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<document id="584167E639E627D1A64BAD4CCC5F6694" ID-DOI="10.5281/zenodo.176361" ID-GBIF-Dataset="cb365895-e857-455a-be07-3e4593e50923" ID-ISSN="1175-5326" ID-Zenodo-Dep="176361" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459832908851" checkinUser="plazi" docAuthor="Boxshall, Geoff A. &amp; Marchenkov, Andrey" docDate="2007" docId="C03D8785045CFFBAFF2DFF49FCFAFA32" docLanguage="en" docName="zt01459p068.pdf" docOrigin="Zootaxa 1459" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Anoplodelphys incerta Lafargue and Laubier 1978" docType="treatment" docVersion="9" lastPageNumber="47" masterDocId="3C04FFFD0456FFB0FFBAFFDEFF9AFF9E" masterDocTitle="A revision of the Brementia - group of genera (Copepoda: Notodelphyidae), with descriptions of a new genus and four new species" masterLastPageNumber="68" masterPageNumber="37" pageNumber="47" updateTime="1698223101261" updateUser="plazi">
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<mods:title id="53BB8CD400CDD303B21627A155E6BFBF">A revision of the Brementia - group of genera (Copepoda: Notodelphyidae), with descriptions of a new genus and four new species</mods:title>
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<mods:namePart id="689D134D1F152F3D87BFF4BE212E473D">Boxshall, Geoff A.</mods:namePart>
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<mods:namePart id="42C60AEEEB90F16CC299D7697A8DB554">Marchenkov, Andrey</mods:namePart>
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<mods:date id="B5BDA5DF00EDC60E08939B6CD1116343">2007</mods:date>
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<treatment id="C03D8785045CFFBAFF2DFF49FCFAFA32" ID-DOI="http://doi.org/10.5281/zenodo.5661723" ID-GBIF-Taxon="119358209" ID-Zenodo-Dep="5661723" LSID="urn:lsid:plazi:treatment:C03D8785045CFFBAFF2DFF49FCFAFA32" httpUri="http://treatment.plazi.org/id/C03D8785045CFFBAFF2DFF49FCFAFA32" lastPageNumber="47" pageId="10" pageNumber="47">
<subSubSection id="008E6518045CFFBAFF2DFF49FC90FF2F" box="[151,778,151,177]" pageId="10" pageNumber="47" type="nomenclature">
<paragraph id="482B3693045CFFBAFF2DFF49FC90FF2F" blockId="10.[151,778,151,177]" box="[151,778,151,177]" pageId="10" pageNumber="47">
<heading id="136381FF045CFFBAFF2DFF49FC90FF2F" bold="true" box="[151,778,151,177]" fontSize="11" level="1" pageId="10" pageNumber="47" reason="1">
<taxonomicName id="8F944D10045CFFBAFF2DFF49FC90FF2F" authority="Lafargue and Laubier, 1978" authorityName="Lafargue and Laubier" authorityYear="1978" box="[151,778,151,177]" class="Maxillopoda" family="Notodelphyidae" genus="Anoplodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="7AE0EA81045CFFBAFF2DFF49FC90FF2F" bold="true" box="[151,778,151,177]" pageId="10" pageNumber="47">
<emphasis id="7AE0EA81045CFFBAFF2DFF49FE3FFF2F" bold="true" box="[151,421,151,177]" italics="true" pageId="10" pageNumber="47">Anoplodelphys incerta</emphasis>
Lafargue and Laubier, 1978
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="008E6518045CFFBAFF2DFF3CFCDCFCCA" pageId="10" pageNumber="47" type="materials_examined">
<paragraph id="482B3693045CFFBAFF2DFF3CFC0CFF62" blockId="10.[151,1436,226,1452]" box="[151,918,226,252]" pageId="10" pageNumber="47">
<emphasis id="7AE0EA81045CFFBAFF2DFF3CFEEFFF65" box="[151,373,226,251]" italics="true" pageId="10" pageNumber="47">Material examined</emphasis>
:
<typeStatus id="972F8831045CFFBAFE3EFF3CFE6BFF62" box="[388,497,226,252]" pageId="10" pageNumber="47" type="holotype">Holotype</typeStatus>
female, reg. no.
<collectionCode id="2E85AE56045CFFBAFD0DFF3CFD62FF62" box="[695,760,226,252]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="10" pageNumber="47">ZMA</collectionCode>
CO.102.602.
</paragraph>
<paragraph id="482B3693045CFFBAFF7CFED4FD34FDFA" blockId="10.[151,1436,226,1452]" pageId="10" pageNumber="47">
<emphasis id="7AE0EA81045CFFBAFF7CFED4FE4AFEBD" box="[198,464,266,291]" italics="true" pageId="10" pageNumber="47">Differential Diagnosis</emphasis>
: Body elongate, with indistinct segmentation. Cephalosome with narrow frontal margin bearing rostrum. Rostrum inflated, forming large ventrally-directed lobe (
<figureCitation id="D0AF2A16045CFFBAFB2EFEECFB79FED2" box="[1172,1251,306,332]" captionStart="FIGURE 4" captionStartId="8.[151,255,1880,1904]" captionTargetBox="[192,1384,361,1834]" captionTargetId="figure@8.[179,1409,354,1856]" captionTargetPageId="8" captionText="FIGURE 4. Anoplodelphys laubieri n. sp. holotype female (MNHN-Cp 2315). A, Cephalosome and first pedigerous somite, lateral; B, urosome, lateral. Anoplodelphys incerta Lafargue &amp; Laubier, 1977, holotype female (ZMA CO. 106.602). C, Cephalosome and first pedigerous somite, ventral, D, Antennule showing surface ornamentation; E, Leg 5. Scale bars: A = 500 μm, B = 50 μm, C D = 200 μm, E = 25 μm." httpUri="https://zenodo.org/record/176365/files/figure.png" pageId="10" pageNumber="47">Fig. 4</figureCitation>
C). Post-rostral median lobe present. Labrum strongly inflated at base, tending distally into tapering lobe (
<figureCitation id="D0AF2A16045CFFBAFB69FE84FA86FEEA" box="[1235,1308,346,372]" captionStart="FIGURE 4" captionStartId="8.[151,255,1880,1904]" captionTargetBox="[192,1384,361,1834]" captionTargetId="figure@8.[179,1409,354,1856]" captionTargetPageId="8" captionText="FIGURE 4. Anoplodelphys laubieri n. sp. holotype female (MNHN-Cp 2315). A, Cephalosome and first pedigerous somite, lateral; B, urosome, lateral. Anoplodelphys incerta Lafargue &amp; Laubier, 1977, holotype female (ZMA CO. 106.602). C, Cephalosome and first pedigerous somite, ventral, D, Antennule showing surface ornamentation; E, Leg 5. Scale bars: A = 500 μm, B = 50 μm, C D = 200 μm, E = 25 μm." httpUri="https://zenodo.org/record/176365/files/figure.png" pageId="10" pageNumber="47">Fig. 4</figureCitation>
C). Lateral margin of cephalosome expanded ventrally to form ridge-like swelling produced into process. Antennomedial lobe lacking. Metasome elongate with about 25% of length extending posterior to level of origin of leg 4: metasomal somites bearing small legs ventrally near lateral margins. No mid-ventral processes present between legs. Urosome small, unsegmented; located terminally; with median anal incision, bearing minute paired caudal rami armed with setae distally. Surface of body, rostrum, labrum, cephalosomic processes, and legs densely ornamented with surface setules.
</paragraph>
<paragraph id="482B3693045CFFBAFF7CFDACFBAAFCB2" blockId="10.[151,1436,226,1452]" pageId="10" pageNumber="47">
Antennule unsegmented, tapering lobe (
<figureCitation id="D0AF2A16045CFFBAFD12FDACFD69FD12" box="[680,755,626,652]" captionStart="FIGURE 4" captionStartId="8.[151,255,1880,1904]" captionTargetBox="[192,1384,361,1834]" captionTargetId="figure@8.[179,1409,354,1856]" captionTargetPageId="8" captionText="FIGURE 4. Anoplodelphys laubieri n. sp. holotype female (MNHN-Cp 2315). A, Cephalosome and first pedigerous somite, lateral; B, urosome, lateral. Anoplodelphys incerta Lafargue &amp; Laubier, 1977, holotype female (ZMA CO. 106.602). C, Cephalosome and first pedigerous somite, ventral, D, Antennule showing surface ornamentation; E, Leg 5. Scale bars: A = 500 μm, B = 50 μm, C D = 200 μm, E = 25 μm." httpUri="https://zenodo.org/record/176365/files/figure.png" pageId="10" pageNumber="47">Fig. 4</figureCitation>
D). Antenna 2-segmented, with triangular basal segment and distal subchela incorporating sclerotised distal claw, armed with 3 vestigial setal elements. Mandible to maxilliped all lacking. Legs 14 biramous; each with protopodal part almost completely incorporated into somite so small lobate rami appearing to originate from somite surface. Ratios of gaps between legs along metasome as in
<tableCitation id="05160328045CFFBAFEE9FCCCFE36FCB2" box="[339,428,786,812]" captionStart="TABLE 2" captionStartId="11.[151,239,1386,1410]" captionTargetBox="[156,1384,1477,1658]" captionTargetPageId="11" captionText="TABLE 2. Distribution of legs along the metasome in Anoplodelphys species (calculated as distance between front edge of leg bases expressed as a percentage of total length of metasome)." httpUri="http://table.plazi.org/id/1CEB661B045DFFBBFF2DFAB4FCF0FA3A" pageId="10" pageNumber="47" tableUuid="1CEB661B045DFFBBFF2DFAB4FCF0FA3A">Table 2</tableCitation>
. Leg 5 represented by tiny sclerotised points (
<figureCitation id="D0AF2A16045CFFBAFC7DFCCCFB94FCB2" box="[967,1038,786,812]" captionStart="FIGURE 4" captionStartId="8.[151,255,1880,1904]" captionTargetBox="[192,1384,361,1834]" captionTargetId="figure@8.[179,1409,354,1856]" captionTargetPageId="8" captionText="FIGURE 4. Anoplodelphys laubieri n. sp. holotype female (MNHN-Cp 2315). A, Cephalosome and first pedigerous somite, lateral; B, urosome, lateral. Anoplodelphys incerta Lafargue &amp; Laubier, 1977, holotype female (ZMA CO. 106.602). C, Cephalosome and first pedigerous somite, ventral, D, Antennule showing surface ornamentation; E, Leg 5. Scale bars: A = 500 μm, B = 50 μm, C D = 200 μm, E = 25 μm." httpUri="https://zenodo.org/record/176365/files/figure.png" pageId="10" pageNumber="47">Fig. 4</figureCitation>
E).
</paragraph>
<paragraph id="482B3693045CFFBAFF7CFCE4FCDCFCCA" blockId="10.[151,1436,226,1452]" box="[198,838,826,852]" pageId="10" pageNumber="47">
Body length of female
<quantity id="8F6C9B76045CFFBAFE6EFCE4FD18FCCA" box="[468,642,826,852]" metricMagnitude="-3" metricUnit="m" metricValue="3.025" metricValueMax="3.65" metricValueMin="2.4" pageId="10" pageNumber="47" unit="mm" value="3.025" valueMax="3.65" valueMin="2.4">2.403.65 mm</quantity>
. Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="008E6518045CFFBAFF7CFCBCFCFAFA32" pageId="10" pageNumber="47" type="discussion">
<paragraph id="482B3693045CFFBAFF7CFCBCFADBFB7A" blockId="10.[151,1436,226,1452]" pageId="10" pageNumber="47">
<emphasis id="7AE0EA81045CFFBAFF7CFCBCFEB7FCE5" box="[198,301,866,891]" italics="true" pageId="10" pageNumber="47">Remarks</emphasis>
: In their paper on
<taxonomicName id="8F944D10045CFFBAFDAAFCBCFBAEFCE2" authority="Lafargue and Laubier (1978a)" authorityName="Lafargue and Laubier" authorityYear="1978" box="[528,1076,866,892]" class="Maxillopoda" family="Notodelphyidae" genus="Anoplodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="10" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="7AE0EA81045CFFBAFDAAFCBCFD24FCE5" box="[528,702,866,891]" italics="true" pageId="10" pageNumber="47">Anoplodelphys</emphasis>
,
<bibRefCitation id="2C054B62045CFFBAFD75FCBCFC4AFCE2" author="Lafargue" box="[719,976,866,892]" pageId="10" pageNumber="68" refString="Lafargue, F. &amp; Laubier, L. (1978 a) Anoplodelphys g. nov. Copepode Notodelphyidae parasite de Didemnidae (Ascidies Aplousobranches) en Mediterranee. Crustaceana, 35 (3), 277 - 293." type="journal article" year="1978" yearSuffix="a">Lafargue and Laubier</bibRefCitation>
(1978a)
</taxonomicName>
comment upon the very wide intraspecific variability between specimens of
<taxonomicName id="8F944D10045CFFBAFD72FC54FC4EFC3D" box="[712,980,906,931]" class="Maxillopoda" family="Notodelphyidae" genus="Anoplodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="7AE0EA81045CFFBAFD72FC54FC4EFC3D" box="[712,980,906,931]" italics="true" pageId="10" pageNumber="47">Anoplodelphys incerta</emphasis>
</taxonomicName>
collected from different host species. They recognised a so-called “typical” form of
<taxonomicName id="8F944D10045CFFBAFD09FC6CFCBCFC55" box="[691,806,946,971]" class="Maxillopoda" family="Notodelphyidae" genus="Anoplodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="7AE0EA81045CFFBAFD09FC6CFCBCFC55" box="[691,806,946,971]" italics="true" pageId="10" pageNumber="47">A. incerta</emphasis>
</taxonomicName>
from
<taxonomicName id="8F944D10045CFFBAFCD6FC6CFAD2FC52" authority="Della Valle, 1877" authorityName="Della Valle" authorityYear="1877" box="[876,1352,946,972]" class="Ascidiacea" family="Didemnidae" genus="Didemnum" kingdom="Animalia" order="Enterogona" pageId="10" pageNumber="47" phylum="Chordata" rank="species" species="commune">
<emphasis id="7AE0EA81045CFFBAFCD6FC6CFBF9FC55" box="[876,1123,946,971]" italics="true" pageId="10" pageNumber="47">Didemnum commune</emphasis>
(Della Valle, 1877)
</taxonomicName>
and an “atypical” form from
<taxonomicName id="8F944D10045CFFBAFE22FC04FC0AFC6A" authority="Milne Edwards, 1841" authorityName="Milne Edwards" authorityYear="1841" box="[408,912,986,1012]" class="Ascidiacea" family="Didemnidae" genus="Didemnum" kingdom="Animalia" order="Enterogona" pageId="10" pageNumber="47" phylum="Chordata" rank="species" species="fulgens">
<emphasis id="7AE0EA81045CFFBAFE22FC04FDEFFC6D" box="[408,629,986,1011]" italics="true" pageId="10" pageNumber="47">Didemnum fulgens</emphasis>
(Milne Edwards, 1841)
</taxonomicName>
,
<taxonomicName id="8F944D10045CFFBAFC25FC04FA84FC6A" authority="Daumezon, 1908" authorityName="Daumezon" authorityYear="1908" box="[927,1310,986,1012]" class="Ascidiacea" family="Didemnidae" genus="Didemnum" kingdom="Animalia" order="Enterogona" pageId="10" pageNumber="47" phylum="Chordata" rank="species" species="protectum">
<emphasis id="7AE0EA81045CFFBAFC25FC04FBA3FC6D" box="[927,1081,986,1011]" italics="true" pageId="10" pageNumber="47">D. protectum</emphasis>
(Daumézon, 1908)
</taxonomicName>
,
<taxonomicName id="8F944D10045CFFBAFA94FC04FE24FB82" authority="Brement, 1913" authorityName="Brement" authorityYear="1913" class="Ascidiacea" family="Didemnidae" genus="Didemnum" kingdom="Animalia" order="Enterogona" pageId="10" pageNumber="47" phylum="Chordata" rank="species" species="peyrefittense">
<emphasis id="7AE0EA81045CFFBAFA94FC04FF76FB85" italics="true" pageId="10" pageNumber="47">D. peyrefittense</emphasis>
(Brément, 1913)
</taxonomicName>
and
<taxonomicName id="8F944D10045CFFBAFDBBFBDCFC40FB82" authority="Milne Edwards, 1841" authorityName="Milne Edwards" authorityYear="1841" box="[513,986,1026,1052]" class="Ascidiacea" family="Didemnidae" genus="Didemnum" kingdom="Animalia" order="Enterogona" pageId="10" pageNumber="47" phylum="Chordata" rank="species" species="maculosum">
<emphasis id="7AE0EA81045CFFBAFDBBFBDCFD35FB85" box="[513,687,1026,1051]" italics="true" pageId="10" pageNumber="47">D. maculosum</emphasis>
(Milne Edwards, 1841)
</taxonomicName>
. According to
<bibRefCitation id="2C054B62045CFFBAFB2FFBDCFF6FFBDA" author="Lafargue" pageId="10" pageNumber="68" refString="Lafargue, F. &amp; Laubier, L. (1978 a) Anoplodelphys g. nov. Copepode Notodelphyidae parasite de Didemnidae (Ascidies Aplousobranches) en Mediterranee. Crustaceana, 35 (3), 277 - 293." type="journal article" year="1978" yearSuffix="a">Lafargue and Laubier (1978a)</bibRefCitation>
, the main distinguishing characters between “typical” and “atypical” forms were: the form and organisation of the elements of surface ornamentation, the general body shape, and the degree of development and position along the body of legs 14.
<bibRefCitation id="2C054B62045CFFBAFDE9FBA4FC23FB0A" author="Lafargue" box="[595,953,1146,1172]" pageId="10" pageNumber="68" refString="Lafargue, F. &amp; Laubier, L. (1978 a) Anoplodelphys g. nov. Copepode Notodelphyidae parasite de Didemnidae (Ascidies Aplousobranches) en Mediterranee. Crustaceana, 35 (3), 277 - 293." type="journal article" year="1978" yearSuffix="a">Lafargue and Laubier (1978a)</bibRefCitation>
selected a specimen of the typical form collected from
<taxonomicName id="8F944D10045CFFBAFEF0FB7DFE44FB22" box="[330,478,1187,1212]" class="Ascidiacea" family="Didemnidae" genus="Didemnum" kingdom="Animalia" order="Enterogona" pageId="10" pageNumber="47" phylum="Chordata" rank="species" species="commune">
<emphasis id="7AE0EA81045CFFBAFEF0FB7DFE44FB22" box="[330,478,1187,1212]" italics="true" pageId="10" pageNumber="47">D. commune</emphasis>
</taxonomicName>
as the
<typeStatus id="972F8831045CFFBAFD89FB7CFD03FB22" box="[563,665,1186,1212]" pageId="10" pageNumber="47" type="holotype">holotype</typeStatus>
(ZMA Co. 102.602) thereby fixing the name. Re-examination of this
<typeStatus id="972F8831045CFFBAFF72FB14FEB5FB7A" box="[200,303,1226,1252]" pageId="10" pageNumber="47" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8F944D10045CFFBAFEECFB15FE50FB7A" box="[342,458,1227,1252]" class="Maxillopoda" family="Notodelphyidae" genus="Anoplodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="7AE0EA81045CFFBAFEECFB15FE50FB7A" box="[342,458,1227,1252]" italics="true" pageId="10" pageNumber="47">A. incerta</emphasis>
</taxonomicName>
leads us to conclude that the typical and atypical forms are not conspecific.
</paragraph>
<paragraph id="482B3693045CFFBAFF7CFB2CFCFAFA32" blockId="10.[151,1436,226,1452]" pageId="10" pageNumber="47">
We propose to restrict the concept of
<taxonomicName id="8F944D10045CFFBAFD3AFB2DFD6FFA92" box="[640,757,1267,1292]" class="Maxillopoda" family="Notodelphyidae" genus="Anoplodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="7AE0EA81045CFFBAFD3AFB2DFD6FFA92" box="[640,757,1267,1292]" italics="true" pageId="10" pageNumber="47">A. incerta</emphasis>
</taxonomicName>
to the typical form (found on
<taxonomicName id="8F944D10045CFFBAFBE1FB2DFB75FA92" box="[1115,1263,1267,1292]" class="Ascidiacea" family="Didemnidae" genus="Didemnum" kingdom="Animalia" order="Enterogona" pageId="10" pageNumber="47" phylum="Chordata" rank="species" species="commune">
<emphasis id="7AE0EA81045CFFBAFBE1FB2DFB75FA92" box="[1115,1263,1267,1292]" italics="true" pageId="10" pageNumber="47">D. commune</emphasis>
</taxonomicName>
) characterised by the elongate body, the reduced size of legs 1 to 4, and the relatively wide spacing of legs 1 and 2 (18%) along the longitudinal axis of the body (
<tableCitation id="05160328045CFFBAFDCAFA9CFD56FAC2" box="[624,716,1346,1372]" captionStart="TABLE 2" captionStartId="11.[151,239,1386,1410]" captionTargetBox="[156,1384,1477,1658]" captionTargetPageId="11" captionText="TABLE 2. Distribution of legs along the metasome in Anoplodelphys species (calculated as distance between front edge of leg bases expressed as a percentage of total length of metasome)." httpUri="http://table.plazi.org/id/1CEB661B045DFFBBFF2DFAB4FCF0FA3A" pageId="10" pageNumber="47" tableUuid="1CEB661B045DFFBBFF2DFAB4FCF0FA3A">Table 2</tableCitation>
). We here treat the atypical form as
<taxonomicName id="8F944D10045CFFBAFB38FA9DFA0FFAC2" box="[1154,1429,1346,1372]" class="Maxillopoda" family="Notodelphyidae" genus="Anoplodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="galli">
<emphasis id="7AE0EA81045CFFBAFB38FA9DFAAAFAC2" box="[1154,1328,1347,1372]" italics="true" pageId="10" pageNumber="47">Anoplodelphys</emphasis>
cf.
<emphasis id="7AE0EA81045CFFBAFAE5FA9DFA0FFAC2" box="[1375,1429,1347,1372]" italics="true" pageId="10" pageNumber="47">galli</emphasis>
</taxonomicName>
, since the only differences from
<taxonomicName id="8F944D10045CFFBAFDAEFAB5FDF7FA1A" box="[532,621,1387,1412]" class="Maxillopoda" family="Notodelphyidae" genus="Anoplodelphys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="galli">
<emphasis id="7AE0EA81045CFFBAFDAEFAB5FDF7FA1A" box="[532,621,1387,1412]" italics="true" pageId="10" pageNumber="47">A. galli</emphasis>
</taxonomicName>
relate to the arrangement of the spinular ornamentation, a character that is difficult to assess with our current state of knowledge.
</paragraph>
</subSubSection>
</treatment>
</document>