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<mods:title id="8B204942F2C8F97E119C9999A8C616C5">The genera Erhaia and Tricula (Gastropoda, Rissooidea, Amnicolidae and Pomatiopsidae) in Bhutan and elsewhere in the eastern Himalaya</mods:title>
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<mods:namePart id="EB269049A4506FB308DB1F97CBCDBE20">Gittenberger, Edmund</mods:namePart>
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<mods:namePart id="34E1F643286430E04A40BFA650F98413">Leda, Pema</mods:namePart>
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<mods:namePart id="C00D3EA220C276AAFF314F6902294277">Wangchuk, Jigme</mods:namePart>
<mods:affiliation id="35C8BE57B606E7244017E9DA2F3C1B55">Ugyen Wangchuck Institute for Conservation and Environmental Research, Bumthang, Bhutan</mods:affiliation>
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<mods:namePart id="4DCB1C84C916A379E750FCDCBDF1BD9F">Gyeltshen, Choki</mods:namePart>
<mods:affiliation id="89DBBE3990C47C0D0326E79799937CA2">National Biodiversity Centre, Serbithang, Thimphu, Bhutan</mods:affiliation>
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<mods:namePart id="EE141E7EAB762DCBBDFBD4170631F5F9">Bjoern Stelbrink,</mods:namePart>
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<treatment id="17AF80062CC55BF8B55476DE58D31D70" ID-GBIF-Taxon="163735481" LSID="urn:lsid:plazi:treatment:17AF80062CC55BF8B55476DE58D31D70" httpUri="http://treatment.plazi.org/id/17AF80062CC55BF8B55476DE58D31D70" lastPageNumber="8" pageId="7" pageNumber="8" scope_class="Gastropoda" scope_family="Amnicolidae" scope_superFamily="Rissooidea">
<subSubSection id="A246ABF9C02D4B11E6B091B51E9F1C02" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="88D806467F4B71893305218DDB9DCAED" pageId="7" pageNumber="8">
<taxonomicName id="E61BC05A72CEDFDEF3518DC840DB7ADE" ID-CoL="6GK72" LSID="17AF8006-2CC5-5BF8-B554-76DE58D31D70" authority="Nesemann &amp; S. Sharma, 2007" authorityName="Nesemann &amp; S. Sharma" authorityYear="2007" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia banepaensis" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="banepaensis">Erhaia banepaensis Nesemann &amp; S. Sharma, 2007</taxonomicName>
<figureCitation id="F76182C8C1684EF05056E58588117724" captionStart="Figures 210" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub &amp; Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann &amp; S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis &amp; Rao, 1997, holotype of E. chandeshwariensis Nesemann &amp; S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah &amp; Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="7" pageNumber="8">Fig. 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="ADE257EAC8ACE135E540181A8A18BE25" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="6D4348EA5FCD7699465B0F2299F1B12F" pageId="7" pageNumber="8">
<taxonomicName id="A56D7792C8804A26C2FBB7E53A8B42DE" authorityName="Nesemann &amp; S. Sharma" authorityYear="2007" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia banepaensis" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="banepaensis">Erhaia banepaensis</taxonomicName>
Nesemann &amp; S. Sharma, in
<bibRefCitation id="A0AF578AF801204669B12BBA60F653C5" author="Nesemann, H" journalOrPublisher="Molecular Biology and Evolution" pageId="14" pageNumber="15" refId="B13" refString="Nesemann, H, Sharma, S, Sharma, G, Khanal, SN, Pradhan, B, Shah, DN, Tachamo, RD, 2007. Class Gastropoda. In: Nesemann H, Sharma S, Sharma G, Khanal SN, Pradhan B, Shah DN, Tachamo RD (Eds) Aquatic Invertebrates of the Ganga River System, Volume 1: Mollusca, Annelida, Crustacea (in part): 57-102." title="Class Gastropoda. In: Nesemann H, Sharma S, Sharma G, Khanal SN, Pradhan B, Shah DN, Tachamo RD (Eds) Aquatic Invertebrates of the Ganga River System, Volume 1: Mollusca, Annelida, Crustacea (in part): 57 - 102." year="2007">Nesemann et al. 2007</bibRefCitation>
: 64 (&quot;Nepal, Central Zone, Kavre District, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa&quot;; the altitude is not indicated).
</paragraph>
</subSubSection>
<subSubSection id="48BD64797CFA02AD15765007E53AD708" pageId="7" pageNumber="8" type="shell">
<paragraph id="73DBAB9109B2F65009DACDBC454FA5E7" pageId="7" pageNumber="8">Shell.</paragraph>
<paragraph id="634A6B52BBEE3B6053ABDD36C4BFEF50" pageId="7" pageNumber="8">
The shells are described by Nesemann and Sharma (2007: 64) as 1.6-2.0 mm high, with 4-
<normalizedToken id="D6422E8F65232975DDAA5F35BB3C3ED4" originalValue="4½">41/2</normalizedToken>
<normalizedToken id="6A4B5264B340E760C746C4B8F4180BB0" originalValue="“convex”">&quot;convex&quot;</normalizedToken>
whorls, an aperture that is &quot;ovate but not widened and not enlarged&quot;, with an inner lip that is &quot;thin and fused to the body whorl&quot;; it can be distinguished from the other Nepalese
<taxonomicName id="AD6048A3BAFD693005D320E0B7180EF5" authorityName="Davis &amp; Kuo" authorityYear="1985" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="AF0E314F9B020F4CC1AFE71EFA60A411" italics="true" pageId="7" pageNumber="8">Erhaia</emphasis>
</taxonomicName>
species by the &quot;conical and compact shape&quot; and
<normalizedToken id="6D44CCDCF71FB4A83185B888637AC542" originalValue="“convex”">&quot;convex&quot;</normalizedToken>
whorls (2007: 65).
</paragraph>
</subSubSection>
<subSubSection id="5E2E2126FDEA0FBC300F9CA7535DB696" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="F83E1A47294B0F2EE22320D069E464C3" pageId="7" pageNumber="8">Material examined</paragraph>
<paragraph id="1CFF69421AAF99EA10BEC032CB903FF6" pageId="7" pageNumber="8">
(photo only).
<emphasis id="90F9BEF3188937A4E8BF812E575C9FDA" bold="true" pageId="7" pageNumber="8">
<emphasis id="22ADF034F817CC0913570DC252F1D62F" italics="true" pageId="7" pageNumber="8">Holotype</emphasis>
.
</emphasis>
(NHMW 1033159).
</paragraph>
</subSubSection>
<subSubSection id="ED6010D9BE7056B22007E87A317D38BC" pageId="7" pageNumber="8" type="notes">
<paragraph id="DFAD5647351942957D21D611044820CC" pageId="7" pageNumber="8">Notes.</paragraph>
<paragraph id="F1B0BBC6E582D78C2C2CA6F9B30AD8C1" pageId="7" pageNumber="8">
Nesemann and S. Sharma are mentioned as authors for this species, without specifying for what part of the text in
<bibRefCitation id="73A5B70C4DDC6EA1F0BBE4752D2222C0" author="Nesemann, H" journalOrPublisher="Molecular Biology and Evolution" pageId="14" pageNumber="15" refId="B13" refString="Nesemann, H, Sharma, S, Sharma, G, Khanal, SN, Pradhan, B, Shah, DN, Tachamo, RD, 2007. Class Gastropoda. In: Nesemann H, Sharma S, Sharma G, Khanal SN, Pradhan B, Shah DN, Tachamo RD (Eds) Aquatic Invertebrates of the Ganga River System, Volume 1: Mollusca, Annelida, Crustacea (in part): 57-102." title="Class Gastropoda. In: Nesemann H, Sharma S, Sharma G, Khanal SN, Pradhan B, Shah DN, Tachamo RD (Eds) Aquatic Invertebrates of the Ganga River System, Volume 1: Mollusca, Annelida, Crustacea (in part): 57 - 102." year="2007">Nesemann et al (2007)</bibRefCitation>
they have responsibility.
</paragraph>
<paragraph id="4353E7B72928A026319E05030015BD6D" pageId="7" pageNumber="8">
According to
<bibRefCitation id="83A290945374C0E72579B43D23040E49" author="Nesemann, H" journalOrPublisher="Molecular Biology and Evolution" pageId="14" pageNumber="15" refId="B13" refString="Nesemann, H, Sharma, S, Sharma, G, Khanal, SN, Pradhan, B, Shah, DN, Tachamo, RD, 2007. Class Gastropoda. In: Nesemann H, Sharma S, Sharma G, Khanal SN, Pradhan B, Shah DN, Tachamo RD (Eds) Aquatic Invertebrates of the Ganga River System, Volume 1: Mollusca, Annelida, Crustacea (in part): 57-102." title="Class Gastropoda. In: Nesemann H, Sharma S, Sharma G, Khanal SN, Pradhan B, Shah DN, Tachamo RD (Eds) Aquatic Invertebrates of the Ganga River System, Volume 1: Mollusca, Annelida, Crustacea (in part): 57 - 102." year="2007">Nesemann et al. (2007</bibRefCitation>
: 64)
<taxonomicName id="3BD630AC4B55F47A6F960861078F0EB0" lsidName="E. banepaensis" pageId="7" pageNumber="8" rank="species" species="banepaensis">
<emphasis id="A733D5FB2AA9A9E92D2D6A685209E324" italics="true" pageId="7" pageNumber="8">E. banepaensis</emphasis>
</taxonomicName>
occurs sympatrically with
<taxonomicName id="DD1ACC29A819C531B43EF5AAA937944E" lsidName="E. chandeshwariensis" pageId="7" pageNumber="8" rank="species" species="chandeshwariensis">
<emphasis id="0093268D23EE3E7EE511FCAC14972374" italics="true" pageId="7" pageNumber="8">E. chandeshwariensis</emphasis>
</taxonomicName>
at the shared type locality of these species.
<taxonomicName id="A4A34565770B48039DE3EEC95CE5D8EC" authorityName="Nesemann &amp; S. Sharma" authorityYear="2007" class="Gastropoda" family="Amnicolidae" genus="Erhaia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erhaia banepaensis" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="banepaensis">
<emphasis id="24B0EBFA255F5C9BD28D4E3E236DC80B" italics="true" pageId="7" pageNumber="8">Erhaia banepaensis</emphasis>
</taxonomicName>
is supposed to be more widely distributed in Nepal between 1400 and 2085 m a.s.l., but the type series is restricted to the holotype (NHMW 103319 [not 1033159]) and a paratype (NHMW 103320). The location of the additional material is not indicated.
</paragraph>
<caption id="E42E7EC3C31A21D7F9617F5E50970907" httpUri="https://binary.pensoft.net/fig/402720" pageId="7" pageNumber="8" start="Figure 13" startId="F5">
<paragraph id="6632E7C0D0B0CADD53599172B3E30B09" pageId="7" pageNumber="8">
<emphasis id="7661034B563CA54E308BFEE5088B3F7C" bold="true" pageId="7" pageNumber="8">Figure 13.</emphasis>
Habitat of
<taxonomicName id="1C835CC4D7C61FF3D2C96E0A291F0D39" baseAuthorityName="Benson" baseAuthorityYear="1843" class="Gastropoda" family="Pomatiopsidae" genus="Tricula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tricula montana" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="montana">
<emphasis id="683F2FD0DF3A9577CD6F429E1170EF17" italics="true" pageId="7" pageNumber="8">Tricula montana</emphasis>
</taxonomicName>
; Bhutan, district Mongar, Chhuburee, 818 m a.s.l. Photo by JW.
</paragraph>
</caption>
</subSubSection>
</treatment>
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