treatments-xml/data/8B/93/1C/8B931CAC5AEE572692BBCB4DE6A19AF9.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.996.54187" ID-GBIF-Dataset="f7b78c80-ff88-49f3-af20-1929402411ce" ID-PMC="PMC7710680" ID-Pensoft-Pub="1313-2970-996-37" ID-Pensoft-UUID="8877ADEC42465EEBB9623B5B724E0F0F" ID-PubMed="33312045" ID-ZooBank="CF98F8DEF863419EB904183D85779CAB" ModsDocID="1313-2970-996-37" checkinTime="1606312752476" checkinUser="pensoft" docAuthor="Wang, Pei, Hu, Mei-Ling, Lin, Jun-Hong, Yang, Hai-Fang, Li, Xiao-Jing &amp; Zhou, Wei-Chuan" docDate="2020" docId="8B931CAC5AEE572692BBCB4DE6A19AF9" docLanguage="en" docName="ZooKeys 996: 37-58" docOrigin="ZooKeys 996" docSource="http://dx.doi.org/10.3897/zookeys.996.54187" docTitle="Camaena funingensis Zhou, Wang &amp; Lin 2020, sp. nov." docType="treatment" docUuid="E94E735E-BAD1-4D8C-AC91-5D50DF90AFE5" docUuidSource="ZooBank" docVersion="5" id="8877ADEC42465EEBB9623B5B724E0F0F" lastPageNumber="37" masterDocId="8877ADEC42465EEBB9623B5B724E0F0F" masterDocTitle="Descriptions of four new dextral land snails of the genus Camaena (Gastropoda, Eupulmonata, Camaenidae) from south China" masterLastPageNumber="58" masterPageNumber="37" pageNumber="37" updateTime="1668169953410" updateUser="ExternalLinkService">
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<mods:title>Descriptions of four new dextral land snails of the genus Camaena (Gastropoda, Eupulmonata, Camaenidae) from south China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Pei</mods:namePart>
<mods:affiliation>Key Laboratory of Molluscan Quarantine and Identification of GACC, Fuzhou Customs District, Fujian 350001, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hu, Mei-Ling</mods:namePart>
<mods:affiliation>Key Laboratory of Molluscan Quarantine and Identification of GACC, Fuzhou Customs District, Fujian 350001, China</mods:affiliation>
<mods:nameIdentifier type="email">93770092@qq.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lin, Jun-Hong</mods:namePart>
<mods:affiliation>Key Laboratory of Molluscan Quarantine and Identification of GACC, Fuzhou Customs District, Fujian 350001, China &amp; College of Plant Protection, Fujian Agriculture and Forestry University, Fuzhou, Fujian 350002, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Hai-Fang</mods:namePart>
<mods:affiliation>National Wetland Museum of China, Hangzhou, Zhejiang, 310013, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Xiao-Jing</mods:namePart>
<mods:affiliation>Key Laboratory of Molluscan Quarantine and Identification of GACC, Fuzhou Customs District, Fujian 350001, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhou, Wei-Chuan</mods:namePart>
<mods:affiliation>Key Laboratory of Molluscan Quarantine and Identification of GACC, Fuzhou Customs District, Fujian 350001, China</mods:affiliation>
<mods:nameIdentifier type="email">wczhou@163.com</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>996</mods:number>
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<mods:start>37</mods:start>
<mods:end>58</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.996.54187</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.996.54187</mods:identifier>
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<treatment ID-GBIF-Taxon="170515900" LSID="urn:lsid:zoobank.org:act:E94E735E-BAD1-4D8C-AC91-5D50DF90AFE5" httpUri="http://treatment.plazi.org/id/8B931CAC5AEE572692BBCB4DE6A19AF9" lastPageNumber="37" pageId="0" pageNumber="37">
<subSubSection pageId="0" pageNumber="37" type="nomenclature">
<paragraph pageId="0" pageNumber="37">
<taxonomicName LSID="8B931CAC-5AEE-5726-92BB-CB4DE6A19AF9" authority="Zhou, Wang &amp; Lin" authorityName="Zhou, Wang &amp; Lin" authorityYear="2020" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena funingensis" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="funingensis" status="sp. nov.">Camaena funingensis Zhou, Wang &amp; Lin</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="37">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Photographs of the four new species A Camaena funingensis sp. nov. (holotype, FJIQBC 19340, Laolida, Funing, Yunnan, China) B Camaena gaolongensis sp. nov. (holotype, FJIQBC 19353, Dayao, Gaolong, Guangxi, China) C Camaena maguanensis sp. nov. (FJIQBC 19405, Huazhige, Maguan, Yunnan, China) D Camaena yulinensis sp. nov. (FJIQBC 19460, Longquan cave, Yulin, Guangxi, China). Scale bars: 10 mm." figureDoi="10.3897/zookeys.996.54187.figure3" httpUri="https://binary.pensoft.net/fig/478745" pageId="0" pageNumber="37">Figures 3A</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Photographs of three camaenids A Camaena vorvonga (Pingxiang, Guangxi, China) B Camaena jinpingensis (Jinping, Yunnan, China) C Camaena longsonensis (Lang-Son, Vietnam). Scale bars: 10 mm." figureDoi="10.3897/zookeys.996.54187.figure4" httpUri="https://binary.pensoft.net/fig/478746" pageId="0" pageNumber="37">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Ecological photographs of snails A Camaena funingensis sp. nov. (Laolida, Funing, Yunnan, China) B Camaena gaolongensis sp. nov. (Dayao, Gaolong, Guangxi, China) C Camaena maguanensis sp. nov. (Huazhige, Maguan, Yunnan, China) D Camaena yulinensis sp. nov. (Longquan cave, Yulin, Guangxi, China)." figureDoi="10.3897/zookeys.996.54187.figure5" httpUri="https://binary.pensoft.net/fig/478747" pageId="0" pageNumber="37">, 5A</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Reproductive system of the snail Camaena funingensis sp. nov. (holotype, FJIQBC 19340, Laolida, Funing, Yunnan, China) A reproductive organ B penis C verge D inner penial wall. The arrow indicates opening position of the verge." figureDoi="10.3897/zookeys.996.54187.figure6" httpUri="https://binary.pensoft.net/fig/478748" pageId="0" pageNumber="37">, 6</figureCitation>
, 
<figureCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Inter and intra-specific P - distances of the COI sequences on dextral Camaena species." pageId="0" pageNumber="37">Tables 3</figureCitation>
, 
<figureCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Adult shell dimensions (mm)." pageId="0" pageNumber="37">4</figureCitation>
, 
<figureCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Diagnostic comparisons of morphological characters of the four new species." pageId="0" pageNumber="37">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="type material">
<paragraph pageId="0" pageNumber="37">Type material.</paragraph>
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">
<emphasis italics="true" pageId="0" pageNumber="37">Holotype</emphasis>
.
</emphasis>
[FJIQBC 19340] Shell height 21.0 mm, shell width 41.0 mm, height of aperture 14.0 mm, width of aperture 18.7 mm, 22 October 2014, collected from the type locality.
</paragraph>
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">
<emphasis italics="true" pageId="0" pageNumber="37">Paratype</emphasis>
.
</emphasis>
[FJIQBC 19341-19343] 3 live specimens: 2 adults, 1 juvenile.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="type locality">
<paragraph pageId="0" pageNumber="37">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="37">
Laolida, Funing, Wenshan, Yunnan, China (
<geoCoordinate degrees="23" direction="north" minutes="31" orientation="latitude" precision="1" seconds="48.88" value="23.530245">23°31'48.88&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="105" direction="east" minutes="32" orientation="longitude" precision="1" seconds="59.70" value="105.54992">105°32'59.70&quot;E</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="etymology">
<paragraph pageId="0" pageNumber="37">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="37">The name of the new species refers to the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="diagnosis">
<paragraph pageId="0" pageNumber="37">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">
<emphasis italics="true" pageId="0" pageNumber="37">Shell</emphasis>
.
</emphasis>
Shell dextral, large, thin, fragile and lucent, low, and flat conical. 4.5 whorls, the front whorls increasing slowly. Spire relatively low. Body whorl rapidly expanded. Shell light yellowish brown with clear growth lines and spiral bands on the surface. Apex quite blunt. Suture shallow. The protoconch surface smooth, and some short clear growth lines near the inner side of suture under 32 
<normalizedToken originalValue="×">x</normalizedToken>
stereomicroscope. Body whorl with carinate periphery, and a thin reddish brown band on the carina and several sparse bands below the carina. Aperture lunate, slightly descending. Peristome reflected, white, thin, sharp. Columellar lip reflected. Umbilicus reddish brown, large, only 1/5 covered. Inner lip attached to the body whorl, forming translucent callus.
</paragraph>
<caption doi="10.3897/zookeys.996.54187.figure3" httpUri="https://binary.pensoft.net/fig/478745" pageId="0" pageNumber="37" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">Figure 3.</emphasis>
Photographs of the four new species 
<emphasis bold="true" pageId="0" pageNumber="37">A</emphasis>
<taxonomicName authorityName="Zhou, Wang &amp; Lin" authorityYear="2020" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena funingensis" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="funingensis">
<emphasis italics="true" pageId="0" pageNumber="37">Camaena funingensis</emphasis>
</taxonomicName>
sp. nov. (holotype, FJIQBC 19340, Laolida, Funing, Yunnan, China) 
<emphasis bold="true" pageId="0" pageNumber="37">B</emphasis>
<taxonomicName authorityName="Zhou, Wang &amp; Lin" authorityYear="2020" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena gaolongensis" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="gaolongensis">
<emphasis italics="true" pageId="0" pageNumber="37">Camaena gaolongensis</emphasis>
</taxonomicName>
sp. nov. (holotype, FJIQBC 19353, Dayao, Gaolong, Guangxi, China) 
<emphasis bold="true" pageId="0" pageNumber="37">C</emphasis>
<taxonomicName authorityName="Zhou, Wang &amp; Hu" authorityYear="2020" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena maguanensis" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="maguanensis">
<emphasis italics="true" pageId="0" pageNumber="37">Camaena maguanensis</emphasis>
</taxonomicName>
sp. nov. (FJIQBC 19405, Huazhige, Maguan, Yunnan, China) 
<emphasis bold="true" pageId="0" pageNumber="37">D</emphasis>
<taxonomicName authorityName="Zhou, Wang &amp; Hu" authorityYear="2020" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena yulinensis" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="yulinensis">
<emphasis italics="true" pageId="0" pageNumber="37">Camaena yulinensis</emphasis>
</taxonomicName>
sp. nov. (FJIQBC 19460, Longquan cave, Yulin, Guangxi, China). Scale bars: 10 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">
<emphasis italics="true" pageId="0" pageNumber="37">Soft body</emphasis>
.
</emphasis>
Yellowish brown with irregular black lines and spots. Tentacles dark.
</paragraph>
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">
<emphasis italics="true" pageId="0" pageNumber="37">Reproductive system</emphasis>
.
</emphasis>
Bursa copulatrix oval and large with long and tapering pedunculus, expanded at the base. Flagellum long, tapering distally. Vas deferens short and thin. Epiphallus long, slightly thick. Penis retractor muscle medium length and slender, becoming wider at the end. Penis swollen and long, with longitudinal, slightly corrugated, strong and widely spaced pilasters internally. Verge ovate, opened terminally, and one clear crack on the verge surface extending from the terminal to the base.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="habitat">
<paragraph pageId="0" pageNumber="37">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="37">The species was found on limestone.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="distribution">
<paragraph pageId="0" pageNumber="37">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="37">Only known from the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="remarks">
<paragraph pageId="0" pageNumber="37">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="37">
<taxonomicName authorityName="Zhou, Wang &amp; Lin" authorityYear="2020" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena funingensis" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="funingensis">
<emphasis italics="true" pageId="0" pageNumber="37">Camaena funingensis</emphasis>
</taxonomicName>
sp. nov. is characterized by a more oblate shape, lower spire, thin and fragile shell, and yellowish brown coloration, which are clearly different from the other dextral camaenids except 
<taxonomicName lsidName="C. longsonensis" pageId="0" pageNumber="37" rank="species" species="longsonensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. longsonensis</emphasis>
</taxonomicName>
(Morlet, 1891), 
<emphasis italics="true" pageId="0" pageNumber="37">
C 
<taxonomicName lsidName="jinpingensis" pageId="0" pageNumber="37" rank="species" species="jinpingensis">jinpingensis</taxonomicName>
</emphasis>
Chen, Zhang &amp; Li, 1990, and 
<taxonomicName lsidName="C. vorvonga" pageId="0" pageNumber="37" rank="species" species="vorvonga">
<emphasis italics="true" pageId="0" pageNumber="37">C. vorvonga</emphasis>
</taxonomicName>
(Bavay &amp; Dautzenberg, 1900) (
<bibRefCitation author="Chen, DN" journalOrPublisher="Zoological Systematics" pageId="0" pageNumber="37" pagination="33 - 35" refId="B8" refString="Chen, DN, Zhang, QG, Li, BH, 1990. A new terrestrial snail species from China. Zoological Systematics 2 (6): 33 - 35" title="A new terrestrial snail species from China." volume="2" year="1990">Chen et al. 1990</bibRefCitation>
; 
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="0" pageNumber="37" pagination="1 - 68" refId="B42" refString="Schileyko, AA, 2011. Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). Ruthenica 21 (1): 1 - 68" title="Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." volume="21" year="2011">Schileyko 2011</bibRefCitation>
). The shells of the above four species are distinct from 
<taxonomicName lsidName="C. funingensis" pageId="0" pageNumber="37" rank="species" species="funingensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. funingensis</emphasis>
</taxonomicName>
in the following ways:
</paragraph>
<paragraph pageId="0" pageNumber="37">
(1) The umbilicus of 
<taxonomicName lsidName="C. funingensis" pageId="0" pageNumber="37" rank="species" species="funingensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. funingensis</emphasis>
</taxonomicName>
sp. nov. is only 1/5 covered, while that of 
<taxonomicName lsidName="C. longsonensis" pageId="0" pageNumber="37" rank="species" species="longsonensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. longsonensis</emphasis>
</taxonomicName>
is almost covered by reflected columellar lip leaving only a narrow slit, and that of 
<taxonomicName lsidName="C. jinpingensis" pageId="0" pageNumber="37" rank="species" species="jinpingensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. jinpingensis</emphasis>
</taxonomicName>
is fully covered.
</paragraph>
<paragraph pageId="0" pageNumber="37">
(2) 
<taxonomicName lsidName="C. funingensis" pageId="0" pageNumber="37" rank="species" species="funingensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. funingensis</emphasis>
</taxonomicName>
sp. nov. has several reddish brown bands at the bottom of the body whorl in addition to those on the carina, while only one thin reddish brown band is present on the carinate periphery of 
<taxonomicName lsidName="C. vorvonga" pageId="0" pageNumber="37" rank="species" species="vorvonga">
<emphasis italics="true" pageId="0" pageNumber="37">C. vorvonga</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="37">
(3) For 
<taxonomicName lsidName="C. funingensis" pageId="0" pageNumber="37" rank="species" species="funingensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. funingensis</emphasis>
</taxonomicName>
sp. nov., the verge is ovate and has one clear crack on the surface extending to the base, which makes it stand out other dextral camaenids.
</paragraph>
<caption doi="10.3897/zookeys.996.54187.figure4" httpUri="https://binary.pensoft.net/fig/478746" pageId="0" pageNumber="37" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">Figure 4.</emphasis>
Photographs of three camaenids 
<emphasis bold="true" pageId="0" pageNumber="37">A</emphasis>
<taxonomicName baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena vorvonga" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="vorvonga">
<emphasis italics="true" pageId="0" pageNumber="37">Camaena vorvonga</emphasis>
</taxonomicName>
(Pingxiang, Guangxi, China) 
<emphasis bold="true" pageId="0" pageNumber="37">B</emphasis>
<taxonomicName authorityName="Chen, Zhang &amp; Li" authorityYear="1990" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena jinpingensis" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="jinpingensis">
<emphasis italics="true" pageId="0" pageNumber="37">Camaena jinpingensis</emphasis>
</taxonomicName>
(Jinping, Yunnan, China) 
<emphasis bold="true" pageId="0" pageNumber="37">C</emphasis>
<taxonomicName baseAuthorityName="Morlet" baseAuthorityYear="1891" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena longsonensis" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="longsonensis">
<emphasis italics="true" pageId="0" pageNumber="37">Camaena longsonensis</emphasis>
</taxonomicName>
(Lang-Son, Vietnam). Scale bars: 10 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="37">
<taxonomicName authorityName="Zhou, Wang &amp; Lin" authorityYear="2020" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena gaolongensis" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="gaolongensis">
<emphasis italics="true" pageId="0" pageNumber="37">Camaena gaolongensis</emphasis>
</taxonomicName>
sp. nov. is distinguishable from 
<taxonomicName lsidName="C. funingensis" pageId="0" pageNumber="37" rank="species" species="funingensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. funingensis</emphasis>
</taxonomicName>
sp. nov. in having no spiral band. For 
<taxonomicName lsidName="C. maguanensis" pageId="0" pageNumber="37" rank="species" species="maguanensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. maguanensis</emphasis>
</taxonomicName>
sp. nov., there is no band on the carinate periphery of the body whorl except for several below the carina. Moreover, the verge of 
<taxonomicName lsidName="C. maguanensis" pageId="0" pageNumber="37" rank="species" species="maguanensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. maguanensis</emphasis>
</taxonomicName>
sp. nov. is small and circular. 
<taxonomicName authorityName="Zhou, Wang &amp; Hu" authorityYear="2020" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena yulinensis" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="yulinensis">
<emphasis italics="true" pageId="0" pageNumber="37">Camaena yulinensis</emphasis>
</taxonomicName>
sp. nov. differs to 
<taxonomicName lsidName="C. funingensis" pageId="0" pageNumber="37" rank="species" species="funingensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. funingensis</emphasis>
</taxonomicName>
sp. nov. in having a conical verge and flesh-colored peristome.
</paragraph>
<paragraph pageId="0" pageNumber="37">
<emphasis italics="true" pageId="0" pageNumber="37">P</emphasis>
-distances of the COI gene between 
<taxonomicName lsidName="C. funingensis" pageId="0" pageNumber="37" rank="species" species="funingensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. funingensis</emphasis>
</taxonomicName>
sp. nov. and the other camaenids are 0.068-0.200 (Table 
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Inter and intra-specific P - distances of the COI sequences on dextral Camaena species." httpUri="http://table.plazi.org/id/31CBB6A16103FB9DF425E69A539081D2" pageId="0" pageNumber="37" tableUuid="31CBB6A16103FB9DF425E69A539081D2">3</tableCitation>
), and those between 
<taxonomicName lsidName="C. funingensis" pageId="0" pageNumber="37" rank="species" species="funingensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. funingensis</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="C. gaolongensis" pageId="0" pageNumber="37" rank="species" species="gaolongensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. gaolongensis</emphasis>
</taxonomicName>
sp. nov., 
<taxonomicName lsidName="C. maguanensis" pageId="0" pageNumber="37" rank="species" species="maguanensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. maguanensis</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="C. yulinensis" pageId="0" pageNumber="37" rank="species" species="yulinensis">
<emphasis italics="true" pageId="0" pageNumber="37">C. yulinensis</emphasis>
</taxonomicName>
sp. nov. are 0.075, 0.068 and 0.094 respectively. All of these 
<emphasis italics="true" pageId="0" pageNumber="37">P</emphasis>
-distances exceed the maximum intra-specific value 0.059 in the family 
<taxonomicName family="Camaenidae" lsidName="" pageId="0" pageNumber="37" rank="family">Camaenidae</taxonomicName>
. On the phylogenetic tree, these four new species are adjacent, hence it is reasonable to designate this as a new species.
</paragraph>
<caption doi="10.3897/zookeys.996.54187.figure5" httpUri="https://binary.pensoft.net/fig/478747" pageId="0" pageNumber="37" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">Figure 5.</emphasis>
Ecological photographs of snails 
<emphasis bold="true" pageId="0" pageNumber="37">A</emphasis>
<taxonomicName authorityName="Zhou, Wang &amp; Lin" authorityYear="2020" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena funingensis" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="funingensis">
<emphasis italics="true" pageId="0" pageNumber="37">Camaena funingensis</emphasis>
</taxonomicName>
sp. nov. (Laolida, Funing, Yunnan, China) 
<emphasis bold="true" pageId="0" pageNumber="37">B</emphasis>
<taxonomicName authorityName="Zhou, Wang &amp; Lin" authorityYear="2020" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena gaolongensis" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="gaolongensis">
<emphasis italics="true" pageId="0" pageNumber="37">Camaena gaolongensis</emphasis>
</taxonomicName>
sp. nov. (Dayao, Gaolong, Guangxi, China) 
<emphasis bold="true" pageId="0" pageNumber="37">C</emphasis>
<taxonomicName authorityName="Zhou, Wang &amp; Hu" authorityYear="2020" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena maguanensis" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="maguanensis">
<emphasis italics="true" pageId="0" pageNumber="37">Camaena maguanensis</emphasis>
</taxonomicName>
sp. nov. (Huazhige, Maguan, Yunnan, China) 
<emphasis bold="true" pageId="0" pageNumber="37">D</emphasis>
<taxonomicName authorityName="Zhou, Wang &amp; Hu" authorityYear="2020" class="Gastropoda" family="Camaenidae" genus="Camaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camaena yulinensis" order="Stylommatophora" pageId="0" pageNumber="37" phylum="Mollusca" rank="species" species="yulinensis">
<emphasis italics="true" pageId="0" pageNumber="37">Camaena yulinensis</emphasis>
</taxonomicName>
sp. nov. (Longquan cave, Yulin, Guangxi, China).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>