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<document id="7A7C864DC63E8B26186337856B85A2AA" ID-DOI="10.11646/zootaxa.5505.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745912" ID-ZooBank="93F5E9F6-9EDC-4ED2-8F2B-1E6D0C18F893" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726052234810" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03E7FF50FFD8502AFF23F946FE04409A" docLanguage="en" docName="zootaxa.5505.1.1.pdf" docOrigin="Zootaxa 5505 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5505.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Anhoraeomorphus moheliensis Jałoszyński 2024, comb. n." docType="treatment" docVersion="2" lastPageNumber="43" masterDocId="FFDE8728FFF05000FFB4FFCCFFC54473" masterDocTitle="The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="41" updateTime="1726053143081" updateUser="ExternalLinkService">
<mods:mods id="0164723C1CE191507BC68C3D86C2F7A9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="66DC41B9DCF0F0469644F652B5B5F6CD">The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="52571CEE4C7844CD53D3768B8AE05C35">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="0DE87AED13E050C649974B146EA5D921">2024</mods:date>
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<mods:number id="07BAE618469BDD1A663E8AB24C0E371F">2024-09-04</mods:number>
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<mods:number id="ABF56669DDFAFBBF46B6365E55B3ABBD">5505</mods:number>
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<mods:classification id="095AB97DCE515CDA0320F2E1B11DF535">journal article</mods:classification>
<mods:identifier id="5F7906C12EF50486F90271FCBD46A6B5" type="DOI">10.11646/zootaxa.5505.1.1</mods:identifier>
<mods:identifier id="2C9977C2FBBD9C7DC5FEBD5C127DD9CA" type="ISSN">1175-5334</mods:identifier>
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<treatment id="03E7FF50FFD8502AFF23F946FE04409A" ID-DOI="http://doi.org/10.5281/zenodo.13746071" ID-Zenodo-Dep="13746071" LSID="urn:lsid:plazi:treatment:03E7FF50FFD8502AFF23F946FE04409A" httpUri="http://treatment.plazi.org/id/03E7FF50FFD8502AFF23F946FE04409A" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
<subSubSection id="C3541DCDFFD85028FF23F946FD3742D6" box="[151,754,1674,1701]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="8BF14E46FFD85028FF23F946FD3742D6" blockId="40.[151,754,1674,1737]" box="[151,754,1674,1701]" pageId="40" pageNumber="41">
<heading id="D0B9F92AFFD85028FF23F946FD3742D6" bold="true" box="[151,754,1674,1701]" fontSize="11" level="2" pageId="40" pageNumber="41" reason="4">
<emphasis id="B93A9254FFD85028FF23F946FD3742D6" bold="true" box="[151,754,1674,1701]" pageId="40" pageNumber="41">
<taxonomicName id="4C4E35C5FFD85028FF23F946FDBC42D7" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,633,1674,1701]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="moheliensis" status="comb. nov.">
<emphasis id="B93A9254FFD85028FF23F946FDD742D6" bold="true" box="[151,530,1674,1701]" italics="true" pageId="40" pageNumber="41">Anhoraeomorphus moheliensis</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFD85028FD33F947FD3742D6" box="[647,754,1675,1701]" pageId="40" pageNumber="41" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD85028FF23F962FE8642BA" box="[151,323,1710,1737]" pageId="40" pageNumber="41" type="description">
<paragraph id="8BF14E46FFD85028FF23F962FE8642BA" blockId="40.[151,754,1674,1737]" box="[151,323,1710,1737]" pageId="40" pageNumber="41">
(
<figureCitation id="137552C3FFD85028FF2BF962FEFF42BA" box="[159,314,1710,1737]" captionStart="FIGURES 151–158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151–158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="40" pageNumber="41">Figs 151–158</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD85028FF23F920FD474377" box="[151,642,1772,1796]" pageId="40" pageNumber="41" type="reference_group">
<paragraph id="8BF14E46FFD85028FF23F920FD474377" blockId="40.[151,642,1772,1796]" box="[151,642,1772,1796]" pageId="40" pageNumber="41">
<treatmentCitationGroup id="AB5E6968FFD85028FF23F920FD474377" box="[151,642,1772,1796]" pageId="40" pageNumber="41">
<treatmentCitation id="0AEF6857FFD85028FF23F920FDBB4377" author="Franz, H." box="[151,638,1772,1796]" page="159" pageId="40" pageNumber="41" year="1986">
<taxonomicName id="4C4E35C5FFD85028FF23F920FDBB4377" ID-CoL="3MNKT" authority="Franz, 1986 b: 159" authorityName="Franz" authorityPageNumber="159" authorityYear="1986" box="[151,638,1772,1796]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="moheliensis">
<emphasis id="B93A9254FFD85028FF23F920FE794377" box="[151,444,1772,1796]" italics="true" pageId="40" pageNumber="41">Horaeomorphus moheliensis</emphasis>
<bibRefCitation id="EFDF33B7FFD85028FE76F921FDBB4377" author="Franz, H." box="[450,638,1772,1796]" pageId="40" pageNumber="41" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 159</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD85029FF23F8E0FDFC43C0" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="8BF14E46FFD85028FF23F8E0FE664389" blockId="40.[151,1436,1835,2042]" pageId="40" pageNumber="41">
<emphasis id="B93A9254FFD85028FF23F8E0FE614335" bold="true" box="[151,420,1836,1862]" pageId="40" pageNumber="41">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFD85028FE1FF8E0FE664389" collectionCode="NHMW" collectorName="G. Gomy" country="Comoros" county="Insel" determinerName="H. Franz" elevation="420" location="Typus" municipality="Moihani" pageId="40" pageNumber="41" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Moheli" typeStatus="holotype">
<emphasis id="B93A9254FFD85028FE1FF8E0FC9A4336" bold="true" box="[427,863,1835,1862]" pageId="40" pageNumber="41">
<typeStatus id="54F5F0E4FFD85028FE1FF8E0FDDD4335" box="[427,536,1836,1862]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFD85028FD93F8E7FD5C4336" box="[551,665,1835,1861]" name="Comoros" pageId="40" pageNumber="41">Comoros</collectingCountry>
:
<collectingRegion id="498A80A4FFD85028FD12F8E7FC974336" box="[678,850,1835,1862]" country="Comoros" name="Moheli" pageId="40" pageNumber="41">Mohéli Island</collectingRegion>
):
</emphasis>
<specimenCount id="9D4885CFFFD85028FCD2F8E0FCB84335" box="[870,893,1836,1862]" count="1" pageId="40" pageNumber="41" type="male">♂</specimenCount>
, four labels: “(
<collectingCounty id="629036CAFFD85028FB97F8E0FB984335" box="[1059,1117,1836,1862]" pageId="40" pageNumber="41">Insel</collectingCounty>
<collectingRegion id="498A80A4FFD85028FBD0F8E0FB704335" box="[1124,1205,1836,1862]" country="Comoros" name="Moheli" pageId="40" pageNumber="41">Moheli</collectingRegion>
/
<collectingMunicipality id="6B95D43CFFD85028FB78F8E0FAEE4335" box="[1228,1323,1836,1862]" pageId="40" pageNumber="41">Moihani</collectingMunicipality>
/ 20.8.69,
<collectorName id="26BB2B90FFD85028FF23F89CFEC04319" box="[151,261,1872,1898]" pageId="40" pageNumber="41">G. Gomy</collectorName>
” [white, handwritten], “
<taxonomicName id="4C4E35C5FFD85028FDABF89CFD134319" box="[543,726,1872,1898]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ moheliensis m. / det.
<determinerName id="7E2DA282FFD85028FC5CF89CFB834319" box="[1000,1094,1872,1898]" pageId="40" pageNumber="41">H.Franz</determinerName>
” [yellowish, handwritten and printed], “
<location id="8E91189DFFD85028FEBFF8B8FE9143FD" LSID="urn:lsid:plazi:treatment:03E7FF50FFD8502AFF23F946FE04409A:8E91189DFFD85028FEBFF8B8FE9143FD" box="[267,340,1908,1934]" country="Comoros" county="Insel" municipality="Moihani" name="Typus" pageId="40" pageNumber="41" stateProvince="Moheli">Typus</location>
” [red, handwritten], “
<specimenCount id="9D4885CFFFD85028FDE5F8B8FDA243FD" box="[593,615,1908,1934]" count="1" pageId="40" pageNumber="41" type="male">♂</specimenCount>
” [white, printed] (
<collectionCode id="ED5FD683FFD85028FC8BF8B8FC6743FD" box="[831,930,1908,1934]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="41" type="Museum">NHMW</collectionCode>
).
<emphasis id="B93A9254FFD85028FC02F8B8FBE843FD" bold="true" box="[950,1069,1908,1934]" pageId="40" pageNumber="41">
<typeStatus id="54F5F0E4FFD85028FC02F8B8FBE243FD" box="[950,1063,1908,1934]" pageId="40" pageNumber="41" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D4885CFFFD85028FB82F8B8FB8E43FD" box="[1078,1099,1908,1934]" count="1" pageId="40" pageNumber="41" type="female">♀</specimenCount>
, “
<collectingRegion id="498A80A4FFD85028FBD2F8B8FB7C43FD" box="[1126,1209,1908,1934]" country="Comoros" name="Moheli" pageId="40" pageNumber="41">Moheli</collectingRegion>
/
<location id="8E91189DFFD85028FB61F8B8FA9643FD" LSID="urn:lsid:plazi:treatment:03E7FF50FFD8502AFF23F946FE04409A:8E91189DFFD85028FB61F8B8FA9643FD" box="[1237,1363,1908,1934]" country="Comoros" county="Insel" municipality="Moihani" name="Bandamale" pageId="40" pageNumber="41" stateProvince="Moheli">Bandamalé</location>
<quantity id="4CB6E3A3FFD85028FAE8F8B8FA5943FD" box="[1372,1436,1908,1934]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="40" pageNumber="41" unit="m" value="420.0">
<elevation id="0063A975FFD85028FAE8F8B8FA5943FD" box="[1372,1436,1908,1934]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="40" pageNumber="41" unit="m" value="420.0">420m</elevation>
</quantity>
/ IX-58 Raharizonina” [white, printed], “INSTITUT / SCIENTIFIQUE / MADAGASCAR” [bluish, printed], “
<specimenCount id="9D4885CFFFD85028FF11F870FF7243A5" box="[165,183,1980,2006]" count="1" pageId="40" pageNumber="41" type="female">♀</specimenCount>
”, “
<taxonomicName id="4C4E35C5FFD85028FF55F870FE5D43A5" box="[225,408,1980,2006]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ moheliensis m. / det.
<determinerName id="7E2DA282FFD85028FD1FF870FCCD43A5" box="[683,776,1980,2006]" pageId="40" pageNumber="41">H.Franz</determinerName>
” [yellowish, handwritten and printed], “
<typeStatus id="54F5F0E4FFD85028FB54F870FA9643A5" box="[1248,1363,1980,2006]" pageId="40" pageNumber="41" type="allotype">Allotypus</typeStatus>
” [red, handwritten] (
<collectionCode id="ED5FD683FFD85028FE81F82CFE5D4389" box="[309,408,2016,2042]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="41" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="DF311ECEFFD95029FF23FA26FE7F4239" ID-DOI="http://doi.org/10.5281/zenodo.13745975" ID-Zenodo-Dep="13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,264,1514,1538]" targetBox="[151,1416,181,1476]" targetPageId="41" targetType="figure">
<paragraph id="8BF14E46FFD95029FF23FA26FE7F4239" blockId="41.[151,1436,1514,1610]" pageId="41" pageNumber="42">
<emphasis id="B93A9254FFD95029FF23FA26FEA84271" bold="true" box="[151,365,1514,1538]" pageId="41" pageNumber="42">FIGURES 151–158.</emphasis>
<taxonomicName id="4C4E35C5FFD95029FEC6FA26FD3A4271" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[370,767,1514,1538]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="moheliensis">
<emphasis id="B93A9254FFD95029FEC6FA26FD694271" box="[370,684,1514,1538]" italics="true" pageId="41" pageNumber="42">Anhoraeomorphus moheliensis</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFD95029FBEAFA26FB414271" bold="true" box="[1118,1156,1514,1538]" pageId="41" pageNumber="42">151</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFD95029FF14F9C2FF034255" bold="true" box="[160,198,1550,1574]" pageId="41" pageNumber="42">152</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFD95029FC23F9C2FC784255" bold="true" box="[919,957,1550,1574]" pageId="41" pageNumber="42">153</emphasis>
); antenna (
<emphasis id="B93A9254FFD95029FB9BF9C2FB904255" bold="true" box="[1071,1109,1550,1574]" pageId="41" pageNumber="42">154</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFD95029FA88F9C2FA514255" bold="true" box="[1340,1428,1550,1574]" pageId="41" pageNumber="42">155, 157</emphasis>
) and lateral (
<emphasis id="B93A9254FFD95029FEA4F9FEFEAC4239" bold="true" box="[272,361,1586,1610]" pageId="41" pageNumber="42">156, 158</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFD95029FF73F950FDFC43C0" blockId="41.[151,1437,1692,2043]" pageId="41" pageNumber="42">
<emphasis id="B93A9254FFD95029FF73F950FE6042C5" bold="true" box="[199,421,1692,1718]" pageId="41" pageNumber="42">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFD95029FDB5F950FDA142C4" box="[513,612,1692,1719]" captionStart="FIGURES 151–158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151–158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="41" pageNumber="42">Fig. 151</figureCitation>
) slender; head (
<figureCitation id="137552C3FFD95029FC90F950FC0142C4" box="[804,964,1692,1719]" captionStart="FIGURES 151–158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151–158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="41" pageNumber="42">Figs 152–153</figureCitation>
) sub-pentagonal, with vertex posteriorly weakly rounded; pronotum (
<figureCitation id="137552C3FFD95029FE55F90CFD8142A8" box="[481,580,1728,1755]" captionStart="FIGURES 151–158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151–158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="41" pageNumber="42">Fig. 151</figureCitation>
) bell-shaped, broadest near anterior third, with one pair of small circular antebasal pits; antennomeres 3–5 (
<figureCitation id="137552C3FFD95029FDADF928FDBC428C" box="[537,633,1764,1791]" captionStart="FIGURES 151–158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151–158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="41" pageNumber="42">Fig. 154</figureCitation>
) each weakly elongate, 6–9 each about as long as broad, 10 indistinctly transverse; profemora slightly thicker than meso- and metafemora; in both sexes pro- and mesotibiae strongly and evenly curved, in males metatibiae curved in proximal third, in female metatibiae evenly curved; aedeagus (
<figureCitation id="137552C3FFD95029FADEF8E0FF3E4318" captionStart="FIGURES 151–158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151–158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="41" pageNumber="42">Figs 155–158</figureCitation>
) in ventral view broadest near middle, apical region of dorsal wall in lateral view weakly curved dorsally, distolateral endophallic sclerites broadly separated, each elongate and asymmetrical, with truncate apices, each paramere with two short apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD9502AFF73F870FA9B4046" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph id="8BF14E46FFD95029FF73F870FE784388" blockId="41.[151,1437,1692,2043]" pageId="41" pageNumber="42">
<emphasis id="B93A9254FFD95029FF73F870FEB043A5" bold="true" box="[199,373,1980,2006]" pageId="41" pageNumber="42">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFD95029FD94F870FDBB43A5" box="[544,638,1980,2006]" captionStart="FIGURES 139–146" captionStartId="38.[151,264,1606,1630]" captionTargetBox="[151,1413,181,1565]" captionTargetId="figure-28@38.[151,1435,181,1582]" captionTargetPageId="38" captionText="FIGURES 139–146. Anhoraeomorphus manangotryanus (Franz), holotype. Habitus in dorsal view (139); head in anterodorsal view (140); head and prothorax in lateral view (only bristles on head are shown) (141); antenna (142); aedeagus in ventral (143, 145) and lateral (144, 146) views." figureDoi="http://doi.org/10.5281/zenodo.13745969" httpUri="https://zenodo.org/record/13745969/files/figure.png" pageId="41" pageNumber="42">Fig. 139</figureCitation>
) flattened, moderately dark brown, tarsi and palps slightly lighter; setae light brown; BL
<quantity id="4CB6E3A3FFD95029FEE6F82CFE7C4388" box="[338,441,2016,2043]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="41" pageNumber="42" unit="mm" value="2.45">2.45 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFDA502AFF73FF5BFD4845DE" blockId="42.[151,1437,150,1257]" pageId="42" pageNumber="43">
Head (
<figureCitation id="137552C3FFDA502AFEA0FF5BFEB344C2" box="[276,374,150,177]" captionStart="FIGURES 151–158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151–158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="42" pageNumber="43">Fig. 152</figureCitation>
) in anterodorsal view sub-pentagonal, broadest at eyes, HL
<quantity id="4CB6E3A3FFDA502AFB9FFF5AFB5144C2" box="[1067,1172,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="42" pageNumber="43" unit="mm" value="0.53">0.53 mm</quantity>
, HW
<quantity id="4CB6E3A3FFDA502AFB6DFF5AFA8744C2" box="[1241,1346,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="42" pageNumber="43" unit="mm" value="0.45">0.45 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFDA502AFEE5FF77FE7644A6" box="[337,435,186,213]" captionStart="FIGURES 151–158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151–158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="42" pageNumber="43">Fig. 153</figureCitation>
) nearly twice as long as eye; vertex and frons confluent, together weakly convex and indistinctly transverse; vertex posteriorly weakly rounded; each eye small and weakly convex, oval, not emarginate posteriorly, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFDA502AFB1AFEEBFACA4532" box="[1198,1295,294,321]" captionStart="FIGURES 151–158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151–158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="42" pageNumber="43">Fig. 154</figureCitation>
) slender but short, indistinctly shorter than half BL, AnL
<quantity id="4CB6E3A3FFDA502AFD3CFE87FD284516" box="[648,749,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="42" pageNumber="43" unit="mm" value="1.1">1.10 mm</quantity>
, antennomeres 1–2 each distinctly elongate, 3–5 each weakly elongate, 6–9 each about as long as broad, 10 indistinctly transverse, 11 slightly shorter than 9 and 10 combined, as wide as 10, about 1.6 times as long as broad.
</paragraph>
<paragraph id="8BF14E46FFDA502AFF73FE7BFEEA4612" blockId="42.[151,1437,150,1257]" pageId="42" pageNumber="43">
Pronotum bell-shaped, broadest near anterior third; PL
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, PW
<quantity id="4CB6E3A3FFDA502AFBB6FE7AFBAB45A2" box="[1026,1134,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="42" pageNumber="43" unit="mm" value="0.55">0.55 mm</quantity>
. Anterior margin strongly rounded, lateral margins strongly rounded in anterior half and sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of small subcircular pits, sublateral carinae short but distinct. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFDA502AFF73FDA7FEAE46BE" blockId="42.[151,1437,150,1257]" pageId="42" pageNumber="43">
Elytra together oval, broadest distinctly in front of middle; EL
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, EW
<quantity id="4CB6E3A3FFDA502AFB87FDA6FB5F46F6" box="[1075,1178,618,645]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="42" pageNumber="43" unit="mm" value="0.85">0.85 mm</quantity>
, EI 1.53; humeral calli weakly elevated, basal elytral foveae barely discernible. Punctures small and shallow, inconspicuous; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFDA502AFF73FD1BFBC04766" blockId="42.[151,1437,150,1257]" pageId="42" pageNumber="43">Legs moderately long, slender, fore femora slightly thicker than meso- and metafemora, pro- and mesotibiae strongly and evenly curved, metatibiae strongly curved only in proximal third.</paragraph>
<paragraph id="8BF14E46FFDA502AFF73FCD3FCBD479E" blockId="42.[151,1437,150,1257]" pageId="42" pageNumber="43">
Aedeagus (
<figureCitation id="137552C3FFDA502AFEFCFCD3FE2D474A" box="[328,488,798,825]" captionStart="FIGURES 151–158" captionStartId="41.[151,264,1514,1538]" captionTargetBox="[151,1416,181,1476]" captionTargetId="figure-16@41.[151,1435,181,1489]" captionTargetPageId="41" captionText="FIGURES 151–158. Anhoraeomorphus moheliensis (Franz), holotype. Habitus in dorsal view (151); head in anterodorsal view (152); head and prothorax in lateral view (only bristles on head are shown) (153); antenna (154); aedeagus in ventral (155, 157) and lateral (156, 158) views." figureDoi="http://doi.org/10.5281/zenodo.13745975" httpUri="https://zenodo.org/record/13745975/files/figure.png" pageId="42" pageNumber="43">Figs 155–158</figureCitation>
) stout, AeL
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; in ventral view pear-shaped, broadest near middle, distally rapidly but weakly narrowing towards subtriangular apex with broadly rounded distal margin hidden behind broad and distally broadly rounded ventral apical plate; in lateral view apical region of dorsal wall weakly curved dorsally; endophallus with all sclerotized structures asymmetrical, distolateral sclerites broadly separated, strongly elongate and each with truncate apex; parameres in lateral view conspicuously broadened and their apices not projecting beyond aedeagal apex, each paramere with two short apical setae.
</paragraph>
<paragraph id="8BF14E46FFDA502AFF73FC3BFA9B4046" blockId="42.[151,1437,150,1257]" pageId="42" pageNumber="43">
Female. Externally similar to male, but with all tibiae, including hind ones, evenly curved. BL
<quantity id="4CB6E3A3FFDA502AFAB7FC3BFAAE4062" box="[1283,1387,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="2.33" pageId="42" pageNumber="43" unit="mm" value="2.33">2.33 mm</quantity>
; HL
<quantity id="4CB6E3A3FFDA502AFF23FBD6FF384046" box="[151,253,1050,1077]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="42" pageNumber="43" unit="mm" value="0.5">0.50 mm</quantity>
, HW
<quantity id="4CB6E3A3FFDA502AFE8AFBD6FE614046" box="[318,420,1050,1077]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="42" pageNumber="43" unit="mm" value="0.45">0.45 mm</quantity>
, AnL
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; PL
<quantity id="4CB6E3A3FFDA502AFD34FBD6FD234046" box="[640,742,1050,1077]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="42" pageNumber="43" unit="mm" value="0.55">0.55 mm</quantity>
, PW
<quantity id="4CB6E3A3FFDA502AFC96FBD6FC4C4046" box="[802,905,1050,1077]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="42" pageNumber="43" unit="mm" value="0.58">0.58 mm</quantity>
; EL
<quantity id="4CB6E3A3FFDA502AFC09FBD7FBE64046" box="[957,1059,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="42" pageNumber="43" unit="mm" value="1.28">1.28 mm</quantity>
, EW
<quantity id="4CB6E3A3FFDA502AFBD4FBD6FB034046" box="[1120,1222,1050,1077]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="42" pageNumber="43" unit="mm" value="0.85">0.85 mm</quantity>
, EI
<quantity id="4CB6E3A3FFDA502AFB47FBD6FA9F4046" box="[1267,1370,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="42" pageNumber="43" unit="mm" value="1.5">1.50 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDA502AFF73FBF2FD45402A" box="[199,640,1086,1113]" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8BF14E46FFDA502AFF73FBF2FD45402A" blockId="42.[151,1437,150,1257]" box="[199,640,1086,1113]" pageId="42" pageNumber="43">
<emphasis id="B93A9254FFDA502AFF73FBF2FEA4402B" bold="true" box="[199,353,1086,1112]" pageId="42" pageNumber="43">Distribution.</emphasis>
<collectingRegion id="498A80A4FFDA502AFED3FBF3FDC3402A" box="[359,518,1087,1113]" country="Comoros" name="Moheli" pageId="42" pageNumber="43">Mohéli Island</collectingRegion>
,
<collectingCountry id="F3590ED6FFDA502AFDA6FBF3FDB9402A" box="[530,636,1087,1113]" name="Comoros" pageId="42" pageNumber="43">Comoros</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDA502AFF73FBAFFE04409A" pageId="42" pageNumber="43" type="discussion">
<paragraph id="8BF14E46FFDA502AFF73FBAFFE04409A" blockId="42.[151,1437,150,1257]" pageId="42" pageNumber="43">
<emphasis id="B93A9254FFDA502AFF73FBAFFEFE400E" bold="true" box="[199,315,1123,1149]" pageId="42" pageNumber="43">Remarks.</emphasis>
This small-bodied and slender species is unique in having all tibiae in both sexes strongly curved, with metatibiae in males curved only in proximal third, and in female evenly curved along entire length. Besides the
<typeStatus id="54F5F0E4FFDA502AFF23FB67FF3C40B6" box="[151,249,1195,1221]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
and
<typeStatus id="54F5F0E4FFDA502AFE84FB67FE5540B6" box="[304,400,1195,1221]" pageId="42" pageNumber="43" type="paratype">paratype</typeStatus>
deposited at NHMW, according to
<bibRefCitation id="EFDF33B7FFDA502AFC94FB67FC7A40B6" author="Franz, H." box="[800,959,1195,1221]" pageId="42" pageNumber="43" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
<specimenCount id="9D4885CFFFDA502AFC73FB67FB5140B6" box="[967,1172,1195,1221]" count="1" pageId="42" pageNumber="43" type="male" typeStatus="paratype">one paratype male</specimenCount>
is in Muséum National d'Histoire Naturelle, Paris.
</paragraph>
</subSubSection>
</treatment>
</document>
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