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<document ID-CLB-Dataset="3754" ID-DOI="http://dx.doi.org/10.3897/zookeys.1067.72779" ID-GBIF-Dataset="39ded5c1-e041-4aa3-bf7e-5efb47e0aee2" ID-PMC="PMC8571249" ID-Pensoft-Pub="1313-2970-1067-57" ID-Pensoft-UUID="921145FDEC60583E831A6266908B4BCD" ID-PubMed="34759719" ID-ZooBank="8DB9C8FE9DF7438D92BA4B40E81F5DE1" ModsDocID="1313-2970-1067-57" checkinTime="1635566822117" checkinUser="pensoft" docAuthor="Kwanboon, Sedtawut, Sartori, Michel &amp; Boonsoong, Boonsatien" docDate="2021" docId="30F70820B88D5541AC88FEC3BF612840" docLanguage="en" docName="ZooKeys 1067: 57-82" docOrigin="ZooKeys 1067" docPubDate="2021-10-29" docSource="http://dx.doi.org/10.3897/zookeys.1067.72779" docTitle="Potamanthus (Potamanthodes) formosus Eaton 1892" docType="treatment" docVersion="6" id="921145FDEC60583E831A6266908B4BCD" lastPageNumber="57" masterDocId="921145FDEC60583E831A6266908B4BCD" masterDocTitle="Behningiidae and Potamanthidae (Insecta, Ephemeroptera) in Thailand" masterLastPageNumber="82" masterPageNumber="57" pageNumber="57" updateTime="1732749980920" updateUser="ExternalLinkService">
<mods:mods id="74019100A78B1C8927A6105A598AE3E3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FA64C051FD27C368BC8C9D39AD3A932D">
<mods:title id="E13DFD75D22BAE06DD8C1F10C3E38E3A">Behningiidae and Potamanthidae (Insecta, Ephemeroptera) in Thailand</mods:title>
</mods:titleInfo>
<mods:name id="D7360FC961F8496DC802E47EEB5DC679" type="personal">
<mods:role id="CE333B9410D1245E588A96B67F4F1D6A">
<mods:roleTerm id="58EC22885027DE1FD48FA9E6ED90418F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6F704D22F2583673E37DEFD7649515AA">Kwanboon, Sedtawut</mods:namePart>
<mods:nameIdentifier id="65FD6877D3A05E4F881846F76BC8E181" type="ORCID">https://orcid.org/0000-0002-3858-9522</mods:nameIdentifier>
<mods:affiliation id="1DF56B39F444422CAADFBB3BFFF478A9">Animal Systematics and Ecology Speciality Research Unit (ASESRU), Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
</mods:name>
<mods:name id="FCCFA3FADE3608104E7AF17B3020D901" type="personal">
<mods:role id="01382A0176AD64A2D7936137A17B7FE5">
<mods:roleTerm id="62E449292A82BBEDE99FB3F0D345DA55">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="85AE48A9BA483BA13A779C7875C6C17E">Sartori, Michel</mods:namePart>
<mods:nameIdentifier id="AD055BBDC9603DF766CD9BECA9C6EA67" type="ORCID">https://orcid.org/0000-0003-3397-3397</mods:nameIdentifier>
<mods:affiliation id="C8D9A289B9994D8341C0753C881E8CB7">Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
</mods:name>
<mods:name id="E96EE0AF4B0E13A45244836EE34549DB" type="personal">
<mods:role id="63DAF67BEE485455310CC202CF008AF2">
<mods:roleTerm id="7507388DB4F3DB88421B1A281C902148">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3D88796F5394E95DD54A72ACA405F254">Boonsoong, Boonsatien</mods:namePart>
<mods:nameIdentifier id="4496F782B50D3BD9115913C1E599DD78" type="ORCID">https://orcid.org/0000-0002-8166-0021</mods:nameIdentifier>
<mods:affiliation id="4D04692736F54DE82D542B1B482ED6F2">Animal Systematics and Ecology Speciality Research Unit (ASESRU), Department of Zoology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand</mods:affiliation>
<mods:nameIdentifier id="2AFE1014C3FAC4D064639FFFFC5A68C6" type="email">fscibtb@ku.ac.th</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="75DF1433A7D94E70C05BF23355996792">text</mods:typeOfResource>
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<mods:title id="F6F41B3300C0448A395CDB0C8C781596">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="A34E76A6FA3140842C95C888420A5320">
<mods:date id="9B233501E8793C9D50133F2B3230DEDA">2021</mods:date>
<mods:detail id="235CB3A47A0A73E77C273E70617D5A1B" type="pubDate">
<mods:number id="5B055F4FAD581F7A2DCA669F765094FF">2021-10-29</mods:number>
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<mods:detail id="2E8FCB798DAE958E709F4D9C5C2B9FC5" type="volume">
<mods:number id="0B11F208426696CB81D3B31E712AFA29">1067</mods:number>
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<mods:start id="3C96849BFD155CC9D9CD7CAA590220CD">57</mods:start>
<mods:end id="F512F9C04C72EF4372BF80C406583596">82</mods:end>
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<mods:url id="DECD36352B6742075BF07A7FD5F944D7">http://dx.doi.org/10.3897/zookeys.1067.72779</mods:url>
</mods:location>
<mods:classification id="6E81995CB716435DBC3863A7D206F851">journal article</mods:classification>
<mods:identifier id="2779C782786E49B5BC6489DECD589831" type="DOI">http://dx.doi.org/10.3897/zookeys.1067.72779</mods:identifier>
<mods:identifier id="7BD90980B86E95D533E97833A8E5DE19" type="Pensoft-Pub">1313-2970-1067-57</mods:identifier>
<mods:identifier id="2CB07F282A9790638CA8CFC7750FF68D" type="ZooBank">8DB9C8FE9DF7438D92BA4B40E81F5DE1</mods:identifier>
<mods:identifier id="3C4D423D41BF0C3406DE19847ED19E5A" type="Pensoft-UUID">921145FDEC60583E831A6266908B4BCD</mods:identifier>
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<treatment id="30F70820B88D5541AC88FEC3BF612840" ID-GBIF-Taxon="189635845" LSID="urn:lsid:plazi:treatment:30F70820B88D5541AC88FEC3BF612840" httpUri="http://treatment.plazi.org/id/30F70820B88D5541AC88FEC3BF612840" lastPageNumber="57" pageId="0" pageNumber="57" scope_class="Insecta" scope_family="Potamanthidae" scope_order="Ephemeroptera" scope_subGenus="Potamanthodes">
<subSubSection id="F418A65BE27923425379947661728FA7" pageId="0" pageNumber="57" type="nomenclature">
<paragraph id="FA68CA885EF59E76A44683F69C339144" pageId="0" pageNumber="57">
<taxonomicName id="6918EB5A09EBAAA89E3E51F728DFB982" ID-CoL="4M389" LSID="30F70820-B88D-5541-AC88-FEC3BF612840" authority="Eaton, 1892" authorityName="Eaton" authorityYear="1892" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus (Potamanthodes) formosus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="formosus" subGenus="Potamanthodes">Potamanthus (Potamanthodes) formosus Eaton, 1892</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="0BD788C9F065A60A9B6D90DF8C30DFC0" pageId="0" pageNumber="57" type="description">
<paragraph id="7CB30F857FD62048B7C081D5C507E925" pageId="0" pageNumber="57">
<figureCitation id="B733AC2D54C99AB87812552DE0742EAB" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larval morphology (head and mandibular tusk) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D angle measurement of mandibular tusk E R. distafurcus Bae &amp; McCafferty, 1991 F R. speciosus Eaton, 1881." figureDoi="10.3897/zookeys.1067.72779.figure10" httpUri="https://binary.pensoft.net/fig/604591" pageId="0" pageNumber="57">Figures 10A</figureCitation>
<figureCitation id="D06954212726EAAD4D0C19B457686F55" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larval morphology (foreleg) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D R. distafurcus Bae &amp; McCafferty, 1991 E R. speciosus Eaton, 1881. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1067.72779.figure11" httpUri="https://binary.pensoft.net/fig/604592" pageId="0" pageNumber="57">, 11A</figureCitation>
<figureCitation id="BFEE29B7BE81EF6984913C5A68728046" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Distribution map of the family Potamanthidae in Thailand." figureDoi="10.3897/zookeys.1067.72779.figure14" httpUri="https://binary.pensoft.net/fig/604595" pageId="0" pageNumber="57">, 14</figureCitation>
<figureCitation id="58EB945FE78667AA415AF4BE9C59763E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. The COI phylogenetic construction based on the maximum likelihood (ML) analysis of family Potamanthidae in Thailand." figureDoi="10.3897/zookeys.1067.72779.figure15" httpUri="https://binary.pensoft.net/fig/604596" pageId="0" pageNumber="57">, 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A10F5C8F8512B41AB3617CC6E8CAC863" pageId="0" pageNumber="57" type="materials_examined">
<paragraph id="5292BCEE71B7D56F7779A2BF7902E1D0" pageId="0" pageNumber="57">Materials examined.</paragraph>
<paragraph id="F022D81061721A775B2AAC460D9A2D27" pageId="0" pageNumber="57">
<materialsCitation id="EA63D8C3D412214D7820B449163A5197" collectingDate="2019-10-05" collectorName="B. Boonsoong" country="Thailand" latitude="2.85" location="Ban Pa Rim Tarn" longLatPrecision="1" longitude="102.20475" municipality="Makham district" specimenCount="1" specimenCount-larva="1" stateProvince="Chanthaburi province">
<specimenCount id="6067457BEA7DCBA74FEE237A03A431FA" type="larva">1 larva</specimenCount>
,
<collectingCountry id="8A962D2B04F723ED0E29DA368CEEF37A" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion id="8DF794E8C18DA90CC7497403C37A4D10" country="Thailand" name="Chanthaburi">Chanthaburi province</collectingRegion>
,
<collectingMunicipality id="B39C13EC233D5E097FCCA67F9B667137">Makham district</collectingMunicipality>
,
<location id="07165B41B0C3808C8D4C25CD9D75E716" LSID="urn:lsid:plazi:treatment:30F70820B88D5541AC88FEC3BF612840:07165B41B0C3808C8D4C25CD9D75E716" country="Thailand" latitude="2.85" longLatPrecision="1" longitude="102.20475" municipality="Makham district" name="Ban Pa Rim Tarn" stateProvince="Chanthaburi province">Ban Pa Rim Tarn</location>
homestay, 1
<geoCoordinate id="DC53274FEF63976650FEA97BF7889DE7" degrees="2" direction="north" minutes="51" orientation="latitude" precision="1" seconds="00.0" value="2.85">2°51'00.0&quot;N</geoCoordinate>
,
<geoCoordinate id="76095DB00C646DF9353F481CC65204CB" degrees="102" direction="east" minutes="12" orientation="longitude" precision="1" seconds="17.1" value="102.20475">102°12'17.1&quot;E</geoCoordinate>
,
<collectingDate id="96A89F1D98F6D9210E5BD306F36CB607" value="2019-10-05">5.X.2019</collectingDate>
,
<collectorName id="0894C7EF90679AFE3C67384D58277150">B. Boonsoong</collectorName>
leg. (ZMKU)
</materialsCitation>
;
<materialsCitation id="DFB6F1458BA154782DEAA28BA36C3107" collectingDate="2019-07-11" collectorName="S. Kwanboon" country="Thailand" elevation="175" latitude="14.659555" location="Huai Pak Kok" longLatPrecision="1" longitude="98.53397" specimenCount="2" specimenCount-larva="2" stateProvince="Kanchanaburi province">
<specimenCount id="7FC1954BE2BDD8A28FB8B2A80D10BE9D" type="larva">2 larvae</specimenCount>
,
<collectingRegion id="11B60DEA503B4FF398B59F61DA2A0BE5" country="Thailand" name="Kanchanaburi">Kanchanaburi province</collectingRegion>
,
<location id="ED6D5F345CE23786A6A6B57A0AA3A531" LSID="urn:lsid:plazi:treatment:30F70820B88D5541AC88FEC3BF612840:ED6D5F345CE23786A6A6B57A0AA3A531" country="Thailand" latitude="14.659555" longLatPrecision="1" longitude="98.53397" name="Huai Pak Kok" stateProvince="Kanchanaburi province">Huai Pak Kok</location>
,
<geoCoordinate id="C4F43B4BC531219AFF459ACA5254E5B9" degrees="14" direction="north" minutes="39" orientation="latitude" precision="1" seconds="34.4" value="14.659555">14°39'34.4&quot;N</geoCoordinate>
,
<geoCoordinate id="8F714EBAEFD7DC314EFAD37E8CDEF610" degrees="98" direction="east" minutes="32" orientation="longitude" precision="1" seconds="02.3" value="98.53397">98°32'02.3&quot;E</geoCoordinate>
,
<quantity id="38D38AD2BD76E1103FFFBB0FAA492A2F" metricMagnitude="2" metricUnit="m" metricValue="1.75" unit="m" value="175.0">
<elevation id="AFB9142D00493F8068744263DB0B5AD6" metricMagnitude="2" metricUnit="m" metricValue="1.75" unit="m" value="175.0">175 m</elevation>
</quantity>
,
<collectingDate id="0E7F25C48EDC16EDD3B08DF30EC5B61C" value="2019-07-11">11.VII.2019</collectingDate>
,
<collectorName id="FB1645223A0884FB38F1FD23EF592000">S. Kwanboon</collectorName>
leg. (ZMKU)
</materialsCitation>
;
<materialsCitation id="6F88644F4A2057A18A9EB1B5C3F873D0" collectingDate="2021-03-05" collectorName="S. Kwanboon" country="Thailand" elevation="519" latitude="20.089334" location="Huai Kang Pla" longLatPrecision="1" longitude="99.779945" specimenCount="2" specimenCount-larva="2" stateProvince="Chiang Rai province">
<specimenCount id="E817A5B468E0320B45B13BA7F9A224A3" type="larva">2 larvae</specimenCount>
,
<collectingRegion id="624F93418BEBEB2209F6D81C8AF3FCAF" country="Thailand" name="Chiang Rai">Chiang Rai province</collectingRegion>
,
<location id="87DF9954CC69935D249092A9EC71E96B" LSID="urn:lsid:plazi:treatment:30F70820B88D5541AC88FEC3BF612840:87DF9954CC69935D249092A9EC71E96B" country="Thailand" latitude="20.089334" longLatPrecision="1" longitude="99.779945" name="Huai Kang Pla" stateProvince="Chiang Rai province">Huai Kang Pla</location>
waterfall,
<geoCoordinate id="78E604DCF451AF2EACD9BD0D64922CBA" degrees="20" direction="north" minutes="05" orientation="latitude" precision="1" seconds="21.6" value="20.089334">20°05'21.6&quot;N</geoCoordinate>
,
<geoCoordinate id="6AE16A748955012F2506AAB8F8D7C2B5" degrees="99" direction="east" minutes="46" orientation="longitude" precision="1" seconds="47.8" value="99.779945">99°46'47.8&quot;E</geoCoordinate>
,
<quantity id="806EC947D416DD16050DD583BE38F150" metricMagnitude="2" metricUnit="m" metricValue="5.1899999999999995" unit="m" value="519.0">
<elevation id="2770E249CB3D6D2DCA011FC21145BF17" metricMagnitude="2" metricUnit="m" metricValue="5.1899999999999995" unit="m" value="519.0">519 m</elevation>
</quantity>
,
<collectingDate id="3A82658066D2248876F18CAF9C304670" value="2021-03-05">5.III.2021</collectingDate>
,
<collectorName id="BDCB0F9C2AC8D9EAD7E05BF3B0A7A703">S. Kwanboon</collectorName>
leg. (ZMKU)
</materialsCitation>
;
<materialsCitation id="8397B4131E0EB65287E80368186F63CA" collectingDate="2021-03-06" collectorName="S. Kwanboon" country="Thailand" latitude="20.164444" location="Klong Mae Salong" longLatPrecision="1" longitude="99.66856" specimenCount="4" specimenCount-larva="4" stateProvince="Chiang Rai province">
<specimenCount id="9D5216F55CE23C755CFA6FD02380E9C5" type="larva">4 larvae</specimenCount>
,
<collectingRegion id="7FA6627F906DFD445B5CAE6B3973D9EA" country="Thailand" name="Chiang Rai">Chiang Rai province</collectingRegion>
,
<location id="29C38EA33B2A689F38243B36C0514F8F" LSID="urn:lsid:plazi:treatment:30F70820B88D5541AC88FEC3BF612840:29C38EA33B2A689F38243B36C0514F8F" country="Thailand" latitude="20.164444" longLatPrecision="1" longitude="99.66856" name="Klong Mae Salong" stateProvince="Chiang Rai province">Klong Mae Salong</location>
,
<geoCoordinate id="FA39DB82FA119801F86A80D88DAABE1A" degrees="20" direction="north" minutes="09" orientation="latitude" precision="1" seconds="52.0" value="20.164444">20°09'52.0&quot;N</geoCoordinate>
,
<geoCoordinate id="8671BB043E8D25E38B99AA4E1C3A1CDA" degrees="99" direction="east" minutes="40" orientation="longitude" precision="1" seconds="06.8" value="99.66856">99°40'06.8&quot;E</geoCoordinate>
,
<collectingDate id="D8195623A28C7ADE80162AE7E54CBAEE" value="2021-03-06">6.III.2021</collectingDate>
,
<collectorName id="1F5EED2845F87F21E829938380675F7D">S. Kwanboon</collectorName>
leg. (ZMKU)
</materialsCitation>
;
<materialsCitation id="C7B2128B2B3C220FDA031ED9AB56F388" collectingDate="2020-11-28" collectorName="B. Boonsoong" country="Thailand" elevation="59" latitude="18.873167" location="Ban Ratsadonsamakkhi" longLatPrecision="1" longitude="100.831696" specimenCount="1" specimenCount-larva="1" stateProvince="Nan province">
<specimenCount id="E234D22B1EEF3697FDC63B3F365CBE6D" type="larva">1 larva</specimenCount>
,
<collectingRegion id="39A117BDFF42855710398E9C792CDE18" country="Thailand" name="Nan">Nan province</collectingRegion>
,
<location id="813F1F24E54ACF808AAD231D5AEB33DB" LSID="urn:lsid:plazi:treatment:30F70820B88D5541AC88FEC3BF612840:813F1F24E54ACF808AAD231D5AEB33DB" country="Thailand" latitude="18.873167" longLatPrecision="1" longitude="100.831696" name="Ban Ratsadonsamakkhi" stateProvince="Nan province">Ban Ratsadonsamakkhi</location>
,
<geoCoordinate id="9ED13210DFCB5F611CACB6F2BFFA7676" degrees="18" direction="north" minutes="52" orientation="latitude" precision="1" seconds="23.4" value="18.873167">18°52'23.4&quot;N</geoCoordinate>
,
<geoCoordinate id="6DEE2179F7EA453F125C0F31CB18B667" degrees="100" direction="east" minutes="49" orientation="longitude" precision="1" seconds="54.1" value="100.831696">100°49'54.1&quot;E</geoCoordinate>
,
<quantity id="940B129A0AE1007B55C50C77C734EBF0" metricMagnitude="1" metricUnit="m" metricValue="5.9" unit="m" value="59.0">
<elevation id="955F2B7BF88C0851BD96F477D560E80F" metricMagnitude="1" metricUnit="m" metricValue="5.9" unit="m" value="59.0">59 m</elevation>
</quantity>
,
<collectingDate id="CF74854A9B9EC5F6D994776951EBEBFA" value="2020-11-28">28.XI.2020</collectingDate>
,
<collectorName id="D976D6D320A8C5320541E65096549D43">B. Boonsoong</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BDBAE3E29F89F7F50F2E2E6B277B840D" pageId="0" pageNumber="57" type="diagnosis">
<paragraph id="9B818A75C05DC9F955D62D23942CE49A" pageId="0" pageNumber="57">Diagnosis.</paragraph>
<paragraph id="333095AC981090473A5F687C894DBA52" pageId="0" pageNumber="57">
The larvae of
<taxonomicName id="AAE1A3CEC343BCF93E1D9C2BE3CE9C9C" authorityName="Eaton" authorityYear="1892" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus formosus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="8DC29952ADC5431BE90A6657B2163D8D" italics="true" pageId="0" pageNumber="57">Potamanthus formosus</emphasis>
</taxonomicName>
Eaton, 1892 can be distinguished from those of other
<taxonomicName id="B46F9356A54296B774F23B0AE5A9E5DE" authority="" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus (Potamanthodes)" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="subGenus" subGenus="Potamanthodes">Potamanthus (Potamanthodes)</taxonomicName>
species based on the following characteristics: i) dorsal forefemora with simple stout setae (Fig.
<figureCitation id="EA9DCE475C5E3E2E3C7F807519FFCA63" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larval morphology (foreleg) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D R. distafurcus Bae &amp; McCafferty, 1991 E R. speciosus Eaton, 1881. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1067.72779.figure11" httpUri="https://binary.pensoft.net/fig/604592" pageId="0" pageNumber="57">11A</figureCitation>
), ii) a subapical cluster of setae on the foretibia, iii) short mandibular tusk (0.10-0.23
<normalizedToken id="19B390AFDA2DF3319156651015040CAA" originalValue="×">x</normalizedToken>
length of the head) (Fig.
<figureCitation id="D916C31491E1838E5261471D039286E5" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larval morphology (head and mandibular tusk) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D angle measurement of mandibular tusk E R. distafurcus Bae &amp; McCafferty, 1991 F R. speciosus Eaton, 1881." figureDoi="10.3897/zookeys.1067.72779.figure10" httpUri="https://binary.pensoft.net/fig/604591" pageId="0" pageNumber="57">10A</figureCitation>
), and iv) relatively small body length.
</paragraph>
<caption id="2F3F8A5630786ED0A9B6EC028ED77CA5" doi="10.3897/zookeys.1067.72779.figure9" httpUri="https://binary.pensoft.net/fig/604590" pageId="0" pageNumber="57" start="Figure 9" startId="F9">
<paragraph id="31F642D3C2433D235A50A2DFBDDB1B04" pageId="0" pageNumber="57">
<emphasis id="17D2FEDCDB858760E601DA7AE55169D4" bold="true" pageId="0" pageNumber="57">Figure 9.</emphasis>
Distribution map of the family
<taxonomicName id="C62BDB2A889100DDCFA7C01333826D8D" class="Insecta" family="Behningiidae" kingdom="Animalia" lsidName="" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="family">Behningiidae</taxonomicName>
in Thailand.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4E14F9A9FA4A5D8DBFC79AE6A9BB4ABA" pageId="0" pageNumber="57" type="distribution">
<paragraph id="9D70116995D71882268C0EB9469CE328" pageId="0" pageNumber="57">Distribution.</paragraph>
<paragraph id="A73DE45403BEE281420E31F5D88E00A8" pageId="0" pageNumber="57">Chanthaburi, Kanchanaburi, Nan, and Chiang Rai provinces.</paragraph>
</subSubSection>
<subSubSection id="AD280320DDE950CE11098E0AEEB55CFB" pageId="0" pageNumber="57" type="remarks">
<paragraph id="9C765164BD8DFEC27B81B512ECE830A1" pageId="0" pageNumber="57">Remark.</paragraph>
<paragraph id="E38BBF93A5B4A2A91D499710BE2EA6BB" pageId="0" pageNumber="57">
The adult of
<taxonomicName id="6ECD3F1852436B1AB56D2D4432C4B13E" lsidName="P. formosus" pageId="0" pageNumber="57" rank="species" species="formosus">
<emphasis id="7D2FD617D31D6DFF43B9B11F7393640E" italics="true" pageId="0" pageNumber="57">P. formosus</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="3777707AB874987FE6795D1502A454BE" author="Eaton, AE" journalOrPublisher="Transactions of the Entomological Society of London" pageId="0" pageNumber="57" pagination="185 - 190" refId="B4" refString="Eaton, AE, 1892. New species of Ephemeridae from the Tenasserim Valley. Transactions of the Entomological Society of London 2: 185 - 190" title="New species of Ephemeridae from the Tenasserim Valley." volume="2" year="1892">Eaton (1892)</bibRefCitation>
based on materials from Myanmar.
<bibRefCitation id="9100A23481DD6FDC20650C503D35C30E" author="Imanishi, K" journalOrPublisher="The Great Lakes Entomologist" pageId="0" pageNumber="57" refId="B9" refString="Imanishi, K, 1940. [Mayflies from Manchuria, Lower Mongolia, and Korea] (in Japanese). Report of the Limnological Society of Kwantung and Manchoukuo. Darien: 169-263." title="[Mayflies from Manchuria, Lower Mongolia, and Korea] (in Japanese). Report of the Limnological Society of Kwantung and Manchoukuo. Darien: 169 - 263." year="1940">Imanishi (1940)</bibRefCitation>
described the species
<taxonomicName id="576C4CAA18662D99A5A8925313A67DDD" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus kamonis" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="kamonis">
<emphasis id="AFC82DC89F0866B7B6267C49BAD1F9DB" italics="true" pageId="0" pageNumber="57">Potamanthus kamonis</emphasis>
</taxonomicName>
based on imaginal and larval materials from Japan, and
<taxonomicName id="EB2C328BED685318520FCB8854F7CB95" lsidName="P. kamonis" pageId="0" pageNumber="57" rank="species" species="kamonis">
<emphasis id="0BFEDDCAAC35671B85011D83614B254C" italics="true" pageId="0" pageNumber="57">P. kamonis</emphasis>
</taxonomicName>
was synonymized with
<taxonomicName id="4C54A1D928657789C6695F88CCB558A4" lsidName="P. formosus" pageId="0" pageNumber="57" rank="species" species="formosus">
<emphasis id="2CC8A44A1E50CD44731A7C05BA5DB50F" italics="true" pageId="0" pageNumber="57">P. formosus</emphasis>
</taxonomicName>
by
<bibRefCitation id="38670ED481C2506355D0C901CD64B5AE" author="Ueno, M" journalOrPublisher="Oriental Insects" pageId="0" pageNumber="57" pagination="221 - 238" refId="B22" refString="Ueno, M, 1969. Mayflies (Ephemeroptera) from various regions of Southeast Asia. Oriental Insects 3: 221 - 238" title="Mayflies (Ephemeroptera) from various regions of Southeast Asia." volume="3" year="1969">Ueno (1969)</bibRefCitation>
.
<taxonomicName id="0734CCF27581A8080199FB73A2402C66" authorityName="Eaton" authorityYear="1892" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus formosus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="3BA7878EA565441A92ED092B69FC7125" italics="true" pageId="0" pageNumber="57">Potamanthus formosus</emphasis>
</taxonomicName>
is widely distributed in East Asia and Southeast Asia (China, Japan, South Korea, Malaysia, Vietnam, Myanmar, and Thailand). In the present study, the specimens were found in eastern, western, and northern Thailand, so
<taxonomicName id="F85EE0C640E1714FAFDA89DE93D92266" lsidName="P. formosus" pageId="0" pageNumber="57" rank="species" species="formosus">
<emphasis id="BE92F7DC96CA010FABD732FBCA5AFF61" italics="true" pageId="0" pageNumber="57">P. formosus</emphasis>
</taxonomicName>
is the most widespread potamanthid in Thailand (Fig.
<figureCitation id="112BD19A88D8138841031FCAABFDE3B9" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Distribution map of the family Potamanthidae in Thailand." figureDoi="10.3897/zookeys.1067.72779.figure14" httpUri="https://binary.pensoft.net/fig/604595" pageId="0" pageNumber="57">14</figureCitation>
).
</paragraph>
<caption id="5CCCFE8A2AE50FBE6EFEFCEF74648C6B" doi="10.3897/zookeys.1067.72779.figure10" httpUri="https://binary.pensoft.net/fig/604591" pageId="0" pageNumber="57" start="Figure 10" startId="F10">
<paragraph id="B7042CB1FA7BBFA7F14C684E1872D977" pageId="0" pageNumber="57">
<emphasis id="398869BE6669BC4994827EF95C0BA5DA" bold="true" pageId="0" pageNumber="57">Figure 10.</emphasis>
Larval morphology (head and mandibular tusk)
<emphasis id="1D2202B7F23F843F6A82C441BCD8E583" bold="true" pageId="0" pageNumber="57">A</emphasis>
<taxonomicName id="39FBC17E005601EB643DA67B7AEF6F40" authorityName="Eaton" authorityYear="1892" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus formosus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="9EDE0FA8ECA491FF116CE7500E26066E" italics="true" pageId="0" pageNumber="57">Potamanthus formosus</emphasis>
</taxonomicName>
Eaton, 1892
<emphasis id="F801A2AE8E164AFCDE3ADD295F97DC56" bold="true" pageId="0" pageNumber="57">B</emphasis>
<taxonomicName id="4D8D1BED984C1E79B861362A100A132E" authorityName="Ulmer" authorityYear="1920" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhoenanthus magnificus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="magnificus">
<emphasis id="54AEAD73066329A2C992B02AA9C4CF88" italics="true" pageId="0" pageNumber="57">Rhoenanthus magnificus</emphasis>
</taxonomicName>
Ulmer, 1920
<emphasis id="00FA5D19F278A4448B8EC7B081F492EE" bold="true" pageId="0" pageNumber="57">C</emphasis>
<taxonomicName id="35060960A12E033661962EE60D826BD9" class="Insecta" kingdom="Animalia" lsidName="R. obscurus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="ADE6F6B38857BF2622984DC9166301F0" italics="true" pageId="0" pageNumber="57">R. obscurus</emphasis>
</taxonomicName>
<normalizedToken id="EFAF57FE1EA92240F71092D1D06CF4F3" originalValue="Navás">Navas</normalizedToken>
, 1922
<emphasis id="E24C5B392CA0733353E10250B3E4AA72" bold="true" pageId="0" pageNumber="57">D</emphasis>
angle measurement of mandibular tusk
<emphasis id="3405A4C9764770E9228A9C830D371EE3" bold="true" pageId="0" pageNumber="57">E</emphasis>
<taxonomicName id="0F40CFC6AAEBC9BF63574747F280E5C0" class="Insecta" kingdom="Animalia" lsidName="R. distafurcus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="distafurcus">
<emphasis id="BB5AFCBC1BAA0A13C2157B50E80D6A61" italics="true" pageId="0" pageNumber="57">R. distafurcus</emphasis>
</taxonomicName>
Bae &amp; McCafferty, 1991
<emphasis id="24E2E6A314B94C9403695029E88EA7AB" bold="true" pageId="0" pageNumber="57">F</emphasis>
<taxonomicName id="E3F78B981A922F1A7078BA88672B2029" class="Insecta" kingdom="Animalia" lsidName="R. speciosus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="speciosus">
<emphasis id="C75CE19778BAD811842C5E208ECD99A9" italics="true" pageId="0" pageNumber="57">R. speciosus</emphasis>
</taxonomicName>
Eaton, 1881.
</paragraph>
</caption>
<caption id="4DBB6ED5A5DAA89A408BB8B8FF28F032" doi="10.3897/zookeys.1067.72779.figure11" httpUri="https://binary.pensoft.net/fig/604592" pageId="0" pageNumber="57" start="Figure 11" startId="F11">
<paragraph id="874C898F8C8B99C40CEBFF8976B0276F" pageId="0" pageNumber="57">
<emphasis id="952EFEECE665A96B740B06C7AD8EFC5C" bold="true" pageId="0" pageNumber="57">Figure 11.</emphasis>
Larval morphology (foreleg)
<emphasis id="70BE0B6A75201C1FB4F4E36D2D6675F1" bold="true" pageId="0" pageNumber="57">A</emphasis>
<taxonomicName id="A301F9FC2E3430C972F14A703D8E93EA" authorityName="Eaton" authorityYear="1892" class="Insecta" family="Potamanthidae" genus="Potamanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamanthus formosus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="AEFA1A76499D217DFBA98A00FC055731" italics="true" pageId="0" pageNumber="57">Potamanthus formosus</emphasis>
</taxonomicName>
Eaton, 1892
<emphasis id="011C88520F14B24BF1AFF997F437C0E7" bold="true" pageId="0" pageNumber="57">B</emphasis>
<taxonomicName id="3E43D7788F3292E24C1800314FDC75F6" authorityName="Ulmer" authorityYear="1920" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhoenanthus magnificus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="magnificus">
<emphasis id="C572277B2D0AF3152102ACA8AF629534" italics="true" pageId="0" pageNumber="57">Rhoenanthus magnificus</emphasis>
</taxonomicName>
Ulmer, 1920
<emphasis id="BB65D29461BF411036B4CC65EC14D4AA" bold="true" pageId="0" pageNumber="57">C</emphasis>
<taxonomicName id="786D437753625F8A3CEB1CBB9F7C5529" class="Insecta" kingdom="Animalia" lsidName="R. obscurus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="D7F145E473336E01491994930CF8B659" italics="true" pageId="0" pageNumber="57">R. obscurus</emphasis>
</taxonomicName>
<normalizedToken id="EEC6EBF0146B0ED9186C2E11BB5E2296" originalValue="Navás">Navas</normalizedToken>
, 1922
<emphasis id="70E4A991A76C7352F2B8E7D8DA5E0DD0" bold="true" pageId="0" pageNumber="57">D</emphasis>
<taxonomicName id="9648E6A9534150AC85844AC54C5D1403" class="Insecta" kingdom="Animalia" lsidName="R. distafurcus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="distafurcus">
<emphasis id="E42229BC3D6E66A2E9A7D3502FD5E208" italics="true" pageId="0" pageNumber="57">R. distafurcus</emphasis>
</taxonomicName>
Bae &amp; McCafferty, 1991
<emphasis id="567698E5B0874DB1BEB8FE7078719302" bold="true" pageId="0" pageNumber="57">E</emphasis>
<taxonomicName id="9A03E4E8F7621DDFB8C359A6CA294458" class="Insecta" kingdom="Animalia" lsidName="R. speciosus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="speciosus">
<emphasis id="20F0A6DDE5ACD7012E332F593B73D65D" italics="true" pageId="0" pageNumber="57">R. speciosus</emphasis>
</taxonomicName>
Eaton, 1881. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="0D4154C6CFBC0538242D572D9BF8BF3B" doi="10.3897/zookeys.1067.72779.figure12" httpUri="https://binary.pensoft.net/fig/604593" pageId="0" pageNumber="57" start="Figure 12" startId="F12">
<paragraph id="6F3986199AC6F4274340075FA1FF1A7E" pageId="0" pageNumber="57">
<emphasis id="0E0D9C11109285018550BDD1DA74FE05" bold="true" pageId="0" pageNumber="57">Figure 12.</emphasis>
Habitus of larva of
<taxonomicName id="5BA2C7C66CCF4A262E546BCA49ADB070" authorityName="Ulmer" authorityYear="1920" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhoenanthus magnificus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="magnificus">
<emphasis id="1331E4BA8ED83FBFF2345B03CAEDA87D" italics="true" pageId="0" pageNumber="57">Rhoenanthus magnificus</emphasis>
</taxonomicName>
Ulmer, 1920.
</paragraph>
</caption>
<caption id="E9CC072D0730BE401202559F835CE9C4" doi="10.3897/zookeys.1067.72779.figure13" httpUri="https://binary.pensoft.net/fig/604594" pageId="0" pageNumber="57" start="Figure 13" startId="F13">
<paragraph id="FF3B16462B733CCBA213C31244356C19" pageId="0" pageNumber="57">
<emphasis id="9987DB2B88A6E009CAEFEABDC57059F8" bold="true" pageId="0" pageNumber="57">Figure 13.</emphasis>
SEMs of egg structures of
<taxonomicName id="E89C1AD5D671AEEF13E7CF8B8C644F4C" authorityName="Eaton" authorityYear="1881" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhoenanthus speciosus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="speciosus">
<emphasis id="5436A44EDBEA961500E33EF749169DB2" italics="true" pageId="0" pageNumber="57">Rhoenanthus speciosus</emphasis>
</taxonomicName>
Eaton, 1881
<emphasis id="B39577F8553F145382B818DAB7EC0A32" bold="true" pageId="0" pageNumber="57">A</emphasis>
general outline of egg
<emphasis id="742FC2E7C7F215457187A1B1B6B54ABD" bold="true" pageId="0" pageNumber="57">B</emphasis>
micropyle.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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