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<document id="99384CECB77893C5ECFFBEFEEF3439FB" ID-CLB-Dataset="29146" ID-DOI="10.11646/zootaxa.4422.2.6" ID-GBIF-Dataset="47816d99-6645-43ad-a52d-33cc68a6a7c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1455619" ID-ZooBank="F6658C2B-9681-430A-8975-7B3AE2C233EE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539185629917" checkinUser="plazi" docAuthor="Shear, William A." docDate="2018" docId="03FF87B7FFC2FF8641EAFC5C1FF3F8D7" docLanguage="en" docName="zootaxa.4422.2.6.pdf" docOrigin="Zootaxa 4422 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anopsobiella dawidoffi Attems 1938" docType="treatment" docVersion="7" lastPageNumber="279" masterDocId="FFC6FFCFFFD6FF92417DFFAA1B3DFFD7" masterDocTitle="The centipede family Anopsobiidae new to North America, with the description of a new genus and species and notes on the Henicopidae of North America and the Anopsobiidae of the Northern Hemisphere (Chilopoda, Lithobiomorpha)" masterLastPageNumber="283" masterPageNumber="259" pageNumber="279" updateTime="1698573810455" updateUser="ExternalLinkService">
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<mods:title id="CF0F788ABE3C2F8C62056628610DA257">The centipede family Anopsobiidae new to North America, with the description of a new genus and species and notes on the Henicopidae of North America and the Anopsobiidae of the Northern Hemisphere (Chilopoda, Lithobiomorpha)</mods:title>
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<mods:namePart id="386839AE1998B679F732AA7F28077354">Shear, William A.</mods:namePart>
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<mods:title id="5C627456D83B515D5E2AD08AEC1A51A1">Zootaxa</mods:title>
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<mods:date id="CA7761ACCA9AF3BB0D5E120637A75D46">2018</mods:date>
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<mods:number id="D9D84494A24FF9BE3DD8790F4899F846">4422</mods:number>
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<mods:identifier id="2E5D7E9EA3AEE68C8AB5F44BBAC61938" type="DOI">10.11646/zootaxa.4422.2.6</mods:identifier>
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<treatment id="03FF87B7FFC2FF8641EAFC5C1FF3F8D7" ID-DOI="http://doi.org/10.5281/zenodo.5969839" ID-GBIF-Taxon="148671750" ID-Zenodo-Dep="5969839" LSID="urn:lsid:plazi:treatment:03FF87B7FFC2FF8641EAFC5C1FF3F8D7" httpUri="http://treatment.plazi.org/id/03FF87B7FFC2FF8641EAFC5C1FF3F8D7" lastPageNumber="279" pageId="20" pageNumber="279">
<subSubSection id="C34C652AFFC2FF8641EAFC5C1965FBC7" box="[151,600,1014,1040]" pageId="20" pageNumber="279" type="nomenclature">
<paragraph id="8BE936A1FFC2FF8641EAFC5C1965FBC7" blockId="20.[151,600,1014,1040]" box="[151,600,1014,1040]" pageId="20" pageNumber="279">
<heading id="D0A181CDFFC2FF8641EAFC5C1965FBC7" bold="true" box="[151,600,1014,1040]" fontSize="11" level="1" pageId="20" pageNumber="279" reason="1">
<taxonomicName id="4C564D22FFC2FF8641EAFC5C1965FBC7" authority="Attems, 1938" authorityName="Attems" authorityYear="1938" box="[151,600,1014,1040]" class="Chilopoda" family="Henicopidae" genus="Anopsobiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="20" pageNumber="279" phylum="Arthropoda" rank="species" species="dawidoffi">
<emphasis id="B922EAB3FFC2FF8641EAFC5C1965FBC7" bold="true" box="[151,600,1014,1040]" pageId="20" pageNumber="279">
<emphasis id="B922EAB3FFC2FF8641EAFC5C1A90FBC7" bold="true" box="[151,429,1014,1040]" italics="true" pageId="20" pageNumber="279">Anopsobiella dawidoffi</emphasis>
<bibRefCitation id="EFC74B50FFC2FF8640C8FC5C1965FBC7" box="[437,600,1014,1040]" pageId="20" pageNumber="279" refId="ref15487" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in franzosisch Indochina gesammelten Myriopoden. Memoires du Museum d'Histoire Naturelle, Paris, N. S., 6, 187 - 353." type="journal article">Attems, 1938</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C34C652AFFC2FF8641EAFB971FF3F8D7" pageId="20" pageNumber="279" type="reference_group">
<paragraph id="8BE936A1FFC2FF8641EAFB97188CFB84" blockId="20.[151,945,1085,1107]" box="[151,945,1085,1107]" pageId="20" pageNumber="279">
<treatmentCitationGroup id="AB46118FFFC2FF8641EAFB97188CFB84" box="[151,945,1085,1107]" pageId="20" pageNumber="279">
<taxonomicName id="4C564D22FFC2FF8641EAFB971907FB84" authority="Attems, 1938: 35" authorityName="Attems" authorityPageNumber="35" authorityYear="1938" box="[151,570,1085,1107]" class="Chilopoda" family="Henicopidae" genus="Anopsobiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="20" pageNumber="279" phylum="Arthropoda" rank="species" species="dawidoffi">
<emphasis id="B922EAB3FFC2FF8641EAFB971ABEFB84" box="[151,387,1085,1107]" italics="true" pageId="20" pageNumber="279">Anopsobiella dawidoffi</emphasis>
<treatmentCitation id="0AF710B0FFC2FF8640F4FB941907FB84" author="" box="[393,570,1085,1107]" page="35" pageId="20" pageNumber="279">
<bibRefCitation id="EFC74B50FFC2FF8640F4FB941929FB84" box="[393,532,1085,1107]" pageId="20" pageNumber="279" refId="ref15487" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in franzosisch Indochina gesammelten Myriopoden. Memoires du Museum d'Histoire Naturelle, Paris, N. S., 6, 187 - 353." type="journal article">Attems, 1938</bibRefCitation>
: 35
</treatmentCitation>
</taxonomicName>
; Zapparoli &amp;
<treatmentCitation id="0AF710B0FFC2FF8643B5FB97188CFB84" author="" box="[712,945,1085,1107]" page="377" pageId="20" pageNumber="279">
<bibRefCitation id="EFC74B50FFC2FF8643B5FB9718BCFB84" box="[712,897,1085,1107]" pageId="20" pageNumber="279" refId="ref17423" refString="Zapporoli, M. &amp; Edgecombe, G. D. (2011) Order Lithobiomorpha. In: Minelli, S. (Ed.), Treatise on Zoology, the Myriapoda. Vol. 1. Brill, Leiden, pp. 371 - 389." type="book">Edgecombe, 2011</bibRefCitation>
: 377
</treatmentCitation>
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</paragraph>
<paragraph id="8BE936A1FFC2FF8641EAFBD51FF3F8D7" blockId="20.[151,1437,1151,1792]" pageId="20" pageNumber="279">
The genus
<taxonomicName id="4C564D22FFC2FF86406EFB2A1A9AFB4F" ID-CoL="62BL9" authorityName="Attems" authorityYear="1938" box="[275,423,1152,1176]" class="Chilopoda" family="Henicopidae" genus="Anopsobiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="20" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAB3FFC2FF86406EFB2A1A9AFB4F" box="[275,423,1152,1176]" italics="true" pageId="20" pageNumber="279">Anopsobiella</emphasis>
</taxonomicName>
was originally described as a subgenus of
<emphasis id="B922EAB3FFC2FF8642F7FB2A1F2EFB4F" box="[906,1043,1152,1176]" italics="true" pageId="20" pageNumber="279">
<taxonomicName id="4C564D22FFC2FF8642F7FB2A1F32FB4F" ID-CoL="X2F" authorityName="Silvestri" authorityYear="1899" box="[906,1039,1152,1176]" class="Chilopoda" family="Henicopidae" genus="Anopsobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="20" pageNumber="279" phylum="Arthropoda" rank="genus">Anopsobius</taxonomicName>
,
</emphasis>
which given the unique characters of the single included species, seems a remarkable decision on Attems’ part. The single species was described from an unspecified number of specimens collected at Caùda, Nha Trang,
<collectingRegion id="4992F843FFC2FF8642E8FB6D1FBFFB37" box="[917,1154,1223,1248]" country="Vietnam" name="Khanh Hoa" pageId="20" pageNumber="279">
Khanh
<collectingRegion id="4992F843FFC2FF86429BFB621FBFFB37" box="[998,1154,1223,1248]" inferredBySuffix="true" pageId="20" pageNumber="279">Hoa Province</collectingRegion>
</collectingRegion>
,
<collectingCountry id="F3417631FFC2FF8645F1FB6D1FD2FB37" box="[1164,1263,1223,1248]" name="Vietnam" pageId="20" pageNumber="279">Vietnam</collectingCountry>
. Nha Trang is a resort city on the coast in southern
<collectingCountry id="F3417631FFC2FF864366FB461940FAD2" box="[539,637,1260,1285]" name="Vietnam" pageId="20" pageNumber="279">Vietnam</collectingCountry>
, and Caùda is a headland projecting into Nha Trang Bay. If
<taxonomicName id="4C564D22FFC2FF864469FB461EA1FAD3" authorityName="Attems" authorityYear="1938" box="[1300,1436,1260,1284]" class="Chilopoda" family="Henicopidae" genus="Anopsobiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="20" pageNumber="279" phylum="Arthropoda" rank="species" species="dawidoffi">
<emphasis id="B922EAB3FFC2FF864469FB461EA1FAD3" box="[1300,1436,1260,1284]" italics="true" pageId="20" pageNumber="279">A. dawidoffi</emphasis>
</taxonomicName>
is in fact an anopsobiid, it is strikingly different from the other members of the family (and all
<taxonomicName id="4C564D22FFC2FF8645CCFAA51E02FAFF" ID-CoL="6254Q" authorityName="Pocock" authorityYear="1901" box="[1201,1343,1295,1320]" class="Chilopoda" family="Henicopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="20" pageNumber="279" phylum="Arthropoda" rank="family">Henicopidae</taxonomicName>
as well) in having very strongly modified penultimate (14th) and ultimate (15th) legs in the male. Both these leg-pairs are shown in Attems’ drawings (his Figs. 319 and 320) as thickened and with the tibiae considerably swollen; in the penultimate leg, the distal tarsal article is attached to a swollen proximal article on its ventral side—about ¼ the length of the first article from its termination—but in the ultimate leg the bisegmented tarsus is attached in this manner to the tibia (or at least that is what the drawings seem to show). The same drawings also show that there are no distal spines on the coxae of the last pair of legs.
<bibRefCitation id="EFC74B50FFC2FF864394FA411F13F9D4" box="[745,1070,1515,1539]" pageId="20" pageNumber="279" refId="ref17059" refString="Prunescu, C. - C. &amp; Prunescu, P. (2004) The male genital system of Anopsobiella dawydoffi Attems, 1938 (Lithobiomorpha, Chilopoda). Annalen des Naturhistorischen Museums in Wien, 105 B, 59 - 65." type="journal article">Prunescu &amp; Prunescu (2004)</bibRefCitation>
also provided drawings of these legs that are perhaps a bit more accurate, but which omit the coxae.
<bibRefCitation id="EFC74B50FFC2FF8642D5F9A51F71F9F0" box="[936,1100,1551,1575]" pageId="20" pageNumber="279" refId="ref15487" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in franzosisch Indochina gesammelten Myriopoden. Memoires du Museum d'Histoire Naturelle, Paris, N. S., 6, 187 - 353." type="journal article">Attems (1938)</bibRefCitation>
states that spiracles occur on pedigerous segments 4, 6, 9, 11 and 13, but this is surely an error, which Zapparoli &amp;
<bibRefCitation id="EFC74B50FFC2FF86452BF9981E16F99C" box="[1110,1323,1586,1611]" pageId="20" pageNumber="279" refId="ref17423" refString="Zapporoli, M. &amp; Edgecombe, G. D. (2011) Order Lithobiomorpha. In: Minelli, S. (Ed.), Treatise on Zoology, the Myriapoda. Vol. 1. Brill, Leiden, pp. 371 - 389." type="book">Edgecombe (2011)</bibRefCitation>
correct to 3, 5, 8, 10, and 14. Single ventrodistal spurs are present on the prefemora, femora, and tibiae of leg-pairs 9–13, and tibial spines occur on legpairs 1–13. Given these departures from the usual characters of
<taxonomicName id="4C564D22FFC2FF8645E9F9D01E0EF944" authorityName="Verhoeff" authorityYear="1907" box="[1172,1331,1658,1683]" class="Chilopoda" family="Anopsobiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiida" pageId="20" pageNumber="279" phylum="Arthropoda" rank="family">Anopsobiidae</taxonomicName>
, it seems likely that
<taxonomicName id="4C564D22FFC2FF864072F9351AA4F960" authorityName="Attems" authorityYear="1938" box="[271,409,1695,1719]" class="Chilopoda" family="Henicopidae" genus="Anopsobiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="20" pageNumber="279" phylum="Arthropoda" rank="species" species="dawidoffi">
<emphasis id="B922EAB3FFC2FF864072F9351AA4F960" box="[271,409,1695,1719]" italics="true" pageId="20" pageNumber="279">A. dawidoffi</emphasis>
</taxonomicName>
is not a member of the family. The
<typeStatus id="54ED8803FFC2FF864253F90A1863F96F" box="[814,862,1696,1720]" pageId="20" pageNumber="279">type</typeStatus>
specimens of
<taxonomicName id="4C564D22FFC2FF86457DF9351FB4F960" authorityName="Attems" authorityYear="1938" box="[1024,1161,1695,1719]" class="Chilopoda" family="Henicopidae" genus="Anopsobiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="20" pageNumber="279" phylum="Arthropoda" rank="species" species="dawidoffi">
<emphasis id="B922EAB3FFC2FF86457DF9351FB4F960" box="[1024,1161,1695,1719]" italics="true" pageId="20" pageNumber="279">A. dawidoffi</emphasis>
</taxonomicName>
can no longer be found, except for one very immature example (N. Akkari, pers. comm. 2018).
<bibRefCitation id="EFC74B50FFC2FF8642B9F9691E36F90C" box="[964,1291,1731,1755]" pageId="20" pageNumber="279" refId="ref17059" refString="Prunescu, C. - C. &amp; Prunescu, P. (2004) The male genital system of Anopsobiella dawydoffi Attems, 1938 (Lithobiomorpha, Chilopoda). Annalen des Naturhistorischen Museums in Wien, 105 B, 59 - 65." type="journal article">Prunescu &amp; Prunescu (2004)</bibRefCitation>
destroyed at least one of the male
<typeStatus id="54ED8803FFC2FF8640FBF9421AFCF8D7" box="[390,449,1768,1792]" pageId="20" pageNumber="279">types</typeStatus>
(NHMW 3815) in their histological study of the male genital system.
</paragraph>
</subSubSection>
</treatment>
</document>
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