317 lines
62 KiB
XML
317 lines
62 KiB
XML
<document id="BB486BB3AA61F9A3C38D4E0D4304579F" ID-DOI="10.11646/zootaxa.5020.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="5224139" ID-ZooBank="CCF5FB8C-2552-4299-B0AF-8673230E7609" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629392338490" checkinUser="plazi" docAuthor="Mielke, Carlos G. C., Grehan, John R. & Koike, Ricardo M." docDate="2021" docId="F17187DFEE3AFFA6FF10EEF421C6F891" docLanguage="en" docName="zootaxa.5020.3.7.pdf" docOrigin="Zootaxa 5020 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mutipialus Mielke & Grehan & Koike 2021, gen. n." docType="treatment" docVersion="3" lastPageNumber="567" masterDocId="0D48FFA7EE3CFFA0FF87EF662528FFD3" masterDocTitle="Descriptions of two new genera and six new species of ghost-moths (Lepidoptera Hepialoidea: Hepialidae) from south-eastern and southern Brazil" masterLastPageNumber="580" masterPageNumber="561" pageNumber="567" updateTime="1699071641799" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="785B9EF484F40D630410CB7E657C15F4">Descriptions of two new genera and six new species of ghost-moths (Lepidoptera Hepialoidea: Hepialidae) from south-eastern and southern Brazil</mods:title>
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<mods:name id="545B3FA35CBE1F0599559450E1597A06" type="personal">
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<mods:namePart id="5665F23A00426F4E77B1D41FFD4DE67A">Mielke, Carlos G. C.</mods:namePart>
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<mods:nameIdentifier id="AD87232CE26FF9F28C61E2248C385DCD" type="ORCID">0000-0003-0091-0840</mods:nameIdentifier>
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<mods:affiliation id="821F3C679CC8FE3A3C336B7FE6AC8795">Caixa postal 1206, 84.145 - 000 Carambeí, Paraná, Brazil. & Research Associate, McGuire Center for Lepidoptera & Biodiversity, Gainesville, FL 32611, USA. & cmielke 1 @ uol. com. br; https: // orcid. org / 0000 - 0003 - 0091 - 0840</mods:affiliation>
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<mods:namePart id="171EC7E3919B5374F7D4D04DCBB65FEB">Grehan, John R.</mods:namePart>
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<mods:nameIdentifier id="8673FFFB4B8F5786F481780E6CA5628F" type="ORCID">0000-0002-3119-1140</mods:nameIdentifier>
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<mods:affiliation id="BC7661A2747D8D3B9C01F220D8DBD410">Research Associate, McGuire Center for Lepidoptera & Biodiversity, Gainesville, FL 32611, USA. & calabar. John @ gmail. com; https: // orcid. org / 0000 - 0002 - 3119 - 1140</mods:affiliation>
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<mods:nameIdentifier id="C9F3BA9F02353763AF1E1178717513DB" type="email">ohn@gmail.com</mods:nameIdentifier>
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<mods:namePart id="3D643AE3A04A160D7CA826130C4F989D">Koike, Ricardo M.</mods:namePart>
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<mods:affiliation id="4C3923F00D1826696428C8AD840D0D98">Estrada das Canjicas, 2500, 07424 - 000, Arujá, São Paulo, Brazil. & ricardokoike. jp @ gmail. com; https: // orcid. org / 0000 - 0002 - 6330 - 6416</mods:affiliation>
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<mods:nameIdentifier id="F396B5EF1AFE6863A0ECE8F11058107B" type="email">ricardokoike.jp@gmail.com</mods:nameIdentifier>
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<mods:date id="FA73A1E08CC02F536B96834756437B33">2021</mods:date>
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<mods:number id="D258FF4B7607E574C540818B308F508E">2021-08-13</mods:number>
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<taxonomicName id="BED84D4AEE3AFFA6FF10EEF42433FE7E" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[151,283,402,429]" class="Insecta" family="Hepialidae" genus="Mutipialus" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="567" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="4BACEADBEE3AFFA6FF10EEF42433FE7E" bold="true" box="[151,283,402,429]" italics="true" pageId="6" pageNumber="567">Mutipialus</emphasis>
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<subSubSection id="31C26542EE3AFFA6FF10EED027DCFDE2" pageId="6" pageNumber="567" type="description">
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<paragraph id="796736C9EE3AFFA6FF10EED02125FE02" blockId="6.[151,1037,402,465]" box="[151,1037,438,465]" pageId="6" pageNumber="567">
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(
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<figureCitation id="E1E32A4CEE3AFFA6FF18EED0243AFE02" box="[159,274,438,465]" captionStart-0="FIGURES 8−12" captionStart-1="FIGURES 13−15" captionStartId-0="7.[151,264,1940,1964]" captionStartId-1="9.[151,264,1548,1572]" captionTargetBox-0="[157,1430,181,1917]" captionTargetBox-1="[184,1402,345,1524]" captionTargetId-0="figure-20@7.[157,1430,181,1917]" captionTargetId-1="figure-107@9.[184,1402,345,1524]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="FIGURES 8−12. Mutipialus dilatus sp. n. Adults. 8, HT ♀: dorsal view (8a), ventral view (8b) (DZUP). 9, PT ♀ (CGCM 7.584): dorsal view (CGCM). 10, PT ♀ (CGCM 18.103): dorsal view (CGCM). 11, PT ♂ (CGCM 41.733): dorsal view (11a), ventral view (11b) (CGCM). 12, PT ♂ (CGCM 23.689): dorsal view (12a), ventral view (12b) (CGCM)." captionText-1="FIGURES 13−15. Mutipialus monticolus sp. n. Adults. 13, HT ♀: dorsal view (13a), ventral view (13b) (DZUP). 14, PT ♂ (CGCM 29.697): dorsal view (14a), ventral view (14b) (CGCM). 15, PT ♀ (CGCM 35.737): dorsal view (15a), ventral view (15b) (CGCM)." figureDoi-0="http://doi.org/10.5281/zenodo.5224145" figureDoi-1="http://doi.org/10.5281/zenodo.5224147" httpUri-0="https://zenodo.org/record/5224145/files/figure.png" httpUri-1="https://zenodo.org/record/5224147/files/figure.png" pageId="6" pageNumber="567">Figs 8–15</figureCitation>
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,
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<figureCitation id="E1E32A4CEE3AFFA6FE99EED024B8FE03" box="[286,400,438,464]" captionStart="FIGURES 16−24" captionStartId="10.[151,264,1874,1898]" captionTargetBox="[242,1332,181,1847]" captionTargetId="figure-21@10.[227,1361,181,1850]" captionTargetPageId="10" captionText="FIGURES 16−24. Transverse scales and legs. Figs 16–19. Transverse scales (arrowed). 16, Mutipialus dilatus sp. n. PT ♂ (CGCM 23.689). 17, M. monticolus sp. n., HT ♀. 18, Parapielus luteicornis ♂ (CGCM 28.544, Argentina, Rio Grande; CGCM). 19, Hepialiscus htayaungi ♂ (CGCM 33.586, Myanmar; CGCM). Figs 20–24. Legs. 20, Agripialus campos sp. n., PT ♀ (CGCM 31.875). 21, A. variabilis sp. n., HT ♀. 22, A. itatiaia sp. n., HT ♂. 23, A. caparao sp. n., HT ♂. 24, Mutipialus dilatus sp. n., PT ♂ (CGCM 27.435)." figureDoi="http://doi.org/10.5281/zenodo.5224149" httpUri="https://zenodo.org/record/5224149/files/figure.png" pageId="6" pageNumber="567">16–17, 24</figureCitation>
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,
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<figureCitation id="E1E32A4CEE3AFFA6FE1BEED02726FE03" box="[412,526,438,464]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="6" pageNumber="567">27–28, 30</figureCitation>
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,
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<figureCitation id="E1E32A4CEE3AFFA6FD9DEED0274AFE02" box="[538,610,438,465]" captionStart="FIGURES 31−35" captionStartId="12.[151,264,1743,1767]" captionTargetBox="[155,1434,184,1706]" captionTargetId="figure-21@12.[151,1435,181,1718]" captionTargetPageId="12" captionText="FIGURES 31−35. Female abdomen (upper side, tergites; lower side, sternites). 31, Agripialus campos sp. n., PT (CGCM 31.875). 32, A. variabilis sp. n., HT. 33, A. itatiaia sp. n., PT (CGCM 25.532). 34, Mutipialus dilatus sp. n., HT (CGCM 5.801). 35, M. monticolus sp. n., HT." figureDoi="http://doi.org/10.5281/zenodo.5224155" httpUri="https://zenodo.org/record/5224155/files/figure.png" pageId="6" pageNumber="567">34–35</figureCitation>
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,
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<figureCitation id="E1E32A4CEE3AFFA6FDE9EED02676FE02" box="[622,862,438,465]" captionStart="FIGURES 36−46" captionStartId="13.[151,264,1743,1767]" captionTargetBox="[251,1340,182,1714]" captionTargetId="figure-20@13.[247,1340,181,1719]" captionTargetPageId="13" captionText="FIGURES 36−46. Abdomen. Figs 36–39. Female segments VI (lower)–VIII (upper). 36, Agripialus campos sp. n., PT (CGCM 31.875); arrows pointing to the sclerotised sternites VII and VIII. 37, A. variabilis sp. n., HT. 38, Mutipialus dilatus sp. n., HT (CGCM 5.801). 39, M. monticolus sp. n., HT; arrow pointing to the membranous sternite VIII. Figs 40–42. Male abdomen (upper side, tergites; lower side, sternites). 40, Agripialus caparao sp. n., HT. 41, M. dilatus sp. n., PT (CGCM 27.435). 42, M. monticolus sp. n., PT (CGCM 29.697). Figs 43–46. Male segments VII (lower)–VIII (upper). 43, Agripialus itatiaia sp. n., HT; arrows pointing to the two plates of the sternite VII. 44, A. caparao sp. n., HT; arrows pointing to the two plates of the sternite VII. 45, Mutipialus dilatus sp. n., PT (CGCM 27.435); arrow pointing to the sternite VIII. 46, M. monticolus sp. n., PT (CGCM 29.697); arrow pointing to the sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.5224157" httpUri="https://zenodo.org/record/5224157/files/figure.png" pageId="6" pageNumber="567">38–39, 41–42, 45–46</figureCitation>
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,
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<figureCitation id="E1E32A4CEE3AFFA6FCEDEED0269BFE02" box="[874,947,438,465]" captionStart="FIGURES 47−50" captionStartId="14.[151,264,1413,1437]" captionTargetBox="[159,1421,185,1378]" captionTargetId="figure-21@14.[151,1436,181,1389]" captionTargetPageId="14" captionText="FIGURES 47−50. Male genitalia. 47, Agripialus itatiaia sp. n., HT: ventral (47a), dorsal (internal; 47b); arrows pointing to the posterior sclerotisation of the tergal lobe and pseudotegumen and the notch in the posterior margin of the saccus. 48, A. caparao sp. n., HT: ventral (48a), dorsal (internal; 48b); arrow pointing to the notch in the posterior margin of the saccus. 49, Mutipialus dilatus sp. n., PT (CGCM 27.435): lateral (49a), ventral (49b), dorsal (internal; 49c); arrow pointing to the entrance of the pocket-like saccus. 50, M. monticolus sp. n., PT (CGCM 29.697): lateral (50a), ventral (50b), dorsal (internal; 50c); arrow pointing to the entrance of the pocket-like caecum." figureDoi="http://doi.org/10.5281/zenodo.5224159" httpUri="https://zenodo.org/record/5224159/files/figure.png" pageId="6" pageNumber="567">49–50</figureCitation>
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,
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<figureCitation id="E1E32A4CEE3AFFA6FC39EED0212DFE02" box="[958,1029,438,465]" captionStart="FIGURES 51−57" captionStartId="15.[151,264,1627,1651]" captionTargetBox="[162,1427,289,1591]" captionTargetId="figure-49@15.[151,1437,272,1603]" captionTargetPageId="15" captionText="FIGURES 51−57. Female genitalia. 51, Agripialus campos sp. n., PT: posterior view and ductus and corpus bursae; arrow pointing to W-shaped lamella antevaginalis. 52, A. variabilis sp. n., HT: posterior view and ductus and corpus bursae. 53, A. itatiaia sp. n., PT (CGCM 25.532): posterior view and ductus and corpus bursae. 54, Mutipialus dilatus sp. n., HT (CGCM 5.801): posterior (54a), lateral (54b); arrow pointing to slight mesal projection of the lamella antevaginalis. 55, M. monticolus sp. n., HT: posterior (55a), lateral (55b); arrow pointing to slight mesal projection of the lamella antevaginalis. 56, Mutipialus dilatus sp. n., HT (CGCM 5.801): ductus and corpus bursae. 57, M. monticolus sp. n., HT: ductus and corpus bursae." figureDoi="http://doi.org/10.5281/zenodo.5224161" httpUri="https://zenodo.org/record/5224161/files/figure.png" pageId="6" pageNumber="567">54–57</figureCitation>
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)
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</paragraph>
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<paragraph id="796736C9EE3AFFA6FF10EE9D2602FDC0" blockId="6.[151,810,507,561]" box="[151,810,507,531]" pageId="6" pageNumber="567">
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<typeStatus id="A663886BEE3AFFA6FF10EE9D25E1FDC0" box="[151,201,507,531]" pageId="6" pageNumber="567">Type</typeStatus>
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species:
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<taxonomicName id="BED84D4AEE3AFFA6FEA1EE9D24F7FDC0" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[294,479,507,531]" class="Insecta" family="Hepialidae" genus="Mutipialus" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="567" phylum="Arthropoda" rank="species" species="dilatus" status="sp. nov.">
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<emphasis id="4BACEADBEE3AFFA6FEA1EE9D24F7FDC0" box="[294,479,507,531]" italics="true" pageId="6" pageNumber="567">Mutipialus dilatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="509F57A0EE3AFFA6FE61EE9D2710FDC1" box="[486,568,507,531]" pageId="6" pageNumber="567" rank="species">
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<emphasis id="4BACEADBEE3AFFA6FE61EE9D24D6FDC0" bold="true" box="[486,510,507,531]" pageId="6" pageNumber="567">sp</emphasis>
|
||
.
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||
<emphasis id="4BACEADBEE3AFFA6FD8DEE9D2719FDC0" bold="true" box="[522,561,507,531]" pageId="6" pageNumber="567">nov</emphasis>
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||
.
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||
</taxonomicNameLabel>
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||
by present designation.
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||
</paragraph>
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||
<paragraph id="796736C9EE3AFFA6FF10ED7F27DCFDE2" blockId="6.[151,810,507,561]" box="[151,756,537,561]" pageId="6" pageNumber="567">
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||
Included species:
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<taxonomicName id="BED84D4AEE3AFFA6FECAED7C249BFDE2" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[333,435,537,561]" class="Insecta" family="Hepialidae" genus="Mutipialus" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="567" phylum="Arthropoda" rank="species" species="dilatus" status="sp. nov.">
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<emphasis id="4BACEADBEE3AFFA6FECAED7C244AFDE2" box="[333,354,538,561]" italics="true" pageId="6" pageNumber="567">M</emphasis>
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||
.
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<emphasis id="4BACEADBEE3AFFA6FEE9ED7F249BFDE2" box="[366,435,537,561]" italics="true" pageId="6" pageNumber="567">dilatus</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="509F57A0EE3AFFA6FE3DED7F24DAFDE2" box="[442,498,537,561]" pageId="6" pageNumber="567" rank="species">
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||
<emphasis id="4BACEADBEE3AFFA6FE3DED7F24FAFDE2" bold="true" box="[442,466,537,561]" pageId="6" pageNumber="567">sp</emphasis>
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||
.
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<emphasis id="4BACEADBEE3AFFA6FE59ED7F24C4FDE2" bold="true" box="[478,492,537,561]" pageId="6" pageNumber="567">n</emphasis>
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||
.
|
||
</taxonomicNameLabel>
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||
and
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||
<taxonomicName id="BED84D4AEE3AFFA6FDA3ED7C279DFDE2" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[548,693,537,561]" class="Insecta" family="Hepialidae" genus="Mutipialus" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="567" phylum="Arthropoda" rank="species" species="monticolus" status="sp. nov.">
|
||
<emphasis id="4BACEADBEE3AFFA6FDA3ED7C2711FDE2" box="[548,569,538,561]" italics="true" pageId="6" pageNumber="567">M</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3AFFA6FDC1ED7F279DFDE2" box="[582,693,537,561]" italics="true" pageId="6" pageNumber="567">monticolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="509F57A0EE3AFFA6FD3CED7F27DCFDE2" box="[699,756,537,561]" pageId="6" pageNumber="567" rank="species">
|
||
<emphasis id="4BACEADBEE3AFFA6FD3CED7F27FBFDE2" bold="true" box="[699,723,537,561]" pageId="6" pageNumber="567">sp</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3AFFA6FD67ED7F27C6FDE2" bold="true" box="[736,750,537,561]" pageId="6" pageNumber="567">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31C26542EE3AFFA6FF10ED39264DFC49" pageId="6" pageNumber="567" type="diagnosis">
|
||
<paragraph id="796736C9EE3AFFA6FF10ED39264DFC49" blockId="6.[151,1437,607,1858]" pageId="6" pageNumber="567">
|
||
<emphasis id="4BACEADBEE3AFFA6FF10ED392422FDAA" bold="true" box="[151,266,607,633]" pageId="6" pageNumber="567">Diagnosis</emphasis>
|
||
.
|
||
<materialsCitation id="C9B03C94EE3AFFA6FE9AED392648FC49" county="South American" location="Distinguished" municipality="Also" pageId="6" pageNumber="567" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
|
||
<location id="7C076012EE3AFFA6FE9AED392491FDAA" LSID="urn:lsid:plazi:treatment:F17187DFEE3AFFA6FF10EEF421C6F891:7C076012EE3AFFA6FE9AED392491FDAA" box="[285,441,607,633]" county="South American" municipality="Also" name="Distinguished" pageId="6" pageNumber="567">Distinguished</location>
|
||
from all other
|
||
<taxonomicName id="BED84D4AEE3AFFA6FDF4ED3927C4FDAA" box="[627,748,607,633]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="567" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
|
||
by the broad dorso-posterior plates of the tergal lobe and pseudotegumen (
|
||
<figureCitation id="E1E32A4CEE3AFFA6FED4EDE524F9FD4D" box="[339,465,643,670]" captionStart="FIGURES 47−50" captionStartId="14.[151,264,1413,1437]" captionTargetBox="[159,1421,185,1378]" captionTargetId="figure-21@14.[151,1436,181,1389]" captionTargetPageId="14" captionText="FIGURES 47−50. Male genitalia. 47, Agripialus itatiaia sp. n., HT: ventral (47a), dorsal (internal; 47b); arrows pointing to the posterior sclerotisation of the tergal lobe and pseudotegumen and the notch in the posterior margin of the saccus. 48, A. caparao sp. n., HT: ventral (48a), dorsal (internal; 48b); arrow pointing to the notch in the posterior margin of the saccus. 49, Mutipialus dilatus sp. n., PT (CGCM 27.435): lateral (49a), ventral (49b), dorsal (internal; 49c); arrow pointing to the entrance of the pocket-like saccus. 50, M. monticolus sp. n., PT (CGCM 29.697): lateral (50a), ventral (50b), dorsal (internal; 50c); arrow pointing to the entrance of the pocket-like caecum." figureDoi="http://doi.org/10.5281/zenodo.5224159" httpUri="https://zenodo.org/record/5224159/files/figure.png" pageId="6" pageNumber="567">Figs 49–50</figureCitation>
|
||
) and from all other
|
||
<location id="7C076012EE3AFFA6FD23EDE2260AFD4E" LSID="urn:lsid:plazi:treatment:F17187DFEE3AFFA6FF10EEF421C6F891:7C076012EE3AFFA6FD23EDE2260AFD4E" box="[676,802,643,670]" county="South American" municipality="Also" name="New World" pageId="6" pageNumber="567">New World</location>
|
||
<taxonomicName id="BED84D4AEE3AFFA6FCA0EDE52688FD4E" box="[807,928,643,669]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="567" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
|
||
by a pocket-like caecum formed by separation of the dorsal and ventral walls of the saccus (
|
||
<figureCitation id="E1E32A4CEE3AFFA6FD17EDC12605FD11" box="[656,813,679,706]" captionStart="FIGURES 47−50" captionStartId="14.[151,264,1413,1437]" captionTargetBox="[159,1421,185,1378]" captionTargetId="figure-21@14.[151,1436,181,1389]" captionTargetPageId="14" captionText="FIGURES 47−50. Male genitalia. 47, Agripialus itatiaia sp. n., HT: ventral (47a), dorsal (internal; 47b); arrows pointing to the posterior sclerotisation of the tergal lobe and pseudotegumen and the notch in the posterior margin of the saccus. 48, A. caparao sp. n., HT: ventral (48a), dorsal (internal; 48b); arrow pointing to the notch in the posterior margin of the saccus. 49, Mutipialus dilatus sp. n., PT (CGCM 27.435): lateral (49a), ventral (49b), dorsal (internal; 49c); arrow pointing to the entrance of the pocket-like saccus. 50, M. monticolus sp. n., PT (CGCM 29.697): lateral (50a), ventral (50b), dorsal (internal; 50c); arrow pointing to the entrance of the pocket-like caecum." figureDoi="http://doi.org/10.5281/zenodo.5224159" httpUri="https://zenodo.org/record/5224159/files/figure.png" pageId="6" pageNumber="567">Figs 49a–50a</figureCitation>
|
||
).
|
||
<collectingMunicipality id="9903ACB3EE3AFFA6FCB9EDC1265BFD12" box="[830,883,679,705]" pageId="6" pageNumber="567">Also</collectingMunicipality>
|
||
distinct among other
|
||
<collectingCounty id="90064E45EE3AFFA6FBE0EDC12034FD12" box="[1127,1308,679,705]" pageId="6" pageNumber="567">South American</collectingCounty>
|
||
<taxonomicName id="BED84D4AEE3AFFA6FAA4EDC120B4FD12" box="[1315,1436,679,705]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="567" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
|
||
by the following combination of characters: i) prominent transverse scale tuft over the eyes emerging between the base of the antenna and the eye margin (
|
||
<figureCitation id="E1E32A4CEE3AFFA6FDDFED8927F2FCD9" box="[600,730,751,778]" captionStart="FIGURES 16−24" captionStartId="10.[151,264,1874,1898]" captionTargetBox="[242,1332,181,1847]" captionTargetId="figure-21@10.[227,1361,181,1850]" captionTargetPageId="10" captionText="FIGURES 16−24. Transverse scales and legs. Figs 16–19. Transverse scales (arrowed). 16, Mutipialus dilatus sp. n. PT ♂ (CGCM 23.689). 17, M. monticolus sp. n., HT ♀. 18, Parapielus luteicornis ♂ (CGCM 28.544, Argentina, Rio Grande; CGCM). 19, Hepialiscus htayaungi ♂ (CGCM 33.586, Myanmar; CGCM). Figs 20–24. Legs. 20, Agripialus campos sp. n., PT ♀ (CGCM 31.875). 21, A. variabilis sp. n., HT ♀. 22, A. itatiaia sp. n., HT ♂. 23, A. caparao sp. n., HT ♂. 24, Mutipialus dilatus sp. n., PT ♂ (CGCM 27.435)." figureDoi="http://doi.org/10.5281/zenodo.5224149" httpUri="https://zenodo.org/record/5224149/files/figure.png" pageId="6" pageNumber="567">Figs 16–17</figureCitation>
|
||
), ii) labial palpus trisegmented (
|
||
<figureCitation id="E1E32A4CEE3AFFA6FBC5ED8921BCFCD9" box="[1090,1172,751,778]" captionStart="FIGURES 16−24" captionStartId="10.[151,264,1874,1898]" captionTargetBox="[242,1332,181,1847]" captionTargetId="figure-21@10.[227,1361,181,1850]" captionTargetPageId="10" captionText="FIGURES 16−24. Transverse scales and legs. Figs 16–19. Transverse scales (arrowed). 16, Mutipialus dilatus sp. n. PT ♂ (CGCM 23.689). 17, M. monticolus sp. n., HT ♀. 18, Parapielus luteicornis ♂ (CGCM 28.544, Argentina, Rio Grande; CGCM). 19, Hepialiscus htayaungi ♂ (CGCM 33.586, Myanmar; CGCM). Figs 20–24. Legs. 20, Agripialus campos sp. n., PT ♀ (CGCM 31.875). 21, A. variabilis sp. n., HT ♀. 22, A. itatiaia sp. n., HT ♂. 23, A. caparao sp. n., HT ♂. 24, Mutipialus dilatus sp. n., PT ♂ (CGCM 27.435)." figureDoi="http://doi.org/10.5281/zenodo.5224149" httpUri="https://zenodo.org/record/5224149/files/figure.png" pageId="6" pageNumber="567">Fig. 17</figureCitation>
|
||
), iii) antenna lamellate, iv) ‘hepialine’ venation, v) forewing CuP distally connected to A vein distally (
|
||
<figureCitation id="E1E32A4CEE3AFFA6FBA0EC752184FCFD" box="[1063,1196,787,814]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="6" pageNumber="567">Figs 27–28</figureCitation>
|
||
), vi)
|
||
<specimenCount id="6FDEFD40EE3AFFA6FB6DEC7221D7FCFD" box="[1258,1279,788,814]" pageId="6" pageNumber="567" type="male">♂</specimenCount>
|
||
hindwing 1A and 2A complete (
|
||
<figureCitation id="E1E32A4CEE3AFFA6FEE1EC512492FC81" box="[358,442,823,850]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="6" pageNumber="567">Fig. 27</figureCitation>
|
||
), vii)
|
||
<specimenCount id="6FDEFD40EE3AFFA6FE7AEC5E273AFC81" box="[509,530,824,850]" pageId="6" pageNumber="567" type="female">♀</specimenCount>
|
||
hindwing 1A, 2A (
|
||
<figureCitation id="E1E32A4CEE3AFFA6FD6BEC512668FC81" box="[748,832,823,850]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="6" pageNumber="567">Fig. 28</figureCitation>
|
||
), viii)
|
||
<specimenCount id="6FDEFD40EE3AFFA6FC0CEC5E2688FC81" box="[907,928,824,850]" pageId="6" pageNumber="567" type="female">♀</specimenCount>
|
||
hindwing 1A and 2A proximate in basal half (
|
||
<figureCitation id="E1E32A4CEE3AFFA6FF27EC3D25DAFCA5" box="[160,242,859,886]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="6" pageNumber="567">Fig. 28</figureCitation>
|
||
), ix) arolium present, x) sternite VIII membranous in females (
|
||
<figureCitation id="E1E32A4CEE3AFFA6FC2CEC3D2103FCA5" box="[939,1067,859,886]" captionStart="FIGURES 31−35" captionStartId="12.[151,264,1743,1767]" captionTargetBox="[155,1434,184,1706]" captionTargetId="figure-21@12.[151,1435,181,1718]" captionTargetPageId="12" captionText="FIGURES 31−35. Female abdomen (upper side, tergites; lower side, sternites). 31, Agripialus campos sp. n., PT (CGCM 31.875). 32, A. variabilis sp. n., HT. 33, A. itatiaia sp. n., PT (CGCM 25.532). 34, Mutipialus dilatus sp. n., HT (CGCM 5.801). 35, M. monticolus sp. n., HT." figureDoi="http://doi.org/10.5281/zenodo.5224155" httpUri="https://zenodo.org/record/5224155/files/figure.png" pageId="6" pageNumber="567">Figs 34–35</figureCitation>
|
||
)and, xi) pseudotegumen unfused dorsally and fused across median ventrally (
|
||
<figureCitation id="E1E32A4CEE3AFFA6FD03EC192670FC49" box="[644,856,895,922]" captionStart="FIGURES 47−50" captionStartId="14.[151,264,1413,1437]" captionTargetBox="[159,1421,185,1378]" captionTargetId="figure-21@14.[151,1436,181,1389]" captionTargetPageId="14" captionText="FIGURES 47−50. Male genitalia. 47, Agripialus itatiaia sp. n., HT: ventral (47a), dorsal (internal; 47b); arrows pointing to the posterior sclerotisation of the tergal lobe and pseudotegumen and the notch in the posterior margin of the saccus. 48, A. caparao sp. n., HT: ventral (48a), dorsal (internal; 48b); arrow pointing to the notch in the posterior margin of the saccus. 49, Mutipialus dilatus sp. n., PT (CGCM 27.435): lateral (49a), ventral (49b), dorsal (internal; 49c); arrow pointing to the entrance of the pocket-like saccus. 50, M. monticolus sp. n., PT (CGCM 29.697): lateral (50a), ventral (50b), dorsal (internal; 50c); arrow pointing to the entrance of the pocket-like caecum." figureDoi="http://doi.org/10.5281/zenodo.5224159" httpUri="https://zenodo.org/record/5224159/files/figure.png" pageId="6" pageNumber="567">Figs 49b–c, 50b–c</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31C26542EE3AFFA6FF40ECC52144F95E" pageId="6" pageNumber="567" type="description">
|
||
<paragraph id="796736C9EE3AFFA6FF40ECC52120FC6D" blockId="6.[151,1437,607,1858]" box="[199,1032,931,958]" pageId="6" pageNumber="567">
|
||
<emphasis id="4BACEADBEE3AFFA6FF40ECC52479FC6E" bold="true" box="[199,337,931,957]" pageId="6" pageNumber="567">Description</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3AFFA6FEE7ECC224B5FC6D" bold="true" box="[352,413,932,958]" pageId="6" pageNumber="567">Male</emphasis>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3AFFA6FE2CECC52703FC6D" box="[427,555,931,958]" captionStart="FIGURES 8−12" captionStartId="7.[151,264,1940,1964]" captionTargetBox="[157,1430,181,1917]" captionTargetId="figure-20@7.[157,1430,181,1917]" captionTargetPageId="7" captionText="FIGURES 8−12. Mutipialus dilatus sp. n. Adults. 8, HT ♀: dorsal view (8a), ventral view (8b) (DZUP). 9, PT ♀ (CGCM 7.584): dorsal view (CGCM). 10, PT ♀ (CGCM 18.103): dorsal view (CGCM). 11, PT ♂ (CGCM 41.733): dorsal view (11a), ventral view (11b) (CGCM). 12, PT ♂ (CGCM 23.689): dorsal view (12a), ventral view (12b) (CGCM)." figureDoi="http://doi.org/10.5281/zenodo.5224145" httpUri="https://zenodo.org/record/5224145/files/figure.png" pageId="6" pageNumber="567">Figs 11–12</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3AFFA6FDB0ECC2277DFC6D" box="[567,597,932,958]" captionStart="FIGURES 13−15" captionStartId="9.[151,264,1548,1572]" captionTargetBox="[184,1402,345,1524]" captionTargetId="figure-107@9.[184,1402,345,1524]" captionTargetPageId="9" captionText="FIGURES 13−15. Mutipialus monticolus sp. n. Adults. 13, HT ♀: dorsal view (13a), ventral view (13b) (DZUP). 14, PT ♂ (CGCM 29.697): dorsal view (14a), ventral view (14b) (CGCM). 15, PT ♀ (CGCM 35.737): dorsal view (15a), ventral view (15b) (CGCM)." figureDoi="http://doi.org/10.5281/zenodo.5224147" httpUri="https://zenodo.org/record/5224147/files/figure.png" pageId="6" pageNumber="567">14</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3AFFA6FDE6ECC52781FC6D" box="[609,681,931,958]" captionStart="FIGURES 16−24" captionStartId="10.[151,264,1874,1898]" captionTargetBox="[242,1332,181,1847]" captionTargetId="figure-21@10.[227,1361,181,1850]" captionTargetPageId="10" captionText="FIGURES 16−24. Transverse scales and legs. Figs 16–19. Transverse scales (arrowed). 16, Mutipialus dilatus sp. n. PT ♂ (CGCM 23.689). 17, M. monticolus sp. n., HT ♀. 18, Parapielus luteicornis ♂ (CGCM 28.544, Argentina, Rio Grande; CGCM). 19, Hepialiscus htayaungi ♂ (CGCM 33.586, Myanmar; CGCM). Figs 20–24. Legs. 20, Agripialus campos sp. n., PT ♀ (CGCM 31.875). 21, A. variabilis sp. n., HT ♀. 22, A. itatiaia sp. n., HT ♂. 23, A. caparao sp. n., HT ♂. 24, Mutipialus dilatus sp. n., PT ♂ (CGCM 27.435)." figureDoi="http://doi.org/10.5281/zenodo.5224149" httpUri="https://zenodo.org/record/5224149/files/figure.png" pageId="6" pageNumber="567">16, 24</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3AFFA6FD32ECC227D5FC6D" box="[693,765,932,958]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="6" pageNumber="567">27, 30</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3AFFA6FC8EECC2268DFC6D" box="[777,933,931,958]" captionStart="FIGURES 36−46" captionStartId="13.[151,264,1743,1767]" captionTargetBox="[251,1340,182,1714]" captionTargetId="figure-20@13.[247,1340,181,1719]" captionTargetPageId="13" captionText="FIGURES 36−46. Abdomen. Figs 36–39. Female segments VI (lower)–VIII (upper). 36, Agripialus campos sp. n., PT (CGCM 31.875); arrows pointing to the sclerotised sternites VII and VIII. 37, A. variabilis sp. n., HT. 38, Mutipialus dilatus sp. n., HT (CGCM 5.801). 39, M. monticolus sp. n., HT; arrow pointing to the membranous sternite VIII. Figs 40–42. Male abdomen (upper side, tergites; lower side, sternites). 40, Agripialus caparao sp. n., HT. 41, M. dilatus sp. n., PT (CGCM 27.435). 42, M. monticolus sp. n., PT (CGCM 29.697). Figs 43–46. Male segments VII (lower)–VIII (upper). 43, Agripialus itatiaia sp. n., HT; arrows pointing to the two plates of the sternite VII. 44, A. caparao sp. n., HT; arrows pointing to the two plates of the sternite VII. 45, Mutipialus dilatus sp. n., PT (CGCM 27.435); arrow pointing to the sternite VIII. 46, M. monticolus sp. n., PT (CGCM 29.697); arrow pointing to the sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.5224157" httpUri="https://zenodo.org/record/5224157/files/figure.png" pageId="6" pageNumber="567">41–42, 45–46</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3AFFA6FC36ECC526D4FC6D" box="[945,1020,931,958]" captionStart="FIGURES 47−50" captionStartId="14.[151,264,1413,1437]" captionTargetBox="[159,1421,185,1378]" captionTargetId="figure-21@14.[151,1436,181,1389]" captionTargetPageId="14" captionText="FIGURES 47−50. Male genitalia. 47, Agripialus itatiaia sp. n., HT: ventral (47a), dorsal (internal; 47b); arrows pointing to the posterior sclerotisation of the tergal lobe and pseudotegumen and the notch in the posterior margin of the saccus. 48, A. caparao sp. n., HT: ventral (48a), dorsal (internal; 48b); arrow pointing to the notch in the posterior margin of the saccus. 49, Mutipialus dilatus sp. n., PT (CGCM 27.435): lateral (49a), ventral (49b), dorsal (internal; 49c); arrow pointing to the entrance of the pocket-like saccus. 50, M. monticolus sp. n., PT (CGCM 29.697): lateral (50a), ventral (50b), dorsal (internal; 50c); arrow pointing to the entrance of the pocket-like caecum." figureDoi="http://doi.org/10.5281/zenodo.5224159" httpUri="https://zenodo.org/record/5224159/files/figure.png" pageId="6" pageNumber="567">49–50</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3AFFA6FF40ECA12440FB9E" blockId="6.[151,1437,607,1858]" pageId="6" pageNumber="567">
|
||
<emphasis id="4BACEADBEE3AFFA6FF40ECA1242CFC32" box="[199,260,967,993]" italics="true" pageId="6" pageNumber="567">Head</emphasis>
|
||
. Clypeus glabrous, projected anteriorly, differentiated from frons. Frons with piliform and porrect scales, transverse scale tuft over eyes. Vertex scales as for frons. Eyes large, occupying 3/5 of head in anterior view. Labial palps trisegmented. Antenna lamellate, sensilla caetica sparse, sensilla trichodea present; scape and pedicel with scales as for frons.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3AFFA6FF40EB3121C3FB46" blockId="6.[151,1437,607,1858]" pageId="6" pageNumber="567">
|
||
<emphasis id="4BACEADBEE3AFFA6FF40EB312408FBA2" box="[199,288,1111,1137]" italics="true" pageId="6" pageNumber="567">Thorax.</emphasis>
|
||
Legs (
|
||
<figureCitation id="E1E32A4CEE3AFFA6FEF6EB3124E0FBA1" box="[369,456,1111,1138]" captionStart="FIGURES 16−24" captionStartId="10.[151,264,1874,1898]" captionTargetBox="[242,1332,181,1847]" captionTargetId="figure-21@10.[227,1361,181,1850]" captionTargetPageId="10" captionText="FIGURES 16−24. Transverse scales and legs. Figs 16–19. Transverse scales (arrowed). 16, Mutipialus dilatus sp. n. PT ♂ (CGCM 23.689). 17, M. monticolus sp. n., HT ♀. 18, Parapielus luteicornis ♂ (CGCM 28.544, Argentina, Rio Grande; CGCM). 19, Hepialiscus htayaungi ♂ (CGCM 33.586, Myanmar; CGCM). Figs 20–24. Legs. 20, Agripialus campos sp. n., PT ♀ (CGCM 31.875). 21, A. variabilis sp. n., HT ♀. 22, A. itatiaia sp. n., HT ♂. 23, A. caparao sp. n., HT ♂. 24, Mutipialus dilatus sp. n., PT ♂ (CGCM 27.435)." figureDoi="http://doi.org/10.5281/zenodo.5224149" httpUri="https://zenodo.org/record/5224149/files/figure.png" pageId="6" pageNumber="567">Fig. 24</figureCitation>
|
||
): epiphysis and arolium present. Venation (
|
||
<figureCitation id="E1E32A4CEE3AFFA6FC3CEB31213AFBA1" box="[955,1042,1111,1138]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="6" pageNumber="567">Fig. 27</figureCitation>
|
||
): forewing without Sc1, CuP connected to A distally; hindwing without Sc1, CuP absent, 1A and 2A complete; Sc and Rs separated.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3AFFA6FF40EBF927DFFAF6" blockId="6.[151,1437,607,1858]" pageId="6" pageNumber="567">
|
||
<emphasis id="4BACEADBEE3AFFA6FF40EBF92419FB6A" box="[199,305,1183,1209]" italics="true" pageId="6" pageNumber="567">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3AFFA6FEC6EBF92734FB69" box="[321,540,1183,1210]" captionStart="FIGURES 36−46" captionStartId="13.[151,264,1743,1767]" captionTargetBox="[251,1340,182,1714]" captionTargetId="figure-20@13.[247,1340,181,1719]" captionTargetPageId="13" captionText="FIGURES 36−46. Abdomen. Figs 36–39. Female segments VI (lower)–VIII (upper). 36, Agripialus campos sp. n., PT (CGCM 31.875); arrows pointing to the sclerotised sternites VII and VIII. 37, A. variabilis sp. n., HT. 38, Mutipialus dilatus sp. n., HT (CGCM 5.801). 39, M. monticolus sp. n., HT; arrow pointing to the membranous sternite VIII. Figs 40–42. Male abdomen (upper side, tergites; lower side, sternites). 40, Agripialus caparao sp. n., HT. 41, M. dilatus sp. n., PT (CGCM 27.435). 42, M. monticolus sp. n., PT (CGCM 29.697). Figs 43–46. Male segments VII (lower)–VIII (upper). 43, Agripialus itatiaia sp. n., HT; arrows pointing to the two plates of the sternite VII. 44, A. caparao sp. n., HT; arrows pointing to the two plates of the sternite VII. 45, Mutipialus dilatus sp. n., PT (CGCM 27.435); arrow pointing to the sternite VIII. 46, M. monticolus sp. n., PT (CGCM 29.697); arrow pointing to the sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.5224157" httpUri="https://zenodo.org/record/5224157/files/figure.png" pageId="6" pageNumber="567">Figs 41–42, 45–46</figureCitation>
|
||
). Tergosternal sclerite with tergosternal bar curving disto-anteriorly, intermediate zone weakly sclerotized with robust posterior edge angled ventrally to, but not fused with posterior margin of tergosternal bar; lateral ridge anterior of tergosternal bar very short, dorsal ridge not fused with anterior ridge of tergum I. Tergite and sternite VII and VIII sclerotized.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3AFFA6FF40EA49278BFA42" blockId="6.[151,1437,607,1858]" pageId="6" pageNumber="567">
|
||
<emphasis id="4BACEADBEE3AFFA6FF40EA492419FA9A" box="[199,305,1327,1353]" italics="true" pageId="6" pageNumber="567">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3AFFA6FEC4EA4924E2FA99" box="[323,458,1327,1354]" captionStart="FIGURES 47−50" captionStartId="14.[151,264,1413,1437]" captionTargetBox="[159,1421,185,1378]" captionTargetId="figure-21@14.[151,1436,181,1389]" captionTargetPageId="14" captionText="FIGURES 47−50. Male genitalia. 47, Agripialus itatiaia sp. n., HT: ventral (47a), dorsal (internal; 47b); arrows pointing to the posterior sclerotisation of the tergal lobe and pseudotegumen and the notch in the posterior margin of the saccus. 48, A. caparao sp. n., HT: ventral (48a), dorsal (internal; 48b); arrow pointing to the notch in the posterior margin of the saccus. 49, Mutipialus dilatus sp. n., PT (CGCM 27.435): lateral (49a), ventral (49b), dorsal (internal; 49c); arrow pointing to the entrance of the pocket-like saccus. 50, M. monticolus sp. n., PT (CGCM 29.697): lateral (50a), ventral (50b), dorsal (internal; 50c); arrow pointing to the entrance of the pocket-like caecum." figureDoi="http://doi.org/10.5281/zenodo.5224159" httpUri="https://zenodo.org/record/5224159/files/figure.png" pageId="6" pageNumber="567">Figs 49–50</figureCitation>
|
||
). Tegumen and tergal lobes fused to pseudotegumen. Saccus with dorsal and ventral walls forming an inflated space. Pseudotegumen unfused dorsally, fused ventrally; anogenital margin with lateral ridge. Fultura inferior and superior sclerotised.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3AFFA6FF40EAFA267BFA65" blockId="6.[151,1437,607,1858]" box="[199,851,1435,1462]" pageId="6" pageNumber="567">
|
||
<emphasis id="4BACEADBEE3AFFA6FF40EAFA2436FA65" bold="true" box="[199,286,1436,1462]" pageId="6" pageNumber="567">Female</emphasis>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3AFFA6FEAAEAFD2488FA65" box="[301,416,1435,1462]" captionStart="FIGURES 8−12" captionStartId="7.[151,264,1940,1964]" captionTargetBox="[157,1430,181,1917]" captionTargetId="figure-20@7.[157,1430,181,1917]" captionTargetPageId="7" captionText="FIGURES 8−12. Mutipialus dilatus sp. n. Adults. 8, HT ♀: dorsal view (8a), ventral view (8b) (DZUP). 9, PT ♀ (CGCM 7.584): dorsal view (CGCM). 10, PT ♀ (CGCM 18.103): dorsal view (CGCM). 11, PT ♂ (CGCM 41.733): dorsal view (11a), ventral view (11b) (CGCM). 12, PT ♂ (CGCM 23.689): dorsal view (12a), ventral view (12b) (CGCM)." figureDoi="http://doi.org/10.5281/zenodo.5224145" httpUri="https://zenodo.org/record/5224145/files/figure.png" pageId="6" pageNumber="567">Figs 8–10</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3AFFA6FE2BEAFA24DCFA65" box="[428,500,1435,1462]" captionStart="FIGURES 13−15" captionStartId="9.[151,264,1548,1572]" captionTargetBox="[184,1402,345,1524]" captionTargetId="figure-107@9.[184,1402,345,1524]" captionTargetPageId="9" captionText="FIGURES 13−15. Mutipialus monticolus sp. n. Adults. 13, HT ♀: dorsal view (13a), ventral view (13b) (DZUP). 14, PT ♂ (CGCM 29.697): dorsal view (14a), ventral view (14b) (CGCM). 15, PT ♀ (CGCM 35.737): dorsal view (15a), ventral view (15b) (CGCM)." figureDoi="http://doi.org/10.5281/zenodo.5224147" httpUri="https://zenodo.org/record/5224147/files/figure.png" pageId="6" pageNumber="567">13, 15</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3AFFA6FD87EAFA2736FA65" box="[512,542,1436,1462]" captionStart="FIGURES 16−24" captionStartId="10.[151,264,1874,1898]" captionTargetBox="[242,1332,181,1847]" captionTargetId="figure-21@10.[227,1361,181,1850]" captionTargetPageId="10" captionText="FIGURES 16−24. Transverse scales and legs. Figs 16–19. Transverse scales (arrowed). 16, Mutipialus dilatus sp. n. PT ♂ (CGCM 23.689). 17, M. monticolus sp. n., HT ♀. 18, Parapielus luteicornis ♂ (CGCM 28.544, Argentina, Rio Grande; CGCM). 19, Hepialiscus htayaungi ♂ (CGCM 33.586, Myanmar; CGCM). Figs 20–24. Legs. 20, Agripialus campos sp. n., PT ♀ (CGCM 31.875). 21, A. variabilis sp. n., HT ♀. 22, A. itatiaia sp. n., HT ♂. 23, A. caparao sp. n., HT ♂. 24, Mutipialus dilatus sp. n., PT ♂ (CGCM 27.435)." figureDoi="http://doi.org/10.5281/zenodo.5224149" httpUri="https://zenodo.org/record/5224149/files/figure.png" pageId="6" pageNumber="567">17</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3AFFA6FDADEAFA2760FA65" box="[554,584,1436,1462]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="6" pageNumber="567">28</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3AFFA6FDD3EAFD27B4FA65" box="[596,668,1435,1462]" captionStart="FIGURES 31−35" captionStartId="12.[151,264,1743,1767]" captionTargetBox="[155,1434,184,1706]" captionTargetId="figure-21@12.[151,1435,181,1718]" captionTargetPageId="12" captionText="FIGURES 31−35. Female abdomen (upper side, tergites; lower side, sternites). 31, Agripialus campos sp. n., PT (CGCM 31.875). 32, A. variabilis sp. n., HT. 33, A. itatiaia sp. n., PT (CGCM 25.532). 34, Mutipialus dilatus sp. n., HT (CGCM 5.801). 35, M. monticolus sp. n., HT." figureDoi="http://doi.org/10.5281/zenodo.5224155" httpUri="https://zenodo.org/record/5224155/files/figure.png" pageId="6" pageNumber="567">34–35</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3AFFA6FD2FEAFA27D9FA65" box="[680,753,1436,1462]" captionStart="FIGURES 36−46" captionStartId="13.[151,264,1743,1767]" captionTargetBox="[251,1340,182,1714]" captionTargetId="figure-20@13.[247,1340,181,1719]" captionTargetPageId="13" captionText="FIGURES 36−46. Abdomen. Figs 36–39. Female segments VI (lower)–VIII (upper). 36, Agripialus campos sp. n., PT (CGCM 31.875); arrows pointing to the sclerotised sternites VII and VIII. 37, A. variabilis sp. n., HT. 38, Mutipialus dilatus sp. n., HT (CGCM 5.801). 39, M. monticolus sp. n., HT; arrow pointing to the membranous sternite VIII. Figs 40–42. Male abdomen (upper side, tergites; lower side, sternites). 40, Agripialus caparao sp. n., HT. 41, M. dilatus sp. n., PT (CGCM 27.435). 42, M. monticolus sp. n., PT (CGCM 29.697). Figs 43–46. Male segments VII (lower)–VIII (upper). 43, Agripialus itatiaia sp. n., HT; arrows pointing to the two plates of the sternite VII. 44, A. caparao sp. n., HT; arrows pointing to the two plates of the sternite VII. 45, Mutipialus dilatus sp. n., PT (CGCM 27.435); arrow pointing to the sternite VIII. 46, M. monticolus sp. n., PT (CGCM 29.697); arrow pointing to the sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.5224157" httpUri="https://zenodo.org/record/5224157/files/figure.png" pageId="6" pageNumber="567">38–39</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3AFFA6FD7BEAFD266EFA65" box="[764,838,1435,1462]" captionStart="FIGURES 51−57" captionStartId="15.[151,264,1627,1651]" captionTargetBox="[162,1427,289,1591]" captionTargetId="figure-49@15.[151,1437,272,1603]" captionTargetPageId="15" captionText="FIGURES 51−57. Female genitalia. 51, Agripialus campos sp. n., PT: posterior view and ductus and corpus bursae; arrow pointing to W-shaped lamella antevaginalis. 52, A. variabilis sp. n., HT: posterior view and ductus and corpus bursae. 53, A. itatiaia sp. n., PT (CGCM 25.532): posterior view and ductus and corpus bursae. 54, Mutipialus dilatus sp. n., HT (CGCM 5.801): posterior (54a), lateral (54b); arrow pointing to slight mesal projection of the lamella antevaginalis. 55, M. monticolus sp. n., HT: posterior (55a), lateral (55b); arrow pointing to slight mesal projection of the lamella antevaginalis. 56, Mutipialus dilatus sp. n., HT (CGCM 5.801): ductus and corpus bursae. 57, M. monticolus sp. n., HT: ductus and corpus bursae." figureDoi="http://doi.org/10.5281/zenodo.5224161" httpUri="https://zenodo.org/record/5224161/files/figure.png" pageId="6" pageNumber="567">54–57</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3AFFA6FF40EAD924B4FA0A" blockId="6.[151,1437,607,1858]" box="[199,412,1471,1498]" pageId="6" pageNumber="567">
|
||
<emphasis id="4BACEADBEE3AFFA6FF40EAD9242CFA0A" box="[199,260,1471,1497]" italics="true" pageId="6" pageNumber="567">Head</emphasis>
|
||
. As for male.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3AFFA6FF40EA8526C6FA2E" blockId="6.[151,1437,607,1858]" box="[199,1006,1507,1534]" pageId="6" pageNumber="567">
|
||
<emphasis id="4BACEADBEE3AFFA6FF40EA852408FA2E" box="[199,288,1507,1533]" italics="true" pageId="6" pageNumber="567">Thorax.</emphasis>
|
||
Legs: as for male. Venation (
|
||
<figureCitation id="E1E32A4CEE3AFFA6FDEFEA852794FA2D" box="[616,700,1507,1534]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="6" pageNumber="567">Fig. 28</figureCitation>
|
||
): 1A, 2A and 3A complete.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3AFFA6FF40E96126C0F9F2" blockId="6.[151,1437,607,1858]" box="[199,1000,1543,1570]" pageId="6" pageNumber="567">
|
||
<emphasis id="4BACEADBEE3AFFA6FF40E9612419F9F2" box="[199,305,1543,1569]" italics="true" pageId="6" pageNumber="567">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3AFFA6FEC7E96124E9F9F1" box="[320,449,1543,1570]" captionStart="FIGURES 31−35" captionStartId="12.[151,264,1743,1767]" captionTargetBox="[155,1434,184,1706]" captionTargetId="figure-21@12.[151,1435,181,1718]" captionTargetPageId="12" captionText="FIGURES 31−35. Female abdomen (upper side, tergites; lower side, sternites). 31, Agripialus campos sp. n., PT (CGCM 31.875). 32, A. variabilis sp. n., HT. 33, A. itatiaia sp. n., PT (CGCM 25.532). 34, Mutipialus dilatus sp. n., HT (CGCM 5.801). 35, M. monticolus sp. n., HT." figureDoi="http://doi.org/10.5281/zenodo.5224155" httpUri="https://zenodo.org/record/5224155/files/figure.png" pageId="6" pageNumber="567">Figs 34–35</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3AFFA6FE4AE96E273FF9F1" box="[461,535,1544,1570]" captionStart="FIGURES 36−46" captionStartId="13.[151,264,1743,1767]" captionTargetBox="[251,1340,182,1714]" captionTargetId="figure-20@13.[247,1340,181,1719]" captionTargetPageId="13" captionText="FIGURES 36−46. Abdomen. Figs 36–39. Female segments VI (lower)–VIII (upper). 36, Agripialus campos sp. n., PT (CGCM 31.875); arrows pointing to the sclerotised sternites VII and VIII. 37, A. variabilis sp. n., HT. 38, Mutipialus dilatus sp. n., HT (CGCM 5.801). 39, M. monticolus sp. n., HT; arrow pointing to the membranous sternite VIII. Figs 40–42. Male abdomen (upper side, tergites; lower side, sternites). 40, Agripialus caparao sp. n., HT. 41, M. dilatus sp. n., PT (CGCM 27.435). 42, M. monticolus sp. n., PT (CGCM 29.697). Figs 43–46. Male segments VII (lower)–VIII (upper). 43, Agripialus itatiaia sp. n., HT; arrows pointing to the two plates of the sternite VII. 44, A. caparao sp. n., HT; arrows pointing to the two plates of the sternite VII. 45, Mutipialus dilatus sp. n., PT (CGCM 27.435); arrow pointing to the sternite VIII. 46, M. monticolus sp. n., PT (CGCM 29.697); arrow pointing to the sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.5224157" httpUri="https://zenodo.org/record/5224157/files/figure.png" pageId="6" pageNumber="567">38–39</figureCitation>
|
||
). As for male. Sternite VIII membranous.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3AFFA6FF40E94D2144F95E" blockId="6.[151,1437,607,1858]" pageId="6" pageNumber="567">
|
||
<emphasis id="4BACEADBEE3AFFA6FF40E94D2419F996" box="[199,305,1579,1605]" italics="true" pageId="6" pageNumber="567">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3AFFA6FEC5E94D24E0F995" box="[322,456,1579,1606]" captionStart="FIGURES 51−57" captionStartId="15.[151,264,1627,1651]" captionTargetBox="[162,1427,289,1591]" captionTargetId="figure-49@15.[151,1437,272,1603]" captionTargetPageId="15" captionText="FIGURES 51−57. Female genitalia. 51, Agripialus campos sp. n., PT: posterior view and ductus and corpus bursae; arrow pointing to W-shaped lamella antevaginalis. 52, A. variabilis sp. n., HT: posterior view and ductus and corpus bursae. 53, A. itatiaia sp. n., PT (CGCM 25.532): posterior view and ductus and corpus bursae. 54, Mutipialus dilatus sp. n., HT (CGCM 5.801): posterior (54a), lateral (54b); arrow pointing to slight mesal projection of the lamella antevaginalis. 55, M. monticolus sp. n., HT: posterior (55a), lateral (55b); arrow pointing to slight mesal projection of the lamella antevaginalis. 56, Mutipialus dilatus sp. n., HT (CGCM 5.801): ductus and corpus bursae. 57, M. monticolus sp. n., HT: ductus and corpus bursae." figureDoi="http://doi.org/10.5281/zenodo.5224161" httpUri="https://zenodo.org/record/5224161/files/figure.png" pageId="6" pageNumber="567">Figs 54–57</figureCitation>
|
||
). Tergite IX slightly projecting posteriorly. Lamella antevaginalis forming continuous plate without dorsal margin forming distinct medial or lateral lobes. Ductus bursae tubular, and corpus bursae similar to ductus, but of variable width, narrower or wider near junction with ductus bursae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31C26542EE3AFFA6FF40E9FE20BDF8CD" pageId="6" pageNumber="567" type="etymology">
|
||
<paragraph id="796736C9EE3AFFA6FF40E9FE275DF92A" blockId="6.[151,1437,607,1858]" pageId="6" pageNumber="567">
|
||
<emphasis id="4BACEADBEE3AFFA6FF40E9FE2460F961" bold="true" box="[199,328,1688,1714]" pageId="6" pageNumber="567">Etymology</emphasis>
|
||
.
|
||
<taxonomicName id="BED84D4AEE3AFFA6FED0E9F124F8F962" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[343,464,1687,1713]" class="Insecta" family="Hepialidae" genus="Mutipialus" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="567" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="4BACEADBEE3AFFA6FED0E9F124F8F962" box="[343,464,1687,1713]" italics="true" pageId="6" pageNumber="567">Mutipialus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="509F57A0EE3AFFA6FE50E9FE270EF961" box="[471,550,1688,1714]" pageId="6" pageNumber="567" rank="genus">
|
||
<emphasis id="4BACEADBEE3AFFA6FE50E9FE2729F961" bold="true" box="[471,513,1688,1714]" pageId="6" pageNumber="567">gen</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3AFFA6FD88E9FE2737F961" bold="true" box="[527,543,1688,1714]" pageId="6" pageNumber="567">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
comes from the Latin prefix
|
||
<emphasis id="4BACEADBEE3AFFA6FCF7E9FE268DF962" box="[880,933,1688,1713]" italics="true" pageId="6" pageNumber="567">Muti</emphasis>
|
||
- (shortened) added to -
|
||
<emphasis id="4BACEADBEE3AFFA6FB2BE9F121D8F962" box="[1196,1264,1687,1713]" italics="true" pageId="6" pageNumber="567">pialus</emphasis>
|
||
extracted from the first described
|
||
<taxonomicName id="BED84D4AEE3AFFA6FEEDE9DD24CBF906" box="[362,483,1723,1749]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="567" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
|
||
genus
|
||
<taxonomicName id="BED84D4AEE3AFFA6FDB3E9DD266CF906" authority="Fabricius. The" authorityName="Fabricius. The" authorityYear="1775" box="[564,836,1723,1749]" class="Insecta" family="Hepialidae" genus="Hepialus" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="567" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4BACEADBEE3AFFA6FDB3E9DD27B1F906" box="[564,665,1723,1749]" italics="true" pageId="6" pageNumber="567">Hepialus</emphasis>
|
||
Fabricius. The
|
||
</taxonomicName>
|
||
name refers to the shortened body appearance of the two new species. The gender is masculine.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3AFFA6FF40E86520BDF8CD" blockId="6.[151,1437,607,1858]" box="[199,1429,1795,1822]" pageId="6" pageNumber="567">
|
||
<emphasis id="4BACEADBEE3AFFA6FF40E86524D6F8CE" bold="true" box="[199,510,1795,1821]" pageId="6" pageNumber="567">Geographical distribution</emphasis>
|
||
. Known from southeastern
|
||
<collectingRegion id="BB1CF82BEE3AFFA6FCB2E86526E3F8CE" box="[821,971,1795,1821]" country="Brazil" name="Minas Gerais" pageId="6" pageNumber="567">Minas Gerais</collectingRegion>
|
||
to northeastern
|
||
<collectingRegion id="BB1CF82BEE3AFFA6FB06E862200DF8CE" box="[1153,1317,1795,1822]" country="Brazil" name="Santa Catarina" pageId="6" pageNumber="567">Santa Catarina</collectingRegion>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3AFFA6FAB3E86520A0F8CD" box="[1332,1416,1795,1822]" captionStart="FIGURE 58" captionStartId="16.[151,250,1100,1124]" captionTargetBox="[169,1335,181,1076]" captionTargetId="figure-21@16.[169,1418,181,1076]" captionTargetPageId="16" captionText="FIGURE 58. Distribution of Agripialus gen. n. spp. and Mutipialus gen. n. spp. within southern southeastern and southern Brazil and records of Agripialus gen. n. in Argentina (see remarks at the description of the genus)." figureDoi="http://doi.org/10.5281/zenodo.5224165" httpUri="https://zenodo.org/record/5224165/files/figure.png" pageId="6" pageNumber="567">Fig. 58</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31C26542EE3AFFA6FF40E84E21C6F891" box="[199,1262,1831,1858]" pageId="6" pageNumber="567" type="discussion">
|
||
<paragraph id="796736C9EE3AFFA6FF40E84E21C6F891" blockId="6.[151,1437,607,1858]" box="[199,1262,1831,1858]" pageId="6" pageNumber="567">
|
||
<emphasis id="4BACEADBEE3AFFA6FF40E84E241CF891" bold="true" box="[199,308,1832,1858]" pageId="6" pageNumber="567">Remarks</emphasis>
|
||
. All
|
||
<taxonomicName id="BED84D4AEE3AFFA6FEECE84124CCF892" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[363,484,1831,1857]" class="Insecta" family="Hepialidae" genus="Mutipialus" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="567" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="4BACEADBEE3AFFA6FEECE84124CCF892" box="[363,484,1831,1857]" italics="true" pageId="6" pageNumber="567">Mutipialus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4BACEADBEE3AFFA6FE6BE84E2713F891" bold="true" box="[492,571,1832,1858]" pageId="6" pageNumber="567">
|
||
<taxonomicNameLabel id="509F57A0EE3AFFA6FE6BE84E2713F891" box="[492,571,1832,1858]" pageId="6" pageNumber="567" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
species are monovoltine with spring and summer emergence.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |