191 lines
19 KiB
XML
191 lines
19 KiB
XML
<document id="845E69633B61E46F224A611237BC5E28" ID-DOI="10.26107/RBZ-2020-0052" ID-ISSN="2345-7600" ID-Zenodo-Dep="4576861" ID-ZooBank="C7D4631B-9B02-4E6E-9665-76CCFECA86A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614780803714" checkinUser="carolina" docAuthor="Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Buczkó, Krisztina & Vermeulen, Jaap J." docDate="2020" docId="9565D05FFFC3FFC3FFE45C54FB679EB9" docLanguage="en" docName="RafflesBZool.68.350-368.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Clostophis platytrochus Pall-Gergely & Hunyadi 2020, new species" docType="treatment" docVersion="3" lastPageNumber="364" masterDocId="695CA827FFCDFFCDFF9A5A29FFDE9B6C" masterDocTitle="Clostophis Benson, 1860, is not a monotypic diplommatinid but a speciose hypselostomatid (Gastropoda: Eupulmonata), with descriptions of six new species" masterLastPageNumber="368" masterPageNumber="350" pageNumber="364" updateTime="1698878356270" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="12B23717A149E962AFD5A8A0C14C680D">Clostophis Benson, 1860, is not a monotypic diplommatinid but a speciose hypselostomatid (Gastropoda: Eupulmonata), with descriptions of six new species</mods:title>
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<mods:namePart id="AFFFC4E374CCC238AFFD64C83C3D9BED">Páll-Gergely, Barna</mods:namePart>
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</mods:role>
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<mods:namePart id="559C1A24C2E8792D6A26B6D2EE8E6B6A">Hunyadi, András</mods:namePart>
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<mods:roleTerm id="D390D846764E98FABF6C96F4D92ADAF0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EFA00C5EFB216F9D12527F1ACDD85D5B">Grego, Jozef</mods:namePart>
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</mods:name>
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<mods:name id="97DB2B30441CDDC36FAAC6A4D005D527" type="personal">
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<mods:roleTerm id="C571AFCAAC2596B7C2758F6AE16B8C64">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="24B698CB36C70FD4CB273D6895DD1024">Reischütz, Alexander</mods:namePart>
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</mods:name>
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<mods:name id="4AABA9599077DD7A8B8B96366097CCEF" type="personal">
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</mods:role>
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<mods:namePart id="4B62D0E5B847E26265D90393A3891229">Buczkó, Krisztina</mods:namePart>
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</mods:name>
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<mods:roleTerm id="99C2FFF498490364B33B69E4AF59B590">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CC4B2FE01F8EB46E5F11611FD4E7DB40">Vermeulen, Jaap J.</mods:namePart>
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<mods:typeOfResource id="EFD42AABA72548DE64F89AF708804305">text</mods:typeOfResource>
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<mods:title id="7189D92FD35C4F0698DCC7E4EE1E6726">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="29FCFA197A28D5DE412EC3CABFA91970">
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<mods:date id="1C1D168ED2A21F1FE7C92F91398DF944">2020</mods:date>
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<mods:number id="22EC400A1FA13F725193E9AD0369F0EA">2020-05-08</mods:number>
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<mods:number id="9578C7D358737F98F816D546ED131F29">68</mods:number>
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<mods:classification id="75084E51CCB0D048769F14AF0DCEEC08">journal article</mods:classification>
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<mods:identifier id="E7D46C137409B6A1E75EFDE551EBEDDA" type="DOI">10.26107/RBZ-2020-0052</mods:identifier>
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<mods:identifier id="42B06E09D45B54F77B1D6604A26B72AE" type="ISSN">2345-7600</mods:identifier>
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<mods:identifier id="D9E6F9E6C62C0034E879EB79996BE360" type="Zenodo-Dep">4576861</mods:identifier>
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</mods:mods>
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<treatment id="9565D05FFFC3FFC3FFE45C54FB679EB9" LSID="urn:lsid:plazi:treatment:9565D05FFFC3FFC3FFE45C54FB679EB9" httpUri="http://treatment.plazi.org/id/9565D05FFFC3FFC3FFE45C54FB679EB9" lastPageNumber="364" pageId="14" pageNumber="364">
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<subSubSection id="55D632C2FFC3FFC3FFE45C54FE079DD9" pageId="14" pageNumber="364" type="nomenclature">
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<paragraph id="1D736149FFC3FFC3FFE45C54FE079DD9" blockId="14.[126,739,1660,1749]" pageId="14" pageNumber="364">
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<heading id="463BD625FFC3FFC3FFE45C54FE079DD9" bold="true" centered="true" fontSize="10" level="2" pageId="14" pageNumber="364" reason="2">
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<emphasis id="2FB8BD5BFFC3FFC3FFE45C54FE079DD9" bold="true" pageId="14" pageNumber="364">
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<taxonomicName id="DACC1ACAFFC3FFC3FFE45C54FD769DF9" ID-CoL="7QQQ6" authority="Pall-Gergely & Hunyadi" authorityName="Pall-Gergely & Hunyadi" authorityYear="2020" box="[126,680,1660,1685]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="364" phylum="Mollusca" rank="species" species="platytrochus" status="sp. nov.">
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<emphasis id="2FB8BD5BFFC3FFC3FFE45C54FE5D9DF9" bold="true" box="[126,387,1661,1685]" italics="true" pageId="14" pageNumber="364">Clostophis platytrochus</emphasis>
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||
Páll-Gergely & Hunyadi
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</taxonomicName>
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,
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<taxonomicNameLabel id="348B0020FFC3FFC3FD2F5C54FE079DD9" pageId="14" pageNumber="364" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="55D632C2FFC3FFC3FE1B5C94FE3F9DB9" box="[385,481,1725,1749]" pageId="14" pageNumber="364" type="description">
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<paragraph id="1D736149FFC3FFC3FE1B5C94FE3F9DB9" blockId="14.[126,739,1660,1749]" box="[385,481,1725,1749]" pageId="14" pageNumber="364">
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(
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<figureCitation id="85F77DCCFFC3FFC3FE135C94FE069DB9" box="[393,472,1725,1749]" captionStart="Fig" captionStartId="16.[113,148,2035,2057]" captionTargetBox="[149,1415,189,2020]" captionTargetId="figure-58@16.[149,1415,189,2020]" captionTargetPageId="16" captionText="Fig. 11. Clostophis platytrochus Páll-Gergely & Hunyadi, new species (HNHM 104408, holotype). A, apertural view; B, lateral view; C, dorsal view; D, ventral view; E, protoconch sculpture; F, G, apertural dentition; H, sculpture of the ventral side; I, sculpture of body whorl from standard apertural view." figureDoi="http://doi.org/10.5281/zenodo.4576886" httpUri="https://zenodo.org/record/4576886/files/figure.png" pageId="14" pageNumber="364">Fig. 11</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="55D632C2FFC3FFC3FFEB5CD4FF289379" pageId="14" pageNumber="364" type="materials_examined">
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<paragraph id="1D736149FFC3FFC3FFEB5CD4FE769CD9" blockId="14.[113,753,1789,1973]" pageId="14" pageNumber="364">
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<emphasis id="2FB8BD5BFFC3FFC3FFEB5CD4FEC19C79" bold="true" box="[113,287,1789,1813]" pageId="14" pageNumber="364">Type material.</emphasis>
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<materialsCitation id="ADA46B14FFC3FFC3FEB35CD4FE169CF9" collectingDate="2019-02-11" collectionCode="HNHM" collectorName="Da N & A. Hunyadi" country="Vietnam" county="Am" elevation="7" latitude="15.999066" location="Ngu Hanh" longLatPrecision="1" longitude="108.25557" municipality="Chua Quan Th" pageId="14" pageNumber="364" specimenCode="HNHM 104408" specimenCount="1" typeStatus="holotype">
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<typeStatus id="C277DFEBFFC3FFC3FEB35CD4FE509C79" box="[297,398,1789,1813]" pageId="14" pageNumber="364" type="holotype">Holotype</typeStatus>
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(SH: 1.00 mm, SW:
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<quantity id="DA34CCACFFC3FFC3FD1E5CD4FD369C79" box="[644,744,1789,1813]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="14" pageNumber="364" unit="mm" value="1.19">1.19 mm</quantity>
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) (
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<specimenCode id="4D6AC932FFC3FFC3FFE35D34FEEA9C59" box="[121,308,1821,1845]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="14" pageNumber="364">HNHM 104408</specimenCode>
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),
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<collectingCountry id="65DB21D9FFC3FFC3FED15D34FE719C59" box="[331,431,1821,1845]" name="Vietnam" pageId="14" pageNumber="364">Vietnam</collectingCountry>
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,
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<collectorName id="B039049FFFC3FFC3FE255D34FDDE9C54" box="[447,512,1821,1848]" pageId="14" pageNumber="364">Ðà N</collectorName>
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ẵng,
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<location id="18133792FFC3FFC3FDA05D37FD6F9C59" LSID="urn:lsid:plazi:treatment:9565D05FFFC3FFC3FFE45C54FB679EB9:18133792FFC3FFC3FDA05D37FD6F9C59" box="[570,689,1821,1846]" country="Vietnam" county="Am" latitude="15.999066" longLatPrecision="1" longitude="108.25557" municipality="Chua Quan Th" name="Ngu Hanh" pageId="14" pageNumber="364">Ngũ Hành</location>
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SƠn,
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<location id="18133792FFC3FFC3FFE85D14FF7D9C39" LSID="urn:lsid:plazi:treatment:9565D05FFFC3FFC3FFE45C54FB679EB9:18133792FFC3FFC3FFE85D14FF7D9C39" box="[114,163,1853,1877]" country="Vietnam" county="Am" latitude="15.999066" longLatPrecision="1" longitude="108.25557" municipality="Chua Quan Th" name="Thai" pageId="14" pageNumber="364">Thái</location>
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SƠn, environment of
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<collectingMunicipality id="FD17FB33FFC3FFC3FE0C5D14FDE89C34" box="[406,566,1853,1880]" pageId="14" pageNumber="364">Chùa Quán Th</collectingMunicipality>
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ế
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<collectingCounty id="F41219C5FFC3FFC3FDD65D17FDAA9C34" box="[588,628,1854,1880]" pageId="14" pageNumber="364">Äm</collectingCounty>
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,
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<elevation id="96E1867AFFC3FFC3FD195D17FD349C39" box="[643,746,1853,1877]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="364" unit="m" value="7.0">
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<quantity id="DA34CCACFFC3FFC3FD195D17FD719C39" box="[643,687,1854,1877]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="364" unit="m" value="7.0">7 m</quantity>
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a.s.l.
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</elevation>
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,
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<geoCoordinate id="78F8078EFFC3FFC3FFEB5D75FF259C19" box="[113,251,1884,1909]" degrees="15" direction="north" minutes="59.944" orientation="latitude" pageId="14" pageNumber="364" precision="1" value="15.999066">15°59.944′N</geoCoordinate>
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,
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<geoCoordinate id="78F8078EFFC3FFC3FE925D75FE449C19" box="[264,410,1884,1909]" degrees="108" direction="east" minutes="15.334" orientation="longitude" pageId="14" pageNumber="364" precision="1" value="108.25557">108°15.334′E</geoCoordinate>
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(locality code: 2019/25a), coll.
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<collectorName id="B039049FFFC3FFC3FFEB5D57FF2A9CF9" box="[113,244,1917,1942]" pageId="14" pageNumber="364">A. Hunyadi</collectorName>
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,
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<date id="69724789FFC3FFC3FE9B5D54FE169CF9" box="[257,456,1917,1941]" pageId="14" pageNumber="364" value="2019-02-11">
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<collectingDate id="7936BE61FFC3FFC3FE9B5D54FE169CF9" box="[257,456,1917,1941]" pageId="14" pageNumber="364" value="2019-02-11">11 February 2019</collectingDate>
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</date>
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</materialsCitation>
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.
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<typeStatus id="C277DFEBFFC3FFC3FE4F5D57FD9F9CFA" box="[469,577,1918,1942]" pageId="14" pageNumber="364" type="paratype">Paratypes</typeStatus>
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: 18 shells (coll. HA), same data as holotype.
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</paragraph>
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<paragraph id="1D736149FFC3FFC3FFEB5DF4FF289379" blockId="14.[113,752,2013,2069]" pageId="14" pageNumber="364">
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<materialsCitation id="ADA46B14FFC3FFC3FFEB5DF4FF2C9379" collectingDate="2019-02-11" collectionCode="HNHM" collectorName="Da N & A. Hunyadi" country="Vietnam" county="Am" elevation="7" latitude="15.999066" location="Ngu Hanh" longLatPrecision="1" longitude="108.25557" municipality="Chua Quan Th" pageId="14" pageNumber="364" specimenCount="5" specimenCount-juvenile="5">
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<emphasis id="2FB8BD5BFFC3FFC3FFEB5DF4FE879C99" bold="true" box="[113,345,2013,2037]" pageId="14" pageNumber="364">Additional material.</emphasis>
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<specimenCount id="0BCAAAC0FFC3FFC3FEFA5DF4FE149C99" box="[352,458,2013,2037]" pageId="14" pageNumber="364" type="juvenile">5 juvenile</specimenCount>
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shells, coll. HA, same data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="55D632C2FFC3FFC3FCB75A94FC119A39" pageId="14" pageNumber="364" type="diagnosis">
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<paragraph id="1D736149FFC3FFC3FCB75A94FC119A39" blockId="14.[813,1452,189,341]" pageId="14" pageNumber="364">
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<emphasis id="2FB8BD5BFFC3FFC3FCB75A94FC799BB9" bold="true" box="[813,935,189,213]" pageId="14" pageNumber="364">Diagnosis.</emphasis>
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A conical
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<taxonomicName id="DACC1ACAFFC3FFC3FBB65A94FB7A9BB9" authorityName="Benson" authorityYear="1860" box="[1068,1188,189,213]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="364" phylum="Mollusca" rank="genus">
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<emphasis id="2FB8BD5BFFC3FFC3FBB65A94FB7A9BB9" box="[1068,1188,189,213]" italics="true" pageId="14" pageNumber="364">Clostophis</emphasis>
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</taxonomicName>
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species with detached terminal part of body whorl, dense spiral striation on the teleoconch except for largest part of ventral side, and a strongly descending aperture having a strong parietal and a palatal tooth.
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</paragraph>
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</subSubSection>
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<subSubSection id="55D632C2FFC3FFC3FCB75B54FC5E9F79" pageId="14" pageNumber="364" type="description">
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<paragraph id="1D736149FFC3FFC3FCB75B54FA7298D9" blockId="14.[811,1454,381,949]" pageId="14" pageNumber="364">
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<emphasis id="2FB8BD5BFFC3FFC3FCB75B54FC649AF9" bold="true" box="[813,954,381,405]" pageId="14" pageNumber="364">Description.</emphasis>
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Shell white, triangular with flat base. Body whorl bluntly keeled. Terminal part of last whorl detached from penultimate whorl and strongly descending, resulting in a nearly horizontal aperture. Protoconch consisting of 1.25 whorls, with a few (ca. 6) widely-spaced spiral striae. Entire shell with 4.25–4.5 whorls. Dorsal part of teleoconch with irregular, fine radial lines, and stronger spiral striation. Spiral striae on the first whorl of teleoconch of similar density to that on the protoconch, and changes to a denser striation afterwards. Number of spiral striae 12–13 on body whorl above the keel in apertural view. Ventral side of shell with a few spiral striae near the keel, otherwise ornamented by irregular, fine, radial lines only. Umbilicus normally wide, occupies ca. one third of shell width. Aperture strongly oblique to the shell axis, nearly horizontal in lateral view. Peristome expanded but not reflected. Parietal tooth short, elevated, strong, situated in some distance from parietal callus. Palatal tooth blunt but strong, sits nearly on peristome.
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</paragraph>
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<paragraph id="1D736149FFC3FFC3FCB759F4FC5E9F79" blockId="14.[813,1451,989,1045]" pageId="14" pageNumber="364">
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<emphasis id="2FB8BD5BFFC3FFC3FCB759F4FBE39899" bold="true" box="[813,1085,989,1013]" pageId="14" pageNumber="364">Measurements (in mm).</emphasis>
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SH = 0.96–1.07, SW = 1.12–1.26 (n = 3).
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</paragraph>
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</subSubSection>
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<subSubSection id="55D632C2FFC3FFC3FCB75E14FBAB9FD9" pageId="14" pageNumber="364" type="diagnosis">
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<paragraph id="1D736149FFC3FFC3FCB75E14FBAB9FD9" blockId="14.[813,1452,1085,1205]" pageId="14" pageNumber="364">
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<emphasis id="2FB8BD5BFFC3FFC3FCB75E14FBF49F39" bold="true" box="[813,1066,1085,1109]" pageId="14" pageNumber="364">Differential diagnosis.</emphasis>
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The unique shell shape, especially the slightly keeled body whorl and the sculpture (lack of spiral striae on the largest part of the ventral side) makes this species easy to recognise.
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</paragraph>
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</subSubSection>
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<subSubSection id="55D632C2FFC3FFC3FCB75EF4FA729E79" pageId="14" pageNumber="364" type="etymology">
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<paragraph id="1D736149FFC3FFC3FCB75EF4FA729E79" blockId="14.[812,1452,1245,1301]" pageId="14" pageNumber="364">
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<emphasis id="2FB8BD5BFFC3FFC3FCB75EF4FC729F99" bold="true" box="[813,940,1245,1269]" pageId="14" pageNumber="364">Etymology.</emphasis>
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The specific epithet is the combination of ‘platus’ (= wide in Greek) and ‘trochus’ (=wheel, Latinised Greek).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="55D632C2FFC3FFC3FCB75F14FB679EB9" pageId="14" pageNumber="364" type="discussion">
|
||
<paragraph id="1D736149FFC3FFC3FCB75F14FB679EB9" blockId="14.[812,1451,1341,1493]" pageId="14" pageNumber="364">
|
||
<emphasis id="2FB8BD5BFFC3FFC3FCB75F14FC7E9E39" bold="true" box="[813,928,1341,1365]" pageId="14" pageNumber="364">Remarks.</emphasis>
|
||
This new species lives sympatrically with a new
|
||
<taxonomicName id="DACC1ACAFFC3FFC3FCFF5F74FC359E19" authorityName="Jochum, Slapnik & Pall-Gergely" authorityYear="2014" box="[869,1003,1373,1397]" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="364" phylum="Mollusca" rank="genus">
|
||
<emphasis id="2FB8BD5BFFC3FFC3FCFF5F74FC359E19" box="[869,1003,1373,1397]" italics="true" pageId="14" pageNumber="364">Angustopila</emphasis>
|
||
</taxonomicName>
|
||
species (unpublished information) and
|
||
<taxonomicName id="DACC1ACAFFC3FFC3FCB75F54FA889EF9" authority="Pall-Gergely & Hunyadi" authorityName="Pall-Gergely & Hunyadi" box="[813,1366,1405,1429]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="364" phylum="Mollusca" rank="species" species="danangensis">
|
||
<emphasis id="2FB8BD5BFFC3FFC3FCB75F54FB999EF9" box="[813,1095,1405,1429]" italics="true" pageId="14" pageNumber="364">Tonkinospira danangensis</emphasis>
|
||
Páll-Gergely & Hunyadi
|
||
</taxonomicName>
|
||
in Páll- Gergely, Grego,
|
||
<bibRefCitation id="795D1CB8FFC3FFC3FC445FB4FCBF9EB9" author="Vermeulen JJ & Luu HT & Theary K & Anker K" pageId="14" pageNumber="364" pagination="7 - 41" refId="ref9235" refString="Vermeulen JJ, Luu HT, Theary K & Anker K (2019) New species of land snails (Mollusca: Gastropoda: Caenogastropoda and Pulmonata) of the Mekong Delta Limestone Hills (Cambodia, Vietnam). Folia Malacologica, 27 (1): 7 - 41." type="journal article" year="2019">Vermeulen, Reischütz, Hunyadi & Jochum, 2019</bibRefCitation>
|
||
(see
|
||
<bibRefCitation id="795D1CB8FFC3FFC3FC075F94FB739EB9" author="Pall-Gergely B & Grego J & Vermeulen JJ & Reischutz A & Hunyadi A & Jochum A" box="[925,1197,1469,1493]" pageId="14" pageNumber="364" pagination="517 - 535" refId="ref8640" refString="Pall-Gergely B, Grego J, Vermeulen JJ, Reischutz A, Hunyadi A & Jochum A (2019) New Tonkinospira Jochum, Slapnik & Pall- Gergely, 2014 species from Laos and Vietnam (Gastropoda: Pulmonata: Hypselostomatidae). Raffles Bulletin of Zoology, 67: 517 - 535." type="journal article" year="2019">Páll-Gergely et al., 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |