480 lines
59 KiB
XML
480 lines
59 KiB
XML
<document id="6A83876EB17A78BF3CBBD25C027E9F78" ID="10.11646/zootaxa.4273.3.4SLASH11253" ID-CLB-Dataset="32904" ID-DOI="10.11646/zootaxa.4273.3.4" ID-GBIF-Dataset="c029f78e-c026-48ac-8c61-8b168fef59ea" ID-ISSN="1175-5326" ID-Zenodo-Dep="803109" ID-ZooBank="ADC167F6-C3A0-43EC-8636-F65F5ABE85AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1496736238170" checkinUser="plazi" docAuthor="Rodriguez, Lily O. & Catenazzi, Alessandro" docDate="2017" docId="ED0DF709FFBB75409EB477D196ACFBA1" docLanguage="en" docName="zootaxa.4273.3.4SLASH11253.pdf" docOrigin="Zootaxa 4273 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phrynopus anancites Rodriguez & Catenazzi, 2017, new species" docType="treatment" docVersion="8" lastPageNumber="389" masterDocId="11348F71FFBE75489E23770F9474FFF9" masterDocTitle="Four new species of terrestrial-breeding frogs of the genus Phrynopus (Anura: Terrarana: Craugastoridae) from Río Abiseo National Park, Peru" masterLastPageNumber="406" masterPageNumber="381" pageNumber="386" updateTime="1698440199461" updateUser="ExternalLinkService">
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<mods:title id="963A350D8538D7E3C286C847AEB2F3CC">Four new species of terrestrial-breeding frogs of the genus Phrynopus (Anura: Terrarana: Craugastoridae) from Río Abiseo National Park, Peru</mods:title>
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<mods:namePart id="BC992B365F71517E22AD4693A39216B4">Rodriguez, Lily O.</mods:namePart>
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<mods:namePart id="C0A225BEA788E70514D3F0445231A846">Catenazzi, Alessandro</mods:namePart>
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<mods:title id="9EE0B134CCB888CFC8C3055A4C04AAE1">Zootaxa</mods:title>
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<mods:date id="FBC7B27773C382D073304ECF646CF2FE">2017</mods:date>
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<treatment id="ED0DF709FFBB75409EB477D196ACFBA1" ID-DOI="http://doi.org/10.5281/zenodo.6033266" ID-GBIF-Taxon="131812330" ID-Zenodo-Dep="6033266" LSID="urn:lsid:plazi:treatment:ED0DF709FFBB75409EB477D196ACFBA1" httpUri="http://treatment.plazi.org/id/ED0DF709FFBB75409EB477D196ACFBA1" lastPageId="8" lastPageNumber="389" pageId="5" pageNumber="386">
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<subSubSection id="2DBE1594FFBB754D9EB477D1952BFEE3" pageId="5" pageNumber="386" type="nomenclature">
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<paragraph id="651B461FFFBB754D9EB477D19659FF01" blockId="5.[151,557,222,282]" box="[151,557,222,248]" pageId="5" pageNumber="386">
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<heading id="3E53F173FFBB754D9EB477D19659FF01" bold="true" box="[151,557,222,248]" fontSize="11" level="1" pageId="5" pageNumber="386" reason="1">
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<emphasis id="57D09A0DFFBB754D9EB477D19659FF01" bold="true" box="[151,557,222,248]" pageId="5" pageNumber="386">
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<taxonomicName id="A2A43D9CFFBB754D9EB477D195E2FF01" box="[151,406,222,248]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="anancites" status="sp. nov.">
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<emphasis id="57D09A0DFFBB754D9EB477D195E2FF01" bold="true" box="[151,406,222,248]" italics="true" pageId="5" pageNumber="386">Phrynopus anancites</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="4CE32776FFBB754D9FBE77D19659FF01" box="[413,557,222,248]" pageId="5" pageNumber="386" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="651B461FFFBB754D9EB4760E952BFEE3" blockId="5.[151,557,222,282]" box="[151,351,257,282]" pageId="5" pageNumber="386">
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(
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<figureCitation id="FD9F5A9AFFBB754D9EBD760E957FFEE0" box="[158,267,257,282]" captionStart="FIGURE 4" captionStartId="6.[151,250,1924,1946]" captionTargetBox="[487,1072,596,1899]" captionTargetId="figure@6.[487,1099,589,1902]" captionTargetPageId="6" captionText="FIGURE 4. Holotype of Phrynopus anancites sp. nov., male MUSM 33168 (SVL = 25.3 mm), showing (A) head in lateral view and (B, C) body in dorsal and ventral views. Specimen collected at Ventanas, 3820 m, near the entrance of Río Abiseo National Park, San Martín, Peru. Photographs by Morris Flecks." httpUri="https://zenodo.org/record/803117/files/figure.png" pageId="5" pageNumber="386">Figures 4</figureCitation>
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,
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<figureCitation id="FD9F5A9AFFBB754D9F34760E9553FEE3" box="[279,295,257,282]" captionStart="FIGURE 5" captionStartId="7.[151,250,834,856]" captionTargetBox="[342,1249,220,784]" captionTargetId="figure@7.[342,1249,220,784]" captionTargetPageId="7" captionText="FIGURE 5. Holotype of Phrynopus anancites sp. nov., male MUSM 33168 (SVL = 25.3 mm), in life. Specimen from Ventanas, 3820 m, near the entrance of Río Abiseo National Park, San Martín, Peru. Photograph by A. Catenazzi." httpUri="https://zenodo.org/record/803119/files/figure.png" pageId="5" pageNumber="386">5</figureCitation>
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,
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<figureCitation id="FD9F5A9AFFBB754D9F10760E9535FEE3" box="[307,321,257,282]" captionStart="FIGURE 6" captionStartId="7.[151,250,1854,1876]" captionTargetBox="[165,1411,981,1831]" captionTargetId="figure@7.[151,1436,972,1832]" captionTargetPageId="7" captionText="FIGURE 6. Palmar and plantar surfaces of the new species of Phrynopus: (A) P. anancites sp. nov. (MUSM 33168), (B) P. capitalis sp. nov. (AMNH 134158), (C) P. dumicola sp. nov. (MUSM 33102), and (D) P. personatus sp. nov. (MUSM 33096). Photographs by M. Flecks (A, C and D) and A. Catenazzi (B)." httpUri="https://zenodo.org/record/803121/files/figure.png" pageId="5" pageNumber="386">6</figureCitation>
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A)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="2DBE1594FFBB754D9EB476469604FE50" pageId="5" pageNumber="386" type="materials_examined">
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<paragraph id="651B461FFFBB754D9EB476469604FE50" blockId="5.[151,1437,329,2010]" pageId="5" pageNumber="386">
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<materialsCitation id="D5CC4C42FFBB754D9EB476469604FE50" ID-GBIF-Occurrence="1563404868" collectingDate="1999-06-29" collectionCode="MUSM" collectorName="Distrito Parcoy & Provincia Pataz & Departamento & A. Catenazzi & L. O. Rodriguez" country="Peru" county="Figure" elevation="3820" latitude="-8.031539" location="Figures" longLatPrecision="1" longitude="-77.4078" municipality="Region" pageId="5" pageNumber="386" specimenCode="MUSM 33168" specimenCount="1" stateProvince="La Libertad" typeStatus="holotype">
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<emphasis id="57D09A0DFFBB754D9EB47646957FFE9B" bold="true" box="[151,267,329,354]" pageId="5" pageNumber="386">
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<typeStatus id="BA1FF8BDFFBB754D9EB476469572FE9B" box="[151,262,329,354]" pageId="5" pageNumber="386" type="holotype">Holotype</typeStatus>
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||
.
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||
</emphasis>
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<specimenCode id="3502EE64FFBB754D9F31764695CDFE9B" box="[274,441,329,354]" collectionCode="MUSM" pageId="5" pageNumber="386">
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<collectionCode id="03B5DEDAFFBB754D9F317646951CFE98" box="[274,360,329,353]" pageId="5" pageNumber="386">MUSM</collectionCode>
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||
33168
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</specimenCode>
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||
, an adult male (
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<figureCitation id="FD9F5A9AFFBB754D9C4F764696AEFE98" box="[620,730,329,354]" captionStart="FIGURE 4" captionStartId="6.[151,250,1924,1946]" captionTargetBox="[487,1072,596,1899]" captionTargetId="figure@6.[487,1099,589,1902]" captionTargetPageId="6" captionText="FIGURE 4. Holotype of Phrynopus anancites sp. nov., male MUSM 33168 (SVL = 25.3 mm), showing (A) head in lateral view and (B, C) body in dorsal and ventral views. Specimen collected at Ventanas, 3820 m, near the entrance of Río Abiseo National Park, San Martín, Peru. Photographs by Morris Flecks." httpUri="https://zenodo.org/record/803117/files/figure.png" pageId="5" pageNumber="386">
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<location id="607B10C4FFBB754D9C4F764696B6FE9B" LSID="urn:lsid:plazi:treatment:ED0DF709FFBB75409EB477D196ACFBA1:607B10C4FFBB754D9C4F764696B6FE9B" box="[620,706,329,354]" country="Peru" county="Figure" latitude="-8.031539" longLatPrecision="1" longitude="-77.4078" municipality="Region" name="Figures" pageId="5" pageNumber="386" stateProvince="La Libertad">Figures</location>
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4
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</figureCitation>
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,
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<figureCitation id="FD9F5A9AFFBB754D9CC476469683FE9B" box="[743,759,329,354]" captionStart="FIGURE 5" captionStartId="7.[151,250,834,856]" captionTargetBox="[342,1249,220,784]" captionTargetId="figure@7.[342,1249,220,784]" captionTargetPageId="7" captionText="FIGURE 5. Holotype of Phrynopus anancites sp. nov., male MUSM 33168 (SVL = 25.3 mm), in life. Specimen from Ventanas, 3820 m, near the entrance of Río Abiseo National Park, San Martín, Peru. Photograph by A. Catenazzi." httpUri="https://zenodo.org/record/803119/files/figure.png" pageId="5" pageNumber="386">5</figureCitation>
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,
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<figureCitation id="FD9F5A9AFFBB754D9D2076469765FE9B" box="[771,785,329,354]" captionStart="FIGURE 6" captionStartId="7.[151,250,1854,1876]" captionTargetBox="[165,1411,981,1831]" captionTargetId="figure@7.[151,1436,972,1832]" captionTargetPageId="7" captionText="FIGURE 6. Palmar and plantar surfaces of the new species of Phrynopus: (A) P. anancites sp. nov. (MUSM 33168), (B) P. capitalis sp. nov. (AMNH 134158), (C) P. dumicola sp. nov. (MUSM 33102), and (D) P. personatus sp. nov. (MUSM 33096). Photographs by M. Flecks (A, C and D) and A. Catenazzi (B)." httpUri="https://zenodo.org/record/803121/files/figure.png" pageId="5" pageNumber="386">6</figureCitation>
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A) collected at
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<location id="607B10C4FFBB754D9D9C76459050FE98" LSID="urn:lsid:plazi:treatment:ED0DF709FFBB75409EB477D196ACFBA1:607B10C4FFBB754D9D9C76459050FE98" box="[959,1060,330,353]" country="Peru" county="Figure" latitude="-8.031539" longLatPrecision="1" longitude="-77.4078" municipality="Region" name="Ventanas" pageId="5" pageNumber="386" stateProvince="La Libertad">Ventanas</location>
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||
(
|
||
<geoCoordinate id="009020D8FFBB754D9A16764690A6FE9B" box="[1077,1234,329,354]" direction="south" orientation="latitude" pageId="5" pageNumber="386" precision="1" value="-8.031539">8°01´53.54”S</geoCoordinate>
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,
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<geoCoordinate id="009020D8FFBB754D9AFD764691ECFE98" box="[1246,1432,329,354]" direction="west" orientation="longitude" pageId="5" pageNumber="386" precision="1" value="-77.4078">77°24´28.06” W</geoCoordinate>
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,
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<quantity id="A25CEBFAFFBB754D9EB476619480FE7F" box="[151,244,366,390]" metricMagnitude="3" metricUnit="m" metricValue="3.82" pageId="5" pageNumber="386" unit="m" value="3820.0">
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<elevation id="EE89A12CFFBB754D9EB476619480FE7F" box="[151,244,366,390]" metricMagnitude="3" metricUnit="m" metricValue="3.82" pageId="5" pageNumber="386" unit="m" value="3820.0">3820 m</elevation>
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</quantity>
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),
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<collectorName id="C85123C9FFBB754D9F2F766295C9FE7F" box="[268,445,365,390]" pageId="5" pageNumber="386">Distrito Parcoy</collectorName>
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,
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<collectorName id="C85123C9FFBB754D9FEE766296F7FE7C" box="[461,643,365,390]" pageId="5" pageNumber="386">Provincia Pataz</collectorName>
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,
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<collectorName id="C85123C9FFBB754D9CB076619746FE7C" box="[659,818,366,389]" pageId="5" pageNumber="386">Departamento</collectorName>
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(
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<collectingMunicipality id="857FDC65FFBB754D9D66766297EDFE7F" box="[837,921,365,390]" pageId="5" pageNumber="386">Región</collectingMunicipality>
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)
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<collectingRegion id="A76088FDFFBB754D9D8E7661904CFE7F" box="[941,1080,365,390]" country="Peru" name="La Libertad" pageId="5" pageNumber="386">La Libertad</collectingRegion>
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,
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<collectingCountry id="1DB3068FFFBB754D9A6B76619008FE7C" box="[1096,1148,366,389]" name="Peru" pageId="5" pageNumber="386">Peru</collectingCountry>
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on
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<date id="111A60DFFFBB754D9A8C76619139FE7F" box="[1199,1357,366,390]" pageId="5" pageNumber="386" value="1999-06-29">
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<collectingDate id="015E9937FFBB754D9A8C76619139FE7F" box="[1199,1357,366,390]" pageId="5" pageNumber="386" value="1999-06-29">29 June 1999</collectingDate>
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||
</date>
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||
by
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||
<collectorName id="C85123C9FFBB754D9BA276619571FE53" pageId="5" pageNumber="386">A. Catenazzi</collectorName>
|
||
and
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||
<collectorName id="C85123C9FFBB754D9F1F769D9586FE53" box="[316,498,401,426]" pageId="5" pageNumber="386">L. O. Rodriguez</collectorName>
|
||
(
|
||
<figureCitation id="FD9F5A9AFFBB754D9C23769E962BFE50" box="[512,607,401,426]" captionStart="FIGURE 1" captionStartId="2.[151,250,1830,1852]" captionTargetBox="[189,1411,235,1774]" captionTargetId="figure@2.[175,1427,193,1813]" captionTargetPageId="2" captionText="FIGURE 1. Map of the Andes of central and northern Peru, with type localities of new species Phrynopus capitalis sp. nov., P. dumicola sp. nov., P. personatus sp. nov. (star) and P. anancites sp. nov. (asterisk) and of congeneric forms: P. valquii (asterisk), P. thompsoni (1), P. daemon (2), P. lechriorhynchus and P. vestigiatus (3), P. horstpauli (4), P. kauneorum and P. kotosh (5), P. dagmarae and P. interstinctus (6), P. heimorum (7), P. barthlenae and P. tautzorum (8), P. miroslawae and P. nicoleae (9), P. badius and P. curator (10), P. auriculatus, P. bracki and P. tribulosus (11), P. pesantesi (12), P. bufoides and P. paucari (13), P. juninensis and P. montium (14), P. oblivious and P. peruanus (15), and P. chaparroi (16)." httpUri="https://zenodo.org/record/803111/files/figure.png" pageId="5" pageNumber="386">
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||
<collectingCounty id="8C7A3E93FFBB754D9C23769E963FFE53" box="[512,587,401,426]" pageId="5" pageNumber="386">Figure</collectingCounty>
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1
|
||
</figureCitation>
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||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2DBE1594FFBB754E9EE476BA95B1FDE1" lastPageId="6" lastPageNumber="387" pageId="5" pageNumber="386" type="diagnosis">
|
||
<paragraph id="651B461FFFBB754D9EE476BA96F6FCCF" blockId="5.[151,1437,329,2010]" pageId="5" pageNumber="386">
|
||
<emphasis id="57D09A0DFFBB754D9EE476BA9535FE37" bold="true" box="[199,321,437,462]" pageId="5" pageNumber="386">Diagnosis.</emphasis>
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A medium size species of
|
||
<taxonomicName id="A2A43D9CFFBB754D9CA576BA968BFE37" box="[646,767,437,462]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="genus">
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||
<emphasis id="57D09A0DFFBB754D9CA576BA968BFE37" box="[646,767,437,462]" italics="true" pageId="5" pageNumber="386">Phrynopus</emphasis>
|
||
</taxonomicName>
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||
characterized by (1) a coarsely aerolate thick dorsal and ventral skin; thoracic fold present, discoidal fold absent; (2) tympanic membrane and tympanic annulus absent, well defined supratympanic fold; (3) snout short, rounded in dorsal view, slightly blunt in profile; (4) upper eyelid without tubercles; width of upper eyelid narrower than IOD, cranial crests absent; (5) dentigerous processes of vomers minute; (6) male lacking nuptial pads, vocal sac and slits; (7) Finger I as long as Finger II; thenar tubercle ovoid, 1.5 times larger than outer palmar tubercle which is hart-shaped, tip of digits rounded, bulbous, (8) fingers without lateral fringes, (9) ulnar tubercle absent, minute tarsal tubercle; (10) heels lacking tubercles, inner tarsal fold absent; (11) inner metatarsal tubercle ovoid; outer metatarsal tubercle slightly smaller, round, protruding; (12) no fringes or webbing between toes, Toe V slightly longer than III; (13) in life, dorsum bluish-olive-green, venter gray, iris yellow with inner orange ring; in preservative, uniform dark brown, lighter brown on upper ventral surfaces; (14) SVL
|
||
<quantity id="A25CEBFAFFBB754D9F50741295A3FCCC" box="[371,471,797,822]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="5" pageNumber="386" unit="mm" value="25.3">25.3 mm</quantity>
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in single male.
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||
</paragraph>
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||
<paragraph id="651B461FFFBB754E9EE4744D95B1FDE1" blockId="5.[151,1437,329,2010]" lastBlockId="6.[151,1437,151,536]" lastPageId="6" lastPageNumber="387" pageId="5" pageNumber="386">
|
||
No other species of
|
||
<taxonomicName id="A2A43D9CFFBB754D9F84744E9654FCA3" box="[423,544,833,858]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="genus">
|
||
<emphasis id="57D09A0DFFBB754D9F84744E9654FCA3" box="[423,544,833,858]" italics="true" pageId="5" pageNumber="386">Phrynopus</emphasis>
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||
</taxonomicName>
|
||
has the combination of lack of tympanun, distinctive supratympanic fold, short snout, small vomerine teeth, and coarsely areolated skin with uniform dorsal and ventral coloration. Of the species lacking a visible tympanum and bearing dentigerous processes of vomers (character condition for
|
||
<taxonomicName id="A2A43D9CFFBB754D9AD674859109FC58" box="[1269,1405,906,929]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="anancites">
|
||
<emphasis id="57D09A0DFFBB754D9AD674859109FC58" box="[1269,1405,906,929]" italics="true" pageId="5" pageNumber="386">P. anancites</emphasis>
|
||
</taxonomicName>
|
||
in parentheses),
|
||
<emphasis id="57D09A0DFFBB754D9F1374A1970DFC3F" box="[304,889,941,966]" italics="true" pageId="5" pageNumber="386">
|
||
<taxonomicName id="A2A43D9CFFBB754D9F1374A195E1FC3F" box="[304,405,941,966]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="bracki">P. bracki</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFBB754D9F8374A19644FC3F" box="[416,560,941,966]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="dagmarae">P. dagmarae</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFBB754D9C1974A19699FC3C" box="[570,749,942,965]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="insterstinctus">P. insterstinctus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFBB754D9CDB74A19701FC3F" box="[760,885,941,966]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="nicoleae">P. nicoleae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="A2A43D9CFFBB754D9D8D74A19035FC3C" box="[942,1089,942,965]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="vestigiatus">
|
||
<emphasis id="57D09A0DFFBB754D9D8D74A19035FC3C" box="[942,1089,942,965]" italics="true" pageId="5" pageNumber="386">P. vestigiatus</emphasis>
|
||
</taxonomicName>
|
||
have bright spots or coloration in the groin (absent). Furthermore,
|
||
<emphasis id="57D09A0DFFBB754D9C3D74DD9704FC10" box="[542,880,977,1002]" italics="true" pageId="5" pageNumber="386">
|
||
<taxonomicName id="A2A43D9CFFBB754D9C3D74DD96DAFC13" box="[542,686,977,1002]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="dagmarae">P. dagmarae</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFBB754D9C9A74DD9718FC10" box="[697,876,978,1001]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="insterstinctus">P. insterstinctus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="A2A43D9CFFBB754D9D8674DD906AFC13" box="[933,1054,977,1002]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="nicoleae">
|
||
<emphasis id="57D09A0DFFBB754D9D8674DD906AFC13" box="[933,1054,977,1002]" italics="true" pageId="5" pageNumber="386">P. nicoleae</emphasis>
|
||
</taxonomicName>
|
||
have either smooth or shagreened dorsal skin (coarsely areolate);
|
||
<taxonomicName id="A2A43D9CFFBB754D9FDF74F99615FBF7" box="[508,609,1013,1038]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="bracki">
|
||
<emphasis id="57D09A0DFFBB754D9FDF74F99615FBF7" box="[508,609,1013,1038]" italics="true" pageId="5" pageNumber="386">P. bracki</emphasis>
|
||
</taxonomicName>
|
||
is smaller with male SVL
|
||
<quantity id="A25CEBFAFFBB754D9DB574FA9048FBF4" box="[918,1084,1013,1038]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" metricValueMax="1.6199999999999999" metricValueMin="1.3199999999999998" pageId="5" pageNumber="386" unit="mm" value="14.7" valueMax="16.2" valueMin="13.2">13.2–16.2 mm</quantity>
|
||
, has a smooth venter (coarsely areolate), ridges forming an “M” on the scapular region (no scapular ridges) and bears lateral fringes on fingers (absent);
|
||
<taxonomicName id="A2A43D9CFFBB754D9F23733195FAFBAF" box="[256,398,1085,1110]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="dagmarae">
|
||
<emphasis id="57D09A0DFFBB754D9F23733195FAFBAF" box="[256,398,1085,1110]" italics="true" pageId="5" pageNumber="386">P. dagmarae</emphasis>
|
||
</taxonomicName>
|
||
bears dorsolateral folds, lateral fringes on all digits, and the first finger is much shorter than the second one;
|
||
<taxonomicName id="A2A43D9CFFBB754D9F6C736D9675FB80" box="[335,513,1122,1145]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="insterstinctus">
|
||
<emphasis id="57D09A0DFFBB754D9F6C736D9675FB80" box="[335,513,1122,1145]" italics="true" pageId="5" pageNumber="386">P. insterstinctus</emphasis>
|
||
</taxonomicName>
|
||
is a smaller frog with longer limbs than
|
||
<taxonomicName id="A2A43D9CFFBB754D9DF2736D902DFB80" box="[977,1113,1122,1145]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="anancites">
|
||
<emphasis id="57D09A0DFFBB754D9DF2736D902DFB80" box="[977,1113,1122,1145]" italics="true" pageId="5" pageNumber="386">P. anancites</emphasis>
|
||
</taxonomicName>
|
||
, and a smooth venter with a distinctive spotted pattern (uniformly gray);
|
||
<taxonomicName id="A2A43D9CFFBB754D9CA47389976EFB64" box="[647,794,1158,1181]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="vestigiatus">
|
||
<emphasis id="57D09A0DFFBB754D9CA47389976EFB64" box="[647,794,1158,1181]" italics="true" pageId="5" pageNumber="386">P. vestigiatus</emphasis>
|
||
</taxonomicName>
|
||
has lateral fringes on digits, bears ridges and dorsolateral folds (absent). Among species of
|
||
<taxonomicName id="A2A43D9CFFBB754D9C2D73A696F3FB3B" box="[526,647,1193,1218]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="genus">
|
||
<emphasis id="57D09A0DFFBB754D9C2D73A696F3FB3B" box="[526,647,1193,1218]" italics="true" pageId="5" pageNumber="386">Phrynopus</emphasis>
|
||
</taxonomicName>
|
||
bearing dentigerous processes of vomers and sharing a supratympanic fold,
|
||
<taxonomicName id="A2A43D9CFFBB754D9EF073C1951AFB1F" box="[211,366,1229,1254]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="kauneorum">
|
||
<emphasis id="57D09A0DFFBB754D9EF073C1951AFB1F" box="[211,366,1229,1254]" italics="true" pageId="5" pageNumber="386">P. kauneorum</emphasis>
|
||
</taxonomicName>
|
||
has a shorter head (HL/SVL= 28–33 % in
|
||
<taxonomicName id="A2A43D9CFFBB754D9D7A73C297A3FB1F" box="[857,983,1229,1254]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="kauneorum">
|
||
<emphasis id="57D09A0DFFBB754D9D7A73C297A3FB1F" box="[857,983,1229,1254]" italics="true" pageId="5" pageNumber="386">kauneorum</emphasis>
|
||
</taxonomicName>
|
||
, 37% in
|
||
<taxonomicName id="A2A43D9CFFBB754D9A1C73C190DEFB1C" box="[1087,1194,1230,1253]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="anancites">
|
||
<emphasis id="57D09A0DFFBB754D9A1C73C190DEFB1C" box="[1087,1194,1230,1253]" italics="true" pageId="5" pageNumber="386">anancites</emphasis>
|
||
</taxonomicName>
|
||
), with smooth dorsal and ventral skin (coarsely areolate) and dark blotches over a pale brown dorsum and a lighter ventral color with some spots (uniform gray in dorsum and venter);
|
||
<taxonomicName id="A2A43D9CFFBB754D9CE872199759FAD7" box="[715,813,1301,1326]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="kotosh">
|
||
<emphasis id="57D09A0DFFBB754D9CE872199759FAD7" box="[715,813,1301,1326]" italics="true" pageId="5" pageNumber="386">P. kotosh</emphasis>
|
||
</taxonomicName>
|
||
has widely separated dentigerous processes of vomers bearing two teeth (minute), dorsal skin shagreened with scattered tubercles, aerolate skin on venter (coarsely aerolate dorsally and ventrally); and
|
||
<taxonomicName id="A2A43D9CFFBB754D9C6F72519756FA8F" box="[588,802,1373,1398]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="lechriorhynchus">
|
||
<emphasis id="57D09A0DFFBB754D9C6F72519756FA8F" box="[588,802,1373,1398]" italics="true" pageId="5" pageNumber="386">P. lechriorhynchus</emphasis>
|
||
</taxonomicName>
|
||
, has a distinctly spatulate snout (rounded and short).
|
||
<taxonomicName id="A2A43D9CFFBB754D9EB4728E9503FA63" box="[151,375,1409,1434]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="capitalis">
|
||
<emphasis id="57D09A0DFFBB754D9EB4728E9503FA63" box="[151,375,1409,1434]" italics="true" pageId="5" pageNumber="386">Phrynopus capitalis</emphasis>
|
||
</taxonomicName>
|
||
has also a thick supratympanic fold, but lacks small vomerine teeth, bears fringes on digits and basal webbing (absent), and its dorsal coloration forms a pattern (absent).
|
||
<taxonomicName id="A2A43D9CFFBB754D9A3D72AA9170FA47" box="[1054,1284,1445,1470]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="bufoides">
|
||
<emphasis id="57D09A0DFFBB754D9A3D72AA9170FA47" box="[1054,1284,1445,1470]" italics="true" pageId="5" pageNumber="386">Phrynopus bufoides</emphasis>
|
||
</taxonomicName>
|
||
shares most characters with
|
||
<taxonomicName id="A2A43D9CFFBB754D9F6472C595B9FA18" box="[327,461,1482,1505]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="anancites">
|
||
<emphasis id="57D09A0DFFBB754D9F6472C595B9FA18" box="[327,461,1482,1505]" italics="true" pageId="5" pageNumber="386">P. anancites</emphasis>
|
||
</taxonomicName>
|
||
, including the texture of the dorsal skin, but differs by bearing large elongated warts on dorsum and flanks over a darker ground color (rounded and continuous), by having first finger longer than second (shorter, or equal in size), and by lacking the dentigerous processes of vomers (present).
|
||
<taxonomicName id="A2A43D9CFFBB754D9AAF711E9113F9D3" box="[1164,1383,1553,1578]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="daemon">
|
||
<emphasis id="57D09A0DFFBB754D9AAF711E9113F9D3" box="[1164,1383,1553,1578]" italics="true" pageId="5" pageNumber="386">Phrynopus daemon</emphasis>
|
||
</taxonomicName>
|
||
also lacks a tympanum and has uniform dorsum coloration, but lacks vomerine teeth, bears a prominent thoracic fold, has a shorter and less pronounced supratympanic fold, subacuminate snout in lateral view (blunt), and lateral fringes in toes (absent).
|
||
<taxonomicName id="A2A43D9CFFBB754D9F8A717296EBF96F" box="[425,671,1661,1686]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="thompsoni">
|
||
<emphasis id="57D09A0DFFBB754D9F8A717296EBF96F" box="[425,671,1661,1686]" italics="true" pageId="5" pageNumber="386">Phrynopus thompsoni</emphasis>
|
||
</taxonomicName>
|
||
lacks a tympanum and has first and second fingers equal in length, but it bears lateral fringes on toes (absent), has pustular dorsal skin (coarsely areolate), and lacks the dentigerous processes of vomers (present).
|
||
<taxonomicName id="A2A43D9CFFBB754D9FD271CA96ABF927" box="[497,735,1733,1758]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="chaparroi">
|
||
<emphasis id="57D09A0DFFBB754D9FD271CA96ABF927" box="[497,735,1733,1758]" italics="true" pageId="5" pageNumber="386">Phrynopus chaparroi</emphasis>
|
||
</taxonomicName>
|
||
shares most characters with
|
||
<taxonomicName id="A2A43D9CFFBB754D9A0371C990D2F924" box="[1056,1190,1734,1757]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="anancites">
|
||
<emphasis id="57D09A0DFFBB754D9A0371C990D2F924" box="[1056,1190,1734,1757]" italics="true" pageId="5" pageNumber="386">P. anancites</emphasis>
|
||
</taxonomicName>
|
||
, but differs by having the skin of the tympanic area covered by elongate, round subconical tubercles instead of a supratympanic fold (fold present), by having a rounded snout in profile (blunt), by lacking the dentigerous processes of vomers (present), and by bearing a brownish-black dorsal coloration scattered with white to yellow spots (uniform olive), and a brown reticulated iris (yellow ring and bronze in
|
||
<taxonomicName id="A2A43D9CFFBB754D9CFA70599710F894" box="[729,868,1878,1901]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="anancites">
|
||
<emphasis id="57D09A0DFFBB754D9CFA70599710F894" box="[729,868,1878,1901]" italics="true" pageId="5" pageNumber="386">P. anancites</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="A2A43D9CFFBB754D9D5D705A90F6F897" box="[894,1154,1877,1902]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="auriculatus">
|
||
<emphasis id="57D09A0DFFBB754D9D5D705A90F6F897" box="[894,1154,1877,1902]" italics="true" pageId="5" pageNumber="386">Phrynopus auriculatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A2A43D9CFFBB754D9A9D70599131F894" box="[1214,1349,1878,1901]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="peruanus">
|
||
<emphasis id="57D09A0DFFBB754D9A9D70599131F894" box="[1214,1349,1878,1901]" italics="true" pageId="5" pageNumber="386">P. peruanus</emphasis>
|
||
</taxonomicName>
|
||
have a visible tympanic annulus (absent), whereas
|
||
<taxonomicName id="A2A43D9CFFBB754D9CA170759774F868" box="[642,768,1914,1937]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="montium">
|
||
<emphasis id="57D09A0DFFBB754D9CA170759774F868" box="[642,768,1914,1937]" italics="true" pageId="5" pageNumber="386">P. montium</emphasis>
|
||
</taxonomicName>
|
||
has a dark dorsum and areolate dorsal and ventral skin, but differs by lacking dentigerous processes of vomers, by having a visible tympanic annulus and vocal slits and nuptial pads on swollen Finger I (absent). Most species of the
|
||
<taxonomicName id="A2A43D9CFFBB754D9D4F70CD9038F820" box="[876,1100,1986,2009]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="386" phylum="Chordata" rank="species" species="orestes">
|
||
<emphasis id="57D09A0DFFBB754D9D4F70CD9038F820" box="[876,1100,1986,2009]" italics="true" pageId="5" pageNumber="386">Pristimantis orestes</emphasis>
|
||
</taxonomicName>
|
||
group superficially resemble
|
||
<taxonomicName id="A2A43D9CFFB8754E9EB4779895F6FF49" box="[151,386,151,176]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="anancites">
|
||
<emphasis id="57D09A0DFFB8754E9EB4779895F6FF49" box="[151,386,151,176]" italics="true" pageId="6" pageNumber="387">Phrynopus anancites</emphasis>
|
||
</taxonomicName>
|
||
; those are
|
||
<emphasis id="57D09A0DFFB8754E9FDE779697A7FF2D" italics="true" pageId="6" pageNumber="387">
|
||
<taxonomicName id="A2A43D9CFFB8754E9FDE7796977FFF49" box="[509,779,151,176]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="atrabracus">Pristimantis atrabracus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9D357796970DFF49" box="[790,889,151,176]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="chimu">P. chimu</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9DA67796907EFF49" box="[901,1034,151,176]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="cordovae">P. cordovae</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9A35779690C9FF49" box="[1046,1213,151,176]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="mariaelenae">P mariaelenae</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9AEA77969108FF49" box="[1225,1404,151,176]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="melanogaster">P. melanogaster</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9BAB7796948DFF2C" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="pataikos">P. pataikos</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9F2477B2950FFF2D" box="[263,379,189,212]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="pinguis">P. pinguis</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9FA977B2958BFF2D" box="[394,511,189,212]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="seorsus">P. seorsus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9C2D77B296F8FF2D" box="[526,652,189,212]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="simonsii">P. simonsii</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9CB877B29712FF2C" box="[667,870,188,213]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="stictoboubonus">P. stictoboubonus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9D5677B297B8FF2D" box="[885,972,189,212]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="stipa">P. stipa</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
all of which, except
|
||
<taxonomicName id="A2A43D9CFFB8754E9AED77B2913EFF2D" box="[1230,1354,189,212]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="simonsii">
|
||
<emphasis id="57D09A0DFFB8754E9AED77B2913EFF2D" box="[1230,1354,189,212]" italics="true" pageId="6" pageNumber="387">P. simonsii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A2A43D9CFFB8754E9BA477B29540FF01" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="attenboroughi">
|
||
<emphasis id="57D09A0DFFB8754E9BA477B29540FF01" italics="true" pageId="6" pageNumber="387">P. attenboroughi</emphasis>
|
||
</taxonomicName>
|
||
, can easily be distinguished from
|
||
<taxonomicName id="A2A43D9CFFB8754E9CCD77D09797FF01" box="[750,995,223,248]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="anancites">
|
||
<emphasis id="57D09A0DFFB8754E9CCD77D09797FF01" box="[750,995,223,248]" italics="true" pageId="6" pageNumber="387">Phrynopus anancites</emphasis>
|
||
</taxonomicName>
|
||
by having a visible tympanum (
|
||
<emphasis id="57D09A0DFFB8754E9BA477EE973AFEE5" italics="true" pageId="6" pageNumber="387">
|
||
<taxonomicName id="A2A43D9CFFB8754E9BA477EE9561FEE4" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="atrabracus">P. atrabracus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9F02760A95F1FEE4" box="[289,389,260,285]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="chimu">P. chimu</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9FB0760A966EFEE4" box="[403,538,260,285]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="cordovae">P. cordovae</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9C04760A96A7FEE4" box="[551,723,260,285]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="mariaelenae">P. mariaelenae</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9CFC760A973AFEE5" box="[735,846,261,284]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="pinguis">P. pinguis</taxonomicName>
|
||
</emphasis>
|
||
), or the tympanic annulus weakly defined and only visible beneath the skin (
|
||
<emphasis id="57D09A0DFFB8754E9F8D762690E3FEB9" box="[430,1175,295,320]" italics="true" pageId="6" pageNumber="387">
|
||
<taxonomicName id="A2A43D9CFFB8754E9F8D76269616FEB9" box="[430,610,295,320]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="melanogaster">P. melanogaster</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9C4E7626969FFEB9" box="[621,747,295,320]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="pataikos">P. pataikos</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9CD5762697C9FEB9" box="[758,957,295,320]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="stictoboubonus">P. stictoboubonus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9DEB7626904FFEB9" box="[968,1083,297,320]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="seorsus">P. seorsus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB8754E9A65762690E3FEB9" box="[1094,1175,297,320]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="stipa">P. stipa</taxonomicName>
|
||
</emphasis>
|
||
).
|
||
<taxonomicName id="A2A43D9CFFB8754E9A8C762691EFFEB9" box="[1199,1435,297,320]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="simonsii">
|
||
<emphasis id="57D09A0DFFB8754E9A8C762691EFFEB9" box="[1199,1435,297,320]" italics="true" pageId="6" pageNumber="387">Pristimantis simonsii</emphasis>
|
||
</taxonomicName>
|
||
lacks the dentigerous processes of vomers (present), and bears lateral fringes and basal webbing on digits (absent), while
|
||
<taxonomicName id="A2A43D9CFFB8754E9EFE767E95D1FE71" box="[221,421,367,392]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="attenbouroughi">
|
||
<emphasis id="57D09A0DFFB8754E9EFE767E95D1FE71" box="[221,421,367,392]" italics="true" pageId="6" pageNumber="387">P. attenbouroughi</emphasis>
|
||
</taxonomicName>
|
||
shares with
|
||
<taxonomicName id="A2A43D9CFFB8754E9C17767E96EBFE71" box="[564,671,369,392]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="anancites">
|
||
<emphasis id="57D09A0DFFB8754E9C17767E96EBFE71" box="[564,671,369,392]" italics="true" pageId="6" pageNumber="387">anancites</emphasis>
|
||
</taxonomicName>
|
||
the lack of tympanic membrane and annulus, and lateral fringes on all digits, and the presence of dentigerous processes of vomers, but differs by bearing dorsal skin shagreened with low scattered tubercles (coarsely areolate with large rounded warts), by having a dark canthal and supratympanic stripe (well defined supratympanic fold, no color stripe), and by being smaller in size, males SVL
|
||
<quantity id="A25CEBFAFFB8754E9ADB76D391E8FE0D" box="[1272,1436,476,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" metricValueMax="1.92" metricValueMin="1.46" pageId="6" pageNumber="387" unit="mm" value="16.9" valueMax="19.2" valueMin="14.6">14.6–19.2 mm</quantity>
|
||
(Lehr and von
|
||
<bibRefCitation id="01353BEEFFB8754E9F1F750F95CDFDE1" author="May" box="[316,441,512,536]" pageId="6" pageNumber="387" refString="Lehr, E. & von May, R. (2017) A new species of terrestrial-breeding frog (Amphibia, Craugastoridae, Pristimantis) from high elevations of the Pui Pui Protected Forest in central Peru. ZooKeys, 660, 17 - 42." type="journal article" year="2017">May, 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="31DB1697FFB8754E9EB4708B9755F82E" httpUri="https://zenodo.org/record/803117/files/figure.png" pageId="6" pageNumber="387" targetBox="[487,1072,596,1899]" targetPageId="6">
|
||
<paragraph id="651B461FFFB8754E9EB4708B9755F82E" blockId="6.[151,1436,1924,2007]" pageId="6" pageNumber="387">
|
||
<emphasis id="57D09A0DFFB8754E9EB4708B9560F860" bold="true" box="[151,276,1924,1946]" pageId="6" pageNumber="387">FIGURE 4.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="A2A43D9CFFB8754E9F83708B9603F863" box="[416,631,1924,1946]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="387" phylum="Chordata" rank="species" species="anancites" status="sp. nov.">
|
||
<emphasis id="57D09A0DFFB8754E9F83708B9603F863" box="[416,631,1924,1946]" italics="true" pageId="6" pageNumber="387">Phrynopus anancites</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="57D09A0DFFB8754E9C5C708B96A5F860" bold="true" box="[639,721,1924,1945]" pageId="6" pageNumber="387">
|
||
<taxonomicNameLabel id="4CE32776FFB8754E9C5C708B96A5F860" box="[639,721,1924,1945]" pageId="6" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male MUSM 33168 (SVL = 25.3 mm), showing (
|
||
<emphasis id="57D09A0DFFB8754E9AC5708B908CF860" bold="true" box="[1254,1272,1924,1945]" pageId="6" pageNumber="387">A</emphasis>
|
||
) head in lateral view and (
|
||
<emphasis id="57D09A0DFFB8754E9F2770AC9541F841" bold="true" box="[260,309,1954,1976]" pageId="6" pageNumber="387">B, C</emphasis>
|
||
) body in dorsal and ventral views. Specimen collected at Ventanas, 3820 m, near the entrance of Río Abiseo National Park, San Martín, Peru. Photographs by Morris Flecks.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="31DB1697FFB9754F9EB4744D9161FC8F" httpUri="https://zenodo.org/record/803119/files/figure.png" pageId="7" pageNumber="388" targetBox="[342,1249,220,784]" targetPageId="7">
|
||
<paragraph id="651B461FFFB9754F9EB4744D9161FC8F" blockId="7.[151,1435,834,886]" pageId="7" pageNumber="388">
|
||
<emphasis id="57D09A0DFFB9754F9EB4744D9563FCAE" bold="true" box="[151,279,834,856]" pageId="7" pageNumber="388">FIGURE 5.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="A2A43D9CFFB9754F9F8D744D96FCFCA1" box="[430,648,834,856]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="anancites" status="sp. nov.">
|
||
<emphasis id="57D09A0DFFB9754F9F8D744D96FCFCA1" box="[430,648,834,856]" italics="true" pageId="7" pageNumber="388">Phrynopus anancites</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="57D09A0DFFB9754F9CB7744D969DFCAE" bold="true" box="[660,745,834,855]" pageId="7" pageNumber="388">
|
||
<taxonomicNameLabel id="4CE32776FFB9754F9CB7744D969DFCAE" box="[660,745,834,855]" pageId="7" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male MUSM 33168 (SVL= 25.3 mm), in life. Specimen from Ventanas, 3820 m, near the entrance of Río Abiseo National Park, San Martín, Peru. Photograph by A. Catenazzi.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="31DB1697FFB9754F9EB470319779F868" httpUri="https://zenodo.org/record/803121/files/figure.png" pageId="7" pageNumber="388" targetBox="[165,1411,981,1831]" targetPageId="7">
|
||
<paragraph id="651B461FFFB9754F9EB470319779F868" blockId="7.[151,1437,1854,1937]" pageId="7" pageNumber="388">
|
||
<emphasis id="57D09A0DFFB9754F9EB470319560F8AA" bold="true" box="[151,276,1854,1876]" pageId="7" pageNumber="388">FIGURE 6.</emphasis>
|
||
Palmar and plantar surfaces of the new species of
|
||
<taxonomicName id="A2A43D9CFFB9754F9D05703197E7F8AD" box="[806,915,1854,1876]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="388" phylum="Chordata" rank="genus">
|
||
<emphasis id="57D09A0DFFB9754F9D05703197E7F8AD" box="[806,915,1854,1876]" italics="true" pageId="7" pageNumber="388">Phrynopus</emphasis>
|
||
</taxonomicName>
|
||
: (
|
||
<emphasis id="57D09A0DFFB9754F9D88703197C9F8AA" bold="true" box="[939,957,1854,1875]" pageId="7" pageNumber="388">A</emphasis>
|
||
)
|
||
<taxonomicName id="A2A43D9CFFB9754F9DED7030903DF8AD" box="[974,1097,1855,1876]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="anancites" status="sp. nov.">
|
||
<emphasis id="57D09A0DFFB9754F9DED7030903DF8AD" box="[974,1097,1855,1876]" italics="true" pageId="7" pageNumber="388">P. anancites</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="57D09A0DFFB9754F9A72703090D7F8AD" bold="true" box="[1105,1187,1855,1876]" pageId="7" pageNumber="388">
|
||
<taxonomicNameLabel id="4CE32776FFB9754F9A72703090D7F8AD" box="[1105,1187,1855,1876]" pageId="7" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(MUSM 33168), (
|
||
<emphasis id="57D09A0DFFB9754F9B4B7030910DF8AD" bold="true" box="[1384,1401,1855,1876]" pageId="7" pageNumber="388">B</emphasis>
|
||
)
|
||
<taxonomicName id="A2A43D9CFFB9754F9BAA7030949AF88B" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="capitalis" status="sp. nov.">
|
||
<emphasis id="57D09A0DFFB9754F9BAA7030949AF88B" italics="true" pageId="7" pageNumber="388">P. capitalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="57D09A0DFFB9754F9ED770529530F88B" bold="true" box="[244,324,1885,1906]" pageId="7" pageNumber="388">
|
||
<taxonomicNameLabel id="4CE32776FFB9754F9ED770529530F88B" box="[244,324,1885,1906]" pageId="7" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(AMNH 134158), (
|
||
<emphasis id="57D09A0DFFB9754F9C3270529657F88A" bold="true" box="[529,547,1885,1907]" pageId="7" pageNumber="388">C</emphasis>
|
||
)
|
||
<taxonomicName id="A2A43D9CFFB9754F9C11705196DDF88B" box="[562,681,1884,1907]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="dumicola" status="sp. nov.">
|
||
<emphasis id="57D09A0DFFB9754F9C11705196DDF88B" box="[562,681,1884,1907]" italics="true" pageId="7" pageNumber="388">P. dumicola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="57D09A0DFFB9754F9C9370529774F88B" bold="true" box="[688,768,1885,1906]" pageId="7" pageNumber="388">
|
||
<taxonomicNameLabel id="4CE32776FFB9754F9C9370529774F88B" box="[688,768,1885,1906]" pageId="7" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(MUSM 33102), and (
|
||
<emphasis id="57D09A0DFFB9754F9DC870529789F88B" bold="true" box="[1003,1021,1885,1906]" pageId="7" pageNumber="388">D</emphasis>
|
||
)
|
||
<taxonomicName id="A2A43D9CFFB9754F9A2F705190E7F88A" box="[1036,1171,1886,1907]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="388" phylum="Chordata" rank="species" species="personatus" status="sp. nov.">
|
||
<emphasis id="57D09A0DFFB9754F9A2F705190E7F88A" box="[1036,1171,1886,1907]" italics="true" pageId="7" pageNumber="388">P. personatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="57D09A0DFFB9754F9AB97052909EF88B" bold="true" box="[1178,1258,1885,1906]" pageId="7" pageNumber="388">
|
||
<taxonomicNameLabel id="4CE32776FFB9754F9AB97052909EF88B" box="[1178,1258,1885,1906]" pageId="7" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(MUSM 33096). Photographs by M. Flecks (A, C and D) and A. Catenazzi (B).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="2DBE1594FFB675409EE4779897C5FC5C" pageId="8" pageNumber="389" type="description">
|
||
<paragraph id="651B461FFFB675409EE47798910AFE54" blockId="8.[151,1437,151,1112]" pageId="8" pageNumber="389">
|
||
<emphasis id="57D09A0DFFB675409EE477989665FF49" bold="true" box="[199,529,151,176]" pageId="8" pageNumber="389">
|
||
Description of the
|
||
<typeStatus id="BA1FF8BDFFB675409F8077989678FF49" box="[419,524,151,176]" pageId="8" pageNumber="389" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Head narrower than body, wider than long; HW 107% of HL, HW 41% of SVL; HL 37% of SVL; snout short, rounded in dorsal view and slightly blunt in lateral view, E-N distance around half the size of eye; nostrils slightly protruding laterally, canthus rostralis rounded; loreal region bearing a few large postrictal tubercles under the supratympanic fold; no tubercles on upper eyelid; choanae small, round, well separated and anterolaterally situated; dentigerous processes of vomers minute, posterior to choanae, slightly oblique, broadly separated; cranial crests absent; upper eyelid smaller than interorbital distance, EW 88% of IOD; tympanic membrane and tympanic annulus absent; supratympanic fold thick, extending from edge of eye to the end of postrictal area; tongue oval, not notched posteriorly, posterior two-third free. Vocal sac and vocal slits absent.
|
||
</paragraph>
|
||
<paragraph id="651B461FFFB675409EE476B890AAFD34" blockId="8.[151,1437,151,1112]" pageId="8" pageNumber="389">
|
||
Skin on dorsum coarsely areolate, with large rounded warts, without lateral folds; outside of arms bearing low rounded tubercles; discoidal fold absent, a thin thoracic fold present; skin on venter and throat coarsely areolated bearing distinctive rounded warts, extending onto the lower parts of arms and legs, and the femoral and perianal areas. Outer surface of arms lacking tubercles; first finger and second fingers equal in length; thenar tubercle ovoid, 1.5 times larger than palmar tubercle which is hart-shaped; fingers lacking lateral fringes, tip of digits rounded, bulbous (
|
||
<figureCitation id="FD9F5A9AFFB675409F2275639510FD7C" box="[257,356,620,645]" captionStart="FIGURE 6" captionStartId="7.[151,250,1854,1876]" captionTargetBox="[165,1411,981,1831]" captionTargetId="figure@7.[151,1436,972,1832]" captionTargetPageId="7" captionText="FIGURE 6. Palmar and plantar surfaces of the new species of Phrynopus: (A) P. anancites sp. nov. (MUSM 33168), (B) P. capitalis sp. nov. (AMNH 134158), (C) P. dumicola sp. nov. (MUSM 33102), and (D) P. personatus sp. nov. (MUSM 33096). Photographs by M. Flecks (A, C and D) and A. Catenazzi (B)." httpUri="https://zenodo.org/record/803121/files/figure.png" pageId="8" pageNumber="389">Figure 6</figureCitation>
|
||
A); TL 34% of SVL; tarsus bearing a small tubercle, no tarsal fold; two metatarsal tubercles, inner slightly elongated, outer slightly smaller, rounded, protruding; toes without lateral fringes, lacking basal webbing; toe tips bulbous, as large as those on fingers; Toe V slightly longer than III (
|
||
<figureCitation id="FD9F5A9AFFB675409A7A75BB90CEFD34" box="[1113,1210,692,717]" captionStart="FIGURE 6" captionStartId="7.[151,250,1854,1876]" captionTargetBox="[165,1411,981,1831]" captionTargetId="figure@7.[151,1436,972,1832]" captionTargetPageId="7" captionText="FIGURE 6. Palmar and plantar surfaces of the new species of Phrynopus: (A) P. anancites sp. nov. (MUSM 33168), (B) P. capitalis sp. nov. (AMNH 134158), (C) P. dumicola sp. nov. (MUSM 33102), and (D) P. personatus sp. nov. (MUSM 33096). Photographs by M. Flecks (A, C and D) and A. Catenazzi (B)." httpUri="https://zenodo.org/record/803121/files/figure.png" pageId="8" pageNumber="389">Figure 6</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<paragraph id="651B461FFFB675409EE475D89523FCED" blockId="8.[151,1437,151,1112]" pageId="8" pageNumber="389">
|
||
<emphasis id="57D09A0DFFB675409EE475D89592FD09" bold="true" box="[199,486,727,752]" pageId="8" pageNumber="389">Measurements (in mm).</emphasis>
|
||
SVL 25.3, HL 9.4, HW 10.1, TL 8.5, THL 9.7, FL 9.9, IOD 2.8, EW 2.5, E-N 2.7, IND 3.1, ED 3.2.
|
||
</paragraph>
|
||
<paragraph id="651B461FFFB675409EE4741097F0FCA4" blockId="8.[151,1437,151,1112]" pageId="8" pageNumber="389">
|
||
<emphasis id="57D09A0DFFB675409EE474109529FCC1" bold="true" box="[199,349,799,824]" pageId="8" pageNumber="389">Color in life.</emphasis>
|
||
The dorsal and lateral surfaces are olive-green (
|
||
<figureCitation id="FD9F5A9AFFB675409D54741097ACFCC1" box="[887,984,799,824]" captionStart="FIGURE 5" captionStartId="7.[151,250,834,856]" captionTargetBox="[342,1249,220,784]" captionTargetId="figure@7.[342,1249,220,784]" captionTargetPageId="7" captionText="FIGURE 5. Holotype of Phrynopus anancites sp. nov., male MUSM 33168 (SVL = 25.3 mm), in life. Specimen from Ventanas, 3820 m, near the entrance of Río Abiseo National Park, San Martín, Peru. Photograph by A. Catenazzi." httpUri="https://zenodo.org/record/803119/files/figure.png" pageId="8" pageNumber="389">Figure 5</figureCitation>
|
||
), whereas the ventral parts are pale gray with darker flecks. The iris is uniformly bronze with a yellow ring.
|
||
</paragraph>
|
||
<paragraph id="651B461FFFB675409EE4746897C5FC5C" blockId="8.[151,1437,151,1112]" pageId="8" pageNumber="389">
|
||
<emphasis id="57D09A0DFFB675409EE4746895B8FC79" bold="true" box="[199,460,871,896]" pageId="8" pageNumber="389">Color in preservative.</emphasis>
|
||
In alcohol (
|
||
<figureCitation id="FD9F5A9AFFB675409C77746896C2FC79" box="[596,694,871,896]" captionStart="FIGURE 4" captionStartId="6.[151,250,1924,1946]" captionTargetBox="[487,1072,596,1899]" captionTargetId="figure@6.[487,1099,589,1902]" captionTargetPageId="6" captionText="FIGURE 4. Holotype of Phrynopus anancites sp. nov., male MUSM 33168 (SVL = 25.3 mm), showing (A) head in lateral view and (B, C) body in dorsal and ventral views. Specimen collected at Ventanas, 3820 m, near the entrance of Río Abiseo National Park, San Martín, Peru. Photographs by Morris Flecks." httpUri="https://zenodo.org/record/803117/files/figure.png" pageId="8" pageNumber="389">Figure 4</figureCitation>
|
||
), the dorsal surfaces are uniformly dark grayish-brown; chest and throat areas are light brown, the remaining areas are as dorsal surfaces.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2DBE1594FFB675409EE474A09742FC14" pageId="8" pageNumber="389" type="etymology">
|
||
<paragraph id="651B461FFFB675409EE474A09742FC14" blockId="8.[151,1437,151,1112]" pageId="8" pageNumber="389">
|
||
<emphasis id="57D09A0DFFB675409EE474A0953AFC31" bold="true" box="[199,334,943,968]" pageId="8" pageNumber="389">Etymology.</emphasis>
|
||
The specific name
|
||
<taxonomicName id="A2A43D9CFFB675409C1274BE96E9FC31" box="[561,669,945,968]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="389" phylum="Chordata" rank="species" species="anancites">
|
||
<emphasis id="57D09A0DFFB675409C1274BE96E9FC31" box="[561,669,945,968]" italics="true" pageId="8" pageNumber="389">anancites</emphasis>
|
||
</taxonomicName>
|
||
is a Latin noun for a
|
||
<typeStatus id="BA1FF8BDFFB675409DB874BF97BFFC31" box="[923,971,944,968]" pageId="8" pageNumber="389">type</typeStatus>
|
||
of diamond used to drive sadness away, given in reference to the frog’s rare dorsal coloration in life.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2DBE1594FFB675409EE474F896ACFBA1" pageId="8" pageNumber="389" type="distribution">
|
||
<paragraph id="651B461FFFB675409EE474F896ACFBA1" blockId="8.[151,1437,151,1112]" pageId="8" pageNumber="389">
|
||
<emphasis id="57D09A0DFFB675409EE474F89586FBE9" bold="true" box="[199,498,1015,1040]" pageId="8" pageNumber="389">Distribution and ecology.</emphasis>
|
||
The species is known from a single specimen that was found near the Park rangers’ station and entrance to Rio Abiseo NP at Ventanas, in wet montane grassland covered with mosses at
|
||
<quantity id="A25CEBFAFFB675409B0573139108FBCD" box="[1318,1404,1052,1076]" metricMagnitude="3" metricUnit="m" metricValue="3.82" pageId="8" pageNumber="389" unit="m" value="3820.0">3820 m</quantity>
|
||
of altitude. Nothing is known about its natural history.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |