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<document id="1F55B3B05A230C56CBE4CA69A9058A80" ID-CLB-Dataset="25263" ID-DOI="10.11646/zootaxa.4683.4.1" ID-GBIF-Dataset="48496621-de1a-4490-b866-1ae69d81ecbc" ID-ISSN="1175-5326" ID-Zenodo-Dep="3479609" ID-ZooBank="982FE3FB-5610-44A1-AAE3-659B28F45307" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570688900916" checkinUser="plazi" docAuthor="Lee, Chi-Feng &amp; Bezdĕk, Jan" docDate="2019" docId="03FF87D6FFBB4464FF6BAF8BFE15F9AA" docLanguage="en" docName="zootaxa.4683.4.1.pdf" docOrigin="Zootaxa 4683 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Theopea tsoui Lee &amp; Bezdĕk 2019, sp. nov." docType="treatment" docVersion="15" lastPageNumber="495" masterDocId="FFC6FFAEFF974448FFFCAF12FFAEFFA1" masterDocTitle="Revision of the genus Theopea Baly (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland and the Philippines: Redefinition of the genus" masterLastPageNumber="507" masterPageNumber="451" pageNumber="495" updateTime="1698758705207" updateUser="ExternalLinkService">
<mods:mods id="B243BA6F4C165D3F47B6422C81CA170D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E9E244F7D4416A7F1E943CD7C1CC1B67">Revision of the genus Theopea Baly (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland and the Philippines: Redefinition of the genus</mods:title>
</mods:titleInfo>
<mods:name id="488FBD5C9D313FB1F00E4F381EAD35C7" type="personal">
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<mods:namePart id="6C730A6A89D454AF0D66D3378615EDF4">Lee, Chi-Feng</mods:namePart>
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<mods:namePart id="9C0C88A5BEF3BB1E8C317869DC3B6457">Bezdĕk, Jan</mods:namePart>
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<mods:title id="673C6A3288A1D79F736888F53A11585B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B5B5ECEF10F7C11B2B2315F3FCBA87DF">
<mods:date id="B5D7248C8D879A4DB3D912D046DF969D">2019</mods:date>
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<mods:number id="64142070697462C7BD4A6F365DE31A81">2019-10-10</mods:number>
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<mods:number id="2EEE23CB35EB99996BBF9F2BF68EC404">4683</mods:number>
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<mods:classification id="0BC445AF35369BF69BDE01195D3F7863">journal article</mods:classification>
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<treatment id="03FF87D6FFBB4464FF6BAF8BFE15F9AA" ID-DOI="http://doi.org/10.5281/zenodo.4505277" ID-GBIF-Taxon="159690567" ID-Zenodo-Dep="4505277" LSID="urn:lsid:plazi:treatment:03FF87D6FFBB4464FF6BAF8BFE15F9AA" httpUri="http://treatment.plazi.org/id/03FF87D6FFBB4464FF6BAF8BFE15F9AA" lastPageNumber="495" pageId="44" pageNumber="495">
<subSubSection id="C34C654BFFBB4464FF6BAF8BFE08FF15" box="[151,422,153,180]" pageId="44" pageNumber="495" type="nomenclature">
<paragraph id="8BE936C0FFBB4464FF6BAF8BFE08FF15" blockId="44.[151,422,153,215]" box="[151,422,153,180]" pageId="44" pageNumber="495">
<heading id="D0A181ACFFBB4464FF6BAF8BFE08FF15" bold="true" box="[151,422,153,180]" fontSize="11" level="1" pageId="44" pageNumber="495" reason="1">
<emphasis id="B922EAD2FFBB4464FF6BAF8BFE08FF15" bold="true" box="[151,422,153,180]" pageId="44" pageNumber="495">
<taxonomicName id="4C564D43FFBB4464FF6BAF8BFEECFF12" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[151,322,153,180]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="tsoui" status="sp. nov.">
<emphasis id="B922EAD2FFBB4464FF6BAF8BFEECFF12" bold="true" box="[151,322,153,180]" italics="true" pageId="44" pageNumber="495">Theopea tsoui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157A9FFBB4464FEB5AF88FE08FF15" box="[329,422,154,180]" pageId="44" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFFBB4464FF6BAFAFFD59FB02" pageId="44" pageNumber="495" type="description">
<paragraph id="8BE936C0FFBB4464FF6BAFAFFEDBFF76" blockId="44.[151,422,153,215]" box="[151,373,189,215]" pageId="44" pageNumber="495">
(
<figureCitation id="136D2A45FFBB4464FF63AFAFFF5DFF76" box="[159,243,189,215]" captionStart="FIGURE 19" captionStartId="37.[151,250,1936,1961]" captionTargetBox="[151,1406,268,1771]" captionTargetId="figure@37.[151,1435,259,1771]" captionTargetPageId="37" captionText="FIGURE 19. Habitus of Theopea kedenburgi Weise and T. tsoui sp. nov. 19A. T. kedenburgi, male, dorsal view; 19B. T. keden- burgi, male, dorsal view, color variation; 19C. T. kedenburgi, male, dorsal view, color variation; 19D. T. tsoui sp. nov., male, dorsal view; 19E. Same, ventral view; 19F. T. tsoui sp. nov., female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3479647" httpUri="https://zenodo.org/record/3479647/files/figure.png" pageId="44" pageNumber="495">Figs 19</figureCitation>
D19F, 23)
</paragraph>
<paragraph id="8BE936C0FFBB4464FF6BAE14FAA0FE2A" blockId="44.[151,1437,261,1548]" pageId="44" pageNumber="495">
<emphasis id="B922EAD2FFBB4464FF6BAE14FE80FE81" bold="true" box="[151,302,261,288]" pageId="44" pageNumber="495">Types (n= 5).</emphasis>
<materialsCitation id="3B3E3C9DFFBB4464FECFAE17FECFFEE2" ID-GBIF-Occurrence="2425473974" collectingDate="1992-02-08" collectionCode="BMNH" collectorName="N. Mawdsley." country="Brunei" latitude="4.5666666" location="Kula Belalong" longLatPrecision="1306" longitude="115.11667" pageId="44" pageNumber="495" specimenCount="1" specimenCount-male="1" stateProvince="Temburong" typeStatus="holotype">
<typeStatus id="54ED8862FFBB4464FECFAE17FE35FEBE" box="[307,411,261,287]" pageId="44" pageNumber="495" type="holotype">Holotype</typeStatus>
<specimenCount id="9D50FD49FFBB4464FE5CAE14FE1BFE81" box="[416,437,262,288]" pageId="44" pageNumber="495" type="male"></specimenCount>
(
<collectionCode id="ED47AE05FFBB4464FE3EAE17FDB2FEBE" box="[450,540,261,287]" pageId="44" pageNumber="495">BMNH</collectionCode>
),
<emphasis id="B922EAD2FFBB4464FDD1AE14FD34FE81" bold="true" box="[557,666,262,288]" pageId="44" pageNumber="495">
<collectingCountry id="F3417650FFBB4464FDD1AE14FD34FE81" box="[557,666,262,288]" name="Brunei" pageId="44" pageNumber="495">BRUNEI</collectingCountry>
</emphasis>
.
<collectingRegion id="4992F822FFBB4464FD59AE17FC2CFEBE" box="[677,898,261,287]" country="Brunei" name="Temburong" pageId="44" pageNumber="495">Temburong District</collectingRegion>
,
<location id="8E89601BFFBB4464FC77AE17FB85FEBE" LSID="urn:lsid:plazi:treatment:03FF87D6FFBB4464FF6BAF8BFE15F9AA:8E89601BFFBB4464FC77AE17FB85FEBE" box="[907,1067,261,287]" country="Brunei" latitude="4.5666666" longLatPrecision="1306" longitude="115.11667" name="Kula Belalong" pageId="44" pageNumber="495" stateProvince="Temburong">Kula Belalong</location>
FSC,
<geoCoordinate id="EE625007FFBB4464FB92AE17FB65FE81" box="[1134,1227,261,288]" degrees="115" direction="east" minutes="7" orientation="longitude" pageId="44" pageNumber="495" precision="925" value="115.11667">115°7E</geoCoordinate>
<geoCoordinate id="EE625007FFBB4464FB2CAE17FA8BFEBE" box="[1232,1317,261,287]" degrees="4" direction="north" minutes="34" orientation="latitude" pageId="44" pageNumber="495" precision="925" value="4.5666666">4°34N</geoCoordinate>
,
<date id="FFE81000FFBB4464FAD3AE17FA39FEBE" box="[1327,1431,261,287]" pageId="44" pageNumber="495" value="1992-02-08">
<collectingDate id="EFACE9E8FFBB4464FAD3AE17FA39FEBE" box="[1327,1431,261,287]" pageId="44" pageNumber="495" value="1992-02-08">8.II.1992</collectingDate>
</date>
, leg.
<collectorName id="26A35316FFBB4464FF3AAE38FECFFEE2" box="[198,353,297,324]" pageId="44" pageNumber="495">N. Mawdsley.</collectorName>
</materialsCitation>
<materialsCitation id="3B3E3C9DFFBB4464FE9AAE38FCDEFEE2" ID-GBIF-Occurrence="2425474186" box="[358,880,297,324]" collectingDate="1992-02-08" collectionCode="BMNH" collectorName="N. Mawdsley." country="Brunei" latitude="4.5666666" location="Kula Belalong" longLatPrecision="1306" longitude="115.11667" pageId="44" pageNumber="495" specimenCount="1" specimenCount-male="1" stateProvince="Temburong" typeStatus="paratype">
<typeStatus id="54ED8862FFBB4464FE9AAE38FE79FEE5" box="[358,471,298,324]" pageId="44" pageNumber="495" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="9D50FD49FFBB4464FE23AE3BFDACFEE2" box="[479,514,297,323]" pageId="44" pageNumber="495" type="male">1♂</specimenCount>
(
<collectionCode id="ED47AE05FFBB4464FDF3AE3BFDC7FEE2" box="[527,617,297,323]" pageId="44" pageNumber="495">BMNH</collectionCode>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="3B3E3C9DFFBB4464FC86AE3BFD4CFEC6" ID-GBIF-Occurrence="2425474155" collectingDate="1991-05-19" collectionCode="BMNH" collectorName="N. Mawdsley" country="Brunei" latitude="4.5666666" location="Kula Belalong" longLatPrecision="1306" longitude="115.11667" pageId="44" pageNumber="495" specimenCount="1" specimenCount-female="1" stateProvince="Temburong" typeStatus="paratype">
<specimenCount id="9D50FD49FFBB4464FC86AE3BFC32FEE2" box="[890,924,297,323]" pageId="44" pageNumber="495" type="female">1♀</specimenCount>
(
<collectionCode id="ED47AE05FFBB4464FC56AE3BFBAAFEE2" box="[938,1028,297,323]" pageId="44" pageNumber="495">BMNH</collectionCode>
),
<collectingRegion id="4992F822FFBB4464FBE9AE3BFB5CFEE2" box="[1045,1266,297,323]" country="Brunei" name="Temburong" pageId="44" pageNumber="495">Temburong District</collectingRegion>
,
<location id="8E89601BFFBB4464FB07AE3BFA32FEE2" LSID="urn:lsid:plazi:treatment:03FF87D6FFBB4464FF6BAF8BFE15F9AA:8E89601BFFBB4464FB07AE3BFA32FEE2" box="[1275,1436,297,323]" country="Brunei" latitude="4.5666666" longLatPrecision="1306" longitude="115.11667" name="Kula Belalong" pageId="44" pageNumber="495" stateProvince="Temburong">Kula Belalong</location>
FSC,
<geoCoordinate id="EE625007FFBB4464FF2AAE5FFE9DFEC9" box="[214,307,333,360]" degrees="115" direction="east" minutes="7" orientation="longitude" pageId="44" pageNumber="495" precision="925" value="115.11667">115°7E</geoCoordinate>
<geoCoordinate id="EE625007FFBB4464FEC5AE5FFE21FEC6" box="[313,399,333,359]" degrees="4" direction="north" minutes="34" orientation="latitude" pageId="44" pageNumber="495" precision="925" value="4.5666666">4°34N</geoCoordinate>
,
<date id="FFE81000FFBB4464FE65AE5FFDA0FEC6" box="[409,526,333,359]" pageId="44" pageNumber="495" value="1991-05-19">
<collectingDate id="EFACE9E8FFBB4464FE65AE5FFDA0FEC6" box="[409,526,333,359]" pageId="44" pageNumber="495" value="1991-05-19">19.V.1991</collectingDate>
</date>
, leg.
<collectorName id="26A35316FFBB4464FDB5AE5CFD4CFEC6" box="[585,738,333,360]" pageId="44" pageNumber="495">N. Mawdsley</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3E3C9DFFBB4464FD11AE5FFB7CFEC6" ID-GBIF-Occurrence="2425474177" box="[749,1234,333,360]" collectingDate="1991-07-26" collectionCode="BMNH" country="Brunei" pageId="44" pageNumber="495" specimenCount="1" specimenCount-female="1" stateProvince="Temburong" typeStatus="paratype">
<specimenCount id="9D50FD49FFBB4464FD11AE5FFCBEFEC6" box="[749,784,333,359]" pageId="44" pageNumber="495" type="female">1♀</specimenCount>
(
<collectionCode id="ED47AE05FFBB4464FCE2AE5FFCD6FEC6" box="[798,888,333,359]" pageId="44" pageNumber="495">BMNH</collectionCode>
), same but with “
<date id="FFE81000FFBB4464FBC4AE5FFB69FEC6" box="[1080,1223,333,359]" pageId="44" pageNumber="495" value="1991-07-26">
<collectingDate id="EFACE9E8FFBB4464FBC4AE5FFB69FEC6" box="[1080,1223,333,359]" pageId="44" pageNumber="495" value="1991-07-26">26.VII.1991</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E3C9DFFBB4464FB21AE5FFAA0FE2A" ID-GBIF-Occurrence="2425474185" collectingDate="2015-02-12" collectingDateMax="2015-02-19" collectingDateMin="2015-02-12" collectionCode="JBCB" collectorName="P. Kocarek." country="Brunei" elevation="130" latitude="-4.5469446" location="Ulu" longLatPrecision="21" longitude="115.156944" municipality="Darussalam" pageId="44" pageNumber="495" specimenCount="1" specimenCount-male="1" stateProvince="Temburong" typeStatus="paratype">
<specimenCount id="9D50FD49FFBB4464FB21AE5FFAAEFEC6" box="[1245,1280,333,359]" pageId="44" pageNumber="495" type="male">1♂</specimenCount>
(
<collectionCode id="ED47AE05FFBB4464FAF2AE5FFAF8FEC6" box="[1294,1366,333,359]" pageId="44" pageNumber="495">JBCB</collectionCode>
), Bru- nei
<collectingMunicipality id="6B8DACBAFFBB4464FF3CAE63FEE9FE2A" box="[192,327,369,395]" pageId="44" pageNumber="495">Darussalam</collectingMunicipality>
,
<location id="8E89601BFFBB4464FEAFAE63FED3FE2A" LSID="urn:lsid:plazi:treatment:03FF87D6FFBB4464FF6BAF8BFE15F9AA:8E89601BFFBB4464FEAFAE63FED3FE2A" box="[339,381,369,395]" country="Brunei" latitude="-4.5469446" longLatPrecision="21" longitude="115.156944" municipality="Darussalam" name="Ulu" pageId="44" pageNumber="495" stateProvince="Temburong">Ulu</location>
<collectingRegion id="4992F822FFBB4464FE7FAE63FDAAFE2A" box="[387,516,369,395]" country="Brunei" name="Temburong" pageId="44" pageNumber="495">Temburong</collectingRegion>
N.P.,
<quantity id="4CAE9B25FFBB4464FDBBAE63FD3EFE2D" box="[583,656,369,396]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="44" pageNumber="495" unit="m" value="130.0">
<elevation id="007BD1F3FFBB4464FDBBAE63FD3EFE2D" box="[583,656,369,396]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="44" pageNumber="495" unit="m" value="130.0">130 m</elevation>
</quantity>
,
<geoCoordinate id="EE625007FFBB4464FD60AE63FCBDFE2A" box="[668,787,369,395]" degrees="4" direction="south" minutes="32" orientation="latitude" pageId="44" pageNumber="495" precision="15" seconds="49" value="-4.5469446">4°3249”S</geoCoordinate>
<geoCoordinate id="EE625007FFBB4464FCE6AE63FC0FFE2D" box="[794,929,369,396]" degrees="115" direction="east" minutes="9" orientation="longitude" pageId="44" pageNumber="495" precision="15" seconds="25" value="115.156944">115°925”E</geoCoordinate>
,
<date id="FFE81000FFBB4464FC51AE63FBE0FE2D" box="[941,1102,369,396]" pageId="44" pageNumber="495" value="2015-02-12" valueMax="2015-02-19" valueMin="2015-02-12">
<collectingDate id="EFACE9E8FFBB4464FC51AE63FBE0FE2D" box="[941,1102,369,396]" pageId="44" pageNumber="495" value="2015-02-12" valueMax="2015-02-19" valueMin="2015-02-12">1219.II.2015</collectingDate>
</date>
, leg.
<collectorName id="26A35316FFBB4464FB76AE60FAA0FE2A" box="[1162,1294,369,396]" pageId="44" pageNumber="495">P. Kočárek.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BE936C0FFBB4464FF3BAE87FD59FB02" blockId="44.[151,1437,261,1548]" pageId="44" pageNumber="495">
<emphasis id="B922EAD2FFBB4464FF3BAE87FEF7FE11" bold="true" box="[199,345,405,432]" pageId="44" pageNumber="495">Description.</emphasis>
Length
<quantity id="4CAE9B25FFBB4464FE45AE87FDEAFE11" box="[441,580,405,432]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="8.4" metricValueMin="8.2" pageId="44" pageNumber="495" unit="mm" value="8.3" valueMax="8.4" valueMin="8.2">8.28.4 mm</quantity>
, width
<quantity id="4CAE9B25FFBB4464FD66AE87FC88FE11" box="[666,806,405,432]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.2" metricValueMin="3.1" pageId="44" pageNumber="495" unit="mm" value="3.15" valueMax="3.2" valueMin="3.1">3.13.2 mm</quantity>
. General color (
<figureCitation id="136D2A45FFBB4464FC21AE87FB9CFE0E" box="[989,1074,405,431]" captionStart="FIGURE 19" captionStartId="37.[151,250,1936,1961]" captionTargetBox="[151,1406,268,1771]" captionTargetId="figure@37.[151,1435,259,1771]" captionTargetPageId="37" captionText="FIGURE 19. Habitus of Theopea kedenburgi Weise and T. tsoui sp. nov. 19A. T. kedenburgi, male, dorsal view; 19B. T. keden- burgi, male, dorsal view, color variation; 19C. T. kedenburgi, male, dorsal view, color variation; 19D. T. tsoui sp. nov., male, dorsal view; 19E. Same, ventral view; 19F. T. tsoui sp. nov., female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3479647" httpUri="https://zenodo.org/record/3479647/files/figure.png" pageId="44" pageNumber="495">Figs 19</figureCitation>
D19F) purplish black; antenna black except antennomeres XXI white or pale yellow. Antennae filiform in males, except antennomeres VIIIX strongly swollen, X with a broad lateral process at middle (
<figureCitation id="136D2A45FFBB4464FCCEAECFFC34FE56" box="[818,922,477,503]" captionStart="FIGURE 23" captionStartId="45.[151,250,1932,1957]" captionTargetBox="[163,1420,179,1908]" captionTargetId="figure@45.[151,1436,174,1912]" captionTargetPageId="45" captionText="FIGURE 23. Diagnostic characters of Theopea tsoui sp. nov. 23A. Antenna, male; 23B. Antenna, female; 23C. Aedeagus, dorsal view; 23D. Aedeagus, lateral view; 23E. Aedeagus, ventral view; 23F. Abdominal ventrite VIII; 23G. Gonocoxae; 23H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479655" httpUri="https://zenodo.org/record/3479655/files/figure.png" pageId="44" pageNumber="495">Fig. 23A</figureCitation>
), length ratios of antennomeres IXI 1.0: 0.3: 0.7: 1.0: 1.0: 1.0: 1.3: 1.1: 1.1: 0.8: 1.1, length to width ratios of antennomeres IXI 3.1: 1.4: 2.6: 3.3: 3.4: 2.7: 2.5: 2.3: 2.3: 2.1: 4.6; filiform in females (
<figureCitation id="136D2A45FFBB4464FD2FAD37FC96FD9E" box="[723,824,549,575]" captionStart="FIGURE 23" captionStartId="45.[151,250,1932,1957]" captionTargetBox="[163,1420,179,1908]" captionTargetId="figure@45.[151,1436,174,1912]" captionTargetPageId="45" captionText="FIGURE 23. Diagnostic characters of Theopea tsoui sp. nov. 23A. Antenna, male; 23B. Antenna, female; 23C. Aedeagus, dorsal view; 23D. Aedeagus, lateral view; 23E. Aedeagus, ventral view; 23F. Abdominal ventrite VIII; 23G. Gonocoxae; 23H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479655" httpUri="https://zenodo.org/record/3479655/files/figure.png" pageId="44" pageNumber="495">Fig. 23B</figureCitation>
), antennomeres VIVIII slightly swollen, length ratios of antennomeres IXI 1.0: 0.3: 0.6: 0.8: 0.8: 0.8: 0.9: 0.8: 0.8: 0.7: 1.0, length to width ratios of antennomeres IXI 3.3: 1.6: 2.6: 3.2: 3.0: 2.6: 3.0: 2.8: 3.1: 3.3: 4.6. Elytra elongate and parallel-sided, 1.8x longer than wide; disc with reduced reticulate microsculpture, dense, coarse punctures, arranged into longitudinal rows, with one weak longitudinal ridge between two longitudinal rows of punctures, ridges near suture abbreviated. Tarsomeres I of front legs swollen in males; subparallel in females. Aedeagus (
<figureCitation id="136D2A45FFBB4464FC2EADCBFB88FD52" box="[978,1062,729,755]" captionStart="FIGURE 23" captionStartId="45.[151,250,1932,1957]" captionTargetBox="[163,1420,179,1908]" captionTargetId="figure@45.[151,1436,174,1912]" captionTargetPageId="45" captionText="FIGURE 23. Diagnostic characters of Theopea tsoui sp. nov. 23A. Antenna, male; 23B. Antenna, female; 23C. Aedeagus, dorsal view; 23D. Aedeagus, lateral view; 23E. Aedeagus, ventral view; 23F. Abdominal ventrite VIII; 23G. Gonocoxae; 23H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479655" httpUri="https://zenodo.org/record/3479655/files/figure.png" pageId="44" pageNumber="495">Figs 23</figureCitation>
C23E) slender, 8.4x longer than wide; parallel-sided, but narrowed at middle; apically narrowed, apex with median shallow, narrow notch, apices narrowly rounded; tectum well sclerotized, as wide as aedeagus, very long, about 0.5x as long as aedeagus, apex broadly rounded; straight from apex to middle in lateral view, and then slightly curved, apex narrowly rounded; ventral surface with deep notch from apex to middle, mesally extending and approximate at apical 1/10; triangular sclerites small; internal sac with one median, elongate sclerite, as wide as aedeagus but basally narrowed, with one pair of small hook-like sclerites near apex, with one pair of large apically tapering sclerites near base. Gonocoxae (
<figureCitation id="136D2A45FFBB4464FF5CACC7FEA9FC4E" box="[160,263,981,1007]" captionStart="FIGURE 23" captionStartId="45.[151,250,1932,1957]" captionTargetBox="[163,1420,179,1908]" captionTargetId="figure@45.[151,1436,174,1912]" captionTargetPageId="45" captionText="FIGURE 23. Diagnostic characters of Theopea tsoui sp. nov. 23A. Antenna, male; 23B. Antenna, female; 23C. Aedeagus, dorsal view; 23D. Aedeagus, lateral view; 23E. Aedeagus, ventral view; 23F. Abdominal ventrite VIII; 23G. Gonocoxae; 23H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479655" httpUri="https://zenodo.org/record/3479655/files/figure.png" pageId="44" pageNumber="495">Fig. 23G</figureCitation>
) elongate, widest at apical 1/7, both gonocoxae combined together from basal 1/5 to apical 1/7; apices narrowly rounded, each gonocoxa with eight setae along outer margin from apex to apical 1/6; with one pair of short lateral processes at basal 2/5. Ventrite VIII (
<figureCitation id="136D2A45FFBB4464FD2DAB0FFC9AFB96" box="[721,820,1053,1079]" captionStart="FIGURE 23" captionStartId="45.[151,250,1932,1957]" captionTargetBox="[163,1420,179,1908]" captionTargetId="figure@45.[151,1436,174,1912]" captionTargetPageId="45" captionText="FIGURE 23. Diagnostic characters of Theopea tsoui sp. nov. 23A. Antenna, male; 23B. Antenna, female; 23C. Aedeagus, dorsal view; 23D. Aedeagus, lateral view; 23E. Aedeagus, ventral view; 23F. Abdominal ventrite VIII; 23G. Gonocoxae; 23H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479655" httpUri="https://zenodo.org/record/3479655/files/figure.png" pageId="44" pageNumber="495">Fig. 23F</figureCitation>
) elongate and well sclerotized; disc with several long setae laterally and near apical margin, and with dense, short setae along apical margin; spiculum extremely slender. Receptacle of spermatheca (
<figureCitation id="136D2A45FFBB4464FE2FAB77FD96FBDE" box="[467,568,1125,1151]" captionStart="FIGURE 23" captionStartId="45.[151,250,1932,1957]" captionTargetBox="[163,1420,179,1908]" captionTargetId="figure@45.[151,1436,174,1912]" captionTargetPageId="45" captionText="FIGURE 23. Diagnostic characters of Theopea tsoui sp. nov. 23A. Antenna, male; 23B. Antenna, female; 23C. Aedeagus, dorsal view; 23D. Aedeagus, lateral view; 23E. Aedeagus, ventral view; 23F. Abdominal ventrite VIII; 23G. Gonocoxae; 23H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479655" httpUri="https://zenodo.org/record/3479655/files/figure.png" pageId="44" pageNumber="495">Fig. 23H</figureCitation>
) strongly swollen; pump slender and strongly curved; proximal spermathecal duct deeply inserted into receptacle, narrow and short.
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFFBB4464FF3BABBFFC2CFA01" pageId="44" pageNumber="495" type="diagnosis">
<paragraph id="8BE936C0FFBB4464FF3BABBFFC2CFA01" blockId="44.[151,1437,261,1548]" pageId="44" pageNumber="495">
<emphasis id="B922EAD2FFBB4464FF3BABBFFEECFB69" bold="true" box="[199,322,1197,1224]" pageId="44" pageNumber="495">Diagnosis.</emphasis>
Specimens of
<taxonomicName id="4C564D43FFBB4464FE15ABBCFD95FB66" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[489,571,1198,1223]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="tsoui" status="sp. nov.">
<emphasis id="B922EAD2FFBB4464FE15ABBCFD95FB66" box="[489,571,1198,1223]" italics="true" pageId="44" pageNumber="495">T. tsoui</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFBB4464FDBFABBCFD33FB69" bold="true" box="[579,669,1198,1224]" pageId="44" pageNumber="495">
<taxonomicNameLabel id="A21157A9FFBB4464FDBFABBCFD33FB69" box="[579,669,1198,1224]" pageId="44" pageNumber="495" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are similar to those of
<taxonomicName id="4C564D43FFBB4464FC5FABBCFB92FB66" box="[931,1084,1197,1223]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="kedenburgi">
<emphasis id="B922EAD2FFBB4464FC5FABBCFB92FB66" box="[931,1084,1197,1223]" italics="true" pageId="44" pageNumber="495">T. kedenburgi</emphasis>
</taxonomicName>
with only two apical white antennomeres, lateral expansion only on antennomere X in males, and reduced ridges near suture on the elytra, but they differ from those of
<taxonomicName id="4C564D43FFBB4464FE53ABE4FDE9FAAE" box="[431,583,1269,1295]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="kedenburgi">
<emphasis id="B922EAD2FFBB4464FE53ABE4FDE9FAAE" box="[431,583,1269,1295]" italics="true" pageId="44" pageNumber="495">T. kedenburgi</emphasis>
</taxonomicName>
by the strongly swollen antennomeres VIIIX in males (moderately swollen in
<taxonomicName id="4C564D43FFBB4464FF48AA08FEE3FA92" box="[180,333,1305,1331]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="kedenburgi">
<emphasis id="B922EAD2FFBB4464FF48AA08FEE3FA92" box="[180,333,1305,1331]" italics="true" pageId="44" pageNumber="495">T. kedenburgi</emphasis>
</taxonomicName>
). Females of
<taxonomicName id="4C564D43FFBB4464FE1BAA08FD96FA92" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[487,568,1306,1331]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="tsoui">
<emphasis id="B922EAD2FFBB4464FE1BAA08FD96FA92" box="[487,568,1306,1331]" italics="true" pageId="44" pageNumber="495">T. tsoui</emphasis>
</taxonomicName>
possess slightly swollen antennomeres VIVIII (no swollen antennomeres in females of
<taxonomicName id="4C564D43FFBB4464FEEFAA2CFE05FAF6" box="[275,427,1341,1367]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="kedenburgi">
<emphasis id="B922EAD2FFBB4464FEEFAA2CFE05FAF6" box="[275,427,1341,1367]" italics="true" pageId="44" pageNumber="495">T. kedenburgi</emphasis>
</taxonomicName>
). The aedeagi of males of
<taxonomicName id="4C564D43FFBB4464FD2FAA2CFC8BFAF6" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[723,805,1342,1367]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="tsoui" status="sp. nov.">
<emphasis id="B922EAD2FFBB4464FD2FAA2CFC8BFAF6" box="[723,805,1342,1367]" italics="true" pageId="44" pageNumber="495">T. tsoui</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FFBB4464FCD0AA2CFC2EFAF9" bold="true" box="[812,896,1342,1368]" pageId="44" pageNumber="495">
<taxonomicNameLabel id="A21157A9FFBB4464FCD0AA2CFC2EFAF9" box="[812,896,1342,1368]" pageId="44" pageNumber="495" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
can be separated from those of
<taxonomicName id="4C564D43FFBB4464FB1EAA2CFAD4FAF6" box="[1250,1402,1341,1367]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="kedenburgi">
<emphasis id="B922EAD2FFBB4464FB1EAA2CFAD4FAF6" box="[1250,1402,1341,1367]" italics="true" pageId="44" pageNumber="495">T. kedenburgi</emphasis>
</taxonomicName>
by the elongate tectum, 0.5x as long as aedeagus (short tectum in
<taxonomicName id="4C564D43FFBB4464FCABAA70FC5EFADA" box="[855,1008,1377,1403]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="kedenburgi">
<emphasis id="B922EAD2FFBB4464FCABAA70FC5EFADA" box="[855,1008,1377,1403]" italics="true" pageId="44" pageNumber="495">T. kedenburgi</emphasis>
</taxonomicName>
, 0.2x as long as aedeagus), and medially narrowed aedeagus (parallel-sided aedeagus in
<taxonomicName id="4C564D43FFBB4464FD26AA94FCDCFA3E" box="[730,882,1413,1439]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="495" phylum="Arthropoda" rank="species" species="kedenburgi">
<emphasis id="B922EAD2FFBB4464FD26AA94FCDCFA3E" box="[730,882,1413,1439]" italics="true" pageId="44" pageNumber="495">T. kedenburgi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFFBB4464FF3BAAB8FE0BFA46" pageId="44" pageNumber="495" type="etymology">
<paragraph id="8BE936C0FFBB4464FF3BAAB8FE0BFA46" blockId="44.[151,1437,261,1548]" pageId="44" pageNumber="495">
<emphasis id="B922EAD2FFBB4464FF3BAAB8FEE0FA65" bold="true" box="[199,334,1450,1476]" pageId="44" pageNumber="495">Etymology.</emphasis>
This new species is named after Mr. Mei-Hua Tsou, a member of TCRT for his contribution to diversity of leaf beetles.
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFFBB4464FF3BAAE3FE15F9AA" box="[199,443,1521,1548]" pageId="44" pageNumber="495" type="distribution">
<paragraph id="8BE936C0FFBB4464FF3BAAE3FE15F9AA" blockId="44.[151,1437,261,1548]" box="[199,443,1521,1548]" pageId="44" pageNumber="495">
<emphasis id="B922EAD2FFBB4464FF3BAAE3FECFF9AD" bold="true" box="[199,353,1521,1548]" pageId="44" pageNumber="495">Distribution.</emphasis>
<collectingCountry id="F3417650FFBB4464FE9BAAE3FE19F9AA" box="[359,439,1521,1547]" name="Brunei" pageId="44" pageNumber="495">Brunei</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>