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<document id="A36FE2588B2B5C37EDA94802CAF18BE9" ID-CLB-Dataset="27557" ID-DOI="10.11646/zootaxa.4555.1.5" ID-GBIF-Dataset="892bcc0c-1516-4a00-bf87-6571cc961061" ID-ISSN="1175-5326" ID-Zenodo-Dep="2624152" ID-ZooBank="869A864E-EC9D-4BA5-9932-3F116B79B298" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554272005453" checkinUser="plazi" docAuthor="Kosterin, Oleg E., Garrison, Rosser, Kompier, Tom &amp; Farrell, Dennis" docDate="2019" docId="03E7A6213E702E7702F9D3FBFD49C7FA" docLanguage="en" docName="zootaxa.4555.1.5.pdf" docOrigin="Zootaxa 4555 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Indolestes anomalus Fraser 1946" docType="treatment" docVersion="10" lastPageNumber="78" masterDocId="FFDEDE593E712E7C026ED36CFF8CC60E" masterDocTitle="Taxonomic notes on Indolestes Fraser, 1922 (Lestidae, Zygoptera). 3. Male and clarified type locality of Indolestes anomalus Fraser, 1946" masterLastPageNumber="78" masterPageNumber="67" pageNumber="68" updateTime="1698716367155" updateUser="ExternalLinkService">
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<mods:title id="99E203F9C1773A8DF0D4BFFEAB525C8E">Taxonomic notes on Indolestes Fraser, 1922 (Lestidae, Zygoptera). 3. Male and clarified type locality of Indolestes anomalus Fraser, 1946</mods:title>
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<mods:namePart id="0FE815179FC8A154BC1862A9CE6CF1AB">Kosterin, Oleg E.</mods:namePart>
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<mods:namePart id="CB1E38C80559D7FC491E4363184B9800">Kompier, Tom</mods:namePart>
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<mods:namePart id="2F748B63B5D2C40032FE4EBF9B23C187">Farrell, Dennis</mods:namePart>
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<treatment id="03E7A6213E702E7702F9D3FBFD49C7FA" ID-DOI="http://doi.org/10.5281/zenodo.5522677" ID-GBIF-Taxon="154533202" ID-Zenodo-Dep="5522677" LSID="urn:lsid:plazi:treatment:03E7A6213E702E7702F9D3FBFD49C7FA" httpUri="http://treatment.plazi.org/id/03E7A6213E702E7702F9D3FBFD49C7FA" lastPageId="11" lastPageNumber="78" pageId="1" pageNumber="68">
<subSubSection id="C35444BC3E702E7D02F9D3FBFDBCC6BC" box="[151,560,151,178]" pageId="1" pageNumber="68" type="nomenclature">
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<heading id="D0B9A05B3E702E7D02F9D3FBFDBCC6BC" bold="true" box="[151,560,151,178]" fontSize="11" level="1" pageId="1" pageNumber="68" reason="1">
<taxonomicName id="4C4E6CB43E702E7D02F9D3FBFDBCC6BC" ID-CoL="6MVY3" authority="Fraser, 1946" authorityName="Fraser" authorityYear="1946" box="[151,560,151,178]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E702E7D02F9D3FBFDBCC6BC" bold="true" box="[151,560,151,178]" pageId="1" pageNumber="68">
<emphasis id="B93ACB253E702E7D02F9D3FBFE03C6BF" bold="true" box="[151,399,151,177]" italics="true" pageId="1" pageNumber="68">Indolestes anomalus</emphasis>
<bibRefCitation id="EFDF6AC63E702E7D03F8D3F4FDBCC6BC" author="Fraser, F. C." box="[406,560,152,178]" pageId="1" pageNumber="68" pagination="41 - 45" refId="ref5050" refString="Fraser, F. C. (1946) On rare and two new species of Odonata from tropical Africa and Asia. Proceedings of Royal Entomological Society of London. Series B, 15 (3 / 4), 41 - 45. https: // doi. org / 10.1111 / j. 1365 - 3113.1946. tb 00815. x" type="journal article" year="1946">Fraser, 1946</bibRefCitation>
</emphasis>
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<subSubSection id="C35444BC3E702E7D02F9D3D6FE8CC6DD" box="[151,256,186,211]" pageId="1" pageNumber="68" type="description">
<paragraph id="8BF117373E702E7D02F9D3D6FE8CC6DD" blockId="1.[151,560,151,211]" box="[151,256,186,211]" pageId="1" pageNumber="68">
<figureCitation id="13750BB23E702E7D02F9D3D6FE8CC6DD" box="[151,256,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="2.[151,250,1507,1529]" captionStartId-1="3.[151,250,1674,1696]" captionStartId-2="5.[151,250,1922,1944]" captionStartId-3="6.[151,250,1919,1941]" captionStartId-4="7.[151,250,1941,1963]" captionStartId-5="8.[151,250,1904,1926]" captionStartId-6="9.[151,250,1434,1456]" captionTargetBox-0="[340,1226,424,1423]" captionTargetBox-1="[276,1312,337,1653]" captionTargetBox-2="[151,1435,282,1922]" captionTargetBox-3="[296,1246,481,1879]" captionTargetBox-4="[445,1150,193,1921]" captionTargetBox-5="[287,1278,511,1836]" captionTargetBox-6="[233,1294,242,1333]" captionTargetId-0="figure@2.[340,1247,267,1483]" captionTargetId-1="figure@3.[276,1312,337,1653]" captionTargetId-2="figure@5.[151,1435,282,1900]" captionTargetId-3="figure@6.[288,1298,481,1903]" captionTargetId-4="figure@7.[445,1150,193,1921]" captionTargetId-5="figure@8.[287,1299,471,1887]" captionTargetId-6="figure@9.[229,1358,208,1405]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionText-0="FIGURE 1. The holotype (female) of Indolestes anomalus as illustrated in Fraser (1946): a—wing dorsoproximal part (after Fraser 1946: Fig. 2e); b—thorax (after Fraser 1946: Fig. 1c). Not to scale." captionText-1="FIGURE 2. The holotype (female) of Indolestes anomalus (NHMUK 013324318) at BMNH: a—habitus, dorsal view; bhabitus, lateral view; c—head and thorax, lateral view; d—labels; e—right fore wing pterostigma; f—right hind wing pterostigma. Not to scale. Courtesy of Benjamin Price." captionText-2="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." captionText-3="FIGURE 4. The female of Indolestes anomalus collected in tandem with a male in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—head top, dorsofrontal view; dsynthorax, ventral view; e—tip of abdomen, lateral view; f—right wings; g—fore wing tip; h—hind wing tip. Not to scale. Photos by Rosser Garrison." captionText-4="FIGURE 5. Indolestes anomalus photographed in nature in Thailand in Chaiyaphum Province, Phu Khieo Wildlife Sanctuary, 12 v 2016 (a) and Phetchabun Province, Nam Nao National Park, Suan Son Ban Baek, 15 viii 2015 (b–d); a, b—mature males; c—mature female; d—ovipositing tandem. Photo by Dennis Farrell." captionText-5="FIGURE 6. The male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison." captionText-6="FIGURE 7. The structure of the genital ligula of a male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996: a—ectal view; b—details of ectal arm, ectolateral view; c—lateral view. (Drawings by Rosser Garrison)." httpUri-0="https://zenodo.org/record/2624154/files/figure.png" httpUri-1="https://zenodo.org/record/2624157/files/figure.png" httpUri-2="https://zenodo.org/record/2624159/files/figure.png" httpUri-3="https://zenodo.org/record/2624161/files/figure.png" httpUri-4="https://zenodo.org/record/2624165/files/figure.png" httpUri-5="https://zenodo.org/record/2624167/files/figure.png" httpUri-6="https://zenodo.org/record/2624169/files/figure.png" pageId="1" pageNumber="68">Figs. 1–7</figureCitation>
</paragraph>
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<treatmentCitationGroup id="AB5E30193E702E7D02F9D26EFCDDC7E1" infered="true" pageId="1" pageNumber="68">
<taxonomicName id="4C4E6CB43E702E7D02F9D26EFD83C716" authority="Fraser 1946: 43" authorityName="Fraser, Dalat" authorityPageNumber="43" authorityYear="1946" box="[151,527,258,280]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B93ACB253E702E7D02F9D26EFEECC716" box="[151,352,258,280]" italics="true" pageId="1" pageNumber="68">Indolestes anomala</emphasis>
<treatmentCitation id="0AEF31263E702E7D0304D26FFD83C716" author="Fraser, F. C." box="[362,527,258,280]" page="43" pageId="1" pageNumber="68" year="1946">
<bibRefCitation id="EFDF6AC63E702E7D0304D26FFE66C716" author="Fraser, F. C." box="[362,490,258,280]" pageId="1" pageNumber="68" pagination="41 - 45" refId="ref5050" refString="Fraser, F. C. (1946) On rare and two new species of Odonata from tropical Africa and Asia. Proceedings of Royal Entomological Society of London. Series B, 15 (3 / 4), 41 - 45. https: // doi. org / 10.1111 / j. 1365 - 3113.1946. tb 00815. x" type="journal article" year="1946">Fraser 1946</bibRefCitation>
: 43
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–44, figs. 1c, 2e—original description of “a single rather teneral female” from “
<collectingCountry id="F35957A73E702E7D070ED26EFA15C716" box="[1376,1433,258,280]" name="Thailand" pageId="1" pageNumber="68">Siam</collectingCountry>
: Doun-moi”, drawings of the pterostigma and thorax in lateral view;
<treatmentCitation id="0AEF31263E702E7D01E1D24DFB7CC739" author="Hamalainen, M. &amp; Pinratana, A." box="[911,1264,289,311]" page="32" pageId="1" pageNumber="68" year="1999">
<bibRefCitation id="EFDF6AC63E702E7D01E1D24DFB40C739" author="Hamalainen, M. &amp; Pinratana, A." box="[911,1228,289,311]" pageId="1" pageNumber="68" refId="ref5150" refString="Hamalainen, M. &amp; Pinratana, A. (1999) Atlas of the Dragonflies of Thailand. Distribution maps by provinces. Brothers of St. Gabriel in Thailand, Bangkok, 176 pp." type="book" year="1999">Hämäläinen &amp; Pinratana 1999</bibRefCitation>
: 32
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—report for
<collectingRegion id="498AD9D53E702E7D0718D24DFA17C739" box="[1398,1435,289,311]" country="Thailand" name="Tak" pageId="1" pageNumber="68">Tak</collectingRegion>
(June) [Mae Sot,
<date id="FFF031F73E702E7D0314D22CFE52C758" box="[378,478,320,342]" pageId="1" pageNumber="68" value="1986-06-01">1 vi 1986</date>
,
<specimenCount id="9D48DCBE3E702E7D0387D22CFD9EC758" box="[489,530,320,342]" pageId="1" pageNumber="68" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D48DCBE3E702E7D0070D22CFDC9C758" box="[542,581,320,342]" pageId="1" pageNumber="68" type="female">1 ♀</specimenCount>
, A. Pinratana’s collectors leg.—M. Hämäläinen, pers. comm.],
<collectingRegion id="498AD9D53E702E7D06BCD22DFABEC758" box="[1234,1330,321,342]" country="Thailand" name="Lampang" pageId="1" pageNumber="68">Lampang</collectingRegion>
(October) [Huay
<collectingRegion id="498AD9D53E702E7D0363D232FEBFC77A" box="[269,307,350,372]" country="Thailand" name="Tak" pageId="1" pageNumber="68">Tak</collectingRegion>
Teak Plantation,
<date id="FFF031F73E702E7D0388D232FDDCC77A" box="[486,592,350,372]" pageId="1" pageNumber="68" value="1986-10-23">23 x 1986</date>
,
<specimenCount id="9D48DCBE3E702E7D0035D232FD08C77A" box="[603,644,350,372]" pageId="1" pageNumber="68" type="male">1 ♂</specimenCount>
, M. Hämäläinen leg.—M. Hämäläinen, pers. comm.] and
<collectingRegion id="498AD9D53E702E7D06B6D232FAD5C77A" box="[1240,1369,350,372]" country="Thailand" name="Chaiyaphum" pageId="1" pageNumber="68">Chaiyaphum</collectingRegion>
(July) [Phu Khieo,
<date id="FFF031F73E702E7D0329D211FE31C79D" box="[327,445,381,403]" pageId="1" pageNumber="68" value="1996-07-20">20 vii 1996</date>
,
<specimenCount id="9D48DCBE3E702E7D03A9D211FE61C79D" box="[455,493,381,403]" pageId="1" pageNumber="68" type="female">1 ♀</specimenCount>
, A. Pinratana leg.—M. Hämäläinen, pers. comm.] Provinces of
<collectingCountry id="F35957A73E702E7D0619D211FB59C79D" box="[1143,1237,381,403]" name="Thailand" pageId="1" pageNumber="68">Thailand</collectingCountry>
;
<treatmentCitation id="0AEF31263E702E7D06B1D212FA1BC79D" author="Ng, J. F. &amp; Dow, R. W. &amp; Choong, C. Y." box="[1247,1431,381,403]" page="12" pageId="1" pageNumber="68" year="2011">
<bibRefCitation id="EFDF6AC63E702E7D06B1D212FAFFC79D" author="Ng, J. F. &amp; Dow, R. W. &amp; Choong, C. Y." box="[1247,1395,381,403]" pageId="1" pageNumber="68" pagination="9 - 16" refId="ref5415" refString="Ng, J. F., Dow, R. W. &amp; Choong, C. Y. (2011) New records of Odonata (Insecta) from the Cameron Highlands, with first records of two species for Malaysia. Journal of Science and Technology in Tropics, 7, 9 - 16." type="journal article" year="2011">
Ng
<emphasis id="B93ACB253E702E7D076AD212FABCC79D" box="[1284,1328,381,403]" italics="true" pageId="1" pageNumber="68">et al</emphasis>
. 2011
</bibRefCitation>
: 12
</treatmentCitation>
, fig. 1—report for the Peninsular
<collectingCountry id="F35957A73E702E7D007FD2F0FDFCC7BC" box="[529,624,412,434]" name="Malaysia" pageId="1" pageNumber="68">Malaysia</collectingCountry>
, Cameron Highlands of
<specimenCount id="9D48DCBE3E702E7D0109D2F0FC13C7BC" box="[871,927,412,434]" pageId="1" pageNumber="68" type="male">2 ♂♂</specimenCount>
collected on
<date id="FFF031F73E702E7D0648D2F0FB1EC7BC" box="[1062,1170,412,434]" pageId="1" pageNumber="68" value="2008-09-20">20 ix 2008</date>
at Sg. Pos Terisu area and of
<specimenCount id="9D48DCBE3E702E7D028DD2D6FE86C7DE" box="[227,266,442,464]" pageId="1" pageNumber="68" type="male">1 ♂</specimenCount>
collected on
<date id="FFF031F73E702E7D03FBD2D6FD88C7DE" box="[405,516,442,464]" pageId="1" pageNumber="68" value="2008-09-24">24 ix 2008</date>
at Sg. Telom area; drawings of the variable pattern of synthorax of males from
<collectingCountry id="F35957A73E702E7D0757D2D6FA14C7DE" box="[1337,1432,442,464]" name="Malaysia" pageId="1" pageNumber="68">Malaysia</collectingCountry>
, Cameron Highlands, and
<collectingCountry id="F35957A73E702E7D03A4D2B5FDABC7E1" box="[458,551,473,495]" name="Thailand" pageId="1" pageNumber="68">Thailand</collectingCountry>
,
<collectingRegion id="498AD9D53E702E7D005ED2B6FD1CC7E1" box="[560,656,474,495]" country="Thailand" name="Lampang" pageId="1" pageNumber="68">Lampang</collectingRegion>
and
<collectingRegion id="498AD9D53E702E7D00AFD2B5FD6AC7E1" box="[705,742,473,495]" country="Thailand" name="Tak" pageId="1" pageNumber="68">Tak</collectingRegion>
Provinces
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</paragraph>
<paragraph id="8BF117373E702E7D02F9D295FBB7C445" blockId="1.[151,1436,258,648]" pageId="1" pageNumber="68">
<treatmentCitationGroup id="AB5E30193E702E7D02F9D295FBB7C445" bridged="true" pageId="1" pageNumber="68">
<taxonomicName id="4C4E6CB43E702E7D02F9D295FD98C400" authority="(Fraser)" authorityName="Fraser" baseAuthorityName="Fraser" box="[151,532,504,526]" class="Insecta" family="Lestidae" genus="Lestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="anomala" subGenus="Indolestes">
<emphasis id="B93ACB253E702E7D02F9D295FF5AC400" box="[151,214,505,526]" italics="true" pageId="1" pageNumber="68">Lestes</emphasis>
(
<emphasis id="B93ACB253E702E7D0289D294FEC1C400" box="[231,333,504,526]" italics="true" pageId="1" pageNumber="68">Indolestes</emphasis>
)
<emphasis id="B93ACB253E702E7D0331D294FE3BC400" box="[351,439,504,526]" italics="true" pageId="1" pageNumber="68">anomala</emphasis>
(Fraser)
</taxonomicName>
:
<treatmentCitation id="0AEF31263E702E7D004ED294FD60C400" author="Kimmins, D. E." box="[544,748,504,526]" page="178" pageId="1" pageNumber="68" year="1966">
<bibRefCitation id="EFDF6AC63E702E7D004ED294FD31C400" author="Kimmins, D. E." box="[544,701,504,526]" pageId="1" pageNumber="68" pagination="173 - 227" refId="ref5187" refString="Kimmins, D. E. (1966) A list of the Odonata types described by F. C. Fraser, now in the British Museum (Natural History). Bulletin of the British Museum (Natural History), Entomology, 18, 173 - 227." type="journal article" year="1966">Kimmins 1966</bibRefCitation>
: 178
</treatmentCitation>
—information on the
<typeStatus id="54F5A9953E702E7D01BCD294FBA7C400" box="[978,1067,504,526]" pageId="1" pageNumber="68" type="holotype">holotype</typeStatus>
in the collection of British Natural History Museum: “
<typeStatus id="54F5A9953E702E7D03E6D17AFE64C422" box="[392,488,534,556]" pageId="1" pageNumber="68" type="holotype">Holotype</typeStatus>
<specimenCount id="9D48DCBE3E702E7D0380D17BFD8EC422" box="[494,514,535,556]" pageId="1" pageNumber="68" type="female">♀</specimenCount>
.
<taxonomicName id="4C4E6CB43E702E7D0063D17AFCD6C422" authority="Fraser, Dalat" authorityName="Fraser, Dalat" authorityYear="1946" box="[525,858,534,556]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="anomala">Indolestes anomala Fraser, Dalat</taxonomicName>
,
<collectingCountry id="F35957A73E702E7D010DD17AFC16C422" box="[867,922,534,556]" name="Thailand" pageId="1" pageNumber="68">Siam</collectingCountry>
,
<date id="FFF031F73E702E7D01CAD17AFC7BC422" box="[932,1015,534,556]" pageId="1" pageNumber="68" value="1920-10-17">17.x. 20</date>
, Type [label F.C.F.]. The locality given in the original description is Doun-moi, without date, but the specimen otherwise agrees.”
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF117373E702E7D02F9D138FE4EC486" blockId="1.[151,1436,258,648]" pageId="1" pageNumber="68">
<treatmentCitationGroup id="AB5E30193E702E7D02F9D138FE4EC486" pageId="1" pageNumber="68">
<taxonomicName id="4C4E6CB43E702E7D02F9D138FDBBC464" ID-CoL="6MVY3" authority=": Day et al. 2012: 18" authorityName="Day" authorityPageNumber="18" authorityYear="2012" box="[151,567,596,618]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E702E7D02F9D138FEEAC464" box="[151,358,596,618]" italics="true" pageId="1" pageNumber="68">Indolestes anomalus</emphasis>
:
<treatmentCitation id="0AEF31263E702E7D031BD139FDBBC464" author="Day, L. &amp; Farrell, D. &amp; Guenther, A. &amp; Hamalainen, M. &amp; Klimsa, E. &amp; Korshunov, A. &amp; Kosterin, O. &amp; Makbun N. &amp; Pelegrin, A. &amp; Roeder, U. &amp; Ruangrong R. &amp; Vikhrev, N." box="[373,567,596,618]" page="18" pageId="1" pageNumber="68" year="2012">
<bibRefCitation id="EFDF6AC63E702E7D031BD139FD94C464" author="Day, L. &amp; Farrell, D. &amp; Guenther, A. &amp; Hamalainen, M. &amp; Klimsa, E. &amp; Korshunov, A. &amp; Kosterin, O. &amp; Makbun N. &amp; Pelegrin, A. &amp; Roeder, U. &amp; Ruangrong R. &amp; Vikhrev, N." box="[373,536,596,618]" pageId="1" pageNumber="68" pagination="16 - 24" refId="ref4865" refString="Day, L., Farrell, D., Guenther, A., Hamalainen, M., Klimsa, E., Korshunov, A., Kosterin, O., Makbun N., Pelegrin, A., Roeder, U., Ruangrong R. &amp; Vikhrev, N. (2012) New provincial records of Odonata from Thailand mostly based on photographs. Agrion, 16 (1), 16 - 24." type="journal article" year="2012">
Day
<emphasis id="B93ACB253E702E7D03C8D139FE56C464" box="[422,474,596,618]" italics="true" pageId="1" pageNumber="68">et al.</emphasis>
2012
</bibRefCitation>
: 18
</treatmentCitation>
</taxonomicName>
—report of
<specimenCount id="9D48DCBE3E702E7D00DDD138FD63C464" box="[691,751,596,618]" pageId="1" pageNumber="68" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D48DCBE3E702E7D0094D138FCADC464" box="[762,801,596,618]" pageId="1" pageNumber="68" type="female">1 ♀</specimenCount>
collected on 09–
<date id="FFF031F73E702E7D01BFD138FBCEC464" box="[977,1090,596,618]" pageId="1" pageNumber="68" value="2011-10-07">10.07.2011</date>
in
<collectingCountry id="F35957A73E702E7D060AD138FB4DC464" box="[1124,1217,596,618]" name="Thailand" pageId="1" pageNumber="68">Thailand</collectingCountry>
, Petchabun Province, Nam Nao National Park.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF117373E702E7D02F9D1D9FB20C518" blockId="1.[151,1437,693,2014]" pageId="1" pageNumber="68">
<emphasis id="B93ACB253E702E7D02F9D1D9FE1AC4C0" bold="true" box="[151,406,693,718]" pageId="1" pageNumber="68">Specimens examined.</emphasis>
<materialsCitation id="3B261D6A3E702E7D03F3D1D9FE2DC4FC" ID-GBIF-Occurrence="2251610950" collectingDate="2016-02-06" collectorName="T. Kompier" country="Vietnam" latitude="11.44" location="Cat Tien National Park" longLatPrecision="77" longitude="107.422" pageId="1" pageNumber="68" specimenCount="1" specimenCount-female="1" stateProvince="Dong Nai Province">
<emphasis id="B93ACB253E702E7D03F3D1D9FD85C4C0" bold="true" box="[413,521,693,718]" pageId="1" pageNumber="68">
<collectingCountry id="F35957A73E702E7D03F3D1D9FD89C4C0" box="[413,517,693,718]" name="Vietnam" pageId="1" pageNumber="68">Vietnam</collectingCountry>
.
</emphasis>
<specimenCount id="9D48DCBE3E702E7D007ED1DAFDB6C4C3" box="[528,570,694,718]" pageId="1" pageNumber="68" type="female">1 ♀</specimenCount>
(
<figureCitation id="13750BB23E702E7D0027D1D9FD00C4C0" box="[585,652,693,718]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="1" pageNumber="68">Fig. 3</figureCitation>
), Đỗng
<collectingRegion id="498AD9D53E702E7D0086D1D9FCF2C4C0" box="[744,894,693,718]" inferredBySuffix="true" pageId="1" pageNumber="68">Nai Province</collectingRegion>
,
<location id="8E9141EC3E702E7D01E6D1DAFB01C4C0" LSID="urn:lsid:plazi:treatment:03E7A6213E702E7702F9D3FBFD49C7FA:8E9141EC3E702E7D01E6D1DAFB01C4C0" box="[904,1165,693,718]" country="Vietnam" latitude="11.44" longLatPrecision="77" longitude="107.422" name="Cat Tien National Park" pageId="1" pageNumber="68" stateProvince="Nai Province">Cát Tiên National Park</location>
,
<geoCoordinate id="EE7A71F03E702E7D06F6D1DAFA8EC4C3" box="[1176,1282,694,718]" direction="north" orientation="latitude" pageId="1" pageNumber="68" precision="55" value="11.44">11.440 N</geoCoordinate>
,
<geoCoordinate id="EE7A71F03E702E7D0763D1DAFA0EC4C3" box="[1293,1410,694,718]" direction="east" orientation="longitude" pageId="1" pageNumber="68" precision="55" value="107.422">107.422 E</geoCoordinate>
,
<collectingDate id="EFB4C81F3E702E7D07E0D1D9FF64C4FC" pageId="1" pageNumber="68" value="2016-02-06">
<date id="FFF031F73E702E7D07E0D1D9FF64C4FC" pageId="1" pageNumber="68" value="2016-02-06">6 ii 2016</date>
</collectingDate>
,
<collectorName id="26BB72E13E702E7D029AD1B6FEF9C4FC" box="[244,373,729,754]" pageId="1" pageNumber="68">T. Kompier</collectorName>
leg
</materialsCitation>
.
<materialsCitation id="3B261D6A3E702E7D03C2D1B5FB20C518" ID-GBIF-Occurrence="2234218740" collectingDate="1996-07-19" collectorName="R. W. &amp; J. A. Garrison" country="Thailand" habitat="small muddy pond" latitude="16.6089" location="Tung Lui Lai" longLatPrecision="7" longitude="101.8906" pageId="1" pageNumber="68" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Chaiyaphum">
<emphasis id="B93ACB253E702E7D03C2D1B5FDAEC4FC" bold="true" box="[428,546,729,754]" pageId="1" pageNumber="68">
<collectingCountry id="F35957A73E702E7D03C2D1B5FD92C4FC" box="[428,542,729,754]" name="Thailand" pageId="1" pageNumber="68">Thailand</collectingCountry>
:
</emphasis>
<specimenCount id="9D48DCBE3E702E7D0047D1B6FDD9C4FF" box="[553,597,730,754]" pageId="1" pageNumber="68" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D48DCBE3E702E7D000FD1B5FD2CC4FF" box="[609,672,729,754]" pageId="1" pageNumber="68" type="female">5 ♀♀</specimenCount>
(
<specimenCount id="9D48DCBE3E702E7D00DDD1B6FD57C4FF" box="[691,731,730,754]" pageId="1" pageNumber="68" type="male">1 ♂</specimenCount>
(
<figureCitation id="13750BB23E702E7D0084D1B5FCA3C4FC" box="[746,815,729,754]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[287,1278,511,1836]" captionTargetId="figure@8.[287,1299,471,1887]" captionTargetPageId="8" captionText="FIGURE 6. The male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison." httpUri="https://zenodo.org/record/2624167/files/figure.png" pageId="1" pageNumber="68">Fig. 6</figureCitation>
),
<specimenCount id="9D48DCBE3E702E7D012AD1B6FCE2C4FF" box="[836,878,730,754]" pageId="1" pageNumber="68" type="female">1 ♀</specimenCount>
(
<figureCitation id="13750BB23E702E7D0110D1B5FC4DC4FC" box="[894,961,729,754]" captionStart="FIGURE 4" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[296,1246,481,1879]" captionTargetId="figure@6.[288,1298,481,1903]" captionTargetPageId="6" captionText="FIGURE 4. The female of Indolestes anomalus collected in tandem with a male in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—head top, dorsofrontal view; dsynthorax, ventral view; e—tip of abdomen, lateral view; f—right wings; g—fore wing tip; h—hind wing tip. Not to scale. Photos by Rosser Garrison." httpUri="https://zenodo.org/record/2624161/files/figure.png" pageId="1" pageNumber="68">Fig. 4</figureCitation>
) in tandem),
<collectingRegion id="498AD9D53E702E7D0639D1B5FAD9C4FC" box="[1111,1365,729,754]" country="Thailand" name="Chaiyaphum" pageId="1" pageNumber="68">Chaiyaphum Province</collectingRegion>
, small muddy pond at
<location id="8E9141EC3E702E7D0328D192FE51C518" LSID="urn:lsid:plazi:treatment:03E7A6213E702E7702F9D3FBFD49C7FA:8E9141EC3E702E7D0328D192FE51C518" box="[326,477,765,790]" country="Thailand" latitude="16.6089" longLatPrecision="7" longitude="101.8906" name="Tung Lui Lai" pageId="1" pageNumber="68" stateProvince="Chaiyaphum">Tung Lui Lai</location>
,
<geoCoordinate id="EE7A71F03E702E7D0386D191FDEDC51B" box="[488,609,765,790]" direction="north" orientation="latitude" pageId="1" pageNumber="68" precision="5" value="16.6089">16.6089 N</geoCoordinate>
,
<geoCoordinate id="EE7A71F03E702E7D0002D191FD7CC51B" box="[620,752,765,790]" direction="east" orientation="longitude" pageId="1" pageNumber="68" precision="5" value="101.8906">101.8906 E</geoCoordinate>
,
<collectingDate id="EFB4C81F3E702E7D0092D192FCF2C518" box="[764,894,765,790]" pageId="1" pageNumber="68" value="1996-07-19">
<date id="FFF031F73E702E7D0092D192FCF2C518" box="[764,894,765,790]" pageId="1" pageNumber="68" value="1996-07-19">19 vii 1996</date>
</collectingDate>
,
<collectorName id="26BB72E13E702E7D01E7D192FBF7C518" box="[905,1147,765,790]" pageId="1" pageNumber="68">R.W. &amp; J.A. Garrison</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35444BC3E702E7502A9D04DFD30C0CD" lastPageId="9" lastPageNumber="76" pageId="1" pageNumber="68" type="description">
<paragraph id="8BF117373E702E7D02A9D04DFB16C5A8" blockId="1.[151,1437,693,2014]" pageId="1" pageNumber="68">
<emphasis id="B93ACB253E702E7D02A9D04DFE1FC534" bold="true" box="[199,403,801,826]" pageId="1" pageNumber="68">The type locality.</emphasis>
The
<typeStatus id="54F5A9953E702E7D03A2D04EFE70C534" box="[460,508,802,826]" pageId="1" pageNumber="68">type</typeStatus>
locality of
<taxonomicName id="4C4E6CB43E702E7D0015D04EFD73C534" box="[635,767,801,826]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E702E7D0015D04EFD73C534" box="[635,767,801,826]" italics="true" pageId="1" pageNumber="68">I. anomalus</emphasis>
</taxonomicName>
was given in the original description as “
<collectingCountry id="F35957A73E702E7D06A2D04DFA87C534" box="[1228,1291,801,826]" name="Thailand" pageId="1" pageNumber="68">Siam</collectingCountry>
: Doun-moi”.
<collectingCountry id="F35957A73E702E7D02F9D029FF5DC550" box="[151,209,837,862]" name="Thailand" pageId="1" pageNumber="68">Siam</collectingCountry>
is the historical name of
<collectingCountry id="F35957A73E702E7D0380D029FDD8C550" box="[494,596,837,862]" name="Thailand" pageId="1" pageNumber="68">Thailand</collectingCountry>
, but no site with this or a similar name could be located in that country, so
<bibRefCitation id="EFDF6AC63E702E7D02F9D005FDEBC58C" author="Hamalainen, M. &amp; Pinratana, A." box="[151,615,873,898]" pageId="1" pageNumber="68" refId="ref5150" refString="Hamalainen, M. &amp; Pinratana, A. (1999) Atlas of the Dragonflies of Thailand. Distribution maps by provinces. Brothers of St. Gabriel in Thailand, Bangkok, 176 pp." type="book" year="1999">Hämäläinen &amp; Pinratana (1999: page 32)</bibRefCitation>
noted that “The whereabouts of the
<typeStatus id="54F5A9953E702E7D066AD006FBB8C58C" box="[1028,1076,874,898]" pageId="1" pageNumber="68">type</typeStatus>
locality is unknown”, and
<bibRefCitation id="EFDF6AC63E702E7D0708D006FF6DC5A8" author="Dow, R. A." pageId="1" pageNumber="68" refId="ref4948" refString="Dow, R. A. (2009) Indolestes anomalus. The IUCN Red List of Threatened Species. Version 2014.3. Available from: www. iucnredlist. org (accessed February 2018)" type="journal volume" year="2009">Dow (2009)</bibRefCitation>
noted that “The
<typeStatus id="54F5A9953E702E7D03F3D0E2FE41C5A8" box="[413,461,910,934]" pageId="1" pageNumber="68">type</typeStatus>
locality, &quot;
<collectingCountry id="F35957A73E702E7D0050D0E1FDF1C5A8" box="[574,637,909,934]" name="Thailand" pageId="1" pageNumber="68">Siam</collectingCountry>
, Doun Moi&quot; (
<bibRefCitation id="EFDF6AC63E702E7D0178D0E2FC12C5A8" author="Fraser, F. C." box="[790,926,909,934]" pageId="1" pageNumber="68" pagination="41 - 45" refId="ref5050" refString="Fraser, F. C. (1946) On rare and two new species of Odonata from tropical Africa and Asia. Proceedings of Royal Entomological Society of London. Series B, 15 (3 / 4), 41 - 45. https: // doi. org / 10.1111 / j. 1365 - 3113.1946. tb 00815. x" type="journal article" year="1946">Fraser 1946</bibRefCitation>
) cannot be relocated.”
</paragraph>
<paragraph id="8BF117373E702E7D02A9D0DEFC63C2AC" blockId="1.[151,1437,693,2014]" pageId="1" pageNumber="68">
“Doun Moi” might be variant spelling of “Than Moi” or “Than Muoi”, a place in northern
<collectingCountry id="F35957A73E702E7D069DD0DDFADEC5C4" box="[1267,1362,945,970]" name="Vietnam" pageId="1" pageNumber="68">Vietnam</collectingCountry>
(or in
<collectingCountry id="F35957A73E702E7D02F9D0B9FF65C5E0" box="[151,233,981,1006]" name="Vietnam" pageId="1" pageNumber="68">Tonkin</collectingCountry>
, as that was referred to at that time), well known for numerous insect specimens collected there by Hans Fruhstorfer in June–July 1901. Presently this location is called Đỗng Mỏ town, the capital of Chi Lăng District, Lạng SƠn Province (
<geoCoordinate id="EE7A71F03E702E7D03EAD771FE7DC23B" box="[388,497,1053,1078]" direction="north" orientation="latitude" pageId="1" pageNumber="68" precision="555" value="21.66">21.66° N</geoCoordinate>
,
<geoCoordinate id="EE7A71F03E702E7D0390D771FDFEC238" box="[510,626,1053,1078]" direction="east" orientation="longitude" pageId="1" pageNumber="68" precision="555" value="106.58">106.58° E</geoCoordinate>
) (
<bibRefCitation id="EFDF6AC63E702E7D00E5D771FCD6C238" author="Hamalainen, M." box="[651,858,1053,1078]" pageId="1" pageNumber="68" pagination="156 - 201" refId="ref5114" refString="Hamalainen, M. (2015) From Echo maxima to Archineura maxima - a slow taxonomic process (Odonata: Calopterygidae). Notulae odonatologicae, 8 (6), 156 - 201." type="journal article" year="2015">Hämäläinen 2015</bibRefCitation>
). Thus “
<collectingCountry id="F35957A73E702E7D01D2D771FC75C238" box="[956,1017,1053,1078]" name="Thailand" pageId="1" pageNumber="68">Siam</collectingCountry>
” could result from a confusion with
<collectingCountry id="F35957A73E702E7D02F9D72DFF6AC254" box="[151,230,1089,1114]" name="Vietnam" pageId="1" pageNumber="68">Tonkin</collectingCountry>
or used in a sense to include all of Indochina. (For instance, the
<typeStatus id="54F5A9953E702E7D01AED72DFBAEC254" box="[960,1058,1089,1114]" pageId="1" pageNumber="68" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C4E6CB43E702E7D0627D72DFEAFC270" authority="Fraser, 1933" authorityName="Fraser" authorityYear="1933" class="Insecta" family="Gomphidae" genus="Macrogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="borikhanensis">
<emphasis id="B93ACB253E702E7D0627D72DFA10C254" box="[1097,1436,1089,1114]" italics="true" pageId="1" pageNumber="68">Macrogomphus borikhanensis</emphasis>
<bibRefCitation id="EFDF6AC63E702E7D02F9D70AFEAFC270" author="Fraser, F. C." box="[151,291,1126,1150]" pageId="1" pageNumber="68" refId="ref5015" refString="Fraser, F. C. (1933) The fauna of British India, including Ceylon and Burma. Odonata. Uol. I. Taylor and Francis, London, 423 pp." type="book" year="1933">Fraser, 1933</bibRefCitation>
</taxonomicName>
is labelled “Borikhane/
<collectingCountry id="F35957A73E702E7D005CD709FDE5C270" box="[562,617,1125,1150]" name="Laos" pageId="1" pageNumber="68">Laos</collectingCountry>
,
<collectingCountry id="F35957A73E702E7D001BD709FD3FC270" box="[629,691,1125,1150]" name="Thailand" pageId="1" pageNumber="68">Siam</collectingCountry>
”, see
<bibRefCitation id="EFDF6AC63E702E7D0098D709FC2EC270" author="Kimmins, D. E." box="[758,930,1125,1150]" pageId="1" pageNumber="68" pagination="173 - 227" refId="ref5187" refString="Kimmins, D. E. (1966) A list of the Odonata types described by F. C. Fraser, now in the British Museum (Natural History). Bulletin of the British Museum (Natural History), Entomology, 18, 173 - 227." type="journal article" year="1966">Kimmins 1966</bibRefCitation>
). But the
<typeStatus id="54F5A9953E702E7D067CD709FBF8C270" box="[1042,1140,1125,1150]" pageId="1" pageNumber="68" type="holotype">holotype</typeStatus>
label has no indication of the collector, which as a rule is present on labels of Fruhstorfer’s specimens.
</paragraph>
<paragraph id="8BF117373E702E7D02A9D7C2FE19C0EC" blockId="1.[151,1437,693,2014]" pageId="1" pageNumber="68">
However,
<bibRefCitation id="EFDF6AC63E702E7D0353D7C1FE77C2C8" author="Kimmins, D. E." box="[317,507,1197,1222]" pageId="1" pageNumber="68" pagination="173 - 227" refId="ref5187" refString="Kimmins, D. E. (1966) A list of the Odonata types described by F. C. Fraser, now in the British Museum (Natural History). Bulletin of the British Museum (Natural History), Entomology, 18, 173 - 227." type="journal article" year="1966">Kimmins (1966)</bibRefCitation>
transcribed the
<typeStatus id="54F5A9953E702E7D00D6D7C1FC96C2C8" box="[696,794,1197,1222]" pageId="1" pageNumber="68" type="holotype">holotype</typeStatus>
label (
<figureCitation id="13750BB23E702E7D0100D7C1FC4FC2C8" box="[878,963,1197,1222]" captionStart="FIGURE 2" captionStartId="3.[151,250,1674,1696]" captionTargetBox="[276,1312,337,1653]" captionTargetId="figure@3.[276,1312,337,1653]" captionTargetPageId="3" captionText="FIGURE 2. The holotype (female) of Indolestes anomalus (NHMUK 013324318) at BMNH: a—habitus, dorsal view; bhabitus, lateral view; c—head and thorax, lateral view; d—labels; e—right fore wing pterostigma; f—right hind wing pterostigma. Not to scale. Courtesy of Benjamin Price." httpUri="https://zenodo.org/record/2624157/files/figure.png" pageId="1" pageNumber="68">Fig. 2d</figureCitation>
) as follows: &quot;
<taxonomicName id="4C4E6CB43E702E7D0606D7C1FF5BC2E4" authority="Fraser, Dalat" authorityName="Fraser, Dalat" authorityYear="1946" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="68" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B93ACB253E702E7D0606D7C1FACAC2C8" box="[1128,1350,1197,1222]" italics="true" pageId="1" pageNumber="68">Indolestes anomala</emphasis>
Fraser, Dalat
</taxonomicName>
,
<collectingCountry id="F35957A73E702E7D028CD7BDFE92C2E4" box="[226,286,1233,1258]" name="Thailand" pageId="1" pageNumber="68">Siam</collectingCountry>
,
<date id="FFF031F73E702E7D0344D7BEFE0CC2E4" box="[298,384,1234,1258]" pageId="1" pageNumber="68" value="1920-10-17">17.x.20</date>
, Type&quot; and added: &quot;Type locality given in the original description is Doun-moi, without date, but the specimen otherwise agrees&quot;.
<bibRefCitation id="EFDF6AC63E702E7D002DD799FC8DC300" author="Kimmins, D. E." box="[579,769,1269,1294]" pageId="1" pageNumber="68" pagination="173 - 227" refId="ref5187" refString="Kimmins, D. E. (1966) A list of the Odonata types described by F. C. Fraser, now in the British Museum (Natural History). Bulletin of the British Museum (Natural History), Entomology, 18, 173 - 227." type="journal article" year="1966">Kimmins (1966)</bibRefCitation>
noted a number of cases where actual labels of the type specimens did not coincide with information given by Fraser in his original descriptions. There are a Dalet or Dalat Town in
<collectingCountry id="F35957A73E702E7D0293D652FEE6C358" box="[253,362,1342,1366]" name="Myanmar" pageId="1" pageNumber="68">Myanmar</collectingCountry>
and Đà Lạt Town in
<collectingCountry id="F35957A73E702E7D0031D651FD4DC358" box="[607,705,1341,1366]" name="Vietnam" pageId="1" pageNumber="68">Vietnam</collectingCountry>
, the capital of Lâm Đỗng Province. The Cát Tiên District of the latter province is the location of the headwaters of the Đỗng Nai River. This river gave rise to the name of Đỗng Nai Thưọng commune in Cát Tiên District and Đỗng Nai Province (with the capital Biên Hòa), which borders Lâm Đỗng Province to the southwest. It is not clear which were the local toponyms and administrative divisions during Fraser’s time but it is very plausible that the
<typeStatus id="54F5A9953E702E7D00E1D6A1FD7DC3E8" box="[655,753,1485,1510]" pageId="1" pageNumber="68" type="holotype">holotype</typeStatus>
originated from the Đỗng Nai River basin in South
<collectingCountry id="F35957A73E702E7D0753D6A1FA10C3E8" box="[1341,1436,1485,1510]" name="Vietnam" pageId="1" pageNumber="68">Vietnam</collectingCountry>
and that the river (or town) name was misspelled as “Doun-moi”. The original label of the specimen (or otherwise communicated information on the locality) could include both ‘Dalat’ and ‘Doun-moi’, two different toponyms from the same area, while Fraser included only the former on the label under the
<typeStatus id="54F5A9953E702E7D0644D555FB00C05C" box="[1066,1164,1593,1618]" pageId="1" pageNumber="68" type="holotype">holotype</typeStatus>
and mentioned only the latter in the original description. “
<collectingCountry id="F35957A73E702E7D007AD531FDDEC078" box="[532,594,1629,1654]" name="Thailand" pageId="1" pageNumber="68">Siam</collectingCountry>
” still was also an error or was used in too broad a sense. So any topotypes should be sought within the Đỗng Nai River basin in Lâm Đỗng Province or Đỗng Nai Province. Cát Tiên National Park is located along the Dong Nai River on the border between these provinces and so would be a good place to search for this species.
</paragraph>
<paragraph id="8BF117373E702E7D02A9D581FCFEC17C" blockId="1.[151,1437,693,2014]" pageId="1" pageNumber="68">
<emphasis id="B93ACB253E702E7D02A9D581FEE0C108" bold="true" box="[199,364,1773,1798]" pageId="1" pageNumber="68">
The
<typeStatus id="54F5A9953E702E7D0290D581FEEBC108" box="[254,359,1773,1798]" pageId="1" pageNumber="68" type="holotype">holotype</typeStatus>
.
</emphasis>
<location id="8E9141EC3E702E7D031BD581FE2DC108" LSID="urn:lsid:plazi:treatment:03E7A6213E702E7702F9D3FBFD49C7FA:8E9141EC3E702E7D031BD581FE2DC108" box="[373,417,1773,1798]" name="The" pageId="1" pageNumber="68">The</location>
<typeStatus id="54F5A9953E702E7D03C7D581FD87C108" box="[425,523,1773,1798]" pageId="1" pageNumber="68" type="holotype">holotype</typeStatus>
is kept in
<location id="8E9141EC3E702E7D00E6D581FC2AC10B" LSID="urn:lsid:plazi:treatment:03E7A6213E702E7702F9D3FBFD49C7FA:8E9141EC3E702E7D00E6D581FC2AC10B" box="[648,934,1773,1798]" country="United Kingdom" name="Natural History Museum" pageId="1" pageNumber="68">Natural History Museum</location>
,
<collectingCountry id="F35957A73E702E7D01DDD581FBFEC108" box="[947,1138,1773,1798]" name="United Kingdom" pageId="1" pageNumber="68">United Kingdom</collectingCountry>
(BMNH) under reference number NHMUK 0 13324318. Photographs of its details were kindly provided by Benjamin Price (
<figureCitation id="13750BB23E702E7D077FD47DFAD5C124" box="[1297,1369,1809,1834]" captionStart="FIGURE 2" captionStartId="3.[151,250,1674,1696]" captionTargetBox="[276,1312,337,1653]" captionTargetId="figure@3.[276,1312,337,1653]" captionTargetPageId="3" captionText="FIGURE 2. The holotype (female) of Indolestes anomalus (NHMUK 013324318) at BMNH: a—habitus, dorsal view; bhabitus, lateral view; c—head and thorax, lateral view; d—labels; e—right fore wing pterostigma; f—right hind wing pterostigma. Not to scale. Courtesy of Benjamin Price." httpUri="https://zenodo.org/record/2624157/files/figure.png" pageId="1" pageNumber="68">Fig. 2</figureCitation>
). The
<typeStatus id="54F5A9953E702E7D02F9D459FF75C140" box="[151,249,1845,1870]" pageId="1" pageNumber="68" type="holotype">holotype</typeStatus>
is a somewhat teneral female in poor condition.
<location id="8E9141EC3E702E7D0172D459FCC4C140" LSID="urn:lsid:plazi:treatment:03E7A6213E702E7702F9D3FBFD49C7FA:8E9141EC3E702E7D0172D459FCC4C140" box="[796,840,1845,1870]" country="United Kingdom" name="The" pageId="1" pageNumber="68">The</location>
left hind wing and abdominal segments [hereafter S] 8–10 are missing (but were described in the original description).
</paragraph>
<paragraph id="8BF117373E702E7E02A9D411FB1DC6DB" blockId="1.[151,1437,693,2014]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="69" pageId="1" pageNumber="68">
There is a pair of pale spots on the occiput lateroposteriorly of the lateral ocelli, surrounding small tubercles, mentioned in the original description (
<bibRefCitation id="EFDF6AC63E702E7D0035D4CEFD67C1B4" author="Fraser, F. C." box="[603,747,1953,1978]" pageId="1" pageNumber="68" pagination="41 - 45" refId="ref5050" refString="Fraser, F. C. (1946) On rare and two new species of Odonata from tropical Africa and Asia. Proceedings of Royal Entomological Society of London. Series B, 15 (3 / 4), 41 - 45. https: // doi. org / 10.1111 / j. 1365 - 3113.1946. tb 00815. x" type="journal article" year="1946">Fraser 1946</bibRefCitation>
). The synthoracic pattern (
<figureCitation id="13750BB23E702E7D0644D4CDFBF3C1B4" box="[1066,1151,1953,1978]" captionStart="FIGURE 2" captionStartId="3.[151,250,1674,1696]" captionTargetBox="[276,1312,337,1653]" captionTargetId="figure@3.[276,1312,337,1653]" captionTargetPageId="3" captionText="FIGURE 2. The holotype (female) of Indolestes anomalus (NHMUK 013324318) at BMNH: a—habitus, dorsal view; bhabitus, lateral view; c—head and thorax, lateral view; d—labels; e—right fore wing pterostigma; f—right hind wing pterostigma. Not to scale. Courtesy of Benjamin Price." httpUri="https://zenodo.org/record/2624157/files/figure.png" pageId="1" pageNumber="68">Fig. 2c</figureCitation>
) fits the drawing in the original description (
<bibRefCitation id="EFDF6AC63E702E7D03E6D4AAFDF4C1D0" author="Fraser, F. C." box="[392,632,1989,2014]" pageId="1" pageNumber="68" pagination="41 - 45" refId="ref5050" refString="Fraser, F. C. (1946) On rare and two new species of Odonata from tropical Africa and Asia. Proceedings of Royal Entomological Society of London. Series B, 15 (3 / 4), 41 - 45. https: // doi. org / 10.1111 / j. 1365 - 3113.1946. tb 00815. x" type="journal article" year="1946">
Fraser 1946:
<figureCitation id="13750BB23E702E7D004BD4A9FDF4C1D0" box="[549,632,1989,2014]" captionStart="FIGURE 1" captionStartId="2.[151,250,1507,1529]" captionTargetBox="[340,1226,424,1423]" captionTargetId="figure@2.[340,1247,267,1483]" captionTargetPageId="2" captionText="FIGURE 1. The holotype (female) of Indolestes anomalus as illustrated in Fraser (1946): a—wing dorsoproximal part (after Fraser 1946: Fig. 2e); b—thorax (after Fraser 1946: Fig. 1c). Not to scale." httpUri="https://zenodo.org/record/2624154/files/figure.png" pageId="1" pageNumber="68">Fig. 1b</figureCitation>
</bibRefCitation>
) where, however, the black spots at the posteroventral corner of the mesepisternum and anteroventral corner of the metepimeron were shown as smaller, while the dark stripe from the latter to the spiracle was shown as distinct although this is barely traceable in the
<typeStatus id="54F5A9953E732E7E0649D3D0FB01C6DB" box="[1063,1165,188,213]" pageId="2" pageNumber="69" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="DF3147BF3E732E7E02F9D68FFC08C016" httpUri="https://zenodo.org/record/2624154/files/figure.png" pageId="2" pageNumber="69" startId="2.[151,250,1507,1529]" targetBox="[340,1226,424,1423]" targetPageId="2">
<paragraph id="8BF117373E732E7E02F9D68FFC08C016" blockId="2.[151,1436,1507,1560]" pageId="2" pageNumber="69">
<emphasis id="B93ACB253E732E7E02F9D68FFE98C3F6" bold="true" box="[151,276,1507,1529]" pageId="2" pageNumber="69">FIGURE 1.</emphasis>
The holotype (female) of
<taxonomicName id="4C4E6CB43E732E7E004BD68FFD7AC3F7" box="[549,758,1507,1529]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E732E7E004BD68FFD7AC3F7" box="[549,758,1507,1529]" italics="true" pageId="2" pageNumber="69">Indolestes anomalus</emphasis>
</taxonomicName>
as illustrated in Fraser (1946): a—wing dorsoproximal part (after Fraser 1946: Fig. 2e); b—thorax (after Fraser 1946: Fig. 1c). Not to scale.
</paragraph>
</caption>
<paragraph id="8BF117373E732E7F02A9D52CFB72C712" blockId="2.[151,1437,1599,2020]" lastBlockId="3.[151,1436,151,285]" lastPageId="3" lastPageNumber="70" pageId="2" pageNumber="69">
In describing the most conspicuous diagnostic character for the species, the pterostigma unusually short for
<taxonomicName id="4C4E6CB43E732E7E02F9D50FFE42C072" authority=", Fraser (1946: 45)" authorityName=", Fraser" authorityPageNumber="45" authorityYear="1946" box="[151,462,1635,1660]" class="Insecta" family="Lestidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="69" phylum="Arthropoda" rank="family">
Lestidae,
<bibRefCitation id="EFDF6AC63E732E7E0368D508FE42C072" author="Fraser, F. C." box="[262,462,1635,1660]" pageId="2" pageNumber="69" pagination="41 - 45" refId="ref5050" refString="Fraser, F. C. (1946) On rare and two new species of Odonata from tropical Africa and Asia. Proceedings of Royal Entomological Society of London. Series B, 15 (3 / 4), 41 - 45. https: // doi. org / 10.1111 / j. 1365 - 3113.1946. tb 00815. x" type="journal article" year="1946">Fraser (1946: 45)</bibRefCitation>
</taxonomicName>
made somewhat confusing statements: “pterostigma ... of fore wing about one-third longer than deep, that of the hind-wing about half as long again as deep” meaning that the pterostigmata length in the fore and hind wings was, respectively, ca 1.33 and 1.5 times its height. Fraser did not mention which pterostigma he figures (
<figureCitation id="13750BB23E732E7E03CBD5A3FE74C0E9" box="[421,504,1743,1768]" captionStart="FIGURE 2" captionStartId="3.[151,250,1674,1696]" captionTargetBox="[276,1312,337,1653]" captionTargetId="figure@3.[276,1312,337,1653]" captionTargetPageId="3" captionText="FIGURE 2. The holotype (female) of Indolestes anomalus (NHMUK 013324318) at BMNH: a—habitus, dorsal view; bhabitus, lateral view; c—head and thorax, lateral view; d—labels; e—right fore wing pterostigma; f—right hind wing pterostigma. Not to scale. Courtesy of Benjamin Price." httpUri="https://zenodo.org/record/2624157/files/figure.png" pageId="2" pageNumber="69">Fig. 2e</figureCitation>
), showing the maximum length comprising 1.68 of the maximum width (
<figureCitation id="13750BB23E732E7E0752D5A3FA01C0E9" box="[1340,1421,1743,1768]" captionStart="FIGURE 1" captionStartId="2.[151,250,1507,1529]" captionTargetBox="[340,1226,424,1423]" captionTargetId="figure@2.[340,1247,267,1483]" captionTargetPageId="2" captionText="FIGURE 1. The holotype (female) of Indolestes anomalus as illustrated in Fraser (1946): a—wing dorsoproximal part (after Fraser 1946: Fig. 2e); b—thorax (after Fraser 1946: Fig. 1c). Not to scale." httpUri="https://zenodo.org/record/2624154/files/figure.png" pageId="2" pageNumber="69">Fig. 1a</figureCitation>
). The actual pterostigmata of the
<typeStatus id="54F5A9953E732E7E006ED59FFDEEC102" box="[512,610,1779,1804]" pageId="2" pageNumber="69" type="holotype">holotype</typeStatus>
are substantially longer: 1.90 times as long as high along the dorsal side and 2.25 times as long as high along the ventral side in the fore wing (
<figureCitation id="13750BB23E732E7E01A2D47BFB93C121" box="[972,1055,1815,1840]" captionStart="FIGURE 2" captionStartId="3.[151,250,1674,1696]" captionTargetBox="[276,1312,337,1653]" captionTargetId="figure@3.[276,1312,337,1653]" captionTargetPageId="3" captionText="FIGURE 2. The holotype (female) of Indolestes anomalus (NHMUK 013324318) at BMNH: a—habitus, dorsal view; bhabitus, lateral view; c—head and thorax, lateral view; d—labels; e—right fore wing pterostigma; f—right hind wing pterostigma. Not to scale. Courtesy of Benjamin Price." httpUri="https://zenodo.org/record/2624157/files/figure.png" pageId="2" pageNumber="69">Fig. 2e</figureCitation>
) and, respectively, 1.75 and 2.20 times as long as high in the hind wing (
<figureCitation id="13750BB23E732E7E0016D457FD47C15A" box="[632,715,1851,1876]" captionStart="FIGURE 2" captionStartId="3.[151,250,1674,1696]" captionTargetBox="[276,1312,337,1653]" captionTargetId="figure@3.[276,1312,337,1653]" captionTargetPageId="3" captionText="FIGURE 2. The holotype (female) of Indolestes anomalus (NHMUK 013324318) at BMNH: a—habitus, dorsal view; bhabitus, lateral view; c—head and thorax, lateral view; d—labels; e—right fore wing pterostigma; f—right hind wing pterostigma. Not to scale. Courtesy of Benjamin Price." httpUri="https://zenodo.org/record/2624157/files/figure.png" pageId="2" pageNumber="69">Fig. 2f</figureCitation>
). In both wings, the pterostigmata covers a bit less than two underlying cells (
<figureCitation id="13750BB23E732E7E0331D433FE41C176" box="[351,461,1887,1912]" captionStart="FIGURE 2" captionStartId="3.[151,250,1674,1696]" captionTargetBox="[276,1312,337,1653]" captionTargetId="figure@3.[276,1312,337,1653]" captionTargetPageId="3" captionText="FIGURE 2. The holotype (female) of Indolestes anomalus (NHMUK 013324318) at BMNH: a—habitus, dorsal view; bhabitus, lateral view; c—head and thorax, lateral view; d—labels; e—right fore wing pterostigma; f—right hind wing pterostigma. Not to scale. Courtesy of Benjamin Price." httpUri="https://zenodo.org/record/2624157/files/figure.png" pageId="2" pageNumber="69">Fig. 2e, f</figureCitation>
), but one and half cells in Fraser’s figure. Fraser clearly stated that he had only one specimen and the BMNH specimen labeled as the
<typeStatus id="54F5A9953E732E7E00A9D4EFFCA5C192" box="[711,809,1923,1948]" pageId="2" pageNumber="69" type="holotype">holotype</typeStatus>
of “
<taxonomicName id="4C4E6CB43E732E7E0137D4EFFBBFC192" authorityName="Fraser, Dalat" authorityYear="1946" box="[857,1075,1923,1948]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="anomala">
<emphasis id="B93ACB253E732E7E0137D4EFFBBFC192" box="[857,1075,1923,1948]" italics="true" pageId="2" pageNumber="69">Indolestes anomala</emphasis>
</taxonomicName>
” fits very well in other respects, e.g. the synthoracic pattern, thus excluding the possibility of other specimens. The left hind wing is missing in the
<typeStatus id="54F5A9953E732E7E02F9D4A7FF75C1EA" box="[151,249,1995,2020]" pageId="2" pageNumber="69" type="holotype">holotype</typeStatus>
(
<figureCitation id="13750BB23E732E7E0366D4A7FED6C1EA" box="[264,346,1995,2020]" captionStart="FIGURE 2" captionStartId="3.[151,250,1674,1696]" captionTargetBox="[276,1312,337,1653]" captionTargetId="figure@3.[276,1312,337,1653]" captionTargetPageId="3" captionText="FIGURE 2. The holotype (female) of Indolestes anomalus (NHMUK 013324318) at BMNH: a—habitus, dorsal view; bhabitus, lateral view; c—head and thorax, lateral view; d—labels; e—right fore wing pterostigma; f—right hind wing pterostigma. Not to scale. Courtesy of Benjamin Price." httpUri="https://zenodo.org/record/2624157/files/figure.png" pageId="2" pageNumber="69">Fig. 2a</figureCitation>
); perhaps it had a shorter pterostigma, which Fraser may have figured and he may have detached that wing for convenience. Some of the discrepancies mentioned above between Fraser’s own figure and his textual description of the pterostigmata proportions indicates that he likely did not aim for high degree of accuracy of his measurements and/or drawings. Other incorrect proportions in Fraser’s drawings are well known (e.g. the wrong shape of the paraprocts shown for the
<typeStatus id="54F5A9953E722E7F002CD268FD25C713" box="[578,681,260,285]" pageId="3" pageNumber="70" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C4E6CB43E722E7F00A3D269FB9BC712" authority="Fraser, 1924" authorityName="Fraser" authorityYear="1924" box="[717,1047,260,284]" class="Insecta" family="Lestidae" genus="Lestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="nigriceps">
<emphasis id="B93ACB253E722E7F00A3D269FC0FC712" box="[717,899,261,284]" italics="true" pageId="3" pageNumber="70">Lestes nigriceps</emphasis>
Fraser, 1924
</taxonomicName>
, see
<bibRefCitation id="EFDF6AC63E722E7F0623D268FB7EC712" author="Kosterin, O. E." box="[1101,1266,260,285]" pageId="3" pageNumber="70" pagination="561 - 575" refId="ref5316" refString="Kosterin, O. E. (2018) Rediscovery of Lestes nigriceps Fraser, 1924 (Odonata: Lestidae) in eastern Cambodia. Zootaxa, 4526 (4), 561 - 575. https: // doi. org / 10.11646 / zootaxa. 4526.4.8" type="journal article" year="2018">Kosterin 2018</bibRefCitation>
).
</paragraph>
<caption id="DF3147BF3E722E7F02F9D5E6FD4EC0D3" httpUri="https://zenodo.org/record/2624157/files/figure.png" pageId="3" pageNumber="70" startId="3.[151,250,1674,1696]" targetBox="[276,1312,337,1653]" targetPageId="3">
<paragraph id="8BF117373E722E7F02F9D5E6FD4EC0D3" blockId="3.[151,1436,1674,1757]" pageId="3" pageNumber="70">
<emphasis id="B93ACB253E722E7F02F9D5E6FE99C091" bold="true" box="[151,277,1674,1696]" pageId="3" pageNumber="70">FIGURE 2.</emphasis>
The holotype (female) of
<taxonomicName id="4C4E6CB43E722E7F0042D5E6FD72C0AE" box="[556,766,1674,1696]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E722E7F0042D5E6FD72C0AE" box="[556,766,1674,1696]" italics="true" pageId="3" pageNumber="70">Indolestes anomalus</emphasis>
</taxonomicName>
(NHMUK 013324318) at BMNH: a—habitus, dorsal view; bhabitus, lateral view; c—head and thorax, lateral view; d—labels; e—right fore wing pterostigma; f—right hind wing pterostigma. Not to scale. Courtesy of Benjamin Price.
</paragraph>
</caption>
<paragraph id="8BF117373E722E7802A9D466FC70C6DB" blockId="3.[151,1436,1802,2007]" lastBlockId="4.[151,1437,151,2015]" lastPageId="4" lastPageNumber="71" pageId="3" pageNumber="70">
<emphasis id="B93ACB253E722E7F02A9D466FD8DC12D" bold="true" box="[199,513,1802,1827]" pageId="3" pageNumber="70">A nearly topotypic female.</emphasis>
On
<date id="FFF031F73E722E7F005ED466FD1EC12D" box="[560,658,1802,1827]" pageId="3" pageNumber="70" value="2016-02-06">6 ii 2016</date>
one of us (T.K.) collected a female in Cát Tiên National Park (
<figureCitation id="13750BB23E722E7F0724D466FA02C12D" box="[1354,1422,1802,1827]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="3" pageNumber="70">Fig. 3</figureCitation>
). It fits, with minor differences, the original description well. It displays the short pterostigmata (
<figureCitation id="13750BB23E722E7F06A7D442FA91C149" box="[1225,1309,1838,1863]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="3" pageNumber="70">Fig. 3b</figureCitation>
), and has a pair of pale-coloured tubercles on the occiput, behind the lateral ocelli, a pair of dark spots on the anteclypeus, and a synthoracic pattern consisting of a posthumeral bronze stripe split into three irregular spots and a blunt projection of the dorsal bronze stripe at ca. three-fourths of its length (
<figureCitation id="13750BB23E722E7F0156D4F6FC06C1BD" box="[824,906,1946,1971]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="3" pageNumber="70">Fig. 3a</figureCitation>
). The central and posterior bronze spots on the mesepimeron are somewhat shorter (
<figureCitation id="13750BB23E722E7F0024D4D2FD13C1D9" box="[586,671,1982,2007]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="3" pageNumber="70">Fig. 3a</figureCitation>
) than in the
<typeStatus id="54F5A9953E722E7F012CD4D2FC28C1D9" box="[834,932,1982,2007]" pageId="3" pageNumber="70" type="holotype">holotype</typeStatus>
(
<figureCitation id="13750BB23E722E7F01D7D4D2FB9DC1D9" box="[953,1041,1982,2007]" captionStart="FIGURE 2" captionStartId="3.[151,250,1674,1696]" captionTargetBox="[276,1312,337,1653]" captionTargetId="figure@3.[276,1312,337,1653]" captionTargetPageId="3" captionText="FIGURE 2. The holotype (female) of Indolestes anomalus (NHMUK 013324318) at BMNH: a—habitus, dorsal view; bhabitus, lateral view; c—head and thorax, lateral view; d—labels; e—right fore wing pterostigma; f—right hind wing pterostigma. Not to scale. Courtesy of Benjamin Price." httpUri="https://zenodo.org/record/2624157/files/figure.png" pageId="3" pageNumber="70">Fig. 2c</figureCitation>
). The pale occipital spots extend lateroposteriorly of the pair of tubercles (
<figureCitation id="13750BB23E752E7800EAD3FBFD57C6BE" box="[644,731,151,176]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="4" pageNumber="71">Fig. 3a</figureCitation>
). The length of the hindwing is
<quantity id="4CB6BAD23E752E780609D3F4FB37C6A1" box="[1127,1211,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="71" unit="mm" value="20.0">20 mm</quantity>
(as in the original description); of the abdomen
<quantity id="4CB6BAD23E752E78038ED3D0FDC8C6DA" box="[480,580,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" pageId="4" pageNumber="71" unit="mm" value="28.5">28.5 mm</quantity>
(vs
<quantity id="4CB6BAD23E752E78001AD3D0FD48C6DA" box="[628,708,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="71" unit="mm" value="27.0">27 mm</quantity>
in the original description).
</paragraph>
<paragraph id="8BF117373E752E7802A9D3B3FB4EC713" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="71">
The hind wing pterostigma of the female from Cát Tiên is 1.63 and 1.78 times as long as high along the dorsal and ventral margins, respectively (
<figureCitation id="13750BB23E752E780077D268FDE0C713" box="[537,620,260,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="4" pageNumber="71">Fig. 3b</figureCitation>
), which is even shorter than in the
<typeStatus id="54F5A9953E752E78019AD268FBDAC713" box="[1012,1110,260,285]" pageId="4" pageNumber="71" type="holotype">holotype</typeStatus>
(
<figureCitation id="13750BB23E752E78060BD268FB39C713" box="[1125,1205,260,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1674,1696]" captionTargetBox="[276,1312,337,1653]" captionTargetId="figure@3.[276,1312,337,1653]" captionTargetPageId="3" captionText="FIGURE 2. The holotype (female) of Indolestes anomalus (NHMUK 013324318) at BMNH: a—habitus, dorsal view; bhabitus, lateral view; c—head and thorax, lateral view; d—labels; e—right fore wing pterostigma; f—right hind wing pterostigma. Not to scale. Courtesy of Benjamin Price." httpUri="https://zenodo.org/record/2624157/files/figure.png" pageId="4" pageNumber="71">Fig. 2f</figureCitation>
).
</paragraph>
<paragraph id="8BF117373E752E7802A9D24BFF4DC432" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="71">
The characters of the end of abdomen differ considerably from the original description, which states “segments 8 to 10 dark brown with obscure indications of large yellow lateral spots. Anal appendages brownish, shortly conical”. This statement cannot be verified since the
<typeStatus id="54F5A9953E752E78016DD203FCE9C786" box="[771,869,367,392]" pageId="4" pageNumber="71" type="holotype">holotype</typeStatus>
lacks S8–10. In the Cát Tiên female, S8 is dark dorsally and pale ventrally and posteriorly, with a pair of smaller dark lateral spots, all colour transitions of which are diffuse; S9 dark becoming paler laterally, while S10 is entirely pale (
<figureCitation id="13750BB23E752E7801D3D2DBFB82C7DE" box="[957,1038,439,464]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="4" pageNumber="71">Fig. 3c</figureCitation>
). Cerci are also pale with black tips and are conical but are as long as S10 (
<figureCitation id="13750BB23E752E78000ED2B0FD38C7FA" box="[608,692,476,501]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="4" pageNumber="71">Fig. 3c</figureCitation>
). The colorational differences could result from the post-mortem changes or from the teneral state of the
<typeStatus id="54F5A9953E752E780037D293FD32C416" box="[601,702,511,536]" pageId="4" pageNumber="71" type="holotype">holotype</typeStatus>
, while the Cát Tiên female was photographed upon capture (
<figureCitation id="13750BB23E752E780701D293FF38C432" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="4" pageNumber="71">Fig. 3c</figureCitation>
).
</paragraph>
<paragraph id="8BF117373E752E7802A9D12BFEE6C4A6" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="71">This was the only specimen collected during many visits to the park, nor has it been found during the extensive explorations of the area by James Holden (pers. comm.), so it may be rare. However, large parts of the park have not been surveyed.</paragraph>
<paragraph id="8BF117373E752E7802A9D1D8FD9FC21E" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="71">
<emphasis id="B93ACB253E752E7802A9D1D8FEEBC4C3" bold="true" box="[199,359,692,717]" pageId="4" pageNumber="71">Thai females.</emphasis>
We have five females from a small muddy pond at Tung Lui Lai,
<collectingRegion id="498AD9D53E752E780627D1D8FACAC4C3" box="[1097,1350,692,717]" country="Thailand" name="Chaiyaphum" pageId="4" pageNumber="71">Chaiyaphum Province</collectingRegion>
, central
<collectingCountry id="F35957A73E752E7802F9D1BBFF77C4FE" box="[151,251,727,752]" name="Thailand" pageId="4" pageNumber="71">Thailand</collectingCountry>
collected by R. W. and J. A. Garrison. One female was collected in tandem with a male and is illustrated here (
<figureCitation id="13750BB23E752E7802B9D190FE91C51A" box="[215,285,764,789]" captionStart="FIGURE 4" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[296,1246,481,1879]" captionTargetId="figure@6.[288,1298,481,1903]" captionTargetPageId="6" captionText="FIGURE 4. The female of Indolestes anomalus collected in tandem with a male in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—head top, dorsofrontal view; dsynthorax, ventral view; e—tip of abdomen, lateral view; f—right wings; g—fore wing tip; h—hind wing tip. Not to scale. Photos by Rosser Garrison." httpUri="https://zenodo.org/record/2624161/files/figure.png" pageId="4" pageNumber="71">Fig. 4</figureCitation>
). It has somewhat longer pterostigmata: 2.21 and 2.14 as long as high along the ventral margin in the fore and hind wings, respectively (
<figureCitation id="13750BB23E752E780070D073FD02C536" box="[542,654,799,824]" captionStart="FIGURE 4" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[296,1246,481,1879]" captionTargetId="figure@6.[288,1298,481,1903]" captionTargetPageId="6" captionText="FIGURE 4. The female of Indolestes anomalus collected in tandem with a male in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—head top, dorsofrontal view; dsynthorax, ventral view; e—tip of abdomen, lateral view; f—right wings; g—fore wing tip; h—hind wing tip. Not to scale. Photos by Rosser Garrison." httpUri="https://zenodo.org/record/2624161/files/figure.png" pageId="4" pageNumber="71">
Fig.
<quantity id="4CB6BAD23E752E78003DD04CFDFDC536" box="[595,625,800,824]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="4" pageNumber="71" unit="g" value="4.0">4g</quantity>
, h
</figureCitation>
), which fits the
<typeStatus id="54F5A9953E752E78012BD073FC27C536" box="[837,939,799,824]" pageId="4" pageNumber="71" type="holotype">holotype</typeStatus>
. The synthoracic pattern and such structural characters as a pair of pale-coloured occipital (postocellar) tubercles, although very slight (
<figureCitation id="13750BB23E752E7806B4D028FADFC552" box="[1242,1363,836,861]" captionStart="FIGURE 4" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[296,1246,481,1879]" captionTargetId="figure@6.[288,1298,481,1903]" captionTargetPageId="6" captionText="FIGURE 4. The female of Indolestes anomalus collected in tandem with a male in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—head top, dorsofrontal view; dsynthorax, ventral view; e—tip of abdomen, lateral view; f—right wings; g—fore wing tip; h—hind wing tip. Not to scale. Photos by Rosser Garrison." httpUri="https://zenodo.org/record/2624161/files/figure.png" pageId="4" pageNumber="71">Fig. 4b, c</figureCitation>
), also correspond to
<emphasis id="B93ACB253E752E780352D005FE45C58E" box="[316,457,871,896]" italics="true" pageId="4" pageNumber="71">
<taxonomicName id="4C4E6CB43E752E780352D005FE4AC58E" box="[316,454,871,896]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="anomalus">I. anomalus</taxonomicName>
,
</emphasis>
and the postocellar light spots are confined to the immediate circumference of those (
<figureCitation id="13750BB23E752E7802F1D0E0FE81C5AA" box="[159,269,908,933]" captionStart="FIGURE 4" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[296,1246,481,1879]" captionTargetId="figure@6.[288,1298,481,1903]" captionTargetPageId="6" captionText="FIGURE 4. The female of Indolestes anomalus collected in tandem with a male in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—head top, dorsofrontal view; dsynthorax, ventral view; e—tip of abdomen, lateral view; f—right wings; g—fore wing tip; h—hind wing tip. Not to scale. Photos by Rosser Garrison." httpUri="https://zenodo.org/record/2624161/files/figure.png" pageId="4" pageNumber="71">Fig. 4b, c</figureCitation>
). The coloration of the end of the abdomen is similar to the Cát Tiên female, with S8 somewhat darker; and the cerci too are as long as S10 (
<figureCitation id="13750BB23E752E780023D0C3FD2EC5C6" box="[589,674,943,968]" captionStart="FIGURE 4" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[296,1246,481,1879]" captionTargetId="figure@6.[288,1298,481,1903]" captionTargetPageId="6" captionText="FIGURE 4. The female of Indolestes anomalus collected in tandem with a male in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—head top, dorsofrontal view; dsynthorax, ventral view; e—tip of abdomen, lateral view; f—right wings; g—fore wing tip; h—hind wing tip. Not to scale. Photos by Rosser Garrison." httpUri="https://zenodo.org/record/2624161/files/figure.png" pageId="4" pageNumber="71">Fig. 4e</figureCitation>
). The posthumeral dark spots are reduced in comparison with the
<typeStatus id="54F5A9953E752E7802F9D0B8FF75C5E3" box="[151,249,980,1005]" pageId="4" pageNumber="71" type="holotype">holotype</typeStatus>
and Cát Tiên female: the posterior one narrow but parallel-sided, without a ‘neck’, the middle one small, the anterior one missing (
<figureCitation id="13750BB23E752E7803DAD09BFD8AC21E" box="[436,518,1015,1040]" captionStart="FIGURE 4" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[296,1246,481,1879]" captionTargetId="figure@6.[288,1298,481,1903]" captionTargetPageId="6" captionText="FIGURE 4. The female of Indolestes anomalus collected in tandem with a male in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—head top, dorsofrontal view; dsynthorax, ventral view; e—tip of abdomen, lateral view; f—right wings; g—fore wing tip; h—hind wing tip. Not to scale. Photos by Rosser Garrison." httpUri="https://zenodo.org/record/2624161/files/figure.png" pageId="4" pageNumber="71">Fig. 4a</figureCitation>
).
</paragraph>
<paragraph id="8BF117373E752E7802A9D771FBA4C23B" blockId="4.[151,1437,151,2015]" box="[199,1064,1052,1077]" pageId="4" pageNumber="71">Measurements (N=5, in mm): Hw: 20–22.5, Abd. 27–31, total length: 35–38.</paragraph>
<paragraph id="8BF117373E752E7802A9D753FAD1C2E6" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="71">
The photos from Nam Nao National Park in neighbouring
<collectingRegion id="498AD9D53E752E78013FD753FBB2C256" box="[849,1086,1087,1112]" country="Thailand" name="Phetchabun" pageId="4" pageNumber="71">Phetchabun Province</collectingRegion>
by Dennis Farrell show mature females (
<figureCitation id="13750BB23E752E78036ED708FEFDC273" box="[256,369,1124,1149]" captionStart="FIGURE 5" captionStartId="7.[151,250,1941,1963]" captionTargetBox="[445,1150,193,1921]" captionTargetId="figure@7.[445,1150,193,1921]" captionTargetPageId="7" captionText="FIGURE 5. Indolestes anomalus photographed in nature in Thailand in Chaiyaphum Province, Phu Khieo Wildlife Sanctuary, 12 v 2016 (a) and Phetchabun Province, Nam Nao National Park, Suan Son Ban Baek, 15 viii 2015 (b–d); a, b—mature males; c—mature female; d—ovipositing tandem. Photo by Dennis Farrell." httpUri="https://zenodo.org/record/2624165/files/figure.png" pageId="4" pageNumber="71">Fig. 5c, d</figureCitation>
) in which the synthorax has a saturated blue ground colour, while that of the prothorax and mesepisternum is brownish. The pale colour of S1–3 and S10 is also blue, while S4–9 is brownish with indistinct darker dorsal spots expanding at the segment joints, with blue present only on the anterior margins of S4–7; the eyes are brownish-grey. Teneral females on his photos (not reproduced) have a brownish-grey ground colour.
</paragraph>
<paragraph id="8BF117373E752E7802A9D798FAB2C33E" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="71">
<emphasis id="B93ACB253E752E7802A9D798FE87C303" bold="true" box="[199,267,1268,1293]" pageId="4" pageNumber="71">Male.</emphasis>
Since the male has not been described and its diagnostic characters remain unknown, we provide below a description of a male (
<figureCitation id="13750BB23E752E7803CBD67BFE79C33E" box="[421,501,1303,1328]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[287,1278,511,1836]" captionTargetId="figure@8.[287,1299,471,1887]" captionTargetPageId="8" captionText="FIGURE 6. The male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison." httpUri="https://zenodo.org/record/2624167/files/figure.png" pageId="4" pageNumber="71">Figs. 6</figureCitation>
,
<figureCitation id="13750BB23E752E78006ED674FD83C33E" box="[512,527,1304,1328]" captionStart="FIGURE 7" captionStartId="9.[151,250,1434,1456]" captionTargetBox="[233,1294,242,1333]" captionTargetId="figure@9.[229,1358,208,1405]" captionTargetPageId="9" captionText="FIGURE 7. The structure of the genital ligula of a male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996: a—ectal view; b—details of ectal arm, ectolateral view; c—lateral view. (Drawings by Rosser Garrison)." httpUri="https://zenodo.org/record/2624169/files/figure.png" pageId="4" pageNumber="71">7</figureCitation>
) collected in tandem with a female at Chayaphum Province of
<collectingCountry id="F35957A73E752E7806BDD67BFAB6C33E" box="[1235,1338,1303,1328]" name="Thailand" pageId="4" pageNumber="71">Thailand</collectingCountry>
.
</paragraph>
<paragraph id="8BF117373E752E7802A9D650FE00C376" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="71">
<emphasis id="B93ACB253E752E7802A9D650FEABC35B" box="[199,295,1340,1365]" italics="true" pageId="4" pageNumber="71">Habitus.</emphasis>
Resembles the female, ground colour pale-bluish (further referred to as ‘pale’), dark pattern bronzeblack (further ‘dark’).
</paragraph>
<paragraph id="8BF117373E752E7802A9D6E8FAEBC3CE" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="71">
<emphasis id="B93ACB253E752E7802A9D6E8FE88C393" box="[199,260,1412,1437]" italics="true" pageId="4" pageNumber="71">Head</emphasis>
. Labrum, base of mandibles, genae and anteclypeus pale; postclypeus pale anteriorly, dark posteriorly, with a pair of large semicircular spots touching each other at base; epicranium and rear of head dark (
<figureCitation id="13750BB23E752E780768D6CBFAD6C3CE" box="[1286,1370,1447,1472]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[287,1278,511,1836]" captionTargetId="figure@8.[287,1299,471,1887]" captionTargetPageId="8" captionText="FIGURE 6. The male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison." httpUri="https://zenodo.org/record/2624167/files/figure.png" pageId="4" pageNumber="71">Fig. 6b</figureCitation>
).
</paragraph>
<paragraph id="8BF117373E752E7802A9D6A0FE9EC023" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="71">
<emphasis id="B93ACB253E752E7802A9D6A0FE93C3EB" box="[199,287,1484,1509]" italics="true" pageId="4" pageNumber="71">Thorax.</emphasis>
Prothorax pale dorsally, with following areas dark: a pair of dorsolateral spots on middle lobe, wash of brown along dorsal margin of pleuron and a pair of ill-defined dorsolateral brown spots on rounded posterior lobe (
<figureCitation id="13750BB23E752E7802F1D578FE89C023" box="[159,261,1556,1581]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[287,1278,511,1836]" captionTargetId="figure@8.[287,1299,471,1887]" captionTargetPageId="8" captionText="FIGURE 6. The male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison." httpUri="https://zenodo.org/record/2624167/files/figure.png" pageId="4" pageNumber="71">Fig. 6a,c</figureCitation>
).
</paragraph>
<paragraph id="8BF117373E752E7802A9D55BFE19C143" blockId="4.[151,1437,151,2015]" pageId="4" pageNumber="71">
Synthorax (
<figureCitation id="13750BB23E752E780325D55BFE30C05E" box="[331,444,1591,1616]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[287,1278,511,1836]" captionTargetId="figure@8.[287,1299,471,1887]" captionTargetPageId="8" captionText="FIGURE 6. The male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison." httpUri="https://zenodo.org/record/2624167/files/figure.png" pageId="4" pageNumber="71">Fig. 6c, d</figureCitation>
). Mesepisternum with dark middorsal stripe occupying over half of mesepisternum, its sides parallel except for a small rounded lobe almost touching mesopleural suture at upper fifth and again at upper margin and with an offshoot extending posteriorly over mesopleural fossa; mesepimeron pale with following areas dark: irregular elongate spot extending posteriorly from mesopleural fossa and curving ventrally to level of semicircular spot of middorsal stripe, isolated irregular spot at middle of mesopleural suture and a small subtriangular spot at juncture of mesinfraepisternum, dorsal carina and upper fifth of interpleural and metapleural sutures dark; a narrow elongate dark spot bordering posterior margin of pale metinfraepisternum (
<figureCitation id="13750BB23E752E780683D463FACCC126" box="[1261,1344,1807,1832]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[287,1278,511,1836]" captionTargetId="figure@8.[287,1299,471,1887]" captionTargetPageId="8" captionText="FIGURE 6. The male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison." httpUri="https://zenodo.org/record/2624167/files/figure.png" pageId="4" pageNumber="71">Fig. 6c</figureCitation>
); venter of thorax as in
<figureCitation id="13750BB23E752E780351D458FE1DC143" box="[319,401,1844,1869]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[287,1278,511,1836]" captionTargetId="figure@8.[287,1299,471,1887]" captionTargetPageId="8" captionText="FIGURE 6. The male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison." httpUri="https://zenodo.org/record/2624167/files/figure.png" pageId="4" pageNumber="71">Fig. 6d</figureCitation>
.
</paragraph>
<paragraph id="8BF117373E752E7802A9D434FAA0C17E" blockId="4.[151,1437,151,2015]" box="[199,1324,1879,1904]" pageId="4" pageNumber="71">
Coxae pale but diffusely brownish anteriorly. Femora black, becoming brown externally (
<figureCitation id="13750BB23E752E7806DCD43BFA92C17E" box="[1202,1310,1879,1904]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[287,1278,511,1836]" captionTargetId="figure@8.[287,1299,471,1887]" captionTargetPageId="8" captionText="FIGURE 6. The male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison." httpUri="https://zenodo.org/record/2624167/files/figure.png" pageId="4" pageNumber="71">Fig. 6a, c</figureCitation>
).
</paragraph>
<paragraph id="8BF117373E752E7902A9D411FD77C6DB" blockId="4.[151,1437,151,2015]" lastBlockId="5.[151,1436,151,213]" lastPageId="5" lastPageNumber="72" pageId="4" pageNumber="71">
<emphasis id="B93ACB253E752E7802A9D411FE86C19A" box="[199,266,1917,1940]" italics="true" pageId="4" pageNumber="71">Wings</emphasis>
hyaline; venation brownish. Postnodals: FW 10/11, Hw 10/10; RP
<subScript id="17CA15723E752E780192D4E7FB88C197" attach="left" box="[1020,1028,1931,1945]" fontSize="6" pageId="4" pageNumber="71">2</subScript>
at Fw starts at postnodal 4/4, at Hw at postnodal 3/3. Pterostigmata: FW: 1.6 2–1.69/1.87–1.95 times as long as high (along dorsal/ventral side); HW 1.72–1.94/1.84–1.98 times as long as high, surmounting surmounting 1.5/cells in both wings. In one male specimen, costal vein at pterostigma is noticeably convex at FW and distinctly convex at HW; in the other male specimen it is only very slightly convex at both wings.
</paragraph>
<caption id="DF3147BF3E742E7902F9D4EEFB14C1B8" httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="5" pageNumber="72" startId="5.[151,250,1922,1944]" targetBox="[151,1435,282,1922]" targetPageId="5">
<paragraph id="8BF117373E742E7902F9D4EEFB14C1B8" blockId="5.[151,1435,1922,1974]" pageId="5" pageNumber="72">
<emphasis id="B93ACB253E742E7902F9D4EEFE9FC199" bold="true" box="[151,275,1922,1944]" pageId="5" pageNumber="72">FIGURE 3.</emphasis>
A nearly topotypical female of
<taxonomicName id="4C4E6CB43E742E790039D4EEFCABC196" box="[599,807,1922,1944]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E742E790039D4EEFCABC196" box="[599,807,1922,1944]" italics="true" pageId="5" pageNumber="72">Indolestes anomalus</emphasis>
</taxonomicName>
from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier.
</paragraph>
</caption>
<paragraph id="8BF117373E772E7A02A9D3FBFDF7C76B" blockId="6.[151,1436,151,429]" pageId="6" pageNumber="73">
<emphasis id="B93ACB253E772E7A02A9D3FBFEBDC6BE" box="[199,305,151,176]" italics="true" pageId="6" pageNumber="73">Abdomen</emphasis>
with the following dark markings (
<figureCitation id="13750BB23E772E7A00A9D3FBFC90C6BE" box="[711,796,151,176]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[287,1278,511,1836]" captionTargetId="figure@8.[287,1299,471,1887]" captionTargetPageId="8" captionText="FIGURE 6. The male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison." httpUri="https://zenodo.org/record/2624167/files/figure.png" pageId="6" pageNumber="73">Fig. 6a</figureCitation>
): S1 with a small dorsal quadrate spot at basal 0.30 and with narrow apical ring; S3–7 with a long dorsal stripe beginning at basal tenth, its base moderately expanded proximally and with a quadrate expansion laterally at distal end (that of S2 shaped as an inverted key-hole), but almost interrupted dorsally by an incomplete apical ring; basal 0.60 of S8 black, distal 0.30 pale, narrowly divided above by thin inverted triangular dark spot; S9 with irregular dark narrow basal ring, otherwise pale; S10 entirely pale, with a deep triangular dorsal incision.
</paragraph>
<paragraph id="8BF117373E772E7A02A9D203FD94C7A3" blockId="6.[151,1436,151,429]" pageId="6" pageNumber="73">
<emphasis id="B93ACB253E772E7A02A9D203FEE8C786" box="[199,356,367,392]" italics="true" pageId="6" pageNumber="73">Genital ligula</emphasis>
(
<figureCitation id="13750BB23E772E7A031AD203FE35C786" box="[372,441,367,392]" captionStart="FIGURE 7" captionStartId="9.[151,250,1434,1456]" captionTargetBox="[233,1294,242,1333]" captionTargetId="figure@9.[229,1358,208,1405]" captionTargetPageId="9" captionText="FIGURE 7. The structure of the genital ligula of a male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996: a—ectal view; b—details of ectal arm, ectolateral view; c—lateral view. (Drawings by Rosser Garrison)." httpUri="https://zenodo.org/record/2624169/files/figure.png" pageId="6" pageNumber="73">Fig. 7</figureCitation>
) of usual lestine form, basal segment ovate, ectal arm with small semihyaline coiled tip that is grooved entally and ectally.
</paragraph>
<caption id="DF3147BF3E772E7A02F9D413FE21C1FC" httpUri="https://zenodo.org/record/2624161/files/figure.png" pageId="6" pageNumber="73" startId="6.[151,250,1919,1941]" targetBox="[296,1246,481,1879]" targetPageId="6">
<paragraph id="8BF117373E772E7A02F9D413FE21C1FC" blockId="6.[151,1437,1919,2034]" pageId="6" pageNumber="73">
<emphasis id="B93ACB253E772E7A02F9D413FE98C19A" bold="true" box="[151,276,1919,1941]" pageId="6" pageNumber="73">FIGURE 4.</emphasis>
The female of
<taxonomicName id="4C4E6CB43E772E7A03DAD413FD09C19B" box="[436,645,1919,1941]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="73" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E772E7A03DAD413FD09C19B" box="[436,645,1919,1941]" italics="true" pageId="6" pageNumber="73">Indolestes anomalus</emphasis>
</taxonomicName>
collected in tandem with a male in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—head top, dorsofrontal view; dsynthorax, ventral view; e—tip of abdomen, lateral view; f—right wings; g—fore wing tip; h—hind wing tip. Not to scale. Photos by Rosser Garrison.
</paragraph>
</caption>
<caption id="DF3147BF3E762E7B02F9D4F9FCC6C1E6" httpUri="https://zenodo.org/record/2624165/files/figure.png" pageId="7" pageNumber="74" startId="7.[151,250,1941,1963]" targetBox="[445,1150,193,1921]" targetPageId="7">
<paragraph id="8BF117373E762E7B02F9D4F9FCC6C1E6" blockId="7.[151,1436,1941,2024]" pageId="7" pageNumber="74">
<emphasis id="B93ACB253E762E7B02F9D4F9FE9FC1A4" bold="true" box="[151,275,1941,1963]" pageId="7" pageNumber="74">FIGURE 5.</emphasis>
<taxonomicName id="4C4E6CB43E762E7B0377D4F9FE65C1A5" box="[281,489,1941,1963]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="74" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E762E7B0377D4F9FE65C1A5" box="[281,489,1941,1963]" italics="true" pageId="7" pageNumber="74">Indolestes anomalus</emphasis>
</taxonomicName>
photographed in nature in Thailand in Chaiyaphum Province, Phu Khieo Wildlife Sanctuary, 12 v 2016 (a) and Phetchabun Province, Nam Nao National Park, Suan Son Ban Baek, 15 viii 2015 (b–d); a, b—mature males; c—mature female; d—ovipositing tandem. Photo by Dennis Farrell.
</paragraph>
</caption>
<paragraph id="8BF117373E792E7402A9D3F4FB5AC6F9" blockId="8.[151,1437,151,429]" pageId="8" pageNumber="75">
<emphasis id="B93ACB253E792E7402A9D3F4FE9AC6A1" box="[199,278,152,175]" italics="true" pageId="8" pageNumber="75">Cercus</emphasis>
(
<figureCitation id="13750BB23E792E740346D3FBFEE2C6BE" box="[296,366,151,176]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[287,1278,511,1836]" captionTargetId="figure@8.[287,1299,471,1887]" captionTargetPageId="8" captionText="FIGURE 6. The male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison." httpUri="https://zenodo.org/record/2624167/files/figure.png" pageId="8" pageNumber="75">Fig. 6</figureCitation>
e–g) with proximal half pale, becoming blackish-brown distally, mesal margin with a small semicircular lobe followed distally by a strong posteromedially directed tooth, apical third of cercus slightly swollen and attenuate distally; paraproct semicircular, black, about one-fifth the length of cercus.
</paragraph>
<paragraph id="8BF117373E792E7402A9D269FBB0C712" blockId="8.[151,1437,151,429]" box="[199,1084,260,285]" pageId="8" pageNumber="75">Measurements (in mm): HW 20, Abdomen (including cerci) 32, total length 38</paragraph>
<paragraph id="8BF117373E792E7402A9D24BFB1CC7A2" blockId="8.[151,1437,151,429]" pageId="8" pageNumber="75">
The other male specimen from the same series shows no significant differences. Mature males photographed by Dennis Farrell (
<figureCitation id="13750BB23E792E740301D220FE77C76B" box="[367,507,332,357]" captionStart="FIGURE 5" captionStartId="7.[151,250,1941,1963]" captionTargetBox="[445,1150,193,1921]" captionTargetId="figure@7.[445,1150,193,1921]" captionTargetPageId="7" captionText="FIGURE 5. Indolestes anomalus photographed in nature in Thailand in Chaiyaphum Province, Phu Khieo Wildlife Sanctuary, 12 v 2016 (a) and Phetchabun Province, Nam Nao National Park, Suan Son Ban Baek, 15 viii 2015 (b–d); a, b—mature males; c—mature female; d—ovipositing tandem. Photo by Dennis Farrell." httpUri="https://zenodo.org/record/2624165/files/figure.png" pageId="8" pageNumber="75">Fig. 5a, b, d</figureCitation>
) have a saturated blue ground colour and dark-blue eyes with a green hue below, while these are brownish-grey in teneral ones (not shown); the expansion of the dorsal synthoracic dark stripe sometimes reaches the mesopleural suture and merges with the posterior posthumeral spot.
</paragraph>
<caption id="DF3147BF3E792E7402F9D41CFE2FC1E2" httpUri="https://zenodo.org/record/2624167/files/figure.png" pageId="8" pageNumber="75" startId="8.[151,250,1904,1926]" targetBox="[287,1278,511,1836]" targetPageId="8">
<paragraph id="8BF117373E792E7402F9D41CFE2FC1E2" blockId="8.[151,1436,1904,2028]" pageId="8" pageNumber="75">
<emphasis id="B93ACB253E792E7402F9D41CFE9EC18B" bold="true" box="[151,274,1904,1926]" pageId="8" pageNumber="75">FIGURE 6.</emphasis>
The male of
<taxonomicName id="4C4E6CB43E792E7403F9D41CFDEBC188" box="[407,615,1904,1926]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="75" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E792E7403F9D41CFDEBC188" box="[407,615,1904,1926]" italics="true" pageId="8" pageNumber="75">Indolestes anomalus</emphasis>
</taxonomicName>
collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison.
</paragraph>
</caption>
<caption id="DF3147BF3E782E7502F9D6F6FCBCC3E3" httpUri="https://zenodo.org/record/2624169/files/figure.png" pageId="9" pageNumber="76" startId="9.[151,250,1434,1456]" targetBox="[233,1294,242,1333]" targetPageId="9">
<paragraph id="8BF117373E782E7502F9D6F6FCBCC3E3" blockId="9.[151,1436,1434,1517]" pageId="9" pageNumber="76">
<emphasis id="B93ACB253E782E7502F9D6F6FE9EC3A1" bold="true" box="[151,274,1434,1456]" pageId="9" pageNumber="76">FIGURE 7.</emphasis>
The structure of the genital ligula of a male of
<taxonomicName id="4C4E6CB43E782E750085D6F6FC36C3BE" box="[747,954,1434,1456]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="76" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E782E750085D6F6FC36C3BE" box="[747,954,1434,1456]" italics="true" pageId="9" pageNumber="76">Indolestes anomalus</emphasis>
</taxonomicName>
collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996: a—ectal view; b—details of ectal arm, ectolateral view; c—lateral view. (Drawings by Rosser Garrison).
</paragraph>
</caption>
<paragraph id="8BF117373E782E7502A9D576FD30C0CD" blockId="9.[151,1436,1562,1911]" pageId="9" pageNumber="76">
<emphasis id="B93ACB253E782E7502A9D576FEAFC03D" bold="true" box="[199,291,1562,1587]" pageId="9" pageNumber="76">Habitat</emphasis>
. In Cát Tiên National Park in
<collectingCountry id="F35957A73E782E7500FED576FD63C03D" box="[656,751,1562,1587]" name="Vietnam" pageId="9" pageNumber="76">Vietnam</collectingCountry>
the female was found in January, which is during the dry season, in seasonally humid tropical forest with dense and thorny undergrowth. In Chayaphum Province of
<collectingCountry id="F35957A73E782E7502F9D50EFF77C075" box="[151,251,1634,1659]" name="Thailand" pageId="9" pageNumber="76">Thailand</collectingCountry>
the species was found by R.G. at a small muddy pond. In
<collectingRegion id="498AD9D53E782E7501CAD50EFB14C075" box="[932,1176,1634,1659]" country="Thailand" name="Phetchabun" pageId="9" pageNumber="76">Phetchabun Province</collectingRegion>
, in Nam Nao National park, the species was regularly found in forest ponds, including a large one with reeds, from January to August, sometimes as extremely abundant (
<bibRefCitation id="EFDF6AC63E782E75004ED5C6FD3DC0CD" author="Farrell, D." box="[544,689,1706,1731]" pageId="9" pageNumber="76" refId="ref4984" refString="Farrell, D. (2018) Dragonflies &amp; Damselflies of Thailand. Available from: http: // thaiodonata. blogspot. ru / (accessed 19 February 2018)" type="url" year="2018">Farrell 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35444BC3E782E7502A9D5A2FC9BC179" pageId="9" pageNumber="76" type="distribution">
<paragraph id="8BF117373E782E7502A9D5A2FC9BC179" blockId="9.[151,1436,1562,1911]" pageId="9" pageNumber="76">
<emphasis id="B93ACB253E782E7502A9D5A2FEECC0E9" bold="true" box="[199,352,1742,1767]" pageId="9" pageNumber="76">Distribution.</emphasis>
The species is presently known from three separated areas: northern
<collectingCountry id="F35957A73E782E750612D5A2FB6CC0E9" box="[1148,1248,1742,1767]" name="Thailand" pageId="9" pageNumber="76">Thailand</collectingCountry>
(
<collectingRegion id="498AD9D53E782E75069ED5A2FA92C0E9" box="[1264,1310,1742,1767]" country="Thailand" name="Tak" pageId="9" pageNumber="76">Tak</collectingRegion>
,
<collectingRegion id="498AD9D53E782E750745D5A3FA14C0E9" box="[1323,1432,1743,1767]" country="Thailand" name="Lampang" pageId="9" pageNumber="76">Lampang</collectingRegion>
,
<collectingRegion id="498AD9D53E782E7502F9D59EFE96C105" box="[151,282,1778,1803]" country="Thailand" name="Phetchabun" pageId="9" pageNumber="76">Phetchabun</collectingRegion>
and
<collectingRegion id="498AD9D53E782E750338D59EFE69C105" box="[342,485,1778,1803]" country="Thailand" name="Chaiyaphum" pageId="9" pageNumber="76">Chaiyaphum</collectingRegion>
Provinces), southern
<collectingCountry id="F35957A73E782E75008AD59EFCCFC105" box="[740,835,1778,1803]" name="Vietnam" pageId="9" pageNumber="76">Vietnam</collectingCountry>
(Đỗng Nai and possibly Lâm Đỗng Provinces) and Cameron Highlands in Peninsular
<collectingCountry id="F35957A73E782E750079D47AFDF1C121" box="[535,637,1814,1839]" name="Malaysia" pageId="9" pageNumber="76">Malaysia</collectingCountry>
(
<bibRefCitation id="EFDF6AC63E782E7500E5D47BFC9AC121" author="Fraser, F. C." box="[651,790,1814,1839]" pageId="9" pageNumber="76" pagination="41 - 45" refId="ref5050" refString="Fraser, F. C. (1946) On rare and two new species of Odonata from tropical Africa and Asia. Proceedings of Royal Entomological Society of London. Series B, 15 (3 / 4), 41 - 45. https: // doi. org / 10.1111 / j. 1365 - 3113.1946. tb 00815. x" type="journal article" year="1946">Fraser 1946</bibRefCitation>
;
<bibRefCitation id="EFDF6AC63E782E75014ED47BFC5CC121" author="Day, L. &amp; Farrell, D. &amp; Guenther, A. &amp; Hamalainen, M. &amp; Klimsa, E. &amp; Korshunov, A. &amp; Kosterin, O. &amp; Makbun N. &amp; Pelegrin, A. &amp; Roeder, U. &amp; Ruangrong R. &amp; Vikhrev, N." box="[800,976,1814,1839]" pageId="9" pageNumber="76" pagination="16 - 24" refId="ref4865" refString="Day, L., Farrell, D., Guenther, A., Hamalainen, M., Klimsa, E., Korshunov, A., Kosterin, O., Makbun N., Pelegrin, A., Roeder, U., Ruangrong R. &amp; Vikhrev, N. (2012) New provincial records of Odonata from Thailand mostly based on photographs. Agrion, 16 (1), 16 - 24." type="journal article" year="2012">
Day
<emphasis id="B93ACB253E782E750138D47BFC02C121" box="[854,910,1814,1839]" italics="true" pageId="9" pageNumber="76">et al.</emphasis>
2012
</bibRefCitation>
;
<bibRefCitation id="EFDF6AC63E782E7501B4D47BFBF7C121" author="Ng, J. F. &amp; Dow, R. W. &amp; Choong, C. Y." box="[986,1147,1814,1839]" pageId="9" pageNumber="76" pagination="9 - 16" refId="ref5415" refString="Ng, J. F., Dow, R. W. &amp; Choong, C. Y. (2011) New records of Odonata (Insecta) from the Cameron Highlands, with first records of two species for Malaysia. Journal of Science and Technology in Tropics, 7, 9 - 16." type="journal article" year="2011">
Ng
<emphasis id="B93ACB253E782E75066DD47BFBB7C121" box="[1027,1083,1814,1839]" italics="true" pageId="9" pageNumber="76">et al.</emphasis>
2011
</bibRefCitation>
;
<bibRefCitation id="EFDF6AC63E782E7506E8D47AFA9FC121" author="Farrell, D." box="[1158,1299,1814,1839]" pageId="9" pageNumber="76" refId="ref4984" refString="Farrell, D. (2018) Dragonflies &amp; Damselflies of Thailand. Available from: http: // thaiodonata. blogspot. ru / (accessed 19 February 2018)" type="url" year="2018">Farrell 2018</bibRefCitation>
; this paper), as shown in
<figureCitation id="13750BB23E782E75034AD456FEEBC15D" box="[292,359,1850,1875]" captionStart="FIGURE 8" captionStartId="10.[151,250,1345,1367]" captionTargetBox="[197,1396,229,1283]" captionTargetId="figure@10.[151,1436,193,1324]" captionTargetPageId="10" captionText="FIGURE 8. The known distribution of Indolestes anomalus. A—the locality of the nearly topotypical female shown in Fig. 3 (Vietnam, Đỗng Nai Province, Cát Tiên National Park) and approximately the presumed type locality; B—the locality of male and female shown in Figs. 4–7 (Thailand, Chaiyaphum Province, at Tung Lui Lai); C—the locality of a males shown in Fig. 6a (Thailand, Chaiyaphum Province, Phu Khieo Wildlife Sanctuary); D—the locality of individuals shown in Fig. 6b–d (Thailand, Phetchabun Province, Nam Nao National Park)." httpUri="https://zenodo.org/record/2624171/files/figure.png" pageId="9" pageNumber="76">Fig. 8</figureCitation>
. It was observed as abundant only in
<collectingRegion id="498AD9D53E782E75017ED456FC72C15D" box="[784,1022,1850,1875]" country="Thailand" name="Phetchabun" pageId="9" pageNumber="76">Phetchabun Province</collectingRegion>
of
<collectingCountry id="F35957A73E782E75064AD456FB04C15D" box="[1060,1160,1850,1875]" name="Thailand" pageId="9" pageNumber="76">Thailand</collectingCountry>
(
<bibRefCitation id="EFDF6AC63E782E7506F9D456FAA4C15D" author="Farrell, D." box="[1175,1320,1850,1875]" pageId="9" pageNumber="76" refId="ref4984" refString="Farrell, D. (2018) Dragonflies &amp; Damselflies of Thailand. Available from: http: // thaiodonata. blogspot. ru / (accessed 19 February 2018)" type="url" year="2018">Farrell 2018</bibRefCitation>
). It can be expected in
<collectingCountry id="F35957A73E782E750370D432FE18C179" box="[286,404,1886,1911]" name="Cambodia" pageId="9" pageNumber="76">Cambodia</collectingCountry>
, which lies between these regions.
</paragraph>
</subSubSection>
<caption id="DF3147BF3E7B2E7602F9D62DFDF1C3DC" httpUri="https://zenodo.org/record/2624171/files/figure.png" pageId="10" pageNumber="77" startId="10.[151,250,1345,1367]" targetBox="[197,1396,229,1283]" targetPageId="10">
<paragraph id="8BF117373E7B2E7602F9D62DFDF1C3DC" blockId="10.[151,1436,1345,1490]" pageId="10" pageNumber="77">
<emphasis id="B93ACB253E7B2E7602F9D62DFE9FC358" bold="true" box="[151,275,1345,1367]" pageId="10" pageNumber="77">FIGURE 8.</emphasis>
The known distribution of
<taxonomicName id="4C4E6CB43E7B2E760042D62DFD71C359" box="[556,765,1345,1367]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E7B2E760042D62DFD71C359" box="[556,765,1345,1367]" italics="true" pageId="10" pageNumber="77">Indolestes anomalus</emphasis>
</taxonomicName>
. A—the locality of the nearly topotypical female shown in Fig. 3 (Vietnam, Đỗng Nai Province, Cát Tiên National Park) and approximately the presumed type locality; B—the locality of male and female shown in Figs. 4–7 (Thailand, Chaiyaphum Province, at Tung Lui Lai); C—the locality of a males shown in Fig. 6a (Thailand, Chaiyaphum Province, Phu Khieo Wildlife Sanctuary); D—the locality of individuals shown in Fig. 6b–d (Thailand, Phetchabun Province, Nam Nao National Park).
</paragraph>
</caption>
<subSubSection id="C35444BC3E7B2E7702A9D692FD49C7FA" lastPageId="11" lastPageNumber="78" pageId="10" pageNumber="77" type="diagnosis">
<paragraph id="8BF117373E7B2E7702A9D692FC59C6F6" blockId="10.[151,1436,1534,2028]" lastBlockId="11.[151,1436,151,501]" lastPageId="11" lastPageNumber="78" pageId="10" pageNumber="77">
<emphasis id="B93ACB253E7B2E7602A9D692FECDC019" bold="true" box="[199,321,1534,1559]" pageId="10" pageNumber="77">Diagnosis.</emphasis>
<taxonomicName id="4C4E6CB43E7B2E760327D692FDA3C019" box="[329,559,1534,1559]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E7B2E760327D692FDA3C019" box="[329,559,1534,1559]" italics="true" pageId="10" pageNumber="77">Indolestes anomalus</emphasis>
</taxonomicName>
is characterized by the synthoracic pattern with (i) the middorsal black stripe having a small extension at ca. three-fourths of its length and posteriorly expanding to the mesopleural suture and (ii) three irregular and variable posthumeral dark spots: an elongate dorsal one followed by a smaller quadrate one and finally by a smaller triangular spot at the margin of mesinfraepisternum (this small spot may be missing) (
<figureCitation id="13750BB23E7B2E76070AD507FF3EC0A9" captionStart="FIGURE 1" captionStartId="2.[151,250,1507,1529]" captionTargetBox="[340,1226,424,1423]" captionTargetId="figure@2.[340,1247,267,1483]" captionTargetPageId="2" captionText="FIGURE 1. The holotype (female) of Indolestes anomalus as illustrated in Fraser (1946): a—wing dorsoproximal part (after Fraser 1946: Fig. 2e); b—thorax (after Fraser 1946: Fig. 1c). Not to scale." httpUri="https://zenodo.org/record/2624154/files/figure.png" pageId="10" pageNumber="77">Figs. 1a</figureCitation>
,
<figureCitation id="13750BB23E7B2E7602AFD5E3FF51C0A9" box="[193,221,1679,1703]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="10" pageNumber="77">3a</figureCitation>
,
<figureCitation id="13750BB23E7B2E760284D5E3FE8BC0A9" box="[234,263,1679,1703]" captionStart="FIGURE 4" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[296,1246,481,1879]" captionTargetId="figure@6.[288,1298,481,1903]" captionTargetPageId="6" captionText="FIGURE 4. The female of Indolestes anomalus collected in tandem with a male in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—head top, dorsofrontal view; dsynthorax, ventral view; e—tip of abdomen, lateral view; f—right wings; g—fore wing tip; h—hind wing tip. Not to scale. Photos by Rosser Garrison." httpUri="https://zenodo.org/record/2624161/files/figure.png" pageId="10" pageNumber="77">4a</figureCitation>
,
<figureCitation id="13750BB23E7B2E76037AD5E3FEA8C0A6" box="[276,292,1679,1704]" captionStart="FIGURE 5" captionStartId="7.[151,250,1941,1963]" captionTargetBox="[445,1150,193,1921]" captionTargetId="figure@7.[445,1150,193,1921]" captionTargetPageId="7" captionText="FIGURE 5. Indolestes anomalus photographed in nature in Thailand in Chaiyaphum Province, Phu Khieo Wildlife Sanctuary, 12 v 2016 (a) and Phetchabun Province, Nam Nao National Park, Suan Son Ban Baek, 15 viii 2015 (b–d); a, b—mature males; c—mature female; d—ovipositing tandem. Photo by Dennis Farrell." httpUri="https://zenodo.org/record/2624165/files/figure.png" pageId="10" pageNumber="77">5</figureCitation>
,
<figureCitation id="13750BB23E7B2E76035FD5E3FEE6C0A9" box="[305,362,1679,1704]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[287,1278,511,1836]" captionTargetId="figure@8.[287,1299,471,1887]" captionTargetPageId="8" captionText="FIGURE 6. The male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison." httpUri="https://zenodo.org/record/2624167/files/figure.png" pageId="10" pageNumber="77">6a, c</figureCitation>
). These spots rarely become confluent thus forming a broad irregular stripe confluent with the middorsal stripe (e. g.
<bibRefCitation id="EFDF6AC63E7B2E7603FBD5D8FDB5C0C5" author="Ng, J. F. &amp; Dow, R. W. &amp; Choong, C. Y." box="[405,569,1715,1740]" pageId="10" pageNumber="77" pagination="9 - 16" refId="ref5415" refString="Ng, J. F., Dow, R. W. &amp; Choong, C. Y. (2011) New records of Odonata (Insecta) from the Cameron Highlands, with first records of two species for Malaysia. Journal of Science and Technology in Tropics, 7, 9 - 16." type="journal article" year="2011">
Ng
<emphasis id="B93ACB253E7B2E7603D0D5D8FE63C0C2" box="[446,495,1715,1740]" italics="true" pageId="10" pageNumber="77">et al</emphasis>
. 2011
</bibRefCitation>
:
<figureCitation id="13750BB23E7B2E76002AD5DFFD1AC0C2" box="[580,662,1715,1740]" captionStart="FIGURE 1" captionStartId="2.[151,250,1507,1529]" captionTargetBox="[340,1226,424,1423]" captionTargetId="figure@2.[340,1247,267,1483]" captionTargetPageId="2" captionText="FIGURE 1. The holotype (female) of Indolestes anomalus as illustrated in Fraser (1946): a—wing dorsoproximal part (after Fraser 1946: Fig. 2e); b—thorax (after Fraser 1946: Fig. 1c). Not to scale." httpUri="https://zenodo.org/record/2624154/files/figure.png" pageId="10" pageNumber="77">Fig. 1b</figureCitation>
). The synthoracic maculation of
<taxonomicName id="4C4E6CB43E7B2E760665D5D8FB03C0C2" box="[1035,1167,1715,1740]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E7B2E760665D5D8FB03C0C2" box="[1035,1167,1715,1740]" italics="true" pageId="10" pageNumber="77">I. anomalus</emphasis>
</taxonomicName>
differs from that in any other Asian
<emphasis id="B93ACB253E7B2E760347D5BBFE16C0FE" box="[297,410,1751,1776]" italics="true" pageId="10" pageNumber="77">Indolestes</emphasis>
spp.
<taxonomicName id="4C4E6CB43E7B2E76038ED5BBFCC0C0E1" authority="Fraser, 1922" authorityName="Fraser" authorityYear="1922" box="[480,844,1751,1776]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B93ACB253E7B2E76038ED5BBFD22C0FE" box="[480,686,1751,1776]" italics="true" pageId="10" pageNumber="77">Indolestes indicus</emphasis>
Fraser, 1922
</taxonomicName>
,
<taxonomicName id="4C4E6CB43E7B2E760132D5B4FAC0C0FE" authority="gracilis (Hagen in Selys, 1862)" authorityName="Hagen in Selys" authorityYear="1862" baseAuthorityName="Hagen in Selys" baseAuthorityYear="1862" box="[860,1356,1751,1776]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="gracilis" subSpecies="gracilis">
<emphasis id="B93ACB253E7B2E760132D5B4FBA3C0FE" box="[860,1071,1751,1776]" italics="true" pageId="10" pageNumber="77">I. gracilis gracilis</emphasis>
(Hagen in Selys, 1862)
</taxonomicName>
and
<taxonomicName id="4C4E6CB43E7B2E7607E2D5B4FE1AC11A" authority="(Ris, 1916)" authorityName="Ris" authorityYear="1916" baseAuthorityName="Ris" baseAuthorityYear="1916" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="B93ACB253E7B2E7607E2D5B4FE83C11D" italics="true" pageId="10" pageNumber="77">I. peregrinus</emphasis>
(Ris, 1916)
</taxonomicName>
also have or may have a number of separate posthumeral spots, but in
<taxonomicName id="4C4E6CB43E7B2E7606D6D590FAAAC11A" authorityName="Hagen in Selys" authorityYear="1862" box="[1208,1318,1787,1812]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B93ACB253E7B2E7606D6D590FAAAC11A" box="[1208,1318,1787,1812]" italics="true" pageId="10" pageNumber="77">I. gracilis</emphasis>
</taxonomicName>
the dorsal dark stripe has straight margins whereas in
<taxonomicName id="4C4E6CB43E7B2E7600EDD44CFD60C136" authorityName="Fraser" authorityYear="1922" box="[643,748,1823,1848]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B93ACB253E7B2E7600EDD44CFD60C136" box="[643,748,1823,1848]" italics="true" pageId="10" pageNumber="77">I. indicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4E6CB43E7B2E76014AD44CFC38C139" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[804,948,1824,1847]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="B93ACB253E7B2E76014AD44CFC38C139" box="[804,948,1824,1847]" italics="true" pageId="10" pageNumber="77">I. peregrinus</emphasis>
</taxonomicName>
this stripe forms long trapezoid extensions medially rather than a small extension as in
<taxonomicName id="4C4E6CB43E7B2E760011D428FC8FC152" box="[639,771,1859,1884]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E7B2E760011D428FC8FC152" box="[639,771,1859,1884]" italics="true" pageId="10" pageNumber="77">I. anomalus</emphasis>
</taxonomicName>
, but also expanding at the posterior end, like in
<taxonomicName id="4C4E6CB43E7B2E760776D428FA10C152" box="[1304,1436,1859,1884]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E7B2E760776D428FA10C152" box="[1304,1436,1859,1884]" italics="true" pageId="10" pageNumber="77">I. anomalus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFDF6AC63E7B2E7602F0D40BFEA5C171" author="Fraser, F. C." box="[158,297,1895,1919]" pageId="10" pageNumber="77" refId="ref5015" refString="Fraser, F. C. (1933) The fauna of British India, including Ceylon and Burma. Odonata. Uol. I. Taylor and Francis, London, 423 pp." type="book" year="1933">Fraser 1933</bibRefCitation>
;
<bibRefCitation id="EFDF6AC63E7B2E76035BD40BFE59C18E" author="Asahina, S." box="[309,469,1895,1920]" pageId="10" pageNumber="77" pagination="1 - 12" refId="ref4802" refString="Asahina, S. (1976) Notes on Chinese Odonata. V. Some Odonata from Hunan and Hupeh Provinces. Kontyu, 44, 1 - 12." type="journal article" year="1976">Asahina 1976</bibRefCitation>
). The male cerci of
<taxonomicName id="4C4E6CB43E7B2E7600D6D404FCB1C18E" box="[696,829,1895,1920]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E7B2E7600D6D404FCB1C18E" box="[696,829,1895,1920]" italics="true" pageId="10" pageNumber="77">I. anomalus</emphasis>
</taxonomicName>
(
<figureCitation id="13750BB23E7B2E760123D40BFC03C18E" box="[845,911,1895,1920]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[287,1278,511,1836]" captionTargetId="figure@8.[287,1299,471,1887]" captionTargetPageId="8" captionText="FIGURE 6. The male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison." httpUri="https://zenodo.org/record/2624167/files/figure.png" pageId="10" pageNumber="77">Fig. 6</figureCitation>
e–g) have attenuated apices similar to those of
<taxonomicName id="4C4E6CB43E7B2E7602F9D4E7FD84C1AA" authority="(Selys, 1862)" authorityName="Selys" authorityYear="1862" baseAuthorityName="Selys" baseAuthorityYear="1862" box="[151,520,1931,1956]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="cyaneus">
<emphasis id="B93ACB253E7B2E7602F9D4E7FEE5C1AD" box="[151,361,1931,1956]" italics="true" pageId="10" pageNumber="77">Indolestes cyaneus</emphasis>
(Selys, 1862)
</taxonomicName>
,
<taxonomicName id="4C4E6CB43E7B2E76007AD4E0FD0DC1AA" authorityName="Hagen in Selys" authorityYear="1862" box="[532,641,1931,1956]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B93ACB253E7B2E76007AD4E0FD0DC1AA" box="[532,641,1931,1956]" italics="true" pageId="10" pageNumber="77">I. gracilis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4E6CB43E7B2E7600E0D4E0FB91C1AA" authority="Zhou &amp; Zhou, 2005" authorityName="Zhou &amp; Zhou" authorityYear="2005" box="[654,1053,1931,1956]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="guizhouensis">
<emphasis id="B93ACB253E7B2E7600E0D4E0FCBAC1AA" box="[654,822,1931,1956]" italics="true" pageId="10" pageNumber="77">I. guizhouensis</emphasis>
Zhou &amp; Zhou, 2005
</taxonomicName>
(if this is a
<emphasis id="B93ACB253E7B2E7606CCD4E7FA87C1AA" box="[1186,1291,1931,1956]" italics="true" pageId="10" pageNumber="77">bona fide</emphasis>
species) and
<taxonomicName id="4C4E6CB43E7B2E7602F9D4DCFEA4C1C9" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[151,296,1968,1991]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="77" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="B93ACB253E7B2E7602F9D4DCFEA4C1C9" box="[151,296,1968,1991]" italics="true" pageId="10" pageNumber="77">I. peregrinus</emphasis>
</taxonomicName>
among Asian species (Kosterin 2015), but differ clearly from those species by the much larger and prominent, rounded inner swellings. Females have a characteristic pair of pale tubercles at the occiput lateroposterior of the lateral ocelli (
<figureCitation id="13750BB23E7A2E770048D3FBFD0FC6BE" box="[550,643,151,176]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="11" pageNumber="78">Figs. 3a</figureCitation>
,
<figureCitation id="13750BB23E7A2E7700E1D3FBFD44C6BE" box="[655,712,151,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[296,1246,481,1879]" captionTargetId="figure@6.[288,1298,481,1903]" captionTargetPageId="6" captionText="FIGURE 4. The female of Indolestes anomalus collected in tandem with a male in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—head top, dorsofrontal view; dsynthorax, ventral view; e—tip of abdomen, lateral view; f—right wings; g—fore wing tip; h—hind wing tip. Not to scale. Photos by Rosser Garrison." httpUri="https://zenodo.org/record/2624161/files/figure.png" pageId="11" pageNumber="78">4b, c</figureCitation>
). Females resemble those of
<taxonomicName id="4C4E6CB43E7A2E77067FD3F5FB2DC6BE" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[1041,1185,153,176]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="78" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="B93ACB253E7A2E77067FD3F5FB2DC6BE" box="[1041,1185,153,176]" italics="true" pageId="11" pageNumber="78">I. peregrinus</emphasis>
</taxonomicName>
by pale S10 and cerci contrasting with dark S9 (
<figureCitation id="13750BB23E7A2E7703D3D3D0FD95C6DA" box="[445,537,188,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="11" pageNumber="78">Figs. 3c</figureCitation>
,
<figureCitation id="13750BB23E7A2E770048D3D0FDD5C6DB" box="[550,601,188,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[296,1246,481,1879]" captionTargetId="figure@6.[288,1298,481,1903]" captionTargetPageId="6" captionText="FIGURE 4. The female of Indolestes anomalus collected in tandem with a male in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—head top, dorsofrontal view; dsynthorax, ventral view; e—tip of abdomen, lateral view; f—right wings; g—fore wing tip; h—hind wing tip. Not to scale. Photos by Rosser Garrison." httpUri="https://zenodo.org/record/2624161/files/figure.png" pageId="11" pageNumber="78">4a,d</figureCitation>
). Pterostigmata at its ventral margin 1.8–2.2 as long as high (
<figureCitation id="13750BB23E7A2E77077CD3D0FAFCC6DB" box="[1298,1392,188,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="11" pageNumber="78">Figs. 3b</figureCitation>
,
<figureCitation id="13750BB23E7A2E770710D3D0FF2BC6F6" captionStart="FIGURE 4" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[296,1246,481,1879]" captionTargetId="figure@6.[288,1298,481,1903]" captionTargetPageId="6" captionText="FIGURE 4. The female of Indolestes anomalus collected in tandem with a male in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—head top, dorsofrontal view; dsynthorax, ventral view; e—tip of abdomen, lateral view; f—right wings; g—fore wing tip; h—hind wing tip. Not to scale. Photos by Rosser Garrison." httpUri="https://zenodo.org/record/2624161/files/figure.png" pageId="11" pageNumber="78">4f, g</figureCitation>
); that is shorter than in the related species (e.g. 3.0–
<quantity id="4CB6BAD23E7A2E770081D38CFCA2C6F6" box="[751,814,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="8.128" pageId="11" pageNumber="78" unit="in" value="3.2">3.2 in</quantity>
<taxonomicName id="4C4E6CB43E7A2E77015BD38DFC48C6F6" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[821,964,225,248]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="78" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="B93ACB253E7A2E77015BD38DFC48C6F6" box="[821,964,225,248]" italics="true" pageId="11" pageNumber="78">I. peregrinus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BF117373E7A2E7702A9D269FD49C7FA" blockId="11.[151,1436,151,501]" pageId="11" pageNumber="78">
In his description,
<bibRefCitation id="EFDF6AC63E7A2E7703F9D269FDA2C713" author="Fraser, F. C." box="[407,558,260,285]" pageId="11" pageNumber="78" pagination="41 - 45" refId="ref5050" refString="Fraser, F. C. (1946) On rare and two new species of Odonata from tropical Africa and Asia. Proceedings of Royal Entomological Society of London. Series B, 15 (3 / 4), 41 - 45. https: // doi. org / 10.1111 / j. 1365 - 3113.1946. tb 00815. x" type="journal article" year="1946">Fraser (1946)</bibRefCitation>
especially stressed the short pterostigmata of the
<typeStatus id="54F5A9953E7A2E770634D268FB30C713" box="[1114,1212,260,285]" pageId="11" pageNumber="78" type="holotype">holotype</typeStatus>
but it is longer than he depicted and becomes a less conspicuous diagnostic character. Nevertheless, it is still relatively shorter than in all other
<emphasis id="B93ACB253E7A2E77036DD220FEF9C76B" box="[259,373,332,357]" italics="true" pageId="11" pageNumber="78">Indolestes</emphasis>
species.
<taxonomicName id="4C4E6CB43E7A2E77038CD220FD40C76B" box="[482,716,332,357]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="78" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E7A2E77038CD220FD40C76B" box="[482,716,332,357]" italics="true" pageId="11" pageNumber="78">Indolestes anomalus</emphasis>
</taxonomicName>
most resembles
<taxonomicName id="4C4E6CB43E7A2E7701FAD221FBABC76A" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[916,1063,333,356]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="78" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="B93ACB253E7A2E7701FAD221FBABC76A" box="[916,1063,333,356]" italics="true" pageId="11" pageNumber="78">I. peregrinus</emphasis>
</taxonomicName>
, with which it shares separated posthumeral spots in both sexes, but differs by its smaller size and a more posterior position of the prominence of the dorsal synthoracic dark stripe. The large rounded inner swellings of the cerci in males (
<figureCitation id="13750BB23E7A2E7706A8D2F8FA87C7A3" box="[1222,1291,404,429]" captionStart="FIGURE 6" captionStartId="8.[151,250,1904,1926]" captionTargetBox="[287,1278,511,1836]" captionTargetId="figure@8.[287,1299,471,1887]" captionTargetPageId="8" captionText="FIGURE 6. The male of Indolestes anomalus collected in tandem with the female of Fig. 3 in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—synthorax, lateral view; dsynthorax, ventral view; e—anal appendages, lateral view; f—ditto, dorsal view; g—ditto, dorsolateral view. Not to scale. Photo by Rosser Garrison." httpUri="https://zenodo.org/record/2624167/files/figure.png" pageId="11" pageNumber="78">Fig. 6</figureCitation>
e–g) (almost absent e.g. in
<taxonomicName id="4C4E6CB43E7A2E77035ED2D5FE32C7DE" baseAuthorityName="Ris" baseAuthorityYear="1916" box="[304,446,441,464]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="78" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="B93ACB253E7A2E77035ED2D5FE32C7DE" box="[304,446,441,464]" italics="true" pageId="11" pageNumber="78">I. peregrinus</emphasis>
</taxonomicName>
) and pair of pale, variably expressed postocellar tubercles in females (
<figureCitation id="13750BB23E7A2E7706A0D2DBFAA4C7DE" box="[1230,1320,439,464]" captionStart="FIGURE 3" captionStartId="5.[151,250,1922,1944]" captionTargetBox="[151,1435,282,1922]" captionTargetId="figure@5.[151,1435,282,1900]" captionTargetPageId="5" captionText="FIGURE 3. A nearly topotypical female of Indolestes anomalus from Vietnam, Đỗng Nai Province, Cát Tiên National Park, 6 ii 2016: a—head and thorax; b—hindwing; c—general habitus. Not to scale. Photo by Tom Kompier." httpUri="https://zenodo.org/record/2624159/files/figure.png" pageId="11" pageNumber="78">Figs. 3a</figureCitation>
,
<figureCitation id="13750BB23E7A2E77075AD2DBFAE7C7DE" box="[1332,1387,439,464]" captionStart="FIGURE 4" captionStartId="6.[151,250,1919,1941]" captionTargetBox="[296,1246,481,1879]" captionTargetId="figure@6.[288,1298,481,1903]" captionTargetPageId="6" captionText="FIGURE 4. The female of Indolestes anomalus collected in tandem with a male in Thailand, Chaiyaphum Province, a small muddy pond at Tung Lui Lai, 19 vii 1996; a—body habitus; b—head, frontal view; c—head top, dorsofrontal view; dsynthorax, ventral view; e—tip of abdomen, lateral view; f—right wings; g—fore wing tip; h—hind wing tip. Not to scale. Photos by Rosser Garrison." httpUri="https://zenodo.org/record/2624161/files/figure.png" pageId="11" pageNumber="78">4b, c</figureCitation>
) are good structural characters to identify
<taxonomicName id="4C4E6CB43E7A2E770054D2B1FD32C7FB" box="[570,702,476,501]" class="Insecta" family="Lestidae" genus="Indolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="78" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B93ACB253E7A2E770054D2B1FD32C7FB" box="[570,702,476,501]" italics="true" pageId="11" pageNumber="78">I. anomalus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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