treatments-xml/data/ED/0F/B5/ED0FB537ED745651A5A4B38F6F77279E.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.935.50492" ID-GBIF-Dataset="a280014e-e72e-4099-ba76-7a6031d1d9b4" ID-PMC="PMC7256067" ID-Pensoft-Pub="1313-2970-935-57" ID-Pensoft-UUID="E3E915D8832354FFABDD77DD100DC24E" ID-PubMed="32508502" ID-ZooBank="3540FBBB2B874908A2EF017E67FE5604" ModsDocID="1313-2970-935-57" checkinTime="1590112787856" checkinUser="pensoft" docAuthor="Padua, Diego Galvao de, Saeaeksjaervi, Ilari Eerikki, Monteiro, Ricardo Ferreira &amp; Oliveira, Marcio Luiz de" docDate="2020" docId="ED0FB537ED745651A5A4B38F6F77279E" docLanguage="en" docName="ZooKeys 935: 57-92" docOrigin="ZooKeys 935" docSource="http://dx.doi.org/10.3897/zookeys.935.50492" docTitle="Hymenoepimecis longilobus Padua &amp; Saeaeksjaervi 2020, sp. nov." docType="treatment" docUuid="8FFC81FF-D55A-4C66-A274-8EA5EAA3164F" docUuidSource="ZooBank" docVersion="4" id="E3E915D8832354FFABDD77DD100DC24E" lastPageNumber="57" masterDocId="E3E915D8832354FFABDD77DD100DC24E" masterDocTitle="Seven new species of spider-attacking Hymenoepimecis Viereck (Hymenoptera, Ichneumonidae, Pimplinae) from Ecuador, French Guiana, and Peru, with an identification key to the world species" masterLastPageNumber="92" masterPageNumber="57" pageNumber="57" updateTime="1668168881236" updateUser="ExternalLinkService">
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<mods:title>Seven new species of spider-attacking Hymenoepimecis Viereck (Hymenoptera, Ichneumonidae, Pimplinae) from Ecuador, French Guiana, and Peru, with an identification key to the world species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Padua, Diego Galvao de</mods:namePart>
<mods:affiliation>Programa de Pos-Graduacao em Entomologia, Instituto Nacional de Pesquisas da Amazonia, Av. Andre Araujo, 2936, Petropolis, 69067 - 375, Manaus, Amazonas, Brazil</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5061-2978</mods:nameIdentifier>
<mods:nameIdentifier type="email">paduadg@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Saeaeksjaervi, Ilari Eerikki</mods:namePart>
<mods:affiliation>Biodiversity Unit, Zoological Museum, University of Turku, FIN- 20014, Turku, Finland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Monteiro, Ricardo Ferreira</mods:namePart>
<mods:affiliation>Laboratorio de Ecologia de Insetos, Depto. de Ecologia, Universidade Federal do Rio de Janeiro, Av. Carlos Chagas Filho, 373, Cidade Universitaria, Ilha do Fundao, 21941 - 971, Rio de Janeiro, Rio de Janeiro, Brazil</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5137-9693</mods:nameIdentifier>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oliveira, Marcio Luiz de</mods:namePart>
<mods:affiliation>Programa de Pos-Graduacao em Entomologia, Instituto Nacional de Pesquisas da Amazonia, Av. Andre Araujo, 2936, Petropolis, 69067 - 375, Manaus, Amazonas, Brazil</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>935</mods:number>
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<mods:start>57</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.935.50492</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.935.50492</mods:identifier>
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<treatment ID-GBIF-Taxon="164248541" LSID="urn:lsid:zoobank.org:act:8FFC81FF-D55A-4C66-A274-8EA5EAA3164F" httpUri="http://treatment.plazi.org/id/ED0FB537ED745651A5A4B38F6F77279E" lastPageNumber="57" pageId="0" pageNumber="57">
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<paragraph pageId="0" pageNumber="57">
<taxonomicName LSID="ED0FB537-ED74-5651-A5A4-B38F6F77279E" authority="Pádua &amp; Sääksjärvi" authorityName="Padua &amp; Saeaeksjaervi" authorityYear="2020" class="Insecta" family="Ichneumonidae" genus="Hymenoepimecis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymenoepimecis longilobus" order="Hymenoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="longilobus" status="sp. nov.">
Hymenoepimecis longilobus 
<normalizedToken originalValue="Pádua">Padua</normalizedToken>
&amp; 
<normalizedToken originalValue="Sääksjärvi">Saeaeksjaervi</normalizedToken>
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="57">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Heads of Hymenoepimecis spp. (females), frontal view 1 H. andina sp. nov. (holotype) 2 H. bicolor 3 H. castilloi sp. nov. (holotype) 4 H. dolichocarinata sp. nov. (holotype) 5 H. duckensis 6 H. ecuatoriana sp. nov. (holotype) 7 H. kleini 8 H. longilobus sp. nov. (holotype) 9 H. manauara 10 H. neotropica 11 H. pucallpina sp. nov. (holotype) 12 H. rafaelmartinezi sp. nov. (holotype) 13 H. ribeiroi 14 H. tedfordi 15 H. uberensis." figureDoi="10.3897/zookeys.935.50492.figures1-15" httpUri="https://binary.pensoft.net/fig/412437" pageId="0" pageNumber="57">Figures 8</figureCitation>
<figureCitation captionStart="Figures 16–30" captionStartId="F2" captionText="Figures 16 - 30. Heads of Hymenoepimecis spp. (females), dorsal view 16 H. andina sp. nov. (holotype) 17 H. bicolor 18 H. castilloi sp. nov. (holotype) 19 H. dolichocarinata sp. nov. (holotype) 20 H. duckensis 21 H. ecuatoriana sp. nov. (holotype) 22 H. kleini 23 H. longilobus sp. nov. (holotype) 24 H. manauara 25 H. neotropica 26 H. pucallpina sp. nov. (holotype) 27 H. rafaelmartinezi sp. nov. (holotype) 28 H. ribeiroi 29 H. tedfordi 30 H. uberensis." figureDoi="10.3897/zookeys.935.50492.figures16-30" httpUri="https://binary.pensoft.net/fig/412438" pageId="0" pageNumber="57">, 23</figureCitation>
<figureCitation captionStart="Figures 31–45" captionStartId="F3" captionText="Figures 31 - 45. Head and part of mesosoma of Hymenoepimecis spp. (females), lateral view 31 H. andina sp. nov. (holotype) 32 H. bicolor 33 H. castilloi sp. nov. (holotype) 34 H. dolichocarinata sp. nov. (holotype), arrow shows the epicnemial carina 35 H. duckensis 36 H. ecuatoriana sp. nov. (holotype) 37 H. kleini 38 H. longilobus sp. nov. (holotype) 39 H. manauara 40 H. neotropica 41 H. pucallpina sp. nov. (holotype) 42 H. rafaelmartinezi sp. nov. (holotype) 43 H. ribeiroi 44 H. tedfordi, arrow shows the epicnemial carina 45 H. uberensis." figureDoi="10.3897/zookeys.935.50492.figures31-45" httpUri="https://binary.pensoft.net/fig/412439" pageId="0" pageNumber="57">, 38</figureCitation>
<figureCitation captionStart="Figures 46–60" captionStartId="F4" captionText="Figures 46 - 60. Sternite I of Hymenoepimecis spp. (females), lateral view 46 H. andina sp. nov. (holotype) 47 H. bicolor 48 H. castilloi sp. nov. (holotype) 49 H. dolichocarinata sp. nov. (holotype) 50 H. duckensis 51 H. ecuatoriana sp. nov. (holotype) 52 H. kleini 53 H. longilobus sp. nov. (holotype) 54 H. manauara 55 H. neotropica 56 H. pucallpina sp. nov. (holotype) 57 H. rafaelmartinezi sp. nov. (holotype) 58 H. ribeiroi 59 H. tedfordi 60 H. uberensis." figureDoi="10.3897/zookeys.935.50492.figures46-60" httpUri="https://binary.pensoft.net/fig/412440" pageId="0" pageNumber="57">, 53</figureCitation>
<figureCitation captionStart="Figures 67–71" captionStartId="F6" captionText="Figures 67 - 71. Habitus of Hymenoepimecis spp. (females), lateral view 67 H. kleini 68 H. longilobus sp. nov. (holotype) 69 H. manauara 70 H. neotropica 71 H. pucallpina sp. nov. (holotype)." figureDoi="10.3897/zookeys.935.50492.figures67-71" httpUri="https://binary.pensoft.net/fig/412442" pageId="0" pageNumber="57">, 68</figureCitation>
<figureCitation captionStart="Figures 76–90" captionStartId="F8" captionText="Figures 76 - 90. Tarsal claws of Hymenoepimecis spp. (females), lateral view 76 H. andina sp. nov. (holotype) 77 H. bicolor 78 H. castilloi sp. nov. (holotype) 79 H. dolichocarinata sp. nov. (holotype) 80 H. duckensis 81 H. ecuatoriana sp. nov. (holotype) 82 H. kleini 83 H. longilobus sp. nov. (holotype) 84 H. manauara 85 H. neotropica 86 H. pucallpina sp. nov. (holotype) 87 H. rafaelmartinezi sp. nov. (holotype) 88 H. ribeiroi 89 H. tedfordi 90 H. uberensis." figureDoi="10.3897/zookeys.935.50492.figures76-90" httpUri="https://binary.pensoft.net/fig/412444" pageId="0" pageNumber="57">, 83</figureCitation>
<figureCitation captionStart="Figures 97–102" captionStartId="F10" captionText="Figures 97 - 102. Habitus of Hymenoepimecis spp. (males), lateral view 97 H. andina sp. nov. (paratype) 98 H. bicolor 99 H. dolichocarinata sp. nov. (paratype) 100 H. duckensis 101 H. kleini 102 H. longilobus sp. nov. (paratype)." figureDoi="10.3897/zookeys.935.50492.figures97-102" httpUri="https://binary.pensoft.net/fig/412446" pageId="0" pageNumber="57">, 102</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="diagnosis">
<paragraph pageId="0" pageNumber="57">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="57">
This species can be distinguished from all other 
<taxonomicName authorityName="Viereck" authorityYear="1912" class="Insecta" family="Ichneumonidae" genus="Hymenoepimecis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymenoepimecis" order="Hymenoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Hymenoepimecis</emphasis>
</taxonomicName>
by the combination of the following characters: 1) fore wing hyaline yellowish, with two blackish bands; 2) pronotum orange (Peruvian specimens with anterior margin of the pronotum blackish); 3) female with tarsal claw with a longitudinally elongated lobe, and with apex overtaking the lobe; 4) female with ovipositor 1.5-1.6 times as long as hind tibia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="description">
<paragraph pageId="0" pageNumber="57">Description.</paragraph>
<paragraph pageId="0" pageNumber="57">
<emphasis bold="true" pageId="0" pageNumber="57">Female.</emphasis>
Body approx. [15.0] 12.0-15.0 mm; face [1.1] 1.0-1.1 times as broad as high, smooth, slightly convex with few spaced bristles; head in dorsal view, with gena strongly narrowed behind eyes; posterior ocelli separated from eyes by approx. [0.9] 0.8-1.0 times its own maximum diameter; occipital carina projected and curved upwards dorsally. Pronotum long, smooth and polished, with distance from tegula to head greater than approx. [0.7] 0.6-0.7 times distance from tegula to hind margin of propodeum, and in anterior part with opening pocket-like structure not reduced longitudinally; mesoscutum smooth and polished; scutellum, in profile, convex; mesopleuron smooth and polished, with anterodorsal and posterodorsal parts bearing sparse, fine setiferous punctures; metapleuron smooth and polished, rather uniformly covered with sparse, fine setiferous punctures; propodeum smooth, polished, with sparse, fine setiferous punctures and with lateral longitudinal carina present only posteriorly. Fore wing approx. [13.0] 9.0-13.0 mm; cu-a interstitial to the base of Rs&amp;M; 2rs-m [0.55] 0.5-0.6 times as long as abscissa of M between 2rs-m and 2m-cu; hind wing with abscissa of Cu1 meeting cu-a equidistant between M and 1A. Hind leg with tibia + tarsus [0.6] times the fore wing length; tarsal claw with longitudinally elongated lobe, with apex of claw overtaking the lobe. Metasoma slender; tergite I [1.6] times as long as posteriorly width, centrally quite strongly convex with lateral carinae present only at extreme anterior end flanking the anterior concavity; sternite I with a low, rounded swelling posteriorly; tergite II approx. [1.2] 1.2-1.4 times as long as posteriorly width; tergites III and IV approx. [1.1] 1.1-1.3 times as long as posteriorly width; ovipositor approx. [1.6] 1.5-1.6 times as long as hind tibia.
</paragraph>
<caption doi="10.3897/zookeys.935.50492.figures103-106" httpUri="https://binary.pensoft.net/fig/412447" pageId="0" pageNumber="57" start="Figures 103–106" startId="F11">
<paragraph pageId="0" pageNumber="57">
<emphasis bold="true" pageId="0" pageNumber="57">Figures 103-106.</emphasis>
Habitus of 
<taxonomicName authorityName="Viereck" authorityYear="1912" class="Insecta" family="Ichneumonidae" genus="Hymenoepimecis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymenoepimecis" order="Hymenoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="57">Hymenoepimecis</emphasis>
</taxonomicName>
spp. (males), lateral view 
<emphasis bold="true" pageId="0" pageNumber="57">103</emphasis>
<taxonomicName lsidName="H. manauara" pageId="0" pageNumber="57" rank="species" species="manauara">
<emphasis italics="true" pageId="0" pageNumber="57">H. manauara</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="57">104</emphasis>
<taxonomicName lsidName="H. rafaelmartinezi" pageId="0" pageNumber="57" rank="species" species="rafaelmartinezi">
<emphasis italics="true" pageId="0" pageNumber="57">H. rafaelmartinezi</emphasis>
</taxonomicName>
sp. nov. (paratype) 
<emphasis bold="true" pageId="0" pageNumber="57">105</emphasis>
<taxonomicName lsidName="H. ribeiroi" pageId="0" pageNumber="57" rank="species" species="ribeiroi">
<emphasis italics="true" pageId="0" pageNumber="57">H. ribeiroi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="57">106</emphasis>
<taxonomicName lsidName="H. uberensis" pageId="0" pageNumber="57" rank="species" species="uberensis">
<emphasis italics="true" pageId="0" pageNumber="57">H. uberensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="57">
<emphasis bold="true" italics="true" pageId="0" pageNumber="57">Colour.</emphasis>
Head black; clypeus black with apex yellowish; mouthparts yellowish, with apex mandible black; antenna brownish. Mesosoma orange. Fore and mid leg orange, the hind entirely blackish brown except base of coxa orange. Fore wing hyaline yellowish, with apex blackish and with a blackish preapical band; pterostigma with basal half black and apical half yellow; hind wing with slightly blackish band in median part. Metasoma orange, with posterior margin black in tergite V and tergites VI+ black; ovipositor and sheath brownish.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="male">
<paragraph pageId="0" pageNumber="57">Male.</paragraph>
<paragraph pageId="0" pageNumber="57">
(Fig. 
<figureCitation captionStart="Figures 97–102" captionStartId="F10" captionText="Figures 97 - 102. Habitus of Hymenoepimecis spp. (males), lateral view 97 H. andina sp. nov. (paratype) 98 H. bicolor 99 H. dolichocarinata sp. nov. (paratype) 100 H. duckensis 101 H. kleini 102 H. longilobus sp. nov. (paratype)." figureDoi="10.3897/zookeys.935.50492.figures97-102" httpUri="https://binary.pensoft.net/fig/412446" pageId="0" pageNumber="57">102</figureCitation>
). Similar to female in structure and colouration, but with hind coxa orange body with 10.5-12.0 mm; face 1.0 times as broad as high; posterior ocelli separated from eyes by 0.8-1.0 times its own maximum diameter. Fore wing 8.0-10.0 mm; 2rs-m 0.3-0.5 times as long as abscissa of M between 2rs-m and 2m-cu. Tarsal claw simple. Metasoma slender; tergite I 1.6-1.7 times as long as posteriorly width; tergite II 1.2-1.5 times as long as posteriorly width; tergites III and IV 1.0-1.3 times as long as posteriorly width.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="variation">
<paragraph pageId="0" pageNumber="57">Variation.</paragraph>
<paragraph pageId="0" pageNumber="57">The Peruvian specimens have anterior margin of pronotum blackish. Some specimens have tergite V entirely black or black with anterior margin orange; hind coxa orange with basal part blackish.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="distribution">
<paragraph pageId="0" pageNumber="57">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="57">
Ecuador**, French Guiana* and Peru* (Fig. 
<figureCitation captionStart="Figures 107–112" captionStartId="F12" captionText="Figures 107 - 112. Geographic distribution of the Hymenoepimecis species in this study." figureDoi="10.3897/zookeys.935.50492.figures107-112" httpUri="https://binary.pensoft.net/fig/412448" pageId="0" pageNumber="57">110</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="notes">
<paragraph pageId="0" pageNumber="57">Biological notes.</paragraph>
<paragraph pageId="0" pageNumber="57">Host unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="etymology">
<paragraph pageId="0" pageNumber="57">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="57">The specific name refers the longitudinally elongated lobe in tarsal claw of the females.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="type material">
<paragraph pageId="0" pageNumber="57">Type material.</paragraph>
<paragraph pageId="0" pageNumber="57">
<emphasis bold="true" italics="true" pageId="0" pageNumber="57">Holotype</emphasis>
♀. French Guiana, M. de Kaw, Patawa, ix.2003, (J. Cerda leg.), ZMUT. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="57">Paratypes</emphasis>
: French Guiana: idem holotype, but vii.2003, 1♀, ZMUT; idem, but xi.2001, 1♂ and 1♀, ZMUT; idem, but xii.2001, 1♀, ZMUT; idem, but ii.2003 (O. Morvan leg.), 1♂, ZMUT. Ecuador: Dept. Orellana, Onkonegare, 
<geoCoordinate degrees="00" direction="south" minutes="39" orientation="latitude" precision="1" seconds="25,7" value="-0.6571389">00°39'25,7&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="76" direction="west" minutes="27" orientation="longitude" precision="1" seconds="10,8" value="-76.452995">76°27'10,8&quot;W</geoCoordinate>
, a.s.l.: 216 m., 23.vi.1996, Fogging, Lot #1616 (T.L. Erwin leg.), 1♂, ZMUT. Peru: Dept. of Loreto, Iquitos area, Allpahuayo, 22.ii-7.iii.2000, white sand, Malaise trap (
<normalizedToken originalValue="Sääksjärvi">Saeaeksjaervi</normalizedToken>
et al. leg.), APHI, E1/2, 1♀, ZMUT; idem, but 14.ix-4.x.2000, clay, APHI, H1/13, 1♀, ZMUT; idem, but G1(15), 8.xi.2000, 1♀, ZMUT.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="comments">
<paragraph pageId="0" pageNumber="57">Comments.</paragraph>
<paragraph pageId="0" pageNumber="57">
<taxonomicName authorityName="Padua &amp; Saeaeksjaervi" authorityYear="2020" class="Insecta" family="Ichneumonidae" genus="Hymenoepimecis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymenoepimecis longilobus" order="Hymenoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="longilobus">
<emphasis italics="true" pageId="0" pageNumber="57">Hymenoepimecis longilobus</emphasis>
</taxonomicName>
sp. nov. closely resembles 
<taxonomicName lsidName="H. neotropica" pageId="0" pageNumber="57" rank="species" species="neotropica">
<emphasis italics="true" pageId="0" pageNumber="57">H. neotropica</emphasis>
</taxonomicName>
(Brues &amp; Richardson, 1913), 
<taxonomicName authorityName="Padua &amp; Saeaeksjaervi" authorityYear="2020" class="Insecta" family="Ichneumonidae" genus="Hymenoepimecis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymenoepimecis ecuatoriana" order="Hymenoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="ecuatoriana">
<emphasis italics="true" pageId="0" pageNumber="57">Hymenoepimecis ecuatoriana</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="H. duckensis" pageId="0" pageNumber="57" rank="species" species="duckensis">
<emphasis italics="true" pageId="0" pageNumber="57">H. duckensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Pádua">Padua</normalizedToken>
&amp; Onody, 2015 mainly by having the fore wing yellowish hyaline with two blackish bands and metasoma orange with last tergites black. It differs from all of these species by having tarsal claw with a longitudinally elongated lobe (tarsal claw with a preapical tooth in 
<taxonomicName lsidName="H. neotropica" pageId="0" pageNumber="57" rank="species" species="neotropica">
<emphasis italics="true" pageId="0" pageNumber="57">H. neotropica</emphasis>
</taxonomicName>
and a more or less square lobe in 
<taxonomicName lsidName="H. ecuatoriana" pageId="0" pageNumber="57" rank="species" species="ecuatoriana">
<emphasis italics="true" pageId="0" pageNumber="57">H. ecuatoriana</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="H. duckensis" pageId="0" pageNumber="57" rank="species" species="duckensis">
<emphasis italics="true" pageId="0" pageNumber="57">H. duckensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
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