221 lines
17 KiB
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221 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.137.46544" ID-PMC="PMC6962254" ID-Pensoft-Pub="1314-2003-137-1" ID-Pensoft-UUID="FB2C14EFA257564AA84CD47711BBC273" ID-PubMed="31969792" ModsDocID="1314-2003-137-1" checkinTime="1578614985992" checkinUser="pensoft" docAuthor="Zaika, Maxim A., Kilian, Norbert, Jones, Katy, Krinitsina, Anastasiya A., Nilova, Maya V., Speranskaya, Anna S. & Sukhorukov, Alexander P." docDate="2020" docId="CC34293805E65C988DBCE81888283F93" docLanguage="en" docName="PhytoKeys 137: 1-85" docOrigin="PhytoKeys 137" docSource="http://dx.doi.org/10.3897/phytokeys.137.46544" docTitle="Scorzonera s. str. clade" docType="treatment" docVersion="5" id="FB2C14EFA257564AA84CD47711BBC273" lastPageNumber="1" masterDocId="FB2C14EFA257564AA84CD47711BBC273" masterDocTitle="Scorzonera sensu lato (Asteraceae, Cichorieae) - taxonomic reassessment in the light of new molecular phylogenetic and carpological analyses" masterLastPageNumber="85" masterPageNumber="1" pageNumber="1" updateTime="1668139594603" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Scorzonera sensu lato (Asteraceae, Cichorieae) - taxonomic reassessment in the light of new molecular phylogenetic and carpological analyses</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zaika, Maxim A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kilian, Norbert</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jones, Katy</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Krinitsina, Anastasiya A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nilova, Maya V.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Speranskaya, Anna S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sukhorukov, Alexander P.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>137</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>85</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.137.46544</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.137.46544</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-137-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FB2C14EFA257564AA84CD47711BBC273</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="161671721" LSID="urn:lsid:plazi:treatment:CC34293805E65C988DBCE81888283F93" httpUri="http://treatment.plazi.org/id/CC34293805E65C988DBCE81888283F93" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="CC342938-05E6-5C98-8DBC-E81888283F93" authority="s. str. clade" class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Scorzonera s.str. clade</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="section" section="Parviflorae">Scorzonera sect. Parviflorae</taxonomicName>
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Lipsch. in Bobrov & Tzvelev, Fl. URSS 29: 79. 1964 ≡
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<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="series" series="Parviflorae">Scorzonera ser. Parviflorae</taxonomicName>
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Lipsch. in Bobrov & Tzvelev, Fl. URSS 29: 720. 1964 [
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<taxonomicName authority="subsect. Parviflora" family="Asteraceae" genus="S." higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="S." order="Asterales" pageId="0" pageNumber="1" rank="genus">S. subsect. Parviflora</taxonomicName>
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Lipsch., Fragm. Monogr. Gen.
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<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Scorzonera</taxonomicName>
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2: 112. 1939, nom. inval. (Art. 39.1,
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<bibRefCitation DOI="https://doi.org/10.12705/Code.2018" author="Turland, NJ" journalOrPublisher="Turkish Journal of Biology" pageId="0" pageNumber="1" refId="B102" refString="Turland, NJ, Wiersema, JH, Barrie, FR, Greuter, W, Hawksworth, DL, Herendeen, PS, Knapp, S, Kusber, W-H, Li, D-Z, Marhold, K, May, TW, McNeill, J, Monro, AM, Pradeo, J, Price, MJ, Smith, G, 2018. Internation Code of Nomenclature for algae, fungi, and plants (Shenzen Code), adopted by the Nineteenth Internatonal Botanical Congress Shenzen, China, July 2017. Regnum Vegetabile 159. https://doi.org/10.12705/Code.2018" title="Internation Code of Nomenclature for algae, fungi, and plants (Shenzen Code), adopted by the Nineteenth Internatonal Botanical Congress Shenzen, China, July 2017. Regnum Vegetabile 159." url="https://doi.org/10.12705/Code.2018" year="2018">Turland et al. 2018</bibRefCitation>
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)]. Type:
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<taxonomicName authorityName="Jacq" authorityYear="1776" class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera parviflora" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="parviflora">Scorzonera parviflora</taxonomicName>
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Jacq.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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=
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<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="section" section="Radiatae">Scorzonera sect. Radiatae</taxonomicName>
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Nakai in Rep. Inst. Sci. Res. Manchoukuo, ser. 1, 6: 169. 1937. Type:
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<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera radiata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="radiata">Scorzonera radiata</taxonomicName>
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Fisch. ex Ledeb.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnostic features">
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<paragraph pageId="0" pageNumber="1">Diagnostic features.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Inner phyllaries apically often with dark red or blackish spot; achene sclerenchyma insular in principal ribs below sclerenchyma and in secondary ribs above sclerenchyma (in
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<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. aristata" order="Asterales" pageId="0" pageNumber="1" rank="species" species="aristata">
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<emphasis italics="true" pageId="0" pageNumber="1">S. aristata</emphasis>
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</taxonomicName>
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, the parenchyma is present in secondary ribs above sclerenchyma).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Habit, life form, subterranean parts:</emphasis>
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perennial or biennial herbs, often with leaf rosettes, rootstock cylindrical.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Leaves:</emphasis>
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rosulate leaves present, petiolate, cauline leaves sessile, all leaves entire, linear to elliptical or ovate, glabrous or glabrescent at maturity.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Stem, synflorescence:</emphasis>
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stems solitary or few, straight, mostly not branched, leafless (scapes) or leafy, glabrous or arachnoid-hairy becoming almost glabrous at the fruiting; capitula terminal, solitary or several.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pollen:</emphasis>
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echinolophate, tricolporate and each colpus divided into 2 lacunae; with 6 abporal lacunae (
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<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. humilis" order="Asterales" pageId="0" pageNumber="1" rank="species" species="humilis">
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<emphasis italics="true" pageId="0" pageNumber="1">S. humilis</emphasis>
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</taxonomicName>
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and
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<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. parviflora" order="Asterales" pageId="0" pageNumber="1" rank="species" species="parviflora">
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<emphasis italics="true" pageId="0" pageNumber="1">S. parviflora</emphasis>
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</taxonomicName>
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:
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<bibRefCitation DOI="https://doi.org/10.1080/00173138209427691" author="Blackmore, S" journalOrPublisher="Grana" pageId="0" pageNumber="1" pagination="149 - 160" refId="B9" refString="Blackmore, S, 1982. Palynology of subtribe Scorzonerinae (Compositae. Lactuceae) and its taxonomic significance. Grana 21 (3): 149 - 160, DOI: https://doi.org/10.1080/00173138209427691" title="Palynology of subtribe Scorzonerinae (Compositae. Lactuceae) and its taxonomic significance." url="https://doi.org/10.1080/00173138209427691" volume="21" year="1982">Blackmore 1982</bibRefCitation>
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;
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<bibRefCitation author="Diaz de la Guardia, C" journalOrPublisher="Anales de la Asociacion de Palinologos de Lengua Espanola" pageId="0" pageNumber="1" pagination="177 - 190" refId="B29" refString="Diaz de la Guardia, C, Blanca, G, 1985. Estudio palinologio del genero Scorzonera L. (Asteraceae) en la peninsula Iberica. Anales de la Asociacion de Palinologos de Lengua Espanola 2: 177 - 190" title="Estudio palinologio del genero Scorzonera L. (Asteraceae) en la peninsula Iberica." volume="2" year="1985">
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<normalizedToken originalValue="Díaz">Diaz</normalizedToken>
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de la Guardia and Blanca 1985
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</bibRefCitation>
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) or 18 (6 abporal, 6 equatorial and 6 interporal) lacunae (
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<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. aristata" order="Asterales" pageId="0" pageNumber="1" rank="species" species="aristata">
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<emphasis italics="true" pageId="0" pageNumber="1">S. aristata</emphasis>
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</taxonomicName>
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:
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<bibRefCitation author="Halbritter, H" journalOrPublisher="Willdenowia" pageId="0" pageNumber="1" publicationUrl="https://www.paldat.org/pub/Scorzonera_aristata/303098;jsessionid=7CA24A4CC451D4B70967A64190B96F55" refId="B37" refString="Halbritter, H, Berger, A, 2017. Scorzoneraaristata. PalDat - A palynological database. https://www.paldat.org/pub/Scorzonera_aristata/303098;jsessionid=7CA24A4CC451D4B70967A64190B96F55" title="Scorzoneraaristata. PalDat - A palynological database." url="https://www.paldat.org/pub/Scorzonera_aristata/303098;jsessionid=7CA24A4CC451D4B70967A64190B96F55" year="2017">Halbritter and Berger 2017</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Capitula:</emphasis>
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glabrous or hairy and glabrescent, phyllaries in several series, outer phyllaries triangular-ovate, inner phyllaries lanceolate to oblong, apically with dark red or blackish spot, outer phyllaries at least
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<normalizedToken originalValue="½">1/2</normalizedToken>
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as long as inner ones, receptacle glabrous, flat, florets more than 12, yellow or whitish, equal in length to involucre or
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<normalizedToken originalValue="1½">11/2</normalizedToken>
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-2 times as long.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Achenes:</emphasis>
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7-12 mm, without carpopodium, glabrous, achene epidermis with papillae, outer achenes more or less curved, with 10 ribs, without emergences or verrucose (
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<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. aristata" order="Asterales" pageId="0" pageNumber="1" rank="species" species="aristata">
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<emphasis italics="true" pageId="0" pageNumber="1">S. aristata</emphasis>
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</taxonomicName>
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); achene wall with parenchyma insular in principal ribs below sclerenchyma and in secondary ribs above sclerenchyma, sclerenchyma continuous, its fibres orientated parallel to the achene axis, air cavities and tannins absent.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pappus:</emphasis>
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11-16 mm, dirty white or rarely ice-white (
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<taxonomicName family="Asteraceae" kingdom="Plantae" lsidName="S. parviflora" order="Asterales" pageId="0" pageNumber="1" rank="species" species="parviflora">
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<emphasis italics="true" pageId="0" pageNumber="1">S. parviflora</emphasis>
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</taxonomicName>
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), bristles plumose or apically scabrid (or 5-10 longer bristles scabrid and other bristles plumose).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="chromosome number">
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<paragraph pageId="0" pageNumber="1">Chromosome number.</paragraph>
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<paragraph pageId="0" pageNumber="1">x = 7, diploids.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="species">
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<paragraph pageId="0" pageNumber="1">Species.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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(1)
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<taxonomicName class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera aristata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="aristata">
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<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera aristata</emphasis>
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</taxonomicName>
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Ramond ex DC.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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(2)
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<taxonomicName class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera humilis" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="humilis">
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<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera humilis</emphasis>
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</taxonomicName>
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L. (Fig.
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Images of living plants. A, B Ramaliella tortuosissima A general view B close up view of the capitula (Jordan, Nakeb, 1250 m alt., 8 May 2019). Photograph by O. Fragman-Sapir C Pseudopodospermum papposum (Jordan, Nakeb, 8 May 2019). Photograph by O. Fragman-Sapir D Takhtajaniantha austriaca (Italy, South Tyrol, Schlanders, 6 May 2010). Photograph by R. Hand E Scorzonera humilis (Germany, Ammersee, 15 May 2009). Photograph by C. Niederbichler F Scorzonera meyeri (Russia, Karachaevo-Cherkessiya, Teberda, 14 July 2005). Photograph by A. S. Zernov." figureDoi="10.3897/phytokeys.137.46544.figure16" httpUri="https://binary.pensoft.net/fig/370364" pageId="0" pageNumber="1">16E</figureCitation>
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)
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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(3)
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<taxonomicName authorityName="Jacq" authorityYear="1776" class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera parviflora" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="parviflora">
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<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera parviflora</emphasis>
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</taxonomicName>
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Jacq.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution area.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Asia-Temperate</emphasis>
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: AF; AM; AR; CN; CY; GE; IR; KG; LB; MN; RU (Asiatic part); SY; TM; TR; UZ.
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<emphasis bold="true" pageId="0" pageNumber="1">Europe</emphasis>
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: AT; BE; BY; CH; CZ; DE; DK; EE; ES; FI; FR; GB; HR; IT; LT; LV; NL; NO; PL; PT; RO; RS; RU; SE; SK; SI; UA.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Magnoliopsida" family="Compositae" genus="Scorzonera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Scorzonera radiata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="radiata">
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<emphasis italics="true" pageId="0" pageNumber="1">Scorzonera radiata</emphasis>
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</taxonomicName>
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Fisch. ex Ledeb. may also belong to this clade on account of its resemblance in morphological and carpological characters.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |