152 lines
11 KiB
XML
152 lines
11 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.130.1422" ID-GBIF-Dataset="405c770a-6a91-4619-9df3-b023ce579055" ID-PMC="PMC3260751" ID-Pensoft-Pub="1313-2970-130-91" ID-PubMed="22259268" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-130-91" ModsDocOrigin="ZooKeys 130" ModsDocTitle="The smallest Neoptera (Baryshnyalidae fam. n.) from Hagen-Vorhalle (early Late Carboniferous: Namurian B; Germany)" checkinTime="1451249891680" checkinUser="pensoft" docAuthor="Ilger, Jan-Michael & Brauckmann, Carsten" docDate="2011" docId="4ADE9D05A5CBEBA787A1FFA033DCF5FC" docLanguage="en" docName="ZooKeys 130: 91-102" docOrigin="ZooKeys 130" docSource="http://dx.doi.org/10.3897/zookeys.130.1422" docTitle="Baryshnyala occulta Ilger & Brauckmann, 2011, gen. et sp. n." docType="treatment" docVersion="5" lastPageNumber="94" masterDocId="A559FF80FFAB1231FF830F57FFD0FF91" masterDocTitle="The smallest Neoptera (Baryshnyalidae fam. n.) from Hagen-Vorhalle (early Late Carboniferous: Namurian B; Germany)" masterLastPageNumber="102" masterPageNumber="91" pageNumber="92" updateTime="1668152295212" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The smallest Neoptera (Baryshnyalidae fam. n.) from Hagen-Vorhalle (early Late Carboniferous: Namurian B; Germany)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ilger, Jan-Michael</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Brauckmann, Carsten</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>130</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>91</mods:start>
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<mods:end>102</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.130.1422</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.130.1422</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-130-91</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152031087" LSID="urn:lsid:zoobank.org:act:A096877A-814A-4E30-B782-5E71ACF0FE5C" httpUri="http://treatment.plazi.org/id/4ADE9D05A5CBEBA787A1FFA033DCF5FC" lastPageId="3" lastPageNumber="94" pageId="1" pageNumber="92">
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<subSubSection pageId="1" pageNumber="92" type="nomenclature">
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<paragraph pageId="1" pageNumber="92">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:A096877A-814A-4E30-B782-5E71ACF0FE5C" class="Insecta" family="Baryshnyalidae" genus="Baryshnyala" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Baryshnyala occulta" pageId="1" pageNumber="92" phylum="Arthropoda" rank="species" species="occulta">Baryshnyala occulta</taxonomicName>
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<taxonomicNameLabel pageId="1" pageNumber="92">gen. et sp. n.</taxonomicNameLabel>
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Fig 1
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<normalizedToken originalValue="A–B2A–F">A-B2A-F</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="92" type="holotype (and only known specimen)">
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<paragraph pageId="1" pageNumber="92">Holotype (and only known specimen).</paragraph>
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<paragraph pageId="1" pageNumber="92">
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Specimen no. WMf.N P.212999, left metathoracic wing, with slightly damaged apex and lacking wing base. Stored in the collection of the LWL-Museum
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<normalizedToken originalValue="für">fuer</normalizedToken>
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Naturkunde,
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<normalizedToken originalValue="Westfälisches">Westfaelisches</normalizedToken>
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Landesmuseum mit Planetarium in
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<normalizedToken originalValue="Münster">Muenster</normalizedToken>
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(Germany).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="92" type="type locality">
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<paragraph pageId="1" pageNumber="92">Type locality.</paragraph>
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<paragraph pageId="1" pageNumber="92">
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Former brickyard quarry near Hagen-Vorhalle, North Rhine-Westphalia, Germany (topographic map 1: 25,000 sheet no. 4610 Hagen/Westfalen;
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="51.381332">51°22.88'N</geoCoordinate>
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;
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="7.4461665">007°26.77'E</geoCoordinate>
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, ~115 m a.s.l.).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="92" type="type stratum">
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<paragraph pageId="1" pageNumber="92">Type stratum.</paragraph>
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<paragraph pageId="1" pageNumber="92">Early Late Carboniferous (Early Pennsylvanian): early Bashkirian, late Namurian B, late Marsdenian, ammonoid zone R2c, Ziegelschiefer Formation.</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="92" type="diagnosis">
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<paragraph pageId="1" pageNumber="92">Diagnosis.</paragraph>
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<paragraph pageId="1" pageNumber="92">As for the genus (due to the temporarily monospecific status). Length ≥9.6 mm.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="93" pageId="1" pageNumber="92" type="preservation">
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<paragraph pageId="1" pageNumber="92">Preservation.</paragraph>
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<paragraph pageId="1" pageNumber="92">
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The isolated wing is quite well preserved, though it lacks the proximal tenth with articulation and most of the anal area. Apical wing margin is also slightly damaged. The corrugation is easily discernible but flattened by diagenetic compression. The anterior midwing is slightly damaged by preparation marks which obscure the exact reconstruction of terminal
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<normalizedToken originalValue="ScP–">ScP-</normalizedToken>
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and RA+ branches-especially whether they do or do not fuse.
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</paragraph>
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<paragraph lastPageId="2" lastPageNumber="93" pageId="1" pageNumber="92">
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Also visible in the apical area (Fig. 1), tiny prodissoconchs of young bivalves are attached to the dorsal wing membrane. These embryonic shells are common in basal
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<taxonomicName class="Insecta" family="Tachinidae" genus="Neoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoptera" order="Diptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="genus">Neoptera</taxonomicName>
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from Hagen-Vorhalle but are absent in all other Pterygota from the same locality. Most recently this has led to new interpretation of the taphonomy of the
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<pageBreakToken pageId="2" pageNumber="93" start="start">
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<normalizedToken originalValue="Lagerstätte">Lagerstaette</normalizedToken>
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</pageBreakToken>
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.-A manuscript by Ilger is currently in review,
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<bibRefCitation author="Ilger, JM" journalOrPublisher="Terra Nostra" pageId="5" pageNumber="96" pagination="55 - 56" title="Ein neues taphonomisches Modell zur Insekten-Lagerstaette Hagen-Vorhalle (" basale Neoptera "; tiefes tiefes Ober-Karbon, aelteres Bashkirium)." volume="2009" year="2009">Ilger and Brauckmann (2009)</bibRefCitation>
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gave a short overview.-The model implies a stage of drifting along a pycnocline in a well stratified water body. During this period the corpses were affected by free-swimming bivalve larvae.
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</paragraph>
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<caption pageId="2" pageNumber="93">
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<paragraph pageId="2" pageNumber="93">
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Figure 1.
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<taxonomicName class="Insecta" family="Baryshnyalidae" genus="Baryshnyala" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Baryshnyala occulta" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="occulta">Baryshnyala occulta</taxonomicName>
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gen. et sp. n.; holotype, isolated left metathoracic wing (WMf.N P.21299); early Late Carboniferous (Early Pennsylvanian: Namurian B, Marsdenian): A superimposed drawing with interpretation of the wing venation pattern; black arrow indicates arculus (arc) B photograph under polarized light; white arrows indicate attached bivalve prodissoconchs. Scale bars: 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="93" type="measurements">
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<paragraph pageId="2" pageNumber="93">Measurements.</paragraph>
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<paragraph pageId="2" pageNumber="93">
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Preserved length: 8.7 mm; estimated length: ≥9.6 mm; maximum width (at most posterior branch of
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<normalizedToken originalValue="MP–">MP-</normalizedToken>
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): 4.2 mm. Approximated ratio length/width: ~2.3.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="94" pageId="2" pageNumber="93" type="description">
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<paragraph pageId="2" pageNumber="93">Description.</paragraph>
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<paragraph lastPageId="3" lastPageNumber="94" pageId="2" pageNumber="93">
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There is only one isolated wing known. Due to its shape and the existence of a cubito-median fold we suppose it is a metathoracic wing (Figs 1-2). Costal margin in mid-wing straight, apex well rounded. Costal area strongly pectinate with wide-standing cross-veins.
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<normalizedToken originalValue="ScP–">ScP-</normalizedToken>
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simple and most likely reaching costal margin. R stem proximally thick and strong, branching in proximal quarter of wing length. RA+ simple and generally straight, terminal development not well preserved but probably reaching costal margin in distal fifth.
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<normalizedToken originalValue="RP–">RP-</normalizedToken>
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well and strongly developed, with 9 terminal branches.
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<normalizedToken originalValue="MP–">MP-</normalizedToken>
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strong, with 5 terminal branches, first branching clearly before mid-wing and fanning at basal third of wing length, posterior branches with strongly curved backwards at posterior wing margin. Strong arculus (= cross-vein between
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<normalizedToken originalValue="MP–">MP-</normalizedToken>
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and CuA+) present in proximal eighth of the wing. CuA+ slightly reduced, slightly concave in the middle part before branching, with 3 terminal branches, all very strongly curved backwards. Well developed and probably sclerotizied cubito-median fold (between
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<normalizedToken originalValue="MP–">MP-</normalizedToken>
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and CuA+) running close to CuA+, especially in the proximal part.
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<normalizedToken originalValue="CuP–">CuP-</normalizedToken>
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with 3 terminal branches, CuP1- very strongly convex. First AA+ straight.
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<pageBreakToken pageId="3" pageNumber="94" start="start">A</pageBreakToken>
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number of thin and straight cross-veins mainly in the sectors of
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<normalizedToken originalValue="RP–">RP-</normalizedToken>
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and
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<normalizedToken originalValue="MP–">MP-</normalizedToken>
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and distal of mid-wing.
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</paragraph>
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<caption pageId="3" pageNumber="94">
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<paragraph pageId="3" pageNumber="94">
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Figure 2.
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<taxonomicName class="Insecta" family="Baryshnyalidae" genus="Baryshnyala" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Baryshnyala occulta" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="occulta">Baryshnyala occulta</taxonomicName>
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gen. et sp. n.; holotype, isolated left metathoracic wing (WMf.N P.21299); early Late Carboniferous (Early Pennsylvanian: Namurian B, Marsdenian):
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<normalizedToken originalValue="A–F">A-F</normalizedToken>
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The same wing in different illumination settings. Scale bar: 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="94" type="etymology">
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<paragraph pageId="3" pageNumber="94">Etymology.</paragraph>
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<paragraph pageId="3" pageNumber="94">Latin occultus, -a, -um (adjective) meaning arcane. The holotype specimen was stored for twenty years in the collection without being identified as a new species.</paragraph>
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</subSubSection>
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</treatment>
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</document> |