
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<mods:end id="1BDD644A5BE177E59C0A8F9A16CC9E79">170</mods:end>
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<mods:location id="3743A0140D2EA5328F540609A465D591">
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<mods:url id="DAA7ADC571D2200D0E10744B6A4D40CB">http://dx.doi.org/10.3897/phytokeys.219.101779</mods:url>
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</mods:location>
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<mods:classification id="283212E01F1F22B42175D5AF87454997">journal article</mods:classification>
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<mods:identifier id="4F049D31AA6C9AE13C9F10AC7D71F02F" type="DOI">http://dx.doi.org/10.3897/phytokeys.219.101779</mods:identifier>
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<mods:identifier id="3CBF1224742970C4D0C72C0BF4990C10" type="Pensoft-Pub">1314-2003-219-145</mods:identifier>
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<mods:identifier id="F7D5068492837916D2DBD837095B8A20" type="Pensoft-UUID">A2ECEB672C4D5027A90252D384FB1795</mods:identifier>
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</mods:mods>
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<treatment id="F2F096E986A450F9843106C6C6B21042" ID-GBIF-Taxon="209339084" LSID="urn:lsid:plazi:treatment:F2F096E986A450F9843106C6C6B21042" httpUri="http://treatment.plazi.org/id/F2F096E986A450F9843106C6C6B21042" lastPageNumber="145" pageId="0" pageNumber="145" scope_family="Ehretiaceae" scope_order="Boraginales">
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<subSubSection id="0F6DB03E91C42344051E7548C2B32CB4" pageId="0" pageNumber="145" type="nomenclature">
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<paragraph id="2B18FA0498B6855F1C64A4CAD7A01D1D" pageId="0" pageNumber="145">
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<taxonomicName id="7F5CB7B5CFB203DAB59779CC07079C6F" ID-CoL="9L9" ID-ENA="1561073" LSID="F2F096E9-86A4-50F9-8431-06C6C6B21042" authorityName="Moonlight & Cardoso" authorityYear="2023" class="Magnoliopsida" family="Ehretiaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="family" status="nom. cons.">Ehretiaceae Mart., Nov. Gen. Sp. Pl. 2: 136, 138. 1827</taxonomicName>
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<taxonomicNameLabel id="3D5374403369A30915AA17005F26E8AC" pageId="0" pageNumber="145">nom. cons.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="B23F4D67C5299ABA311F4D39F2835966" pageId="0" pageNumber="145" type="type genus">
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<paragraph id="1DFA008F053F971C030DC3D60D47E800" pageId="0" pageNumber="145">Type genus.</paragraph>
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<paragraph id="6181ADFCC8FA49FBFFB775770411E7B9" pageId="0" pageNumber="145">
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<taxonomicName id="38B0F85560A7B9D90D6EA1C3794386B7" class="Magnoliopsida" family="Ehretiaceae" genus="Ehretia" kingdom="Plantae" lsidName="Ehretia" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="05749783949C5849F960A22398927BD6" italics="true" pageId="0" pageNumber="145">Ehretia</emphasis>
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</taxonomicName>
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P.Browne.
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</paragraph>
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</subSubSection>
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<subSubSection id="C7FC4136A76ED5C5D3AE8CDC1981729A" pageId="0" pageNumber="145" type="description">
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<paragraph id="9EA2049934C6F9DE0D42F877EBE5F2BE" pageId="0" pageNumber="145">Description.</paragraph>
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<paragraph id="49BFBA1D3F239746B3D49FF0CCF7794F" pageId="0" pageNumber="145">
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[differences from
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<bibRefCitation id="8A7EF20F84DCF74EF48AAE4AF1C05E33" DOI="https://doi.org/10.12705/653.5" author="Luebert, F" journalOrPublisher="Taxon" pageId="0" pageNumber="145" pagination="502 - 522" refId="B14" refString="Luebert, F, Cecchi, L, Frohlich, MW, Gottschling, M, Guilliams, M, Hasenstab-Lehman, KE, Hilger, HH, Miller, JS, Mittelbach, M, Nazaire, M, Nepi, M, Ober, D, Olmstead, RG, Selvi, F, Simpson, MG, Sutory, K, Valdes, B, Walden, GK, Weigend, M, 2016. Familial classification of the Boraginales. Taxon 65 (3): 502 - 522, DOI: https://doi.org/10.12705/653.5" title="Familial classification of the Boraginales." url="https://doi.org/10.12705/653.5" volume="65" year="2016">Luebert et al. (2016)</bibRefCitation>
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in bold] Trees, shrubs,
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<emphasis id="5DDC6101C6C7FCAD81C313AB4B8E186F" bold="true" pageId="0" pageNumber="145">lianas</emphasis>
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, perennial herbs, rarely with thorns (
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<taxonomicName id="955A751965044FDD1EFCB176BA095899" class="Magnoliopsida" family="Ehretiaceae" genus="Rochefortia" kingdom="Plantae" lsidName="Rochefortia" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="3E7631A75E7B648C87A16087D29C9A2E" italics="true" pageId="0" pageNumber="145">Rochefortia</emphasis>
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</taxonomicName>
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Sw.); indumentum variable, hirsute to glabrescent. Leaves alternate, entire, petiolate; lamina variable in shape, strongly dissected in the halophytic
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<taxonomicName id="949047571555575B1619FAF60AB57374" class="Magnoliopsida" family="Ehretiaceae" genus="Cortesia" kingdom="Plantae" lsidName="Cortesia" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="674CD5842789EA1F4F3FA9F006416313" italics="true" pageId="0" pageNumber="145">Cortesia</emphasis>
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</taxonomicName>
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Cav. Inflorescences terminal or axillary thyrses, sometimes congested,
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<emphasis id="167833A9138A33BC1CBD5B4699BE8FF3" bold="true" pageId="0" pageNumber="145">
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or few-flowered corymbs (
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<taxonomicName id="5B24DB0115CF72CCA540B53E72B0AD3A" authorityName="Moonlight & Cardoso" authorityYear="2023" class="Magnoliopsida" family="Convolvulaceae" genus="Keraunea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Keraunea" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="DB501D5129CC50B9F963F4A7084CE18C" italics="true" pageId="0" pageNumber="145">Keraunea</emphasis>
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</taxonomicName>
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)
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</emphasis>
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. Flowers pentamerous, cosexual or unisexual and dioecious in
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<taxonomicName id="8D16B151B3DE694AE1D9F64A11CBDCAD" authorityName="Ducke" authorityYear="1925" class="Magnoliopsida" family="Ehretiaceae" genus="Lepidocordia" kingdom="Plantae" lsidName="Lepidocordia" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="E4E3D8F23549ACEA081BF5E700E32A5C" italics="true" pageId="0" pageNumber="145">Lepidocordia</emphasis>
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</taxonomicName>
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Ducke and
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<taxonomicName id="B0472D9A77D3F520526A8E16A04A741D" class="Magnoliopsida" family="Ehretiaceae" genus="Rochefortia" kingdom="Plantae" lsidName="Rochefortia" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="990664493D5D4511EBFD37B4601D67D9" italics="true" pageId="0" pageNumber="145">Rochefortia</emphasis>
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</taxonomicName>
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,
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<emphasis id="11EA590730568B5F2803AB66B5E867D9" bold="true" pageId="0" pageNumber="145">
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sometimes inserted on the centre of an accrescent bracteole (
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<taxonomicName id="82602E743C6CAF0304EE05FEA2527CD0" authorityName="Moonlight & Cardoso" authorityYear="2023" class="Magnoliopsida" family="Convolvulaceae" genus="Keraunea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Keraunea" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="0B9F4D876B8B24026842935673EE60CD" italics="true" pageId="0" pageNumber="145">Keraunea</emphasis>
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</taxonomicName>
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)
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</emphasis>
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; calyx lobes united in a tube or distinct nearly to the base, tubular to campanulate; aestivation imbricate (mostly quincuncial); corolla sympetalous, generally tubular with spreading lobes, rotate, or campanulate to urceolate, white, red or blue (
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<taxonomicName id="E443AD109152AAE12838A1EFD74439EE" class="Magnoliopsida" family="Ehretiaceae" genus="Halgania" kingdom="Plantae" lsidName="Halgania" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="6E9A2EF4BCC22FF7C2F2C1ACEAC3CE7E" italics="true" pageId="0" pageNumber="145">Halgania</emphasis>
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</taxonomicName>
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, some species of
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<taxonomicName id="8CE0D9512652CD91039B4D3259C205F1" class="Magnoliopsida" family="Ehretiaceae" genus="Bourreria" kingdom="Plantae" lsidName="Bourreria" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="7B25D8CBC4636A17C52D872B436B917B" italics="true" pageId="0" pageNumber="145">Bourreria</emphasis>
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</taxonomicName>
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P.Browne); stamens 5, the filaments generally adnate to the corolla tube at least at the base, sometimes puberulent at the point of insertion, the anthers usually exerted; gynoecium bicarpellate, the ovary uni- to tetralocular from secondary subdivision, style terminal, the stigma clavate to capitate with 1(2) branches; nectar disc usually present at base of the ovary. Fruits drupaceous, often drying and separating into two two-seeded pyrenes, or 4 1-seeded pyrenes or schizocarps, or 4 nutlets.
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</paragraph>
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</subSubSection>
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<subSubSection id="570023DB9BFFEAD63CC3B7F05F53CD6A" pageId="0" pageNumber="145" type="distribution">
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<paragraph id="F57A8657A90FF2699B39BDE8E4D9240C" pageId="0" pageNumber="145">Distribution.</paragraph>
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<paragraph id="87722C542F33EF7EA7E71C9E8310B370" pageId="0" pageNumber="145">
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<taxonomicName id="36A6B43541EE3E29EA1E934F40B374F6" class="Magnoliopsida" family="Ehretiaceae" kingdom="Plantae" lsidName="" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="family">Ehretiaceae</taxonomicName>
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is a broadly distributed family found throughout tropical and subtropical Asia, Australia, sub-Saharan Africa. In the Americas, its distribution encompasses the eastern United States, Florida, Central America, the Caribbean, the Guyana shield and the Andes. In Brazil, the
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<taxonomicName id="004A7EA4D0595FA65393DB24BE28711F" class="Magnoliopsida" family="Ehretiaceae" kingdom="Plantae" lsidName="" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="family">Ehretiaceae</taxonomicName>
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was previously only known from the single species
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<taxonomicName id="34150B4DBF200A96DFFDFB53C73E22AF" class="Magnoliopsida" family="Ehretiaceae" genus="Lepidocordia" kingdom="Plantae" lsidName="Lepidocordia punctata" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="punctata">
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<emphasis id="7FED3AE1119B868E616B2FB93D2A6503" italics="true" pageId="0" pageNumber="145">Lepidocordia punctata</emphasis>
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</taxonomicName>
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Ducke (
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<bibRefCitation id="0CF3F9CC7BD5C8CB0C05DF785F6856B8" author="Stapf, MNS" journalOrPublisher="Global Ecology and Biogeography" pageId="0" pageNumber="145" publicationUrl="https://floradobrasil.jbrj.gov.br/FB78014" refId="B21" refString="Stapf, MNS, 2023. Lepidocordia. In: Flora e Funga do Brasil. Jardim Botanico do Rio de Janeiro. https://floradobrasil.jbrj.gov.br/FB78014" title="Lepidocordia. In: Flora e Funga do Brasil. Jardim Botanico do Rio de Janeiro." url="https://floradobrasil.jbrj.gov.br/FB78014" year="2023">Stapf 2023</bibRefCitation>
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), found in lowland Amazonian forests in
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<normalizedToken id="04FE29807DABDF0E5CDBA94A41F0484C" originalValue="Pará">Para</normalizedToken>
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and Roraima states. Our treatment therefore represents new records of the family from the Caatinga and Mata
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<normalizedToken id="3FF49AE8AAC77897C4E5B8848C967F45" originalValue="Atlântica">Atlantica</normalizedToken>
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phytogeographic regions in Brazil, and from the states of Bahia, Minas Gerais,
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<normalizedToken id="CDE1EB0A6259C447905B187FC7D77616" originalValue="Espírito">Espirito</normalizedToken>
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Santo and Rio de Janeiro (Fig.
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<figureCitation id="E244DFCB26CF091B04E6AF89222E929D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. RAxML bootstrap tree based on nuclear ribosomal (ITS) and plastid (matK, rbcL, trnL-F) DNA sequence data resolving Keraunea within the Ehretiaceae. Node labels indicate nodes with bootstrap support values above 50. Inset phylogram (bottom left) indicates branch lengths. Outgroup taxa from representatives of other Boraginales families are in light grey. Illustrations are (top to bottom): Keraunea, Halgania, Ehretia, Bourreria, Rochefortia and Tiquilia." figureDoi="10.3897/phytokeys.219.101779.figure1" httpUri="https://binary.pensoft.net/fig/813025" pageId="0" pageNumber="145">1</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="008FD5A972CF758A488A3947375E614F" pageId="0" pageNumber="145" type="included genera">
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<paragraph id="077C9F435BB68A51CF2F2F851CDA8CDA" pageId="0" pageNumber="145">Included genera.</paragraph>
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<paragraph id="D8379055E558DB1FB346D78F0C0BDB9F" pageId="0" pageNumber="145">
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The family includes the following eight genera:
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<taxonomicName id="637667B7AC3C270C531577F449525B47" class="Magnoliopsida" family="Ehretiaceae" genus="Bourreria" kingdom="Plantae" lsidName="Bourreria" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="FDAAB8E5F10085B6BCFCA5A11064D77E" italics="true" pageId="0" pageNumber="145">Bourreria</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="FAB37CCBA77FE15A81205CDB33CDC253" class="Magnoliopsida" family="Ehretiaceae" genus="Cortesia" kingdom="Plantae" lsidName="Cortesia" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="5DFE90551ED753258B17DFF089EFBE08" italics="true" pageId="0" pageNumber="145">Cortesia</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="9724F588791CA44BC7E6978DF66D4D21" class="Magnoliopsida" family="Ehretiaceae" genus="Ehretia" kingdom="Plantae" lsidName="Ehretia" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="71D7280C266BC781146B13296B567FE7" italics="true" pageId="0" pageNumber="145">Ehretia</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="F95D6E0145666BEB57DBFC7647A617D5" class="Magnoliopsida" family="Ehretiaceae" genus="Halgania" kingdom="Plantae" lsidName="Halgania" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="2FDF221ACC6F7127B68AA0B2BABD9629" italics="true" pageId="0" pageNumber="145">Halgania</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="711E70E0EEC8E35E8B36C6321BFA04E8" authorityName="Moonlight & Cardoso" authorityYear="2023" class="Magnoliopsida" family="Convolvulaceae" genus="Keraunea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Keraunea" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="4A38C4BA29230A6AD111C72097375323" italics="true" pageId="0" pageNumber="145">Keraunea</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="F196202C379519D0793259F3A3EB546E" authorityName="Ducke" authorityYear="1925" class="Magnoliopsida" family="Ehretiaceae" genus="Lepidocordia" kingdom="Plantae" lsidName="Lepidocordia" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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<emphasis id="8DD8276F5B6C8EC46F6A765F79AA220F" italics="true" pageId="0" pageNumber="145">Lepidocordia</emphasis>
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</taxonomicName>
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|
Ducke,
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<taxonomicName id="EB7205D0D8BA3C6DD90DA8BA8FE97185" class="Magnoliopsida" family="Ehretiaceae" genus="Rochefortia" kingdom="Plantae" lsidName="Rochefortia" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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|
<emphasis id="73671BBBCDB0FA68EBEFECE47989E17B" italics="true" pageId="0" pageNumber="145">Rochefortia</emphasis>
|
|
</taxonomicName>
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|
and
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|
<taxonomicName id="EEFCE7B9380F88E8557938057A652332" authorityName="Persoon" authorityYear="1805" class="Magnoliopsida" family="Ehretiaceae" genus="Tiquilia" kingdom="Plantae" lsidName="Tiquilia" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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|
<emphasis id="B92FF99DAE81ADF5964BF8E5A0AE702C" italics="true" pageId="0" pageNumber="145">Tiquilia</emphasis>
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</taxonomicName>
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|
Pers. (
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|
<bibRefCitation id="25DDBB0F61468550F354EFB25FF9AD4A" DOI="https://doi.org/10.12705/653.5" author="Luebert, F" journalOrPublisher="Taxon" pageId="0" pageNumber="145" pagination="502 - 522" refId="B14" refString="Luebert, F, Cecchi, L, Frohlich, MW, Gottschling, M, Guilliams, M, Hasenstab-Lehman, KE, Hilger, HH, Miller, JS, Mittelbach, M, Nazaire, M, Nepi, M, Ober, D, Olmstead, RG, Selvi, F, Simpson, MG, Sutory, K, Valdes, B, Walden, GK, Weigend, M, 2016. Familial classification of the Boraginales. Taxon 65 (3): 502 - 522, DOI: https://doi.org/10.12705/653.5" title="Familial classification of the Boraginales." url="https://doi.org/10.12705/653.5" volume="65" year="2016">Luebert et al. 2016</bibRefCitation>
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).
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</paragraph>
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|
</subSubSection>
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|
<subSubSection id="28CDE697932C294B454CBBE3E6495F56" pageId="0" pageNumber="145" type="notes">
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<paragraph id="7285071326978F8486D461B96B1A4BD4" pageId="0" pageNumber="145">Notes.</paragraph>
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<paragraph id="34158D1AE54EFDCC90AA9CB2D47C0DF4" pageId="0" pageNumber="145">
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Our morphological concept of the
|
|
<taxonomicName id="33D71184B6125F0F6AB2039F667CC7A0" authorityName="Persoon" authorityYear="1805" class="Magnoliopsida" family="Ehretiaceae" kingdom="Plantae" lsidName="" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="family">Ehretiaceae</taxonomicName>
|
|
is little changed from that of
|
|
<bibRefCitation id="5F2D305284174D42A2D390A5C35C83A4" DOI="https://doi.org/10.12705/653.5" author="Luebert, F" journalOrPublisher="Taxon" pageId="0" pageNumber="145" pagination="502 - 522" refId="B14" refString="Luebert, F, Cecchi, L, Frohlich, MW, Gottschling, M, Guilliams, M, Hasenstab-Lehman, KE, Hilger, HH, Miller, JS, Mittelbach, M, Nazaire, M, Nepi, M, Ober, D, Olmstead, RG, Selvi, F, Simpson, MG, Sutory, K, Valdes, B, Walden, GK, Weigend, M, 2016. Familial classification of the Boraginales. Taxon 65 (3): 502 - 522, DOI: https://doi.org/10.12705/653.5" title="Familial classification of the Boraginales." url="https://doi.org/10.12705/653.5" volume="65" year="2016">Luebert et al. (2016)</bibRefCitation>
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. The characters that differ are included in bold in the description above. The first of these is that the four species of
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|
<taxonomicName id="BF01A8FEFC2CFFBE0994F653211CFDA5" authorityName="Moonlight & Cardoso" authorityYear="2023" class="Magnoliopsida" family="Convolvulaceae" genus="Keraunea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Keraunea" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
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|
<emphasis id="7BD9F439AF0050B9DE2BC96A41415986" italics="true" pageId="0" pageNumber="145">Keraunea</emphasis>
|
|
</taxonomicName>
|
|
are the first lianescent species included within the family (versus perennial herbs, shrubs, or trees). Lianas are elsewhere found in the
|
|
<taxonomicName id="836B787B6343F27FA715BCB7388813BB" authorityName="Moonlight & Cardoso" authorityYear="2023" class="Magnoliopsida" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="order">Boraginales</taxonomicName>
|
|
in the
|
|
<taxonomicName id="1D59087C0C5FEE8ABB923F5A5BB8EBA2" authorityName="Moonlight & Cardoso" authorityYear="2023" class="Magnoliopsida" family="Cordiaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="family">Cordiaceae</taxonomicName>
|
|
(
|
|
<taxonomicName id="C8E12FC4434695AD36EE3CF62A35A816" class="Magnoliopsida" family="Heliotropiaceae" genus="Cordia" kingdom="Plantae" lsidName="Cordia" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="9D8E17344099BA2645B64BEBFB9F98E5" italics="true" pageId="0" pageNumber="145">Cordia</emphasis>
|
|
</taxonomicName>
|
|
L.) and
|
|
<taxonomicName id="A3A95EF3A5A7DEA190E86CCF39EE701D" class="Magnoliopsida" family="Heliotropiaceae" kingdom="Plantae" lsidName="" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="family">Heliotropiaceae</taxonomicName>
|
|
(
|
|
<taxonomicName id="013F191EF2246523DB0BBD8AB60027EC" class="Magnoliopsida" family="Ehretiaceae" genus="Tournefortia" kingdom="Plantae" lsidName="Tournefortia" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="0F55756584D31F60A254C890AD69917B" italics="true" pageId="0" pageNumber="145">Tournefortia</emphasis>
|
|
</taxonomicName>
|
|
L.). Secondly, we have expanded the concept of the
|
|
<taxonomicName id="C279881332E8EB898236B9C4ACDE7C28" class="Magnoliopsida" family="Ehretiaceae" kingdom="Plantae" lsidName="" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="family">Ehretiaceae</taxonomicName>
|
|
to include species with a few-flowered corymb inflorescence structure and where the flower and later fruit are inserted at the centre of an accrescent bracteole. To our knowledge, these characters are unique among not just the
|
|
<taxonomicName id="ADA711DEDC55346F76164AA20E95A221" class="Magnoliopsida" family="Ehretiaceae" kingdom="Plantae" lsidName="" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="family">Ehretiaceae</taxonomicName>
|
|
but the
|
|
<taxonomicName id="3E48722900B6E8F3400B83D6D35CE5F3" class="Magnoliopsida" kingdom="Plantae" lsidName="" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="order">Boraginales</taxonomicName>
|
|
, whose members are known for their characteristic scorpioid cymose inflorescences. We suggest the few-flowered inflorescence of
|
|
<taxonomicName id="FD392F350411E89331E2764AB8E27D2C" authorityName="Moonlight & Cardoso" authorityYear="2023" class="Magnoliopsida" family="Convolvulaceae" genus="Keraunea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Keraunea" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="C618B82AF3A09149D8BEC581EAFBEFD6" italics="true" pageId="0" pageNumber="145">Keraunea</emphasis>
|
|
</taxonomicName>
|
|
is the result of secondary reduction rather than a retained ancestral form. Within the
|
|
<taxonomicName id="8C3C93A1B0F44C0C76B24429EC8B6220" authorityName="Moonlight & Cardoso" authorityYear="2023" class="Magnoliopsida" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="order">Boraginales</taxonomicName>
|
|
, reductions to few-flowered inflorescences are present elsewhere in the
|
|
<taxonomicName id="71D00F9241D983FB7E923E0C92F34052" authorityName="Moonlight & Cardoso" authorityYear="2023" class="Magnoliopsida" family="Boraginaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="family">Boraginaceae</taxonomicName>
|
|
,
|
|
<taxonomicName id="E09B3FC306FA7B0B10DE6DFA32B9A8CA" authorityName="Moonlight & Cardoso" authorityYear="2023" class="Magnoliopsida" family="Codonaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="family">Codonaceae</taxonomicName>
|
|
and
|
|
<taxonomicName id="7F9F75BDB7B66277865D6645EC69CC28" authorityName="Moonlight & Cardoso" authorityYear="2023" class="Magnoliopsida" family="Wellstediaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Boraginales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="family">Wellstediaceae</taxonomicName>
|
|
(
|
|
<bibRefCitation id="55ABD81F695BF6DEF133D75DF8DD2507" DOI="https://doi.org/10.12705/653.5" author="Luebert, F" journalOrPublisher="Taxon" pageId="0" pageNumber="145" pagination="502 - 522" refId="B14" refString="Luebert, F, Cecchi, L, Frohlich, MW, Gottschling, M, Guilliams, M, Hasenstab-Lehman, KE, Hilger, HH, Miller, JS, Mittelbach, M, Nazaire, M, Nepi, M, Ober, D, Olmstead, RG, Selvi, F, Simpson, MG, Sutory, K, Valdes, B, Walden, GK, Weigend, M, 2016. Familial classification of the Boraginales. Taxon 65 (3): 502 - 522, DOI: https://doi.org/10.12705/653.5" title="Familial classification of the Boraginales." url="https://doi.org/10.12705/653.5" volume="65" year="2016">Luebert et al. 2016</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |