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<mods:title id="59EC9B7A466B64802F606DE1009EE389">Morphological revision of the Subgroup 1 Fauchald, 1970 of Marphysa de Quatrefages, 1865 (Eunicidae: Polychaeta)</mods:title>
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<paragraph id="466E7552FFC02172FF5BA674FD5BF9D9" blockId="8.[151,767,1619,1679]" box="[151,767,1619,1646]" pageId="8" pageNumber="9">
<heading id="1D26C23EFFC02172FF5BA674FD5BF9D9" bold="true" box="[151,767,1619,1646]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="74A5A940FFC02172FF5BA674FD5BF9D9" bold="true" box="[151,767,1619,1646]" pageId="8" pageNumber="9">
<taxonomicName id="81D10ED1FFC02172FF5BA674FD29F9D9" ID-CoL="75TQL" authority="(Gravier, 1900)" authorityName="Molina-Acevedo" authorityYear="2018" baseAuthorityName="Gravier" baseAuthorityYear="1900" box="[151,653,1619,1646]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="adenensis" status="comb. nov.">
<emphasis id="74A5A940FFC02172FF5BA674FE6CF9DA" bold="true" box="[151,456,1619,1645]" italics="true" pageId="8" pageNumber="9">Paucibranchia adenensis</emphasis>
(
<bibRefCitation id="224008A3FFC02172FE1BA673FD20F9D9" box="[471,644,1620,1646]" pageId="8" pageNumber="9" refId="ref54865" refString="Gravier, C. (1900) Contribution a l'etude des annelids polychetes de la Mer Rouge. Premiere partie. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 4 e Serie, 2 (2), 137 - 282." type="journal article">Gravier, 1900</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC02172FD58A673FD5BF9D9" box="[660,767,1620,1646]" pageId="8" pageNumber="9">n. comb.</taxonomicNameLabel>
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<paragraph id="466E7552FFC02172FF5BA651FE07F938" blockId="8.[151,767,1619,1679]" box="[151,419,1654,1679]" pageId="8" pageNumber="9">
<figureCitation id="DEEA69D7FFC02172FF5BA651FEBBF938" box="[151,287,1654,1679]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="10.[151,250,1908,1930]" captionStartId-1="11.[151,250,1932,1954]" captionStartId-2="12.[151,250,1960,1982]" captionStartId-3="13.[151,250,765,787]" captionTargetBox-0="[219,1375,193,1892]" captionTargetBox-1="[336,1232,245,1870]" captionTargetBox-2="[182,1414,193,1947]" captionTargetBox-3="[160,1402,193,743]" captionTargetId-0="figure@10.[219,1375,193,1892]" captionTargetId-1="figure@11.[336,1250,245,1870]" captionTargetId-2="figure@12.[182,1414,193,1947]" captionTargetId-3="figure@13.[151,1436,193,743]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionText-0="FIGURE 3. Paucibranchia adenensis (Gravier, 1900) n. comb. A. Anterior end with branchiae, dorsal view; B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, dorsal view; E. Anterior end with branchiae, dorsal view; F. Anterior end, ventral view. A, B, C from holotype MNHN-531; D, E, F from non-type MNHN-A148. Scale bars: A, 2 mm; B, 1 mm, C, 2.6 mm, D, F, 0.7 mm; E, 1.3 mm." captionText-1="FIGURE 4. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Maxillary apparatus, dorsal view; B. Mandibles; C. Left MII-III-IV-V, dorsal view; D. Maxilla I, dorsal view. Apparatus stained with methyl green. Scale bars: A, C, 50 µm; B, 150 µm; D, 130 µm. al-MIV= attachment lamellae in MIV, LMI-MII= ligament between MI and MII." captionText-2="FIGURE 5. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Parapodium 3; B. Parapodium 11; C. Parapodium 26; D. Parapodium 70; E. Parapodium 158; F. Parapodium 216. All in anterior view. Scale bars: A–F, 0.7 mm." captionText-3="FIGURE 6. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Compound falciger, long blade, chaetiger, 11; B. Compound falciger, median blade, chaetiger, 11; C. Compound falciger, short blade, chaetiger, 216; D. Isodonts pectinate narrow with long and slender teeth, chaetiger 3; E. Isodonts pectinate narrow with short and slender teeth, chaetiger 216; F. Subacicular hook, distal end, chaetiger 158. Scale bars: A, 12.5 µm; B, 20 µm; C, F, 9.8 µm; D, 14 µm, E, 4.9 µm." httpUri-0="https://zenodo.org/record/1453037/files/figure.png" httpUri-1="https://zenodo.org/record/1453039/files/figure.png" httpUri-2="https://zenodo.org/record/1453041/files/figure.png" httpUri-3="https://zenodo.org/record/1453043/files/figure.png" pageId="8" pageNumber="9">Figures 3–6</figureCitation>
, 
<tableCitation id="0B5340E9FFC02172FEE6A651FE07F938" box="[298,419,1654,1679]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="111.[151,239,152,175]" captionStartId-1="112.[147,226,502,523]" captionText-0="TABLE 1. Morphometric data on the pectinate chaetae of Paucibranchia n. gen. species. The species P. stragula n. comb. was not included in this list because its pectinate chaetae were broken. Abbreviations: WBB: width at base of blade; WPB: widest part of blade; BL: blade length; WIT: width of internal teeth; LIT: length of internal teeth." captionText-1="TABLE 2. Morphological features of the Paucibranchia n. gen. species with compound falcigers. Abbreviations: MF: Maxillary formula, roman numerals refer to number of maxilla; MxC: maxillary carriers; CIS: closing system; Cop: cavity opening; PR-I: first peristomial ring; PR-II: second peristomial ring; Chaet: chaetiger; p/a: present/absent; DC: dorsal cirri; PrBr: pre-branchial region; PoBr: postbranchial region; Bran: branchial region; Ant: anterior region; Post: posterior region." pageId="8" pageNumber="9">Tables 1–2</tableCitation>
</paragraph>
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<paragraph id="466E7552FFC02172FF5BA699FBAFF945" blockId="8.[151,1437,1726,1779]" pageId="8" pageNumber="9">
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<taxonomicName id="81D10ED1FFC02172FF5BA699FD83F963" ID-CoL="3Y88M" authority="Gravier, 1900: 270" authorityName="Gravier" authorityPageNumber="270" authorityYear="1900" box="[151,551,1726,1748]" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="adenensis">
<emphasis id="74A5A940FFC02172FF5BA699FECCF963" box="[151,360,1726,1748]" italics="true" pageId="8" pageNumber="9">Marphysa adenensis</emphasis>
<treatmentCitation id="C7705343FFC02172FEA3A699FD83F963" author="" box="[367,551,1726,1748]" page="270" pageId="8" pageNumber="9">
<bibRefCitation id="224008A3FFC02172FEA3A699FE59F963" box="[367,509,1726,1748]" pageId="8" pageNumber="9" refId="ref54865" refString="Gravier, C. (1900) Contribution a l'etude des annelids polychetes de la Mer Rouge. Premiere partie. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 4 e Serie, 2 (2), 137 - 282." type="journal article">Gravier, 1900</bibRefCitation>
:270
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–272, Pl. XIV, Figs. 91–92, text-fig. 140–143; 
<treatmentCitation id="C7705343FFC02172FBC9A698FB31F963" author="" box="[1029,1173,1726,1748]" page="644" pageId="8" pageNumber="9">
<bibRefCitation id="224008A3FFC02172FBC9A698FBCEF963" box="[1029,1130,1726,1748]" pageId="8" pageNumber="9" refId="ref54003" refString="Day, J. H. (1962) Polychaeta from several localities in the western Indian Ocean. Proceedings of the Zoological Society of London, 139 (4), 627 - 656. https: // doi. org / 10.1111 / j. 1469 - 7998.1962. tb 01597. x" type="journal article">Day 1962</bibRefCitation>
:644
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–645; 
<treatmentCitation id="C7705343FFC02172FB1AA698FACDF963" author="" box="[1238,1385,1726,1748]" page="399" pageId="8" pageNumber="9">
<bibRefCitation id="224008A3FFC02172FB1AA698FA98F963" box="[1238,1340,1726,1748]" pageId="8" pageNumber="9" refId="ref54110" refString="Day, J. H. (1967) A monograph on the Polychaeta of Southern Africa. Part I. Errantia. British Museum (Natural History), London, 458 pp." type="book">Day 1967</bibRefCitation>
:399
</treatmentCitation>
, Fig. 17.6 p–s; 
<treatmentCitation id="C7705343FFC02172FEE6A6FBFD83F945" author="" box="[298,551,1756,1779]" page="203" pageId="8" pageNumber="9">
<bibRefCitation id="224008A3FFC02172FEE6A6FBFE59F945" box="[298,509,1756,1779]" pageId="8" pageNumber="9" refId="ref55476" refString="Katsiaras, N., Simboura, N. &amp; Koutsoubas, D. (2014) The rare subgroup C 1 of Marphysa (Polychaeta, Eunicidae): redescription of species and first records in the Mediterranean Sea. Zootaxa, 3873 (3), 201 - 217. https: // doi. org / 10.11646 / zootaxa. 3873.3.1" type="journal article">
Katsiaras 
<emphasis id="74A5A940FFC02172FE42A6F9FE66F945" box="[398,450,1756,1779]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2014
</bibRefCitation>
:203
</treatmentCitation>
–207, 
<figureCitation id="DEEA69D7FFC02172FDAAA6FBFD76F945" box="[614,722,1756,1778]" captionStart="FIGURE 1" captionStartId="6.[151,250,1846,1868]" captionTargetBox="[239,1357,218,1789]" captionTargetId="figure@6.[226,1361,195,1827]" captionTargetPageId="6" captionText="FIGURE 1. Maxillary apparatus. A. Paucibranchia disjuncta n. comb. (Hartman, 1961), LACM-AHF POLY 6447, dorsal view; B. Marphysa viridis Treadwell, 1917, ECOSUR-P2678, dorsal view; C. Treadwellphysa yucatanensis Molina-Acevedo &amp; Carrera-Parra, 2017, ECOSUR-P2811, dorsal view; D. Left MI-II-III-IV-V of P. disjuncta n. comb., dorsal view; E. Left MI- II-III-IV-V of M. viridis, dorsal view; F. Left MI-II-III-IV-V of T. yucatanensis, dorsal view. Scale bars: A, D, 0.4 mm; B, E, 0.45 mm; C, F, 0.25 mm. Dark and thin lines in D, E and F indicate the different forms in the MI of the apparatus of each species. fa= falcal arch, boe= basal outer edge, bie= basal inner edge, rp= rounded projection in base of MII." httpUri="https://zenodo.org/record/1453033/files/figure.png" pageId="8" pageNumber="9">Figs. 1a–h</figureCitation>
, 
<figureCitation id="DEEA69D7FFC02172FD10A6FBFCB4F945" box="[732,784,1756,1778]" captionStart="FIGURE 2" captionStartId="8.[151,250,1486,1508]" captionTargetBox="[276,1310,213,1430]" captionTargetId="figure@8.[276,1310,213,1430]" captionTargetPageId="8" captionText="FIGURE 2. Marphysa bonhardi McIntosh, 1885, holotype BMNH 1885.12.1.210. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Aciculae, anterior chaetigers. Scale bars: A, 0.8 mm; B, C, 0.67 mm." httpUri="https://zenodo.org/record/1453035/files/figure.png" pageId="8" pageNumber="9">2a–d</figureCitation>
, 
<figureCitation id="DEEA69D7FFC02172FCD6A6FBFC8CF945" box="[794,808,1756,1778]" captionStart="FIGURE 3" captionStartId="10.[151,250,1908,1930]" captionTargetBox="[219,1375,193,1892]" captionTargetId="figure@10.[219,1375,193,1892]" captionTargetPageId="10" captionText="FIGURE 3. Paucibranchia adenensis (Gravier, 1900) n. comb. A. Anterior end with branchiae, dorsal view; B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, dorsal view; E. Anterior end with branchiae, dorsal view; F. Anterior end, ventral view. A, B, C from holotype MNHN-531; D, E, F from non-type MNHN-A148. Scale bars: A, 2 mm; B, 1 mm, C, 2.6 mm, D, F, 0.7 mm; E, 1.3 mm." httpUri="https://zenodo.org/record/1453037/files/figure.png" pageId="8" pageNumber="9">3</figureCitation>
, 
<figureCitation id="DEEA69D7FFC02172FCFFA6FBFCC3F945" box="[819,871,1756,1778]" captionStart="FIGURE 4" captionStartId="11.[151,250,1932,1954]" captionTargetBox="[336,1232,245,1870]" captionTargetId="figure@11.[336,1250,245,1870]" captionTargetPageId="11" captionText="FIGURE 4. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Maxillary apparatus, dorsal view; B. Mandibles; C. Left MII-III-IV-V, dorsal view; D. Maxilla I, dorsal view. Apparatus stained with methyl green. Scale bars: A, C, 50 µm; B, 150 µm; D, 130 µm. al-MIV= attachment lamellae in MIV, LMI-MII= ligament between MI and MII." httpUri="https://zenodo.org/record/1453039/files/figure.png" pageId="8" pageNumber="9">4a–d</figureCitation>
, 
<tableCitation id="0B5340E9FFC02172FCBDA6FBFC14F945" box="[881,944,1756,1778]" captionStart="TABLE 2" captionStartId="112.[147,226,502,523]" captionText="TABLE 2. Morphological features of the Paucibranchia n. gen. species with compound falcigers. Abbreviations: MF: Maxillary formula, roman numerals refer to number of maxilla; MxC: maxillary carriers; CIS: closing system; Cop: cavity opening; PR-I: first peristomial ring; PR-II: second peristomial ring; Chaet: chaetiger; p/a: present/absent; DC: dorsal cirri; PrBr: pre-branchial region; PoBr: postbranchial region; Bran: branchial region; Ant: anterior region; Post: posterior region." pageId="8" pageNumber="9">Tab. 2</tableCitation>
(
<emphasis id="74A5A940FFC02172FC73A6F9FBA7F944" box="[959,1027,1758,1779]" italics="true" pageId="8" pageNumber="9">partim</emphasis>
)
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</paragraph>
</subSubSection>
<subSubSection id="0ECB26D9FFC02172FF5BA738FE68F837" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="466E7552FFC02172FF5BA738FE68F837" blockId="8.[151,1437,1823,2028]" pageId="8" pageNumber="9">
<materialsCitation id="F6B97F0FFFC02172FF5BA738FE68F837" ID-GBIF-Occurrence="1915273773" collectingDate="1904-02-10" collectionCode="MNHN-A" collectorName="Residence &amp; C. Gravier." country="Yemen" location="Red Sea" municipality="Gulf of Aden" pageId="8" pageNumber="9" specimenCode="MNHN-A148" specimenCount="2" stateProvince="Tadjoura" typeStatus="holotype">
<emphasis id="74A5A940FFC02172FF5BA738FE26F88F" bold="true" box="[151,386,1823,1848]" pageId="8" pageNumber="9">Material examined.</emphasis>
<emphasis id="74A5A940FFC02172FE46A707FD8BF88F" box="[394,559,1823,1848]" italics="true" pageId="8" pageNumber="9">
<typeStatus id="996ACBF0FFC02172FE46A707FE1AF880" box="[394,446,1824,1847]" pageId="8" pageNumber="9">Type</typeStatus>
material:
</emphasis>
<typeStatus id="996ACBF0FFC02172FDF4A738FD04F88F" box="[568,672,1823,1848]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
MNHN-531, 
<collectingCountry id="3EC635C2FFC02172FC8DA707FC36F880" box="[833,914,1824,1847]" name="Yemen" pageId="8" pageNumber="9">Yemen</collectingCountry>
, 
<collectingMunicipality id="A60AEF28FFC02172FC53A738FB9EF88F" box="[927,1082,1823,1848]" pageId="8" pageNumber="9">Gulf of Aden</collectingMunicipality>
, Red Sea. 
<emphasis id="74A5A940FFC02172FB79A738FA38F88F" box="[1205,1436,1823,1848]" italics="true" pageId="8" pageNumber="9">Additional material:</emphasis>
<specimenCode id="1677DD29FFC02172FF5BA763FE90F8EB" box="[151,308,1860,1884]" collectionCode="MNHN-A" pageId="8" pageNumber="9">MNHN-A148</specimenCode>
(5), 
<collectingCountry id="3EC635C2FFC02172FEB9A764FE71F8EB" box="[373,469,1859,1884]" name="Djibouti" pageId="8" pageNumber="9">Djibouti</collectingCountry>
, Gulf of 
<collectingRegion id="8415BBB0FFC02172FD85A764FD0BF8EB" box="[585,687,1859,1884]" country="Djibouti" name="Tadjourah" pageId="8" pageNumber="9">Tadjoura</collectingRegion>
, 
<location id="430E2389FFC02172FD0CA764FC81F8EB" LSID="urn:lsid:plazi:treatment:CE78C444FFC02174FF5BA674FE27FE67:430E2389FFC02172FD0CA764FC81F8EB" box="[704,805,1859,1884]" country="Yemen" municipality="Gulf of Aden" name="Red Sea" pageId="8" pageNumber="9" stateProvince="Tadjoura">Red Sea</location>
, 
<collectingDate id="222BAA7AFFC02172FCFAA763FC74F8EB" box="[822,976,1859,1884]" pageId="8" pageNumber="9" value="1904-02-10">
<date id="326F5392FFC02172FCFAA763FC74F8EB" box="[822,976,1859,1884]" pageId="8" pageNumber="9" value="1904-02-10">10 Feb 1904</date>
</collectingDate>
, in the muddy sand to the east of the 
<collectorName id="EB241084FFC02172FF5BA740FEA9F837" box="[151,269,1895,1920]" pageId="8" pageNumber="9">Residence</collectorName>
, coll. 
<collectorName id="EB241084FFC02172FE9CA74FFE68F837" box="[336,460,1895,1920]" pageId="8" pageNumber="9">C. Gravier.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0ECB26D9FFC02177FF0BA7ACFC42FB44" lastPageId="13" lastPageNumber="14" pageId="8" pageNumber="9" type="description">
<paragraph id="466E7552FFC02173FF0BA7ACFDE0FF07" blockId="8.[151,1437,1823,2028]" lastBlockId="9.[151,1437,151,1832]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="74A5A940FFC02172FF0BA7ACFEFDF813" bold="true" box="[199,345,1931,1956]" pageId="8" pageNumber="9">Description.</emphasis>
<typeStatus id="996ACBF0FFC02172FEADA7ACFE6DF813" box="[353,457,1931,1956]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
incomplete, with 115 chaetigers, L10= 
<quantity id="8129D8B7FFC02172FC5CA7ABFC71F814" box="[912,981,1932,1956]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="9" unit="mm" value="4.0">4 mm</quantity>
, W10= 
<quantity id="8129D8B7FFC02172FBFDA7ABFB2EF814" box="[1073,1162,1932,1956]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="9" unit="mm" value="0.7">0.7 mm</quantity>
, the fragment with TL= 
<quantity id="8129D8B7FFC02172FF5BA797FF4EF870" box="[151,234,1968,1992]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="9" unit="mm" value="40.0">40 mm</quantity>
. Anterior region with convex dorsum, and flat ventrum, without groove; body depressed from chaetiger 9, widest at chaetiger 25, tapering after chaetiger 57. In non-type specimens (MNHN-A148) some areas of the anterior region with brown coloration.
</paragraph>
<paragraph id="466E7552FFC12173FF0BA09BFAE3FED3" blockId="9.[151,1437,151,1832]" pageId="9" pageNumber="10">
Prostomium in poor condition, 
<quantity id="8129D8B7FFC12173FDE8A09BFDDEFF63" box="[548,634,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="10" unit="mm" value="0.5">0.5 mm</quantity>
long, 
<quantity id="8129D8B7FFC12173FD73A09BFCB1FF63" box="[703,789,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="10" unit="mm" value="0.5">0.5 mm</quantity>
wide, frontally rounded, without median sulcus (
<figureCitation id="DEEA69D7FFC12173FAF6A09BFF0AFF4F" captionStart="FIGURE 3" captionStartId="10.[151,250,1908,1930]" captionTargetBox="[219,1375,193,1892]" captionTargetId="figure@10.[219,1375,193,1892]" captionTargetPageId="10" captionText="FIGURE 3. Paucibranchia adenensis (Gravier, 1900) n. comb. A. Anterior end with branchiae, dorsal view; B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, dorsal view; E. Anterior end with branchiae, dorsal view; F. Anterior end, ventral view. A, B, C from holotype MNHN-531; D, E, F from non-type MNHN-A148. Scale bars: A, 2 mm; B, 1 mm, C, 2.6 mm, D, F, 0.7 mm; E, 1.3 mm." httpUri="https://zenodo.org/record/1453037/files/figure.png" pageId="9" pageNumber="10">Fig. 3B– D</figureCitation>
), ventral sulcus deep (
<figureCitation id="DEEA69D7FFC12173FE0EA0F8FDB9FF4F" box="[450,541,223,248]" captionStart="FIGURE 3" captionStartId="10.[151,250,1908,1930]" captionTargetBox="[219,1375,193,1892]" captionTargetId="figure@10.[219,1375,193,1892]" captionTargetPageId="10" captionText="FIGURE 3. Paucibranchia adenensis (Gravier, 1900) n. comb. A. Anterior end with branchiae, dorsal view; B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, dorsal view; E. Anterior end with branchiae, dorsal view; F. Anterior end, ventral view. A, B, C from holotype MNHN-531; D, E, F from non-type MNHN-A148. Scale bars: A, 2 mm; B, 1 mm, C, 2.6 mm, D, F, 0.7 mm; E, 1.3 mm." httpUri="https://zenodo.org/record/1453037/files/figure.png" pageId="9" pageNumber="10">Fig. 3F</figureCitation>
). Prostomial appendages in a semicircle, equidistant (MNHN-A148). Palps reaching second peristomial ring; lateral antennae and median antenna broken, in specimen MNHN-A148 reaching third and four chaetiger, respectively. Palpophores and ceratophores ring-shaped, short, slender; palpostyles and ceratostyles tapering, thick, without articulation. Eyes rounded, brown, between palps and lateral antennae.
</paragraph>
<paragraph id="466E7552FFC12173FF0BA148FDD1FE67" blockId="9.[151,1437,151,1832]" pageId="9" pageNumber="10">
Peristomium slightly wider than prostomium (
<quantity id="8129D8B7FFC12173FD16A157FC90FE30" box="[730,820,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="10" unit="mm" value="0.4">0.4 mm</quantity>
long, 
<quantity id="8129D8B7FFC12173FC4CA148FC7DFE30" box="[896,985,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="10" unit="mm" value="0.6">0.6 mm</quantity>
wide), first ring two times longer than second ring; separation between rings distinct on all sides (
<figureCitation id="DEEA69D7FFC12173FCF7A1B3FC11FE1B" box="[827,949,404,429]" captionStart="FIGURE 3" captionStartId="10.[151,250,1908,1930]" captionTargetBox="[219,1375,193,1892]" captionTargetId="figure@10.[219,1375,193,1892]" captionTargetPageId="10" captionText="FIGURE 3. Paucibranchia adenensis (Gravier, 1900) n. comb. A. Anterior end with branchiae, dorsal view; B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, dorsal view; E. Anterior end with branchiae, dorsal view; F. Anterior end, ventral view. A, B, C from holotype MNHN-531; D, E, F from non-type MNHN-A148. Scale bars: A, 2 mm; B, 1 mm, C, 2.6 mm, D, F, 0.7 mm; E, 1.3 mm." httpUri="https://zenodo.org/record/1453037/files/figure.png" pageId="9" pageNumber="10">Fig. 3A–F</figureCitation>
). Inferior lip broken, in specimen MNHN- A148 without central depression (
<figureCitation id="DEEA69D7FFC12173FDDFA190FDCCFE67" box="[531,616,439,464]" captionStart="FIGURE 3" captionStartId="10.[151,250,1908,1930]" captionTargetBox="[219,1375,193,1892]" captionTargetId="figure@10.[219,1375,193,1892]" captionTargetPageId="10" captionText="FIGURE 3. Paucibranchia adenensis (Gravier, 1900) n. comb. A. Anterior end with branchiae, dorsal view; B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, dorsal view; E. Anterior end with branchiae, dorsal view; F. Anterior end, ventral view. A, B, C from holotype MNHN-531; D, E, F from non-type MNHN-A148. Scale bars: A, 2 mm; B, 1 mm, C, 2.6 mm, D, F, 0.7 mm; E, 1.3 mm." httpUri="https://zenodo.org/record/1453037/files/figure.png" pageId="9" pageNumber="10">Fig. 3F</figureCitation>
).
</paragraph>
<paragraph id="466E7552FFC12173FF0BA1FBFCD9FC8F" blockId="9.[151,1437,151,1832]" pageId="9" pageNumber="10">
<materialsCitation id="F6B97F0FFFC12173FF0BA1FBFC0EFDD7" ID-GBIF-Occurrence="1915273797" collectionCode="MII" location="Maxillary" pageId="9" pageNumber="10" specimenCode="MNHN-A148" specimenCount="1" stateProvince="MII">
Maxillary apparatus in poor condition, according to Gravier’s drawings with MF= 1+1, 7+?, 10+0,?+?,?+?, (
<bibRefCitation id="224008A3FFC12173FF52A1D8FE5FFDAF" box="[158,507,511,536]" pageId="9" pageNumber="10" refId="ref54865" refString="Gravier, C. (1900) Contribution a l'etude des annelids polychetes de la Mer Rouge. Premiere partie. Nouvelles Archives du Museum d'Histoire Naturelle, Paris, 4 e Serie, 2 (2), 137 - 282." type="journal article">Gravier 1900, Pl. XIV, Fig. 92</bibRefCitation>
). Non-type specimen 
<specimenCode id="1677DD29FFC12173FD3DA227FC2AFDAF" box="[753,910,512,536]" collectionCode="MNHN-A" pageId="9" pageNumber="10">MNHN-A148</specimenCode>
with MF= 1+1, 7+7, 7+0, 5+9, 1+1 (
<figureCitation id="DEEA69D7FFC12173FAF9A1D8FA34FDAF" box="[1333,1424,511,536]" captionStart="FIGURE 4" captionStartId="11.[151,250,1932,1954]" captionTargetBox="[336,1232,245,1870]" captionTargetId="figure@11.[336,1250,245,1870]" captionTargetPageId="11" captionText="FIGURE 4. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Maxillary apparatus, dorsal view; B. Mandibles; C. Left MII-III-IV-V, dorsal view; D. Maxilla I, dorsal view. Apparatus stained with methyl green. Scale bars: A, C, 50 µm; B, 150 µm; D, 130 µm. al-MIV= attachment lamellae in MIV, LMI-MII= ligament between MI and MII." httpUri="https://zenodo.org/record/1453039/files/figure.png" pageId="9" pageNumber="10">Fig. 4A</figureCitation>
). 
<location id="430E2389FFC12173FF5BA203FEA1FD8A" LSID="urn:lsid:plazi:treatment:CE78C444FFC02174FF5BA674FE27FE67:430E2389FFC12173FF5BA203FEA1FD8A" box="[151,261,548,573]" name="Maxillary" pageId="9" pageNumber="10" stateProvince="MII">Maxillary</location>
carriers 1.6 times shorter than length of MI. MI forceps-like; closing system 8 times shorter than length of MI; ligament between MI and 
<collectionCode id="20C0ED97FFC12173FDE8A26FFDF6FDE8" box="[548,594,584,607]" country="Ireland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13666" name="Museum of Irish Industry" pageId="9" pageNumber="10">
<collectingRegion id="8415BBB0FFC12173FDE8A26FFDF6FDE8" box="[548,594,584,607]" country="Japan" name="Mie" pageId="9" pageNumber="10">MII</collectingRegion>
</collectionCode>
, not sclerotized (
<figureCitation id="DEEA69D7FFC12173FCD1A260FC05FDD7" box="[797,929,583,608]" captionStart="FIGURE 4" captionStartId="11.[151,250,1932,1954]" captionTargetBox="[336,1232,245,1870]" captionTargetId="figure@11.[336,1250,245,1870]" captionTargetPageId="11" captionText="FIGURE 4. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Maxillary apparatus, dorsal view; B. Mandibles; C. Left MII-III-IV-V, dorsal view; D. Maxilla I, dorsal view. Apparatus stained with methyl green. Scale bars: A, C, 50 µm; B, 150 µm; D, 130 µm. al-MIV= attachment lamellae in MIV, LMI-MII= ligament between MI and MII." httpUri="https://zenodo.org/record/1453039/files/figure.png" pageId="9" pageNumber="10">Fig. 4A, D</figureCitation>
)
</materialsCitation>
. 
<materialsCitation id="F6B97F0FFFC12173FC75A26FFB8DFDD7" ID-GBIF-Occurrence="1915273771" box="[953,1065,583,608]" collectionCode="MII" location="MII" pageId="9" pageNumber="10" specimenCount="1" stateProvince="MII">
<collectionCode id="20C0ED97FFC12173FC75A26FFC41FDE8" box="[953,997,584,607]" country="Ireland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13666" name="Museum of Irish Industry" pageId="9" pageNumber="10">
<collectingRegion id="8415BBB0FFC12173FC75A26FFC41FDE8" box="[953,997,584,607]" country="Japan" name="Mie" pageId="9" pageNumber="10">MII</collectingRegion>
</collectionCode>
wide
</materialsCitation>
; 
<materialsCitation id="F6B97F0FFFC12173FBF5A260FD61FD33" ID-GBIF-Occurrence="1915273781" collectionCode="MII" location="MII" pageId="9" pageNumber="10" specimenCount="1" stateProvince="MII">
teeth recurved, sharped; cavity opening oval, 6 times shorter than length of 
<collectionCode id="20C0ED97FFC12173FD5AA24AFD61FD33" box="[662,709,621,644]" country="Ireland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13666" name="Museum of Irish Industry" pageId="9" pageNumber="10">
<collectingRegion id="8415BBB0FFC12173FD5AA24AFD61FD33" box="[662,709,621,644]" country="Japan" name="Mie" pageId="9" pageNumber="10">MII</collectingRegion>
</collectionCode>
</materialsCitation>
; 
<materialsCitation id="F6B97F0FFFC12173FD1EA24BFE87FD1F" ID-GBIF-Occurrence="1915273790" collectionCode="MII, MIII, MIV" location="MII" pageId="9" pageNumber="10" specimenCount="1" stateProvince="MII">
ligament between 
<collectionCode id="20C0ED97FFC12173FC68A24AFC74FD33" box="[932,976,621,644]" country="Ireland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13666" name="Museum of Irish Industry" pageId="9" pageNumber="10">
<collectingRegion id="8415BBB0FFC12173FC68A24AFC74FD33" box="[932,976,621,644]" country="Japan" name="Mie" pageId="9" pageNumber="10">MII</collectingRegion>
</collectionCode>
and 
<collectionCode id="20C0ED97FFC12173FBC6A24AFB9BFD33" box="[1034,1087,621,644]" pageId="9" pageNumber="10">MIII</collectionCode>
and right 
<collectionCode id="20C0ED97FFC12173FB7BA24AFB49FD33" box="[1207,1261,621,644]" country="Italy" httpUri="http://grbio.org/cool/mev6-t33n" name="Universita' degli Studi di Milano, Medicina Veterinaria, Sez. Parassitologia" pageId="9" pageNumber="10">MIV</collectionCode>
not sclerotized (
<figureCitation id="DEEA69D7FFC12173FF53A2A8FEBFFD1F" box="[159,283,655,680]" captionStart="FIGURE 4" captionStartId="11.[151,250,1932,1954]" captionTargetBox="[336,1232,245,1870]" captionTargetId="figure@11.[336,1250,245,1870]" captionTargetPageId="11" captionText="FIGURE 4. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Maxillary apparatus, dorsal view; B. Mandibles; C. Left MII-III-IV-V, dorsal view; D. Maxilla I, dorsal view. Apparatus stained with methyl green. Scale bars: A, C, 50 µm; B, 150 µm; D, 130 µm. al-MIV= attachment lamellae in MIV, LMI-MII= ligament between MI and MII." httpUri="https://zenodo.org/record/1453039/files/figure.png" pageId="9" pageNumber="10">Fig. 4A, C</figureCitation>
)
</materialsCitation>
. MIII short; 
<materialsCitation id="F6B97F0FFFC12173FE7FA2A8FCD9FC8F" ID-GBIF-Occurrence="1915273768" collectionCode="MIV" location="Left" pageId="9" pageNumber="10" specimenCount="1" stateProvince="MII">
with triangular teeth; with attachment lamella slightly sclerotized (
<figureCitation id="DEEA69D7FFC12173FB6CA2A8FABFFD1F" box="[1184,1307,655,680]" captionStart="FIGURE 4" captionStartId="11.[151,250,1932,1954]" captionTargetBox="[336,1232,245,1870]" captionTargetId="figure@11.[336,1250,245,1870]" captionTargetPageId="11" captionText="FIGURE 4. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Maxillary apparatus, dorsal view; B. Mandibles; C. Left MII-III-IV-V, dorsal view; D. Maxilla I, dorsal view. Apparatus stained with methyl green. Scale bars: A, C, 50 µm; B, 150 µm; D, 130 µm. al-MIV= attachment lamellae in MIV, LMI-MII= ligament between MI and MII." httpUri="https://zenodo.org/record/1453039/files/figure.png" pageId="9" pageNumber="10">Fig. 4A, C</figureCitation>
). 
<location id="430E2389FFC12173FAFCA2A8FAFBFD1F" LSID="urn:lsid:plazi:treatment:CE78C444FFC02174FF5BA674FE27FE67:430E2389FFC12173FAFCA2A8FAFBFD1F" box="[1328,1375,655,680]" name="Left" pageId="9" pageNumber="10" stateProvince="MII">Left</location>
<collectionCode id="20C0ED97FFC12173FAAAA2B7FA38FD10" box="[1382,1436,656,679]" country="Italy" httpUri="http://grbio.org/cool/mev6-t33n" name="Universita' degli Studi di Milano, Medicina Veterinaria, Sez. Parassitologia" pageId="9" pageNumber="10">MIV</collectionCode>
with basal tooth smaller; attachment lamella semicircular, wide, situated along length of posterior edge of maxilla. Right 
<collectionCode id="20C0ED97FFC12173FF11A2FFFEB7FD58" box="[221,275,728,751]" country="Italy" httpUri="http://grbio.org/cool/mev6-t33n" name="Universita' degli Studi di Milano, Medicina Veterinaria, Sez. Parassitologia" pageId="9" pageNumber="10">MIV</collectionCode>
with small teeth; attachment lamella wide, semicircular, better developed in the middle, situated along length of posterior edge of maxilla (
<figureCitation id="DEEA69D7FFC12173FDF3A2DBFD65FCA3" box="[575,705,764,789]" captionStart="FIGURE 4" captionStartId="11.[151,250,1932,1954]" captionTargetBox="[336,1232,245,1870]" captionTargetId="figure@11.[336,1250,245,1870]" captionTargetPageId="11" captionText="FIGURE 4. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Maxillary apparatus, dorsal view; B. Mandibles; C. Left MII-III-IV-V, dorsal view; D. Maxilla I, dorsal view. Apparatus stained with methyl green. Scale bars: A, C, 50 µm; B, 150 µm; D, 130 µm. al-MIV= attachment lamellae in MIV, LMI-MII= ligament between MI and MII." httpUri="https://zenodo.org/record/1453039/files/figure.png" pageId="9" pageNumber="10">Fig. 4A, C</figureCitation>
). MV square, with a short rounded tooth (
<figureCitation id="DEEA69D7FFC12173FB7FA2DBFAABFCA3" box="[1203,1295,764,789]" captionStart="FIGURE 4" captionStartId="11.[151,250,1932,1954]" captionTargetBox="[336,1232,245,1870]" captionTargetId="figure@11.[336,1250,245,1870]" captionTargetPageId="11" captionText="FIGURE 4. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Maxillary apparatus, dorsal view; B. Mandibles; C. Left MII-III-IV-V, dorsal view; D. Maxilla I, dorsal view. Apparatus stained with methyl green. Scale bars: A, C, 50 µm; B, 150 µm; D, 130 µm. al-MIV= attachment lamellae in MIV, LMI-MII= ligament between MI and MII." httpUri="https://zenodo.org/record/1453039/files/figure.png" pageId="9" pageNumber="10">Fig. 4C</figureCitation>
). Mandibles translucent; cutting plates whitish, with 12 growth rings (
<figureCitation id="DEEA69D7FFC12173FCDBA338FCD4FC8F" box="[791,880,799,824]" captionStart="FIGURE 4" captionStartId="11.[151,250,1932,1954]" captionTargetBox="[336,1232,245,1870]" captionTargetId="figure@11.[336,1250,245,1870]" captionTargetPageId="11" captionText="FIGURE 4. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Maxillary apparatus, dorsal view; B. Mandibles; C. Left MII-III-IV-V, dorsal view; D. Maxilla I, dorsal view. Apparatus stained with methyl green. Scale bars: A, C, 50 µm; B, 150 µm; D, 130 µm. al-MIV= attachment lamellae in MIV, LMI-MII= ligament between MI and MII." httpUri="https://zenodo.org/record/1453039/files/figure.png" pageId="9" pageNumber="10">Fig. 4B</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="466E7552FFC12173FF0BA363FDA9FC12" blockId="9.[151,1437,151,1832]" pageId="9" pageNumber="10">
Branchiae pectinate with up to 11 filaments, in chaetigers 15–30 (
<figureCitation id="DEEA69D7FFC12173FC1DA363FBF8FCEB" box="[977,1116,836,861]" captionStart="FIGURE 3" captionStartId="10.[151,250,1908,1930]" captionTargetBox="[219,1375,193,1892]" captionTargetId="figure@10.[219,1375,193,1892]" captionTargetPageId="10" captionText="FIGURE 3. Paucibranchia adenensis (Gravier, 1900) n. comb. A. Anterior end with branchiae, dorsal view; B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, dorsal view; E. Anterior end with branchiae, dorsal view; F. Anterior end, ventral view. A, B, C from holotype MNHN-531; D, E, F from non-type MNHN-A148. Scale bars: A, 2 mm; B, 1 mm, C, 2.6 mm, D, F, 0.7 mm; E, 1.3 mm." httpUri="https://zenodo.org/record/1453037/files/figure.png" pageId="9" pageNumber="10">Figs. 3A, E</figureCitation>
; 
<figureCitation id="DEEA69D7FFC12173FBA0A363FB37FCEA" box="[1132,1171,836,861]" captionStart="FIGURE 5" captionStartId="12.[151,250,1960,1982]" captionTargetBox="[182,1414,193,1947]" captionTargetId="figure@12.[182,1414,193,1947]" captionTargetPageId="12" captionText="FIGURE 5. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Parapodium 3; B. Parapodium 11; C. Parapodium 26; D. Parapodium 70; E. Parapodium 158; F. Parapodium 216. All in anterior view. Scale bars: A–F, 0.7 mm." httpUri="https://zenodo.org/record/1453041/files/figure.png" pageId="9" pageNumber="10">5A</figureCitation>
). Number of branchial filaments per chaetiger in order anterior-posterior: 8, 8, 8, 9, 10, 10, 10, 11, 11, 10, 10, 10, 9, 9, 8, 6. Branchial filaments longer than dorsal cirri.
</paragraph>
<paragraph id="466E7552FFC12173FF0BA388FC63FABA" blockId="9.[151,1437,151,1832]" pageId="9" pageNumber="10">
First two parapodia smallest; most developed in chaetigers 3–16, following ones becoming gradually smaller. Notopodial cirri conical, increasing in size from chaetiger 3 (Ldc3: 
<quantity id="8129D8B7FFC12173FC47A3F3FC56FC5B" box="[907,1010,980,1004]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="10" unit="mm" value="0.17">0.17 mm</quantity>
; Ldc26: 
<quantity id="8129D8B7FFC12173FB9FA3F3FB1FFC5B" box="[1107,1211,980,1004]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="mm" value="0.2">0.20 mm</quantity>
), from chaetiger 40, gradually decreasing in size (Ldc216: 
<quantity id="8129D8B7FFC12173FD98A3DFFD64FBA7" box="[596,704,1016,1040]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="10" unit="mm" value="0.13">0.13 mm</quantity>
), in posterior region almost the same length than pre-branchial region ones; Hayashi &amp; Yamane’s organ present (
<figureCitation id="DEEA69D7FFC12173FD0EA43BFC9FFB82" box="[706,827,1052,1077]" captionStart="FIGURE 5" captionStartId="12.[151,250,1960,1982]" captionTargetBox="[182,1414,193,1947]" captionTargetId="figure@12.[182,1414,193,1947]" captionTargetPageId="12" captionText="FIGURE 5. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Parapodium 3; B. Parapodium 11; C. Parapodium 26; D. Parapodium 70; E. Parapodium 158; F. Parapodium 216. All in anterior view. Scale bars: A–F, 0.7 mm." httpUri="https://zenodo.org/record/1453041/files/figure.png" pageId="9" pageNumber="10">Fig. 5A–F</figureCitation>
). Prechaetal lobes as a transverse fold in all chaetigers (
<figureCitation id="DEEA69D7FFC12173FF53A418FEB3FBEF" box="[159,279,1087,1112]" captionStart="FIGURE 5" captionStartId="12.[151,250,1960,1982]" captionTargetBox="[182,1414,193,1947]" captionTargetId="figure@12.[182,1414,193,1947]" captionTargetPageId="12" captionText="FIGURE 5. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Parapodium 3; B. Parapodium 11; C. Parapodium 26; D. Parapodium 70; E. Parapodium 158; F. Parapodium 216. All in anterior view. Scale bars: A–F, 0.7 mm." httpUri="https://zenodo.org/record/1453041/files/figure.png" pageId="9" pageNumber="10">Fig. 5A–F</figureCitation>
). Chaetal lobe in chaetigers 1–28, rounded, shorter than postchaetal lobe, with aciculae emerging dorsal to midline; from chaetiger 29, triangular, longer than other lobes, with acicula emerging in midline (
<figureCitation id="DEEA69D7FFC12173FADAA443FA34FBCA" box="[1302,1424,1124,1149]" captionStart="FIGURE 5" captionStartId="12.[151,250,1960,1982]" captionTargetBox="[182,1414,193,1947]" captionTargetId="figure@12.[182,1414,193,1947]" captionTargetPageId="12" captionText="FIGURE 5. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Parapodium 3; B. Parapodium 11; C. Parapodium 26; D. Parapodium 70; E. Parapodium 158; F. Parapodium 216. All in anterior view. Scale bars: A–F, 0.7 mm." httpUri="https://zenodo.org/record/1453041/files/figure.png" pageId="9" pageNumber="10">Fig. 5A–F</figureCitation>
). Postchaetal lobes well developed in chaetigers 1–48, bluntly conical in pre-branchial chaetigers, elongated and thinner in branchial region; decreasing in size in chaetigers 31–48, following ones inconspicuous (
<figureCitation id="DEEA69D7FFC12173FAD9A48BFA28FB72" box="[1301,1420,1196,1221]" captionStart="FIGURE 5" captionStartId="12.[151,250,1960,1982]" captionTargetBox="[182,1414,193,1947]" captionTargetId="figure@12.[182,1414,193,1947]" captionTargetPageId="12" captionText="FIGURE 5. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Parapodium 3; B. Parapodium 11; C. Parapodium 26; D. Parapodium 70; E. Parapodium 158; F. Parapodium 216. All in anterior view. Scale bars: A–F, 0.7 mm." httpUri="https://zenodo.org/record/1453041/files/figure.png" pageId="9" pageNumber="10">Fig. 5C–F</figureCitation>
). Ventral cirri digitiform in chaetigers 1–7; in chaetigers 8–38 with oval swollen base and digitiform tip; from chaetiger 39, conical, gradually reducing in size posteriorly (
<figureCitation id="DEEA69D7FFC12173FCF3A4D3FC1FFABA" box="[831,955,1268,1293]" captionStart="FIGURE 5" captionStartId="12.[151,250,1960,1982]" captionTargetBox="[182,1414,193,1947]" captionTargetId="figure@12.[182,1414,193,1947]" captionTargetPageId="12" captionText="FIGURE 5. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Parapodium 3; B. Parapodium 11; C. Parapodium 26; D. Parapodium 70; E. Parapodium 158; F. Parapodium 216. All in anterior view. Scale bars: A–F, 0.7 mm." httpUri="https://zenodo.org/record/1453041/files/figure.png" pageId="9" pageNumber="10">Fig. 5A–C</figureCitation>
).
</paragraph>
<paragraph id="466E7552FFC12173FF0BA530FF4AFAE2" blockId="9.[151,1437,151,1832]" pageId="9" pageNumber="10">
Aciculae blunt, translucent (
<figureCitation id="DEEA69D7FFC12173FDCAA530FD24FA87" box="[518,640,1303,1328]" captionStart="FIGURE 5" captionStartId="12.[151,250,1960,1982]" captionTargetBox="[182,1414,193,1947]" captionTargetId="figure@12.[182,1414,193,1947]" captionTargetPageId="12" captionText="FIGURE 5. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Parapodium 3; B. Parapodium 11; C. Parapodium 26; D. Parapodium 70; E. Parapodium 158; F. Parapodium 216. All in anterior view. Scale bars: A–F, 0.7 mm." httpUri="https://zenodo.org/record/1453041/files/figure.png" pageId="9" pageNumber="10">Fig. 5A–F</figureCitation>
). First 14 chaetigers with 2 aciculae; from chaetiger 15, with only one acicula.
</paragraph>
<paragraph id="466E7552FFC12173FF0BA578FEC2F957" blockId="9.[151,1437,151,1832]" pageId="9" pageNumber="10">
Limbate chaetae of two sizes in same chaetiger, larger in anterior region, reduced in number around chaetiger 27. Two 
<typeStatus id="996ACBF0FFC12173FF31A5A2FE9CFA2A" box="[253,312,1413,1437]" pageId="9" pageNumber="10">types</typeStatus>
of pectinate chaetae, in anterior chaetigers, isodonts narrow with long and slender teeth, with 1–2 pectinate, with 5 teeth, with transverse distal edge (
<figureCitation id="DEEA69D7FFC12173FD16A580FC91FA77" box="[730,821,1447,1472]" captionStart="FIGURE 6" captionStartId="13.[151,250,765,787]" captionTargetBox="[160,1402,193,743]" captionTargetId="figure@13.[151,1436,193,743]" captionTargetPageId="13" captionText="FIGURE 6. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Compound falciger, long blade, chaetiger, 11; B. Compound falciger, median blade, chaetiger, 11; C. Compound falciger, short blade, chaetiger, 216; D. Isodonts pectinate narrow with long and slender teeth, chaetiger 3; E. Isodonts pectinate narrow with short and slender teeth, chaetiger 216; F. Subacicular hook, distal end, chaetiger 158. Scale bars: A, 12.5 µm; B, 20 µm; C, F, 9.8 µm; D, 14 µm, E, 4.9 µm." httpUri="https://zenodo.org/record/1453043/files/figure.png" pageId="9" pageNumber="10">Fig. 6D</figureCitation>
); in median-posterior chaetigers, isodonts narrow with short and slender teeth, 2–4 pectinate, with 10–12 teeth, with transverse distal edge (
<figureCitation id="DEEA69D7FFC12173FB81A5EBFB00FA52" box="[1101,1188,1484,1509]" captionStart="FIGURE 6" captionStartId="13.[151,250,765,787]" captionTargetBox="[160,1402,193,743]" captionTargetId="figure@13.[151,1436,193,743]" captionTargetPageId="13" captionText="FIGURE 6. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Compound falciger, long blade, chaetiger, 11; B. Compound falciger, median blade, chaetiger, 11; C. Compound falciger, short blade, chaetiger, 216; D. Isodonts pectinate narrow with long and slender teeth, chaetiger 3; E. Isodonts pectinate narrow with short and slender teeth, chaetiger 216; F. Subacicular hook, distal end, chaetiger 158. Scale bars: A, 12.5 µm; B, 20 µm; C, F, 9.8 µm; D, 14 µm, E, 4.9 µm." httpUri="https://zenodo.org/record/1453043/files/figure.png" pageId="9" pageNumber="10">Fig. 6E</figureCitation>
). Compound falcigers present in all chaetigers; in anterior region with blades of two sizes (longer 105 µm, 
<figureCitation id="DEEA69D7FFC12173FB9BA5C8FB14F9BF" box="[1111,1200,1519,1544]" captionStart="FIGURE 6" captionStartId="13.[151,250,765,787]" captionTargetBox="[160,1402,193,743]" captionTargetId="figure@13.[151,1436,193,743]" captionTargetPageId="13" captionText="FIGURE 6. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Compound falciger, long blade, chaetiger, 11; B. Compound falciger, median blade, chaetiger, 11; C. Compound falciger, short blade, chaetiger, 216; D. Isodonts pectinate narrow with long and slender teeth, chaetiger 3; E. Isodonts pectinate narrow with short and slender teeth, chaetiger 216; F. Subacicular hook, distal end, chaetiger 158. Scale bars: A, 12.5 µm; B, 20 µm; C, F, 9.8 µm; D, 14 µm, E, 4.9 µm." httpUri="https://zenodo.org/record/1453043/files/figure.png" pageId="9" pageNumber="10">Fig. 6A</figureCitation>
; smaller 90 µm, 
<figureCitation id="DEEA69D7FFC12173FABCA5C8FF1EF99A" captionStart="FIGURE 6" captionStartId="13.[151,250,765,787]" captionTargetBox="[160,1402,193,743]" captionTargetId="figure@13.[151,1436,193,743]" captionTargetPageId="13" captionText="FIGURE 6. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Compound falciger, long blade, chaetiger, 11; B. Compound falciger, median blade, chaetiger, 11; C. Compound falciger, short blade, chaetiger, 216; D. Isodonts pectinate narrow with long and slender teeth, chaetiger 3; E. Isodonts pectinate narrow with short and slender teeth, chaetiger 216; F. Subacicular hook, distal end, chaetiger 158. Scale bars: A, 12.5 µm; B, 20 µm; C, F, 9.8 µm; D, 14 µm, E, 4.9 µm." httpUri="https://zenodo.org/record/1453043/files/figure.png" pageId="9" pageNumber="10">Fig. 6B</figureCitation>
), smaller more abundant; all with triangular teeth, of similar size, distal tooth directed upward, proximal tooth directed laterally; in median-posterior chaetigers with all blades of similar size, slightly shorter than blades of anterior chaetigers (84 µm, 
<figureCitation id="DEEA69D7FFC12173FE17A67BFD92F9C2" box="[475,566,1628,1653]" captionStart="FIGURE 6" captionStartId="13.[151,250,765,787]" captionTargetBox="[160,1402,193,743]" captionTargetId="figure@13.[151,1436,193,743]" captionTargetPageId="13" captionText="FIGURE 6. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Compound falciger, long blade, chaetiger, 11; B. Compound falciger, median blade, chaetiger, 11; C. Compound falciger, short blade, chaetiger, 216; D. Isodonts pectinate narrow with long and slender teeth, chaetiger 3; E. Isodonts pectinate narrow with short and slender teeth, chaetiger 216; F. Subacicular hook, distal end, chaetiger 158. Scale bars: A, 12.5 µm; B, 20 µm; C, F, 9.8 µm; D, 14 µm, E, 4.9 µm." httpUri="https://zenodo.org/record/1453043/files/figure.png" pageId="9" pageNumber="10">Fig. 6C</figureCitation>
), with triangular teeth, distal tooth shorter than proximal, directed upward, proximal tooth directed laterally. Subacicular hooks bidentate, translucent, starting from chaetigers 29L–30R, one per chaetiger; with triangular teeth, distal tooth smaller than proximal, directed upward; proximal tooth directed laterally (
<figureCitation id="DEEA69D7FFC12173FECEA6E0FEFDF957" box="[258,345,1735,1760]" captionStart="FIGURE 6" captionStartId="13.[151,250,765,787]" captionTargetBox="[160,1402,193,743]" captionTargetId="figure@13.[151,1436,193,743]" captionTargetPageId="13" captionText="FIGURE 6. Paucibranchia adenensis (Gravier, 1900) n. comb., non-type MNHN-A148. A. Compound falciger, long blade, chaetiger, 11; B. Compound falciger, median blade, chaetiger, 11; C. Compound falciger, short blade, chaetiger, 216; D. Isodonts pectinate narrow with long and slender teeth, chaetiger 3; E. Isodonts pectinate narrow with short and slender teeth, chaetiger 216; F. Subacicular hook, distal end, chaetiger 158. Scale bars: A, 12.5 µm; B, 20 µm; C, F, 9.8 µm; D, 14 µm, E, 4.9 µm." httpUri="https://zenodo.org/record/1453043/files/figure.png" pageId="9" pageNumber="10">Fig. 6E</figureCitation>
).
</paragraph>
<paragraph id="466E7552FFC12173FF0BA6CBFECBF89F" blockId="9.[151,1437,151,1832]" pageId="9" pageNumber="10">Pygidium with two pairs of anal cirri; dorsal pair as long as last four chaetigers; ventral pair short, as long as two last chaetigers.</paragraph>
<caption id="12AE25DAFFC22170FF5BA753FDFCF852" httpUri="https://zenodo.org/record/1453037/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,250,1908,1930]" targetBox="[219,1375,193,1892]" targetPageId="10">
<paragraph id="466E7552FFC22170FF5BA753FDFCF852" blockId="10.[151,1436,1908,2021]" pageId="10" pageNumber="11">
<emphasis id="74A5A940FFC22170FF5BA753FEB7F83D" bold="true" box="[151,275,1908,1930]" pageId="10" pageNumber="11">FIGURE 3.</emphasis>
<taxonomicName id="81D10ED1FFC22170FED6A753FD1AF83D" authority="(Gravier, 1900)" authorityName="Molina-Acevedo" authorityYear="2018" baseAuthorityName="Gravier" baseAuthorityYear="1900" box="[282,702,1908,1930]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="10" pageNumber="11" phylum="Annelida" rank="species" species="adenensis" status="comb. nov.">
<emphasis id="74A5A940FFC22170FED6A753FDBFF83D" box="[282,539,1908,1930]" italics="true" pageId="10" pageNumber="11">Paucibranchia adenensis</emphasis>
(Gravier, 1900)
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC22170FD09A753FC86F83D" box="[709,802,1908,1930]" pageId="10" pageNumber="11">
<emphasis id="74A5A940FFC22170FD09A753FC86F83D" bold="true" box="[709,802,1908,1930]" pageId="10" pageNumber="11">n. comb.</emphasis>
</taxonomicNameLabel>
A. Anterior end with branchiae, dorsal view; B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, dorsal view; E. Anterior end with branchiae, dorsal view; F. Anterior end, ventral view. A, B, C from holotype MNHN-531; D, E, F from non-type MNHN-A148. Scale bars: A, 2 mm; B, 1 mm, C, 2.6 mm, D, F, 0.7 mm; E, 1.3 mm.
</paragraph>
</caption>
<caption id="12AE25DAFFC32171FF5BA7ABFA8CF868" httpUri="https://zenodo.org/record/1453039/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1932,1954]" targetBox="[336,1232,245,1870]" targetPageId="11">
<paragraph id="466E7552FFC32171FF5BA7ABFA8CF868" blockId="11.[151,1436,1932,2015]" pageId="11" pageNumber="12">
<emphasis id="74A5A940FFC32171FF5BA7ABFEB7F815" bold="true" box="[151,275,1932,1954]" pageId="11" pageNumber="12">FIGURE 4.</emphasis>
<taxonomicName id="81D10ED1FFC32171FED5A7ABFD19F815" authority="(Gravier, 1900)" authorityName="Molina-Acevedo" authorityYear="2018" baseAuthorityName="Gravier" baseAuthorityYear="1900" box="[281,701,1932,1954]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="adenensis" status="comb. nov.">
<emphasis id="74A5A940FFC32171FED5A7ABFDBEF815" box="[281,538,1932,1954]" italics="true" pageId="11" pageNumber="12">Paucibranchia adenensis</emphasis>
(Gravier, 1900)
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC32171FD08A7ABFC84F815" box="[708,800,1932,1954]" pageId="11" pageNumber="12">
<emphasis id="74A5A940FFC32171FD08A7ABFC84F815" bold="true" box="[708,800,1932,1954]" pageId="11" pageNumber="12">n. comb.</emphasis>
</taxonomicNameLabel>
, non-type MNHN-A148. A. Maxillary apparatus, dorsal view; B. Mandibles; C. Left MII-III-IV-V, dorsal view; D. Maxilla I, dorsal view. Apparatus stained with methyl green. Scale bars: A, C, 50 µm; B, 150 µm; D, 130 µm. al-MIV= attachment lamellae in MIV, LMI-MII= ligament between MI and MII.
</paragraph>
</caption>
<caption id="12AE25DAFFC42176FF5BA78FFA35F86A" httpUri="https://zenodo.org/record/1453041/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,250,1960,1982]" targetBox="[182,1414,193,1947]" targetPageId="12">
<paragraph id="466E7552FFC42176FF5BA78FFA35F86A" blockId="12.[151,1436,1960,2013]" pageId="12" pageNumber="13">
<emphasis id="74A5A940FFC42176FF5BA78FFEB6F809" bold="true" box="[151,274,1960,1982]" pageId="12" pageNumber="13">FIGURE 5.</emphasis>
<taxonomicName id="81D10ED1FFC42176FED5A78FFD1FF809" authority="(Gravier, 1900)" authorityName="Molina-Acevedo" authorityYear="2018" baseAuthorityName="Gravier" baseAuthorityYear="1900" box="[281,699,1960,1982]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="13" phylum="Annelida" rank="species" species="adenensis" status="comb. nov.">
<emphasis id="74A5A940FFC42176FED5A78FFDBDF809" box="[281,537,1960,1982]" italics="true" pageId="12" pageNumber="13">Paucibranchia adenensis</emphasis>
(Gravier, 1900)
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC42176FD0EA78FFCBAF809" box="[706,798,1960,1982]" pageId="12" pageNumber="13">
<emphasis id="74A5A940FFC42176FD0EA78FFCBAF809" bold="true" box="[706,798,1960,1982]" pageId="12" pageNumber="13">n. comb.</emphasis>
</taxonomicNameLabel>
, non-type MNHN-A148. A. Parapodium 3; B. Parapodium 11; C. Parapodium 26; D. Parapodium 70; E. Parapodium 158; F. Parapodium 216. All in anterior view. Scale bars: A–F, 0.7 mm.
</paragraph>
</caption>
<caption id="12AE25DAFFC52177FF5BA2DAFF64FC3A" httpUri="https://zenodo.org/record/1453043/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,765,787]" targetBox="[160,1402,193,743]" targetPageId="13">
<paragraph id="466E7552FFC52177FF5BA2DAFF64FC3A" blockId="13.[151,1436,765,909]" pageId="13" pageNumber="14">
<emphasis id="74A5A940FFC52177FF5BA2DAFEB0FCA4" bold="true" box="[151,276,765,787]" pageId="13" pageNumber="14">FIGURE 6.</emphasis>
<taxonomicName id="81D10ED1FFC52177FED0A2DAFD60FCA4" authority="(Gravier, 1900)" authorityName="Molina-Acevedo" authorityYear="2018" baseAuthorityName="Gravier" baseAuthorityYear="1900" box="[284,708,765,787]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="14" phylum="Annelida" rank="species" species="adenensis" status="comb. nov.">
<emphasis id="74A5A940FFC52177FED0A2DAFDBBFCA4" box="[284,543,765,787]" italics="true" pageId="13" pageNumber="14">Paucibranchia adenensis</emphasis>
(Gravier, 1900)
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC52177FD00A2DAFC8EFCA4" box="[716,810,765,787]" pageId="13" pageNumber="14">
<emphasis id="74A5A940FFC52177FD00A2DAFC8EFCA4" bold="true" box="[716,810,765,787]" pageId="13" pageNumber="14">n. comb.</emphasis>
</taxonomicNameLabel>
, non-type MNHN-A148. A. Compound falciger, long blade, chaetiger, 11; B. Compound falciger, median blade, chaetiger, 11; C. Compound falciger, short blade, chaetiger, 216; D. Isodonts pectinate narrow with long and slender teeth, chaetiger 3; E. Isodonts pectinate narrow with short and slender teeth, chaetiger 216; F. Subacicular hook, distal end, chaetiger 158. Scale bars: A, 12.5 µm; B, 20 µm; C, F, 9.8 µm; D, 14 µm, E, 4.9 µm.
</paragraph>
</caption>
<paragraph id="466E7552FFC52177FF0BA3F9FC42FB44" blockId="13.[151,1437,990,1843]" pageId="13" pageNumber="14">
<emphasis id="74A5A940FFC52177FF0BA3F9FEE0FC40" bold="true" box="[199,324,990,1015]" pageId="13" pageNumber="14">Variation.</emphasis>
Material examined varied in the following features: L10= 
<quantity id="8129D8B7FFC52177FBEDA3F9FB3BFC41" box="[1057,1183,990,1015]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="13" pageNumber="14" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">
2.5– 
<quantity id="8129D8B7FFC52177FB98A3F9FB3BFC41" box="[1108,1183,990,1015]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="14" unit="mm" value="4.0">4 mm</quantity>
</quantity>
, W10= 
<quantity id="8129D8B7FFC52177FAC8A3F8FA3DFC41" box="[1284,1433,991,1015]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="13" pageNumber="14" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 mm</quantity>
. Palps reaching second peristomial ring or first chaetiger. Lateral antennae reaching first or third chaetigers. Median antenna reaching third or fourth chaetiger. The maxillary formula varies as follows: 
<collectingRegion id="8415BBB0FFC52177FB3AA400FA87FB89" box="[1270,1315,1063,1086]" country="Japan" name="Mie" pageId="13" pageNumber="14">MII</collectingRegion>
7–8+7–8, MIII 6–7, MIV 4–5+7–9. The proportion of maxillary apparatus varies as follows: maxillary carriers shorter with respect to MI varies 1.6–2 times; cavity opening shorter with respect to 
<collectingRegion id="8415BBB0FFC52177FBEDA448FBEAFB31" box="[1057,1102,1135,1158]" country="Japan" name="Mie" pageId="13" pageNumber="14">MII</collectingRegion>
varies 5–6 times. Branchiae from chaetigers 14–17 to chaetigers 28–36. Maximum number of branchial filaments varied from 10 to 11. Well developed postchaetal lobe in the first 40–52 chaetigers. Ventral cirri with swollen base from chaetigers 6–7 to chaetigers 36–49. Start of subacicular hooks in chaetigers 29–35.
</paragraph>
</subSubSection>
<subSubSection id="0ECB26D9FFC52177FF0BA4D9FDD6FAA0" box="[199,626,1278,1303]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="466E7552FFC52177FF0BA4D9FDD6FAA0" blockId="13.[151,1437,990,1843]" box="[199,626,1278,1303]" pageId="13" pageNumber="14">
<emphasis id="74A5A940FFC52177FF0BA4D9FECCFAA0" bold="true" box="[199,360,1278,1303]" pageId="13" pageNumber="14">Distribution.</emphasis>
Red Sea, 
<collectingCountry id="3EC635C2FFC52177FE13A4D9FDCBFAA0" box="[479,623,1278,1303]" name="Madagascar" pageId="13" pageNumber="14">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="0ECB26D9FFC52174FF0BA505FE27FE67" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="discussion">
<paragraph id="466E7552FFC52177FF0BA505FBC5F9C8" blockId="13.[151,1437,990,1843]" pageId="13" pageNumber="14">
<emphasis id="74A5A940FFC52177FF0BA505FEE4FA8C" bold="true" box="[199,320,1314,1339]" pageId="13" pageNumber="14">Remarks.</emphasis>
<taxonomicName id="81D10ED1FFC52177FE80A505FDDEFA8C" authorityName="Molina-Acevedo" authorityYear="2018" baseAuthorityName="Gravier" baseAuthorityYear="1900" box="[332,634,1314,1339]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="14" phylum="Annelida" rank="species" species="adenensis" status="comb. nov.">
<emphasis id="74A5A940FFC52177FE80A505FDDEFA8C" box="[332,634,1314,1339]" italics="true" pageId="13" pageNumber="14">Paucibranchia adenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC52177FD4AA505FD52FA8C" box="[646,758,1314,1339]" pageId="13" pageNumber="14">
<emphasis id="74A5A940FFC52177FD4AA505FD52FA8C" bold="true" box="[646,758,1314,1339]" pageId="13" pageNumber="14">n. comb.</emphasis>
</taxonomicNameLabel>
was redescribed based on the 
<typeStatus id="996ACBF0FFC52177FBB4A505FB7AFA8C" box="[1144,1246,1314,1339]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
and specimens collected from 
<collectingCountry id="3EC635C2FFC52177FE86A560FE39FAE8" box="[330,413,1351,1375]" name="Greece" pageId="13" pageNumber="14">Greece</collectingCountry>
(Mediterranean Sea) by 
<bibRefCitation id="224008A3FFC52177FD0EA561FC6AFAE8" box="[706,974,1350,1375]" pageId="13" pageNumber="14" refId="ref55476" refString="Katsiaras, N., Simboura, N. &amp; Koutsoubas, D. (2014) The rare subgroup C 1 of Marphysa (Polychaeta, Eunicidae): redescription of species and first records in the Mediterranean Sea. Zootaxa, 3873 (3), 201 - 217. https: // doi. org / 10.11646 / zootaxa. 3873.3.1" type="journal article">
Katsiaras 
<emphasis id="74A5A940FFC52177FCF4A560FCD0FAE8" box="[824,884,1350,1375]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
(2014)
</bibRefCitation>
. They indicated that the Mediterranean specimens differ from the 
<typeStatus id="996ACBF0FFC52177FE1CA54DFD93FA34" box="[464,567,1386,1411]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
in the length of the falcigers blade since in the former the longest blades are twice as long as the shorter ones; while in the 
<typeStatus id="996ACBF0FFC52177FD12A5A9FCECFA10" box="[734,840,1422,1447]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
, the blades differ by only a few micrometers. They attributed this difference to the loss or breakage of falcigers in the 
<typeStatus id="996ACBF0FFC52177FC7AA595FBB8FA7C" box="[950,1052,1458,1483]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
due to the time of preservation. Nevertheless, as in the 
<typeStatus id="996ACBF0FFC52177FE7CA5F1FDBEFA58" box="[432,538,1494,1519]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
, the non-type specimens (MNHN-A148) do not have falcigers with such a marked difference in their blade's length. 
<bibRefCitation id="224008A3FFC52177FD5AA5DDFC02F9A4" box="[662,934,1530,1555]" pageId="13" pageNumber="14" refId="ref55476" refString="Katsiaras, N., Simboura, N. &amp; Koutsoubas, D. (2014) The rare subgroup C 1 of Marphysa (Polychaeta, Eunicidae): redescription of species and first records in the Mediterranean Sea. Zootaxa, 3873 (3), 201 - 217. https: // doi. org / 10.11646 / zootaxa. 3873.3.1" type="journal article">
Katsiaras 
<emphasis id="74A5A940FFC52177FCC2A5DCFCE9F9A4" box="[782,845,1530,1555]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
(2014)
</bibRefCitation>
also noted that the dorsal cirri are twice longer in the posterior region; while, in the 
<typeStatus id="996ACBF0FFC52177FD7AA639FCB8F980" box="[694,796,1566,1591]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
and additional material, the length of dorsal cirri is similar in pre- and postbranchial region. Based on the above, it is possible that the specimens from the Mediterranean may belong to an undescribed species, not to 
<taxonomicName id="81D10ED1FFC52177FC93A640FC54F9C8" authorityName="Molina-Acevedo" authorityYear="2018" baseAuthorityName="Gravier" baseAuthorityYear="1900" box="[863,1008,1638,1663]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="14" phylum="Annelida" rank="species" species="adenensis" status="comb. nov.">
<emphasis id="74A5A940FFC52177FC93A640FC54F9C8" box="[863,1008,1638,1663]" italics="true" pageId="13" pageNumber="14">P. adenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC52177FC3BA641FBC5F9C8" box="[1015,1121,1638,1663]" pageId="13" pageNumber="14">
<emphasis id="74A5A940FFC52177FC3BA641FBC5F9C8" bold="true" box="[1015,1121,1638,1663]" pageId="13" pageNumber="14">n. comb.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="466E7552FFC52177FF0BA6ACFF57F885" blockId="13.[151,1437,990,1843]" pageId="13" pageNumber="14">
<bibRefCitation id="224008A3FFC52177FF0BA6ACFEEAF914" box="[199,334,1674,1699]" pageId="13" pageNumber="14" refId="ref54003" refString="Day, J. H. (1962) Polychaeta from several localities in the western Indian Ocean. Proceedings of the Zoological Society of London, 139 (4), 627 - 656. https: // doi. org / 10.1111 / j. 1469 - 7998.1962. tb 01597. x" type="journal article">Day (1962</bibRefCitation>
, 
<bibRefCitation id="224008A3FFC52177FEAFA6ADFE3BF914" box="[355,415,1674,1699]" pageId="13" pageNumber="14" refId="ref54110" refString="Day, J. H. (1967) A monograph on the Polychaeta of Southern Africa. Part I. Errantia. British Museum (Natural History), London, 458 pp." type="book">1967</bibRefCitation>
) reported 
<taxonomicName id="81D10ED1FFC52177FDE5A6ACFD67F914" authorityName="Molina-Acevedo" authorityYear="2018" baseAuthorityName="Gravier" baseAuthorityYear="1900" box="[553,707,1674,1699]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="14" phylum="Annelida" rank="species" species="adenensis" status="comb. nov.">
<emphasis id="74A5A940FFC52177FDE5A6ACFD67F914" box="[553,707,1674,1699]" italics="true" pageId="13" pageNumber="14">P. adenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC52177FD1FA6ADFCE2F914" box="[723,838,1674,1699]" pageId="13" pageNumber="14">
<emphasis id="74A5A940FFC52177FD1FA6ADFCE2F914" bold="true" box="[723,838,1674,1699]" pageId="13" pageNumber="14">n. comb.</emphasis>
</taxonomicNameLabel>
from Nosy Be, 
<collectingCountry id="3EC635C2FFC52177FBE8A6ADFB10F914" box="[1060,1204,1674,1699]" name="Madagascar" pageId="13" pageNumber="14">Madagascar</collectingCountry>
. According to his description, the two specimens agree with the 
<typeStatus id="996ACBF0FFC52177FD0CA689FC83F970" box="[704,807,1710,1735]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
in the principal features, such as prostomium shape, branchial distribution and filaments number, the presence of only compound falcigers, and color and shape of aciculae and subacicular hook. Due to this similarity, it is possible that Day’s material belongs to the same species.
</paragraph>
<paragraph id="466E7552FFC62174FF0BA0B0FE27FE67" blockId="14.[151,1436,151,464]" pageId="14" pageNumber="15">
<taxonomicName id="81D10ED1FFC62174FF0BA0B0FE5EFF07" authorityName="Molina-Acevedo" authorityYear="2018" baseAuthorityName="Gravier" baseAuthorityYear="1900" box="[199,506,151,176]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="15" phylum="Annelida" rank="species" species="adenensis" status="comb. nov.">
<emphasis id="74A5A940FFC62174FF0BA0B0FE5EFF07" box="[199,506,151,176]" italics="true" pageId="14" pageNumber="15">Paucibranchia adenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC62174FDC0A0B0FD24FF07" box="[524,640,151,176]" pageId="14" pageNumber="15">
<emphasis id="74A5A940FFC62174FDC0A0B0FD24FF07" bold="true" box="[524,640,151,176]" pageId="14" pageNumber="15">n. comb.</emphasis>
</taxonomicNameLabel>
is similar to 
<taxonomicName id="81D10ED1FFC62174FC89A0BEFC73FF07" authorityName="Molina-Acevedo" authorityYear="2018" baseAuthorityName="Mohammad" baseAuthorityYear="1973" box="[837,983,153,176]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="15" phylum="Annelida" rank="species" species="gemmata" status="comb. nov.">
<emphasis id="74A5A940FFC62174FC89A0BEFC73FF07" box="[837,983,153,176]" italics="true" pageId="14" pageNumber="15">P. gemmata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC62174FC25A0B0FBF9FF07" box="[1001,1117,151,176]" pageId="14" pageNumber="15">
<emphasis id="74A5A940FFC62174FC25A0B0FBF9FF07" bold="true" box="[1001,1117,151,176]" pageId="14" pageNumber="15">n. comb.</emphasis>
</taxonomicNameLabel>
<taxonomicName id="81D10ED1FFC62174FBA3A0BEFAA5FF07" authority="Molina-Acevedo, 2018" authorityName="Molina-Acevedo" authorityYear="2018" box="[1135,1281,153,176]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="15" phylum="Annelida" rank="species" species="patriciae" status="sp. nov.">
<emphasis id="74A5A940FFC62174FBA3A0BEFAA5FF07" box="[1135,1281,153,176]" italics="true" pageId="14" pageNumber="15">P. patriciae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC62174FADFA0B0FAC4FF07" box="[1299,1376,151,176]" pageId="14" pageNumber="15" rank="species">
<emphasis id="74A5A940FFC62174FADFA0B0FAC4FF07" bold="true" box="[1299,1376,151,176]" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="81D10ED1FFC62174FF5BA09BFEE1FF62" authorityName="Molina-Acevedo" authorityYear="2018" box="[151,325,188,213]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="15" phylum="Annelida" rank="genus">
<emphasis id="74A5A940FFC62174FF5BA09BFEE1FF62" box="[151,325,188,213]" italics="true" pageId="14" pageNumber="15">Paucibranchia</emphasis>
</taxonomicName>
sp. 2 based on the presence of only compound falcigers and translucent color of aciculae; however, 
<taxonomicName id="81D10ED1FFC62174FEC6A0C6FE3FFF4F" authorityName="Molina-Acevedo" authorityYear="2018" baseAuthorityName="Gravier" baseAuthorityYear="1900" box="[266,411,223,248]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="15" phylum="Annelida" rank="species" species="adenensis" status="comb. nov.">
<emphasis id="74A5A940FFC62174FEC6A0C6FE3FFF4F" box="[266,411,223,248]" italics="true" pageId="14" pageNumber="15">P. adenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC62174FE6FA0F8FDAAFF4F" box="[419,526,223,248]" pageId="14" pageNumber="15">
<emphasis id="74A5A940FFC62174FE6FA0F8FDAAFF4F" bold="true" box="[419,526,223,248]" pageId="14" pageNumber="15">n. comb.</emphasis>
</taxonomicNameLabel>
differs from 
<taxonomicName id="81D10ED1FFC62174FD60A0C6FC91FF4F" authorityName="Molina-Acevedo" authorityYear="2018" baseAuthorityName="Mohammad" baseAuthorityYear="1973" box="[684,821,225,248]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="15" phylum="Annelida" rank="species" species="gemmata" status="comb. nov.">
<emphasis id="74A5A940FFC62174FD60A0C6FC91FF4F" box="[684,821,225,248]" italics="true" pageId="14" pageNumber="15">P. gemmata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC62174FCF0A0F8FC03FF4F" box="[828,935,223,248]" pageId="14" pageNumber="15">
<emphasis id="74A5A940FFC62174FCF0A0F8FC03FF4F" bold="true" box="[828,935,223,248]" pageId="14" pageNumber="15">n. comb.</emphasis>
</taxonomicNameLabel>
in the presence of eyes, dorsal cirri of the same length in pre- and postbranchial region, and falcigers with blades of two sizes in anterior region; while 
<taxonomicName id="81D10ED1FFC62174FF5BA10EFEBBFEF7" authorityName="Molina-Acevedo" authorityYear="2018" baseAuthorityName="Mohammad" baseAuthorityYear="1973" box="[151,287,297,320]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="15" phylum="Annelida" rank="species" species="gemmata" status="comb. nov.">
<emphasis id="74A5A940FFC62174FF5BA10EFEBBFEF7" box="[151,287,297,320]" italics="true" pageId="14" pageNumber="15">P. gemmata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC62174FEEAA100FE35FEF7" box="[294,401,295,320]" pageId="14" pageNumber="15">
<emphasis id="74A5A940FFC62174FEEAA100FE35FEF7" bold="true" box="[294,401,295,320]" pageId="14" pageNumber="15">n. comb.</emphasis>
</taxonomicNameLabel>
lacks eyes, posterior dorsal cirri are two times longer than anterior ones, and by having falcigers with blades of three sizes in the same region. The morphological differentiation of 
<taxonomicName id="81D10ED1FFC62174FB25A16AFADEFED2" authorityName="Molina-Acevedo" authorityYear="2018" baseAuthorityName="Gravier" baseAuthorityYear="1900" box="[1257,1402,332,357]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="15" phylum="Annelida" rank="species" species="adenensis" status="comb. nov.">
<emphasis id="74A5A940FFC62174FB25A16AFADEFED2" box="[1257,1402,332,357]" italics="true" pageId="14" pageNumber="15">P. adenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC62174FA48A16BFF46FE3F" pageId="14" pageNumber="15">
<emphasis id="74A5A940FFC62174FA48A16BFF46FE3F" bold="true" pageId="14" pageNumber="15">n. comb.</emphasis>
</taxonomicNameLabel>
with 
<taxonomicName id="81D10ED1FFC62174FEE5A156FE15FE3F" authority="Molina-Acevedo, 2018" authorityName="Molina-Acevedo" authorityYear="2018" box="[297,433,369,392]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="15" phylum="Annelida" rank="species" species="patriciae" status="sp. nov.">
<emphasis id="74A5A940FFC62174FEE5A156FE15FE3F" box="[297,433,369,392]" italics="true" pageId="14" pageNumber="15">P. patriciae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC62174FE77A148FDA4FE3F" box="[443,512,367,392]" pageId="14" pageNumber="15" rank="species">
<emphasis id="74A5A940FFC62174FE77A148FDA4FE3F" bold="true" box="[443,512,367,392]" pageId="14" pageNumber="15">n. sp.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="81D10ED1FFC62174FDF0A148FD4EFE3F" authorityName="Molina-Acevedo" authorityYear="2018" box="[572,746,367,392]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="15" phylum="Annelida" rank="genus">
<emphasis id="74A5A940FFC62174FDF0A148FD4EFE3F" box="[572,746,367,392]" italics="true" pageId="14" pageNumber="15">Paucibranchia</emphasis>
</taxonomicName>
sp. 2 is provided in the Remarks section of each species. The comparison with other 
<taxonomicName id="81D10ED1FFC62174FE35A1B3FD03FE1A" authority="Molina-Acevedo, 2018" authorityName="Molina-Acevedo" authorityYear="2018" box="[505,679,404,429]" class="Polychaeta" family="Eunicidae" genus="Paucibranchia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="15" phylum="Annelida" rank="genus" status="gen. nov.">
<emphasis id="74A5A940FFC62174FE35A1B3FD03FE1A" box="[505,679,404,429]" italics="true" pageId="14" pageNumber="15">Paucibranchia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6F96143BFFC62174FD7AA1B3FCB4FE1A" box="[694,784,404,429]" pageId="14" pageNumber="15" rank="genus">
<emphasis id="74A5A940FFC62174FD7AA1B3FCB4FE1A" bold="true" box="[694,784,404,429]" pageId="14" pageNumber="15">n. gen.</emphasis>
</taxonomicNameLabel>
species having only compound falcigers present is provided in 
<tableCitation id="0B5340E9FFC62174FEE9A190FEDBFE67" box="[293,383,439,464]" captionStart="TABLE 2" captionStartId="112.[147,226,502,523]" captionText="TABLE 2. Morphological features of the Paucibranchia n. gen. species with compound falcigers. Abbreviations: MF: Maxillary formula, roman numerals refer to number of maxilla; MxC: maxillary carriers; CIS: closing system; Cop: cavity opening; PR-I: first peristomial ring; PR-II: second peristomial ring; Chaet: chaetiger; p/a: present/absent; DC: dorsal cirri; PrBr: pre-branchial region; PoBr: postbranchial region; Bran: branchial region; Ant: anterior region; Post: posterior region." pageId="14" pageNumber="15">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>