542 lines
66 KiB
XML
542 lines
66 KiB
XML
<document id="5B1B7047E5B91B1210C9CA38E6AA3DCE" ID-DOI="10.1080/00222933.2017.1391959" ID-ISSN="1464-5262" ID-Zenodo-Dep="5174516" ID-ZooBank="9E21390E-3FD4-40EB-9442-31BC92A76B4F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628558890842" checkinUser="felipe" docAuthor="Ruane, Sara, Richards, Stephen J., McVay, John D., Tjaturadi, Burhan, Krey, Keliopas & Austin, Christopher C." docDate="2017" docId="03DA87E1D418457F04AEE5D086B099BC" docLanguage="en" docName="JNatHist.52.13-16.917-944.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 52 (13 - 16)" docSource="http://dx.doi.org/10.1080/00222933.2017.1391959" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Stegonotus derooijae Ruane & Richards & McVay & Tjaturadi & Krey & Austin 2017, sp. nov." docType="treatment" docUuid="400B69E0-73E1-4D0B-8DCD-32CA7BEA6D3F" docUuidSource="ZooBank" docVersion="4" lastPageNumber="933" masterDocId="FFE3FF99D415456D0575E55D851E9975" masterDocTitle="Cryptic and non-cryptic diversity in New Guinea ground snakes of the genus Stegonotus Duméril, Bibron and Duméril, 1854: a description of four new species (Squamata: Colubridae)" masterLastPageNumber="944" masterPageNumber="917" pageNumber="929" updateTime="1699046540698" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="70C024D2010E28C8D509E1E0B0A9C620">Cryptic and non-cryptic diversity in New Guinea ground snakes of the genus Stegonotus Duméril, Bibron and Duméril, 1854: a description of four new species (Squamata: Colubridae)</mods:title>
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<mods:namePart id="BB7A2E88A597584746386A9DA5E5AE95">Ruane, Sara</mods:namePart>
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<mods:namePart id="A96A15BC58F2B10036B6736E01A0FA31">Richards, Stephen J.</mods:namePart>
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<mods:namePart id="1331BD46A93E20D92252BAD70A4397C5">McVay, John D.</mods:namePart>
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<mods:namePart id="C74F9D8222F6801BF6A18021B5487532">Tjaturadi, Burhan</mods:namePart>
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<mods:namePart id="29EB494DBECE3345A55E167BD8B42C59">Krey, Keliopas</mods:namePart>
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<mods:namePart id="819D9C0D44891DDB33DCB11EA192110E">Austin, Christopher C.</mods:namePart>
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<mods:title id="713A10B122D3663DC76AE86ACF6D200D">Journal of Natural History</mods:title>
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<mods:date id="1ACBDE7A83FD34461BF1F2686AC8702F">2017</mods:date>
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<mods:title id="1E9BC1AE3C218A742EC1781134CB4EE5">J. Nat. Hist.</mods:title>
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<mods:number id="61BFF6E58792429174593189FB431A6A">2017-11-27</mods:number>
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<mods:number id="8669920B28814607686EB62FFCA4BCF3">52</mods:number>
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<mods:number id="FBFF56ECCA719981E8FA5D6B9BF54B85">13 - 16</mods:number>
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<mods:url id="DB8922AAC2F9E4F66E5C86FC4B6DBC65">http://dx.doi.org/10.1080/00222933.2017.1391959</mods:url>
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<mods:identifier id="226E5F535388D8B5744DDFAF92EDABBB" type="DOI">10.1080/00222933.2017.1391959</mods:identifier>
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<mods:identifier id="7535DDECFBC8032A6C9EC4627971AAD7" type="ISSN">1464-5262</mods:identifier>
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<heading id="D084819BD418456004AEE5D0865699D2" box="[475,840,141,167]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="929" reason="2">
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<taxonomicName id="4C734D74D418456004AEE5D087FC99D2" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[475,738,141,167]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="929" phylum="Chordata" rank="species" species="derooijae" status="sp. nov.">
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<emphasis id="B907EAE5D418456004AEE5D087FC99D2" bold="true" box="[475,738,141,167]" italics="true" pageId="13" pageNumber="929">Stegonotus derooijae</emphasis>
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<taxonomicNameLabel id="A234579ED41845600799E5D3865699D2" box="[748,840,142,167]" pageId="13" pageNumber="929" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C369657CD418456004C0E5EC867099BF" box="[437,878,177,202]" pageId="13" pageNumber="929" type="description">
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<paragraph id="8BCC36F7D418456004C0E5EC867099BF" blockId="13.[437,878,141,202]" box="[437,878,177,202]" pageId="13" pageNumber="929">
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(
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<figureCitation id="13482A72D418456004C8E5EC875799BF" box="[445,585,177,202]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStartId-0="6.[160,225,1511,1534]" captionStartId-1="9.[160,225,1677,1700]" captionStartId-2="14.[160,225,1613,1636]" captionTargetBox-0="[194,1121,142,1480]" captionTargetBox-1="[217,1128,144,1646]" captionTargetBox-2="[334,981,142,1582]" captionTargetId-0="figure-1@6.[193,1121,142,1480]" captionTargetId-1="graphics-2@9.[256,1128,623,1646]" captionTargetId-2="figure-1@14.[333,981,142,1582]" captionTargetPageId-0="6" captionTargetPageId-1="9" captionTargetPageId-2="14" captionText-0="Figure 2. Photographs of Stegonotus species with dorsal, ventral and lateral head views from left to right: (a) Stegonotus batjanensis USNM 237129, (b) Stegonotus borneensis FMNH 251054, (c) Stegonotus muelleri LSUMZ 41802, (d) Stegonotus florensis WAM 104606, (e) Stegonotus guentheri AM 129712, (f) Stegonotus heterurus BPBM 22556, (g) Stegonotus cucullatus LSUMZ 94371. Photos from SR. (h) Stegonotus modestus LSUMZ 92339, (i) Stegonotus iridis sp. nov. SJR 7514, (j) Stegonotus diehli LSUMZ 92344 (k) Stegonotus melanolabiatus sp. nov. AMS R115361 (l) Stegonotus derooijae sp. nov. SAMA R70467 (m) Stegonotus admiraltiensis sp. nov. LSUMZ 93597, (n) Stegonotus parvus LSUMZ 92333. Photos from Sara Ruane and Stephen J. Richards." captionText-1="Figure 3. Map of New Guinea sampling localities for Stegonotus (a) and mtDNA + nucDNA maximum likelihood gene tree (b). Symbols for species/clades match those on the map; * indicates bootstrap values ≥ 70. For some localities, symbols were adjusted for better visibility and so localities are approximate; see Appendix 1. for exact locality information." captionText-2="Figure 4. Photographs of three Stegonotus species described here in life; (a) Stegonotus iridis sp. nov. SAMA R70466 (b) Stegonotus derooijae sp. nov. MZB.Ophi.3288 (c) Stegonotus admiraltiensis sp. nov. from Manus Island, unregistered. Photos from Stephen J. Richards." figureDoi-0="http://doi.org/10.5281/zenodo.5174520" figureDoi-1="http://doi.org/10.5281/zenodo.5174522" figureDoi-2="http://doi.org/10.5281/zenodo.5174525" httpUri-0="https://zenodo.org/record/5174520/files/figure.png" httpUri-1="https://zenodo.org/record/5174522/files/figure.png" httpUri-2="https://zenodo.org/record/5174525/files/figure.png" pageId="13" pageNumber="929">Figures 2−4</figureCitation>
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,
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<figureCitation id="13482A72D4184560072DE5EC877899BF" box="[600,614,177,202]" captionStart="Figure 6" captionStartId="16.[160,225,1385,1408]" captionTargetBox="[163,1153,142,1353]" captionTargetId="figure-71@16.[163,1153,142,1354]" captionTargetPageId="16" captionText="Figure 6. Photographs of paratype Stegonotus derooijae sp. nov. MZB.Ophi.3293 showing (clockwise from top left) dorsal of body, ventral of body, dorsal of head, lateral of head. Photos from Stephen J. Richards." figureDoi="http://doi.org/10.5281/zenodo.5174529" httpUri="https://zenodo.org/record/5174529/files/figure.png" pageId="13" pageNumber="929">6</figureCitation>
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,
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<tableCitation id="C6F1034CD41845600703E5EC87D399BF" box="[630,717,177,202]" captionStart="Table 1" captionStartId="10.[133,189,620,643]" captionText="Table 1. Morphological characters of Stegonotus examined." pageId="13" pageNumber="929">Table 1</tableCitation>
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, Appendix 1)
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</paragraph>
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</subSubSection>
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<subSubSection id="C369657CD418456005D5E5A9870E9879" box="[160,528,244,268]" pageId="13" pageNumber="929" type="reference_group">
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<paragraph id="8BCC36F7D418456005D5E5A9870E9879" blockId="13.[160,528,244,268]" box="[160,528,244,268]" pageId="13" pageNumber="929">
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<taxonomicName id="4C734D74D418456005D5E5A9870E9879" authority="Lindholm 1905" authorityName="Lindholm. It" authorityYear="1905" box="[160,528,244,268]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="929" phylum="Chordata" rank="species" species="diehli">
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<emphasis id="B907EAE5D418456005D5E5A984439879" box="[160,349,244,268]" italics="true" pageId="13" pageNumber="929">Stegonotus diehli</emphasis>
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<bibRefCitation id="EFE24B06D41845600413E5A9870E9879" author="Lindholm WA" box="[358,528,244,268]" pageId="13" pageNumber="929" pagination="227 - 240" refId="ref12249" refString="Lindholm WA. 1905. Uber einige Eidechsen und Schlangen aus Deutsch-Neuguinea. Jahrbucher des Nassau Vereins fur Naturkd [About some lizards and snakes from German New Guinea]. 58: 227 - 240." type="journal article" year="1905">Lindholm 1905</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="8BCC36F7D418456005D5E46584159827" blockId="13.[160,1156,312,477]" box="[160,267,312,338]" pageId="13" pageNumber="929">
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.
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3288 (field number
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, tissue
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), adult male. Collected by
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,
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and
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on
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, from
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,
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,
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elevation
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asl
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<figureCitation id="13482A72D4184560043BE49884B298A8" box="[334,428,453,477]" captionStart="Figure 4" captionStartId="14.[160,225,1613,1636]" captionTargetBox="[334,981,142,1582]" captionTargetId="figure-1@14.[333,981,142,1582]" captionTargetPageId="14" captionText="Figure 4. Photographs of three Stegonotus species described here in life; (a) Stegonotus iridis sp. nov. SAMA R70466 (b) Stegonotus derooijae sp. nov. MZB.Ophi.3288 (c) Stegonotus admiraltiensis sp. nov. from Manus Island, unregistered. Photos from Stephen J. Richards." figureDoi="http://doi.org/10.5281/zenodo.5174525" httpUri="https://zenodo.org/record/5174525/files/figure.png" pageId="13" pageNumber="929">Figure 4</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D418456005D5E754840B9B56" blockId="13.[160,1156,521,720]" box="[160,277,521,547]" pageId="13" pageNumber="929">
|
||
<emphasis id="B907EAE5D418456005D5E754840B9B56" bold="true" box="[160,277,521,547]" italics="true" pageId="13" pageNumber="929">
|
||
<typeStatus id="54C88855D418456005D5E754840B9B56" box="[160,277,521,547]" pageId="13" pageNumber="929" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D418456005D5E77384299BA5" blockId="13.[160,1156,521,720]" pageId="13" pageNumber="929">
|
||
<collectionCode id="ED62AE32D418456005D5E77385C89B33" box="[160,214,558,582]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="13" pageNumber="929" type="Museum">MZB</collectionCode>
|
||
.Ophi.
|
||
<materialsCitation id="3B1B3CAAD41845600464E773819E9B1D" accessionNumber="R70467" collectingDate="2005-06-26" collectionCode="SAMA, R" collectorName="Ophi. & S. Richards & B. Tjaturadi & K. Krey" country="Indonesia" county="Raja Ampat Regency" elevation="75" latitude="-0.956383" location="Waibya Camp" longLatPrecision="1" longitude="130.78433" municipality="Salawati Island" pageId="13" pageNumber="929" specimenCode="SJR 7789, ABTC 90226, SJR 7586, ABTC 90123" specimenCount="1" stateProvince="Papua Barat" typeStatus="paratype">
|
||
3293 (field number
|
||
<specimenCode id="DBD59E8CD41845600776E77387709B33" box="[515,622,558,582]" collectionCode="SJR" pageId="13" pageNumber="929">SJR 7789</specimenCode>
|
||
, tissue
|
||
<specimenCode id="DBD59E8CD418456007BEE77386449B33" box="[715,858,558,582]" collectionCode="ABTC" country="Australia" httpUri="http://grbio.org/cool/y82c-h1ev" name="Australian Biological Tissue Collection, South Australian Museum" pageId="13" pageNumber="929" type="Museum">ABTC 90226</specimenCode>
|
||
;
|
||
<figureCitation id="13482A72D4184560061FE77386D29B33" box="[874,972,558,582]" captionStart="Figure 6" captionStartId="16.[160,225,1385,1408]" captionTargetBox="[163,1153,142,1353]" captionTargetId="figure-71@16.[163,1153,142,1354]" captionTargetPageId="16" captionText="Figure 6. Photographs of paratype Stegonotus derooijae sp. nov. MZB.Ophi.3293 showing (clockwise from top left) dorsal of body, ventral of body, dorsal of head, lateral of head. Photos from Stephen J. Richards." figureDoi="http://doi.org/10.5281/zenodo.5174529" httpUri="https://zenodo.org/record/5174529/files/figure.png" pageId="13" pageNumber="929">Figure 6</figureCitation>
|
||
), same data as holotype;
|
||
<collectionCode id="ED62AE32D41845600463E70D84479B1D" box="[278,345,592,616]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="13" pageNumber="929" type="Museum">SAMA</collectionCode>
|
||
<collectionCode id="ED62AE32D41845600413E70D846B9B1D" box="[358,373,592,616]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="929">R</collectionCode>
|
||
70467 (field number
|
||
<specimenCode id="DBD59E8CD4184560071AE70D87C49B1D" box="[623,730,592,616]" collectionCode="SJR" pageId="13" pageNumber="929">SJR 7586</specimenCode>
|
||
, tissue
|
||
<specimenCode id="DBD59E8CD41845600643E70D86DA9B1D" box="[822,964,592,616]" collectionCode="ABTC" country="Australia" httpUri="http://grbio.org/cool/y82c-h1ev" name="Australian Biological Tissue Collection, South Australian Museum" pageId="13" pageNumber="929" type="Museum">ABTC 90123</specimenCode>
|
||
) collected by S
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1B3CAAD418456001F5E70D84039BFE" accessionNumber="R70467" collectingDate="2005-06-26" collectionCode="SAMA, R" collectorName="Richards, B" country="Indonesia" county="Raja Ampat Regency" elevation="75" isTailEnumeration="true" latitude="-0.956383" location="Waibya Camp" longLatPrecision="1" longitude="130.78433" municipality="Salawati Island" pageId="13" pageNumber="929" specimenCount="1" stateProvince="Papua Barat" typeStatus="paratype">
|
||
.
|
||
<collectorName id="26865321D418456005D5E72E84039BFE" box="[160,285,627,651]" pageId="13" pageNumber="929">Richards, B</collectorName>
|
||
</materialsCitation>
|
||
. Tjaturadi and K
|
||
<materialsCitation id="3B1B3CAAD418456004A3E72E842D9BA5" collectingDate="2005-06-09" country="Indonesia" county="Raja Ampat Regency" elevation="50" latitude="-0.836942" location="Krey" longLatPrecision="1" longitude="130.72162" municipality="Batanta Island" pageId="13" pageNumber="929" specimenCount="1" stateProvince="Papua Barat" typeStatus="paratype">
|
||
.
|
||
<location id="8EAC602CD41845600494E72E870C9BFE" LSID="urn:lsid:plazi:treatment:03DA87E1D418457F04AEE5D086B099BC:8EAC602CD41845600494E72E870C9BFE" box="[481,530,627,651]" country="Indonesia" county="Raja Ampat Regency" latitude="-0.836942" longLatPrecision="1" longitude="130.72162" municipality="Batanta Island" name="Krey" pageId="13" pageNumber="929" stateProvince="Papua Barat">Krey</location>
|
||
on
|
||
<date id="FFCD1037D4184560074AE72E87D69BFE" box="[575,712,627,651]" pageId="13" pageNumber="929" value="2005-06-09">
|
||
<collectingDate id="EF89E9DFD4184560074AE72E87D69BFE" box="[575,712,627,651]" pageId="13" pageNumber="929" value="2005-06-09">9 June 2005</collectingDate>
|
||
</date>
|
||
from
|
||
<location id="8EAC602CD41845600679E72E86839BFE" LSID="urn:lsid:plazi:treatment:03DA87E1D418457F04AEE5D086B099BC:8EAC602CD41845600679E72E86839BFE" box="[780,925,627,651]" country="Indonesia" county="Raja Ampat Regency" latitude="-0.836942" longLatPrecision="1" longitude="130.72162" municipality="Batanta Island" name="Warinkabom" pageId="13" pageNumber="929" stateProvince="Papua Barat">Warinkabom</location>
|
||
,
|
||
<collectingMunicipality id="6BA8AC8DD418456006DDE72E81549BFE" box="[936,1098,627,651]" pageId="13" pageNumber="929">Batanta Island</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62AD4E7BD41845600120E72E84439BDB" pageId="13" pageNumber="929">Raja Ampat Regency</collectingCounty>
|
||
,
|
||
<collectingRegion id="49B7F815D41845600405E7CB876E9BDB" box="[368,624,662,686]" country="Indonesia" name="Papua Barat" pageId="13" pageNumber="929">West Papua Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3647667D418456007F6E7CB87EC9BDB" box="[643,754,662,686]" name="Indonesia" pageId="13" pageNumber="929">Indonesia</collectingCountry>
|
||
,
|
||
<elevation id="005ED1C4D41845600670E7CB86F59BDB" box="[773,1003,662,686]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="929" unit="m" value="50.0">
|
||
elevation
|
||
<quantity id="4C8B9B12D41845600609E7CB86A39BDB" box="[892,957,662,686]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="929" unit="m" value="50.0">50 m</quantity>
|
||
asl
|
||
</elevation>
|
||
(
|
||
<geoCoordinate id="EE475030D41845600174E7CB819F9BDB" box="[1025,1153,662,686]" degrees="0.836942" direction="south" orientation="latitude" pageId="13" pageNumber="929" precision="1" value="-0.836942">0.836942°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE475030D418456005D5E7E584339BA5" box="[160,301,696,720]" degrees="130.72162" direction="east" orientation="longitude" pageId="13" pageNumber="929" precision="1" value="130.72162">130.72162°E</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D418456005D5E7A184989A63" blockId="13.[160,1157,764,894]" box="[160,390,764,790]" pageId="13" pageNumber="929">
|
||
<emphasis id="B907EAE5D418456005D5E7A184989A63" bold="true" box="[160,390,764,790]" italics="true" pageId="13" pageNumber="929">Referred specimens</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D418456005D5E67C819B9A0B" blockId="13.[160,1157,764,894]" pageId="13" pageNumber="929">
|
||
<collectionCode id="ED62AE32D418456005D5E67C85C89A4C" box="[160,214,801,825]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="13" pageNumber="929" type="Museum">MZB</collectionCode>
|
||
.Ophi.
|
||
<materialsCitation id="3B1B3CAAD41845600464E67C86ED9A4C" box="[273,1011,801,825]" collectionCode="SJR" pageId="13" pageNumber="929" specimenCode="SJR 7650" specimenCount="1">
|
||
3292 (field number
|
||
<specimenCode id="DBD59E8CD41845600488E67C87789A4C" box="[509,614,801,825]" collectionCode="SJR" pageId="13" pageNumber="929">SJR 7650</specimenCode>
|
||
, morphology only) collected by S
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1B3CAAD41845600686E67C819E9A4C" box="[1011,1152,801,825]" collectionCode="SJR" collectorName="Richards, B" isTailEnumeration="true" pageId="13" pageNumber="929" specimenCount="1">
|
||
.
|
||
<collectorName id="26865321D41845600175E67C819E9A4C" box="[1024,1152,801,825]" pageId="13" pageNumber="929">Richards, B</collectorName>
|
||
</materialsCitation>
|
||
. Tjaturadi and K
|
||
<materialsCitation id="3B1B3CAAD41845600421E61E819F9A0B" collectingDate="2005-06-13" country="Indonesia" county="Raja Ampat Regency" elevation="45" latitude="-0.337183" location="Krey" longLatPrecision="1" longitude="131.2588" municipality="Waigeo Island" pageId="13" pageNumber="929" specimenCount="1" stateProvince="Papua Barat">
|
||
.
|
||
<location id="8EAC602CD41845600417E61E848D9A2E" LSID="urn:lsid:plazi:treatment:03DA87E1D418457F04AEE5D086B099BC:8EAC602CD41845600417E61E848D9A2E" box="[354,403,835,859]" country="Indonesia" county="Raja Ampat Regency" latitude="-0.337183" longLatPrecision="1" longitude="131.2588" municipality="Waigeo Island" name="Krey" pageId="13" pageNumber="929" stateProvince="Papua Barat">Krey</location>
|
||
on
|
||
<date id="FFCD1037D418456004B3E61E877C9A2E" box="[454,610,835,859]" pageId="13" pageNumber="929" value="2005-06-13">
|
||
<collectingDate id="EF89E9DFD418456004B3E61E877C9A2E" box="[454,610,835,859]" pageId="13" pageNumber="929" value="2005-06-13">13 June 2005</collectingDate>
|
||
</date>
|
||
from
|
||
<location id="8EAC602CD418456007D8E61E86219A2E" LSID="urn:lsid:plazi:treatment:03DA87E1D418457F04AEE5D086B099BC:8EAC602CD418456007D8E61E86219A2E" box="[685,831,835,859]" country="Indonesia" county="Raja Ampat Regency" latitude="-0.337183" longLatPrecision="1" longitude="131.2588" municipality="Waigeo Island" name="Urbinasopen" pageId="13" pageNumber="929" stateProvince="Papua Barat">Urbinasopen</location>
|
||
,
|
||
<collectingMunicipality id="6BA8AC8DD41845600638E61E86EF9A2E" box="[845,1009,835,859]" pageId="13" pageNumber="929">Waigeo Island</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62AD4E7BD4184560068BE61E841C9A0B" pageId="13" pageNumber="929">Raja Ampat Regency</collectingCounty>
|
||
,
|
||
<collectingRegion id="49B7F815D41845600479E63B84E49A0B" box="[268,506,870,894]" country="Indonesia" name="Papua Barat" pageId="13" pageNumber="929">West Papua Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3647667D41845600770E63B876A9A0B" box="[517,628,870,894]" name="Indonesia" pageId="13" pageNumber="929">Indonesia</collectingCountry>
|
||
,
|
||
<elevation id="005ED1C4D4184560070BE63B86549A0B" box="[638,842,870,894]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="929" unit="m" value="45.0">
|
||
elevation
|
||
<quantity id="4C8B9B12D41845600799E63B863B9A0B" box="[748,805,870,894]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="929" unit="m" value="45.0">45 m</quantity>
|
||
asl
|
||
</elevation>
|
||
(
|
||
<geoCoordinate id="EE475030D41845600623E63B86C89A0B" box="[854,982,870,894]" degrees="0.337183" direction="south" orientation="latitude" pageId="13" pageNumber="929" precision="1" value="-0.337183">0.337183°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE475030D41845600695E63B81659A0B" box="[992,1147,870,894]" degrees="131.258800" direction="east" orientation="longitude" pageId="13" pageNumber="929" precision="1" value="131.2588">131.258800°E</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C369657CD418456005D5E6F784219D89" pageId="13" pageNumber="929" type="diagnosis">
|
||
<paragraph id="8BCC36F7D418456005D5E6F7840A9AB1" blockId="13.[160,1156,938,1276]" box="[160,276,938,964]" pageId="13" pageNumber="929">
|
||
<emphasis id="B907EAE5D418456005D5E6F7840A9AB1" bold="true" box="[160,276,938,964]" italics="true" pageId="13" pageNumber="929">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D418456005D5E69284219D89" blockId="13.[160,1156,938,1276]" pageId="13" pageNumber="929">
|
||
A new species of
|
||
<taxonomicName id="4C734D74D41845600412E69284FE9A92" authorityName="Dumeril, Bibron and Dumeril" authorityYear="1854" box="[359,480,975,999]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="929" phylum="Chordata" rank="genus">
|
||
<emphasis id="B907EAE5D41845600412E69284FE9A92" box="[359,480,975,999]" italics="true" pageId="13" pageNumber="929">Stegonotus</emphasis>
|
||
</taxonomicName>
|
||
that can be diagnosed from its congeners by the following combination of characters: a dark grey or brown unpatterned dorsum with a white or cream venter with dorsal scale pigmentation continuing onto some or many of the ventral scales, dark grey pigmentation on the subcaudal scales extending from the dorsal scale pigmentation and with grey on the anterior edges of some subcaudal scales, 17 dorsal scale rows at midbody, 178−197 ventral scales and 90−94 divided subcaudal scales, two preocular scales, seven (rarely nine) supralabial scales, eight or nine infralabial scales, with the supralabial and infralabial scales cream and with grey pigmentation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C369657CD418457C05D5E07584FE9A92" lastPageId="17" lastPageNumber="933" pageId="13" pageNumber="929" type="description">
|
||
<paragraph id="8BCC36F7D418456005D5E07584A69C37" blockId="13.[160,1156,1320,1727]" box="[160,440,1320,1346]" pageId="13" pageNumber="929">
|
||
<emphasis id="B907EAE5D418456005D5E07584A69C37" bold="true" box="[160,440,1320,1346]" italics="true" pageId="13" pageNumber="929">
|
||
Description of
|
||
<typeStatus id="54C88855D41845600425E07584A69C37" box="[336,440,1320,1346]" pageId="13" pageNumber="929" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D418457D05D5E010813D9F78" blockId="13.[160,1156,1320,1727]" lastBlockId="16.[160,1155,1490,1549]" lastPageId="16" lastPageNumber="932" pageId="13" pageNumber="929">
|
||
Adult male in excellent state of preservation, with a short (<
|
||
<quantity id="4C8B9B12D41845600621E01086929C10" box="[852,908,1357,1381]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="929" unit="cm" value="5.0">5 cm</quantity>
|
||
) ventral slit midbody. Body shape relatively gracile with a narrow head. Snout–vent length
|
||
<quantity id="4C8B9B12D418456006CCE03281089CF2" box="[953,1046,1391,1415]" metricMagnitude="-1" metricUnit="m" metricValue="4.55" pageId="13" pageNumber="929" unit="mm" value="455.0">455 mm</quantity>
|
||
with 181 ventral scales, tail length
|
||
<quantity id="4C8B9B12D418456004BDE0CF87399CDF" box="[456,551,1426,1450]" metricMagnitude="-1" metricUnit="m" metricValue="1.93" pageId="13" pageNumber="929" unit="mm" value="193.0">193 mm</quantity>
|
||
with 94 divided subcaudal scales. Anal scale single. Head length 17.0 mm, as measured from the snout to the rear of the quadrate, width
|
||
<quantity id="4C8B9B12D418456005D5E08A85EB9C9A" box="[160,245,1495,1519]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="13" pageNumber="929" unit="mm" value="8.7">8.7 mm</quantity>
|
||
as measured across the widest point of the head anterior to the quadrate. Eyes
|
||
<quantity id="4C8B9B12D418456005D5E0A785EA9F67" box="[160,244,1530,1554]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="13" pageNumber="929" unit="mm" value="2.6">2.6 mm</quantity>
|
||
horizontal diameter, pupil shape is subcircular. Supralabials seven with the third and fourth in contact with the eye. Infralabials eight, with first pair in contact behind mental, infralabials one to five in contact with inframaxillaries. Rostral broader than high,
|
||
<quantity id="4C8B9B12D418456005D5E33F85E69F0F" box="[160,248,1634,1658]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="929" unit="mm" value="3.2">3.2 mm</quantity>
|
||
wide/
|
||
<quantity id="4C8B9B12D41845600435E33F84849F0F" box="[320,410,1634,1658]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="929" unit="mm" value="2.2">2.2 mm</quantity>
|
||
high, visible from above. Nasal in contact with first supralabial. Single loreal, in contact with nasal, preoculars, prefrontal, and supralabials one, two and three. Circumoculars five: one supraocular, two preoculars, zero suboculars and two postoculars. Anterior temporals two. Dorsal surface of head includes pair of internasals (width
|
||
<quantity id="4C8B9B12D41A45620586E0AA84549F7A" box="[243,330,1527,1551]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="931" unit="mm" value="2.1">2.1 mm</quantity>
|
||
/length of suture
|
||
<quantity id="4C8B9B12D41A4562076FE0AA876B9F7A" box="[538,629,1527,1551]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="931" unit="mm" value="1.6">1.6 mm</quantity>
|
||
), pair of prefrontals (width
|
||
<quantity id="4C8B9B12D41A456206CDE0AA81119F7A" box="[952,1039,1527,1551]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="15" pageNumber="931" unit="mm" value="3.4">3.4 mm</quantity>
|
||
/length of suture
|
||
<quantity id="4C8B9B12D41A4562059AE34784599F47" box="[239,327,1562,1586]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="15" pageNumber="931" unit="mm" value="2.8">2.8 mm</quantity>
|
||
), pair of supraoculars (width 2.0 mm/length
|
||
<quantity id="4C8B9B12D41A45620637E34786849F47" box="[834,922,1562,1586]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="15" pageNumber="931" unit="mm" value="2.7">2.7 mm</quantity>
|
||
), frontal longer than wide (length
|
||
<quantity id="4C8B9B12D41A45620443E36184969F21" box="[310,392,1596,1620]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="15" pageNumber="931" unit="mm" value="4.6">4.6 mm</quantity>
|
||
/anterior width 4.0 mm), pair of parietals (length of suture
|
||
<quantity id="4C8B9B12D41A45620154E36181679F21" box="[1057,1145,1596,1620]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="15" pageNumber="931" unit="mm" value="5.1">5.1 mm</quantity>
|
||
). Two pairs of inframaxillaries, (anterior inframaxillary suture length
|
||
<quantity id="4C8B9B12D41A456206C6E30281119F02" box="[947,1039,1631,1655]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="15" pageNumber="931" unit="mm" value="2.6">2.6 mm</quantity>
|
||
, posterior inframaxillary suture length
|
||
<quantity id="4C8B9B12D41A4562049AE3DF87529FEF" box="[495,588,1666,1690]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="931" unit="mm" value="1.5">1.5 mm</quantity>
|
||
), posterior inframaxillaries in contact with each other. Dorsal scale rows 17−17−15 (15th ventral from anterior, midbody and 15th ventral anterior to cloaca). A distinct pair of apical pits on most scales on the dorsum.
|
||
</paragraph>
|
||
<caption id="DF0C667FD41B456305D5E31086209FEA" ID-DOI="http://doi.org/10.5281/zenodo.5174525" ID-Zenodo-Dep="5174525" httpUri="https://zenodo.org/record/5174525/files/figure.png" pageId="14" pageNumber="930" startId="14.[160,225,1613,1636]" targetBox="[334,981,142,1582]" targetPageId="14">
|
||
<paragraph id="8BCC36F7D41B456305D5E31086209FEA" blockId="14.[160,1156,1612,1695]" pageId="14" pageNumber="930">
|
||
<emphasis id="B907EAE5D41B456305D5E31085E49F11" bold="true" box="[160,250,1613,1636]" pageId="14" pageNumber="930">Figure 4.</emphasis>
|
||
Photographs of three
|
||
<taxonomicName id="4C734D74D41B456304AEE311878D9F11" box="[475,659,1612,1636]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="930" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B907EAE5D41B456304AEE311875B9F16" box="[475,581,1612,1635]" italics="true" pageId="14" pageNumber="930">Stegonotus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
described here in life; (a)
|
||
<taxonomicName id="4C734D74D41B456306E6E311812F9F16" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[915,1073,1612,1635]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="930" phylum="Chordata" rank="species" species="iridis" status="sp. nov.">
|
||
<emphasis id="B907EAE5D41B456306E6E311812F9F16" box="[915,1073,1612,1635]" italics="true" pageId="14" pageNumber="930">Stegonotus iridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED41B45630142E310819A9F11" box="[1079,1156,1613,1636]" pageId="14" pageNumber="930" rank="species">sp. nov.</taxonomicNameLabel>
|
||
SAMA R70466 (b)
|
||
<taxonomicName id="4C734D74D41B45630420E33787019FF4" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[341,543,1642,1665]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="930" phylum="Chordata" rank="species" species="derooijae" status="sp. nov.">
|
||
<emphasis id="B907EAE5D41B45630420E33787019FF4" box="[341,543,1642,1665]" italics="true" pageId="14" pageNumber="930">Stegonotus derooijae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED41B45630752E337876B9FF4" box="[551,629,1642,1665]" pageId="14" pageNumber="930" rank="species">sp. nov.</taxonomicNameLabel>
|
||
MZB.Ophi.3288 (c)
|
||
<taxonomicName id="4C734D74D41B4563064DE33781309FF4" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[824,1070,1642,1665]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="930" phylum="Chordata" rank="species" species="admiraltiensis" status="sp. nov.">
|
||
<emphasis id="B907EAE5D41B4563064DE33781309FF4" box="[824,1070,1642,1665]" italics="true" pageId="14" pageNumber="930">Stegonotus admiraltiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED41B45630143E337819A9FF4" box="[1078,1156,1642,1665]" pageId="14" pageNumber="930" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from Manus Island, unregistered. Photos from Stephen J. Richards.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0C667FD41A456205D5E03B819B9CEF" ID-DOI="http://doi.org/10.5281/zenodo.5174527" ID-Zenodo-Dep="5174527" httpUri="https://zenodo.org/record/5174527/files/figure.png" pageId="15" pageNumber="931" startId="15.[160,225,1382,1405]" targetBox="[163,1153,142,1351]" targetPageId="15">
|
||
<paragraph id="8BCC36F7D41A456205D5E03B819B9CEF" blockId="15.[160,1157,1381,1434]" pageId="15" pageNumber="931">
|
||
<emphasis id="B907EAE5D41A456205D5E03B85E39C08" bold="true" box="[160,253,1382,1405]" pageId="15" pageNumber="931">Figure 5.</emphasis>
|
||
Photographs of holotype
|
||
<taxonomicName id="4C734D74D41A45620774E03887829C09" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[513,668,1381,1404]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="931" phylum="Chordata" rank="species" species="iridis" status="sp. nov.">
|
||
<emphasis id="B907EAE5D41A45620774E03887829C09" box="[513,668,1381,1404]" italics="true" pageId="15" pageNumber="931">Stegonotus iridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED41A456207D6E03B87EF9C08" box="[675,753,1382,1405]" pageId="15" pageNumber="931" rank="species">sp. nov.</taxonomicNameLabel>
|
||
MZB.Ophi.3306 showing (clockwise from top left) dorsal of body, ventral of body, dorsal of head, lateral of head. Photos from Stephen J. Richards.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0C667FD405457D05D5E03481679CE8" ID-DOI="http://doi.org/10.5281/zenodo.5174529" ID-Zenodo-Dep="5174529" httpUri="https://zenodo.org/record/5174529/files/figure.png" pageId="16" pageNumber="932" startId="16.[160,225,1385,1408]" targetBox="[163,1153,142,1353]" targetPageId="16">
|
||
<paragraph id="8BCC36F7D405457D05D5E03481679CE8" blockId="16.[160,1155,1384,1437]" pageId="16" pageNumber="932">
|
||
<emphasis id="B907EAE5D405457D05D5E03485E59CF5" bold="true" box="[160,251,1385,1408]" pageId="16" pageNumber="932">Figure 6.</emphasis>
|
||
Photographs of paratype
|
||
<taxonomicName id="4C734D74D405457D0481E03587AD9C0A" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[500,691,1384,1407]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="932" phylum="Chordata" rank="species" species="derooijae" status="sp. nov.">
|
||
<emphasis id="B907EAE5D405457D0481E03587AD9C0A" box="[500,691,1384,1407]" italics="true" pageId="16" pageNumber="932">Stegonotus derooijae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED405457D07CDE034861F9CF5" box="[696,769,1385,1408]" pageId="16" pageNumber="932" rank="species">sp. nov.</taxonomicNameLabel>
|
||
MZB.Ophi.3293 showing (clockwise from top left) dorsal of body, ventral of body, dorsal of head, lateral of head. Photos from Stephen J. Richards.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC36F7D405457D05D5E36084749F22" blockId="16.[160,1155,1597,1727]" box="[160,362,1597,1623]" pageId="16" pageNumber="932">
|
||
<emphasis id="B907EAE5D405457D05D5E36084749F22" bold="true" box="[160,362,1597,1623]" italics="true" pageId="16" pageNumber="932">Coloration in life</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D405457D05D5E33F870A9FCA" blockId="16.[160,1155,1597,1727]" pageId="16" pageNumber="932">
|
||
Overall coloration in life similar to that described for preserved specimen except the white mottling on dorsal scales near the venter and lateral head scales has distinctly yellow pigment in life (
|
||
<figureCitation id="13482A72D405457D04DFE3FA87169FCA" box="[426,520,1703,1727]" captionStart="Figure 4" captionStartId="14.[160,225,1613,1636]" captionTargetBox="[334,981,142,1582]" captionTargetId="figure-1@14.[333,981,142,1582]" captionTargetPageId="14" captionText="Figure 4. Photographs of three Stegonotus species described here in life; (a) Stegonotus iridis sp. nov. SAMA R70466 (b) Stegonotus derooijae sp. nov. MZB.Ophi.3288 (c) Stegonotus admiraltiensis sp. nov. from Manus Island, unregistered. Photos from Stephen J. Richards." figureDoi="http://doi.org/10.5281/zenodo.5174525" httpUri="https://zenodo.org/record/5174525/files/figure.png" pageId="16" pageNumber="932">Figure 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D404457C05D5E5D084C999D2" blockId="17.[160,1156,141,583]" box="[160,471,141,167]" pageId="17" pageNumber="933">
|
||
<emphasis id="B907EAE5D404457C05D5E5D084C999D2" bold="true" box="[160,471,141,167]" italics="true" pageId="17" pageNumber="933">Coloration in preservative</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D404457C05D5E5EF87C29B32" blockId="17.[160,1156,141,583]" pageId="17" pageNumber="933">Overall dorsal coloration dark grey-brown with no pattern down the dorsal side of the body and tail but some dorsal scales nearest the venter whitish-cream. Colour of iris opaque grey. Dorsum of head, including rostral, internasals, prefrontals, frontal, supraoculars and parietals brownish-grey and showing iridescence, with extensive white mottling on dorsal and lateral head scales including temporal scales. Supralabials with grey pigmentation that is darker on the dorsal edge and becoming almost entirely cream approaching the ventral edge of the scales. Infralabials cream with limited mottled grey pigment on the edges of each scale. Ventral scales cream with limited grey pigmentation from the dorsal scales extending on to some ventral scales in small patches but not crossing the ventral scales. Anal plate cream, subcaudals all with grey pigmentation concentrated on the anterior edge of each scale and becoming greyer moving posteriorly. Ventral scales show iridescence under lights.</paragraph>
|
||
<paragraph id="8BCC36F7D404457C05D5E72E84109BF8" blockId="17.[160,1156,627,999]" box="[160,270,627,653]" pageId="17" pageNumber="933">
|
||
<emphasis id="B907EAE5D404457C05D5E72E84109BF8" bold="true" box="[160,270,627,653]" italics="true" pageId="17" pageNumber="933">Variation</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D404457C05D5E7CA84FE9A92" blockId="17.[160,1156,627,999]" pageId="17" pageNumber="933">
|
||
The
|
||
<typeStatus id="54C88855D404457C05A6E7CA84299BDA" box="[211,311,663,687]" pageId="17" pageNumber="933" type="holotype">holotype</typeStatus>
|
||
is the only sexually mature individual in the type series.
|
||
<materialsCitation id="3B1B3CAAD404457C06BAE7CA86B49BA7" accessionNumber="R70467" collectionCode="SAMA, R, MZB" pageId="17" pageNumber="933" specimenCount="1">
|
||
At just
|
||
<quantity id="4C8B9B12D404457C0151E7CA819D9BDA" box="[1060,1155,663,687]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" pageId="17" pageNumber="933" unit="mm" value="290.0">290 mm</quantity>
|
||
snout–vent length,
|
||
<collectionCode id="ED62AE32D404457C040AE7E784DC9BA7" box="[383,450,698,722]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="17" pageNumber="933" type="Museum">SAMA</collectionCode>
|
||
<collectionCode id="ED62AE32D404457C04BEE7E784C49BA7" box="[459,474,698,722]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="933">R</collectionCode>
|
||
70467 is clearly a juvenile, whereas
|
||
<collectionCode id="ED62AE32D404457C0601E7E786B49BA7" box="[884,938,698,722]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="17" pageNumber="933" type="Museum">MZB</collectionCode>
|
||
</materialsCitation>
|
||
.Ophi.3293 is possibly a juvenile female because it lacks any sign of gonadal development under dissection despite being around the same size as the sexually mature male
|
||
<typeStatus id="54C88855D404457C0600E7A286C29A62" box="[885,988,767,791]" pageId="17" pageNumber="933" type="holotype">holotype</typeStatus>
|
||
.
|
||
<materialsCitation id="3B1B3CAAD404457C0690E7A284439A29" accessionNumber="R70467" collectionCode="MZB, SAMA, R" pageId="17" pageNumber="933" specimenCount="1" typeStatus="paratype">
|
||
Apical pits are absent in the
|
||
<specimenCount id="9D75FD7ED404457C0439E67F84EC9A4F" box="[332,498,802,826]" pageId="17" pageNumber="933" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
(
|
||
<collectionCode id="ED62AE32D404457C0771E67F87259A4F" box="[516,571,802,826]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="17" pageNumber="933" type="Museum">MZB</collectionCode>
|
||
.Ophi.3293,
|
||
<collectionCode id="ED62AE32D404457C07B6E67F86189A4F" box="[707,774,802,826]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="17" pageNumber="933" type="Museum">SAMA</collectionCode>
|
||
<collectionCode id="ED62AE32D404457C0666E67F863C9A4F" box="[787,802,802,826]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="933">R</collectionCode>
|
||
70467) and referred specimen (
|
||
<collectionCode id="ED62AE32D404457C05D3E61985C09A29" box="[166,222,836,860]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="17" pageNumber="933" type="Museum">MZB</collectionCode>
|
||
.Ophi.3292)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B1B3CAAD404457C0418E619877E9A0A" collectionCode="MZB" pageId="17" pageNumber="933" specimenCount="2" typeStatus="paratype">
|
||
In overall colour pattern, scalation and body shape
|
||
<typeStatus id="54C88855D404457C0691E619814C9A29" box="[996,1106,836,860]" pageId="17" pageNumber="933" type="paratype">paratypes</typeStatus>
|
||
are similar to the
|
||
<typeStatus id="54C88855D404457C044AE63A84BD9A0A" box="[319,419,871,895]" pageId="17" pageNumber="933" type="holotype">holotype</typeStatus>
|
||
excepting:
|
||
<collectionCode id="ED62AE32D404457C075FE63A877E9A0A" box="[554,608,871,895]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="17" pageNumber="933" type="Museum">MZB</collectionCode>
|
||
</materialsCitation>
|
||
.Ophi.
|
||
<materialsCitation id="3B1B3CAAD404457C07EEE63A86259AD7" accessionNumber="R70467" collectionCode="SAMA, R, MZB" pageId="17" pageNumber="933" specimenCount="1" typeStatus="paratype">
|
||
3293 and
|
||
<collectionCode id="ED62AE32D404457C0664E63A864A9A0A" box="[785,852,871,895]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="17" pageNumber="933" type="Museum">SAMA</collectionCode>
|
||
<collectionCode id="ED62AE32D404457C0629E63A86759A0A" box="[860,875,871,895]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="933">R</collectionCode>
|
||
70467 lack the extensive white mottling on the dorsal and lateral head scales;
|
||
<collectionCode id="ED62AE32D404457C0670E6D786259AD7" box="[773,827,906,930]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="17" pageNumber="933" type="Museum">MZB</collectionCode>
|
||
</materialsCitation>
|
||
.Ophi.
|
||
<materialsCitation id="3B1B3CAAD404457C0603E6D784C29A92" accessionNumber="R70467" collectionCode="SAMA, R" pageId="17" pageNumber="933" specimenCount="1" typeStatus="paratype">
|
||
3293 with 178 ventrals, 81 subcaudals (tail incomplete), nine rather than eight infralabials;
|
||
<collectionCode id="ED62AE32D404457C06DDE6F186F59AB1" box="[936,1003,940,964]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="17" pageNumber="933" type="Museum">SAMA</collectionCode>
|
||
<collectionCode id="ED62AE32D404457C0680E6F1811A9AB1" box="[1013,1028,940,964]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="933">R</collectionCode>
|
||
70467 with 178 ventrals, 90 subcaudals
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C369657CD404457F05D5E14E86B099BC" lastPageId="18" lastPageNumber="934" pageId="17" pageNumber="933" type="etymology">
|
||
<paragraph id="8BCC36F7D404457C05D5E14E84039D58" blockId="17.[160,1155,1043,1173]" box="[160,285,1043,1069]" pageId="17" pageNumber="933">
|
||
<emphasis id="B907EAE5D404457C05D5E14E84039D58" bold="true" box="[160,285,1043,1069]" italics="true" pageId="17" pageNumber="933">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D404457C05D5E16A849E9DE0" blockId="17.[160,1155,1043,1173]" pageId="17" pageNumber="933">
|
||
The species epithet is a noun in the genitive case and is a patronym in honour of Dr Nelly De Rooij, who described many taxa from New
|
||
<collectingCountry id="F3647667D404457C0678E10786429D07" box="[781,860,1114,1138]" name="Guinea-Bissau" pageId="17" pageNumber="933">Guinea</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3647667D404457C06E8E10781129D07" box="[925,1036,1114,1138]" name="Indonesia" pageId="17" pageNumber="933">Indonesia</collectingCountry>
|
||
, including
|
||
<taxonomicName id="4C734D74D404457C05D5E12084649DE0" box="[160,378,1148,1173]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="933" phylum="Chordata" rank="species" species="florensis">
|
||
<emphasis id="B907EAE5D404457C05D5E12084649DE0" box="[160,378,1148,1173]" italics="true" pageId="17" pageNumber="933">Stegonotus florensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D404457C05D5E19D87039DAF" blockId="17.[160,1155,1216,1381]" box="[160,541,1216,1242]" pageId="17" pageNumber="933">
|
||
<emphasis id="B907EAE5D404457C05D5E19D87039DAF" bold="true" box="[160,541,1216,1242]" italics="true" pageId="17" pageNumber="933">Natural history and distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D404457C05D5E1B886849C10" blockId="17.[160,1155,1216,1381]" pageId="17" pageNumber="933">
|
||
Currently known only from the Raja Ampat islands of Batanta, Salawati and Waigeo in eastern
|
||
<collectingCountry id="F3647667D404457C0588E05A84729C6A" box="[253,364,1287,1311]" name="Indonesia" pageId="17" pageNumber="933">Indonesia</collectingCountry>
|
||
. This small, slender species was active on the forest floor in moderately to heavily disturbed lowland (<
|
||
<quantity id="4C8B9B12D404457C073DE07787919C37" box="[584,655,1322,1346]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="933" unit="m" value="100.0">100 m</quantity>
|
||
above sea level) rainforest at night. At each site it occurred in sympatry with
|
||
<taxonomicName id="4C734D74D404457C0769E01087749C10" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[540,618,1357,1381]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="933" phylum="Chordata" rank="species" species="iridis" status="sp. nov.">
|
||
<emphasis id="B907EAE5D404457C0769E01087749C10" box="[540,618,1357,1381]" italics="true" pageId="17" pageNumber="933">S. iridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED404457C0707E01087D39C10" box="[626,717,1357,1381]" pageId="17" pageNumber="933" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, described above.
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D404457C05D5E0CD84249CDF" blockId="17.[160,1156,1424,1727]" box="[160,314,1424,1450]" pageId="17" pageNumber="933">
|
||
<emphasis id="B907EAE5D404457C05D5E0CD84249CDF" bold="true" box="[160,314,1424,1450]" italics="true" pageId="17" pageNumber="933">Comparisons</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D404457F05D5E0E886B099BC" blockId="17.[160,1156,1424,1727]" lastBlockId="18.[160,1156,142,201]" lastPageId="18" lastPageNumber="934" pageId="17" pageNumber="933">
|
||
<taxonomicName id="4C734D74D404457C05D5E0E884969CB8" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[160,392,1461,1485]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="933" phylum="Chordata" rank="species" species="derooijae" status="sp. nov.">
|
||
<emphasis id="B907EAE5D404457C05D5E0E884969CB8" box="[160,392,1461,1485]" italics="true" pageId="17" pageNumber="933">Stegonotus derooijae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED404457C04E7E0E884F29CB8" box="[402,492,1461,1485]" pageId="17" pageNumber="933" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is part of the
|
||
<taxonomicName id="4C734D74D404457C07EDE0E887EE9CB8" authorityName="Lindholm. It" authorityYear="1905" box="[664,752,1461,1485]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="933" phylum="Chordata" rank="species" species="diehli">
|
||
<emphasis id="B907EAE5D404457C07EDE0E887EE9CB8" box="[664,752,1461,1485]" italics="true" pageId="17" pageNumber="933">S. diehli</emphasis>
|
||
</taxonomicName>
|
||
complex (
|
||
<figureCitation id="13482A72D404457C0619E0E886D59CB8" box="[876,971,1461,1485]" captionStart="Figure 3" captionStartId="9.[160,225,1677,1700]" captionTargetBox="[217,1128,144,1646]" captionTargetId="graphics-2@9.[256,1128,623,1646]" captionTargetPageId="9" captionText="Figure 3. Map of New Guinea sampling localities for Stegonotus (a) and mtDNA + nucDNA maximum likelihood gene tree (b). Symbols for species/clades match those on the map; * indicates bootstrap values ≥ 70. For some localities, symbols were adjusted for better visibility and so localities are approximate; see Appendix 1. for exact locality information." figureDoi="http://doi.org/10.5281/zenodo.5174522" httpUri="https://zenodo.org/record/5174522/files/figure.png" pageId="17" pageNumber="933">Figure 3</figureCitation>
|
||
) and this taxon would have been formerly considered
|
||
<taxonomicName id="4C734D74D404457C0724E08A87BB9C9A" authorityName="Lindholm. It" authorityYear="1905" box="[593,677,1495,1519]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="933" phylum="Chordata" rank="species" species="diehli">
|
||
<emphasis id="B907EAE5D404457C0724E08A87BB9C9A" box="[593,677,1495,1519]" italics="true" pageId="17" pageNumber="933">S. diehli</emphasis>
|
||
</taxonomicName>
|
||
. It differs morphologically from
|
||
<taxonomicName id="4C734D74D404457C0178E08A817F9C9A" authorityName="Lindholm. It" authorityYear="1905" box="[1037,1121,1495,1519]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="933" phylum="Chordata" rank="species" species="diehli">
|
||
<emphasis id="B907EAE5D404457C0178E08A817F9C9A" box="[1037,1121,1495,1519]" italics="true" pageId="17" pageNumber="933">S. diehli</emphasis>
|
||
</taxonomicName>
|
||
by having 17 dorsal scale rows at midbody versus 15 dorsal scale rows in
|
||
<emphasis id="B907EAE5D404457C0157E0A784949F40" italics="true" pageId="17" pageNumber="933">
|
||
<taxonomicName id="4C734D74D404457C0157E0A7819E9F67" authorityName="Lindholm. It" authorityYear="1905" box="[1058,1152,1530,1554]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="933" phylum="Chordata" rank="species" species="diehli">S. diehli</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C734D74D404457C05D5E34084949F40" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[160,394,1565,1589]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="933" phylum="Chordata" rank="species" species="derooijae" status="sp. nov.">Stegonotus derooijae</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A234579ED404457C04E3E34084EC9F40" box="[406,498,1565,1589]" pageId="17" pageNumber="933" rank="species">sp. nov.</taxonomicNameLabel>
|
||
also differs from
|
||
<taxonomicName id="4C734D74D404457C07BCE340863C9F40" authorityName="Lindholm. It" authorityYear="1905" box="[713,802,1565,1589]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="933" phylum="Chordata" rank="species" species="diehli">
|
||
<emphasis id="B907EAE5D404457C07BCE340863C9F40" box="[713,802,1565,1589]" italics="true" pageId="17" pageNumber="933">S. diehli</emphasis>
|
||
</taxonomicName>
|
||
in having higher numbers of ventral scales (
|
||
<quantity id="4C8B9B12D404457C0436E36284DD9F22" box="[323,451,1599,1623]" metricMagnitude="0" metricUnit="m" metricValue="4.7625" metricValueMax="5.0038" metricValueMin="4.5212" pageId="17" pageNumber="933" unit="in" value="187.5" valueMax="197.0" valueMin="178.0">178−197 in</quantity>
|
||
<taxonomicName id="4C734D74D404457C04BCE36287599F22" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[457,583,1599,1623]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="933" phylum="Chordata" rank="species" species="derooijae" status="sp. nov.">
|
||
<emphasis id="B907EAE5D404457C04BCE36287599F22" box="[457,583,1599,1623]" italics="true" pageId="17" pageNumber="933">S. derooijae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED404457C073BE36287BB9F22" box="[590,677,1599,1623]" pageId="17" pageNumber="933" rank="species">sp. nov.</taxonomicNameLabel>
|
||
versus
|
||
<quantity id="4C8B9B12D404457C078DE36286689F22" box="[760,886,1599,1623]" metricMagnitude="0" metricUnit="m" metricValue="4.2672" metricValueMax="4.4196" metricValueMin="4.1148" pageId="17" pageNumber="933" unit="in" value="168.0" valueMax="174.0" valueMin="162.0">162−174 in</quantity>
|
||
<taxonomicName id="4C734D74D404457C0608E36286CF9F22" authorityName="Lindholm. It" authorityYear="1905" box="[893,977,1599,1623]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="933" phylum="Chordata" rank="species" species="diehli">
|
||
<emphasis id="B907EAE5D404457C0608E36286CF9F22" box="[893,977,1599,1623]" italics="true" pageId="17" pageNumber="933">S. diehli</emphasis>
|
||
</taxonomicName>
|
||
) and subcaudal scales (
|
||
<quantity id="4C8B9B12D404457C0584E33F84469F0F" box="[241,344,1634,1658]" metricMagnitude="0" metricUnit="m" metricValue="2.3368" metricValueMax="2.3876" metricValueMin="2.286" pageId="17" pageNumber="933" unit="in" value="92.0" valueMax="94.0" valueMin="90.0">90−94 in</quantity>
|
||
<taxonomicName id="4C734D74D404457C0417E33F84FD9F0F" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[354,483,1634,1658]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="933" phylum="Chordata" rank="species" species="derooijae">
|
||
<emphasis id="B907EAE5D404457C0417E33F84709F0F" box="[354,366,1634,1658]" italics="true" pageId="17" pageNumber="933">S</emphasis>
|
||
.
|
||
<emphasis id="B907EAE5D404457C040BE33F84FD9F0F" box="[382,483,1634,1658]" italics="true" pageId="17" pageNumber="933">derooijae</emphasis>
|
||
</taxonomicName>
|
||
versus
|
||
<quantity id="4C8B9B12D404457C0749E33F87BC9F0F" box="[572,674,1634,1658]" metricMagnitude="0" metricUnit="m" metricValue="2.0065999999999997" metricValueMax="2.159" metricValueMin="1.8542" pageId="17" pageNumber="933" unit="in" value="79.0" valueMax="85.0" valueMin="73.0">73−85 in</quantity>
|
||
<taxonomicName id="4C734D74D404457C07DEE33F861C9F0F" authorityName="Lindholm. It" authorityYear="1905" box="[683,770,1634,1658]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="933" phylum="Chordata" rank="species" species="diehli">
|
||
<emphasis id="B907EAE5D404457C07DEE33F87A99F0F" box="[683,695,1634,1658]" italics="true" pageId="17" pageNumber="933">S</emphasis>
|
||
.
|
||
<emphasis id="B907EAE5D404457C07B2E33F861C9F0F" box="[711,770,1634,1658]" italics="true" pageId="17" pageNumber="933">diehli</emphasis>
|
||
</taxonomicName>
|
||
) for the specimens we examined (excluding specimens with incomplete tails; Appendix 1). It differs from the new species (described below) from
|
||
<collectingRegion id="49B7F815D404457C04D8E3FA87929FCA" box="[429,652,1703,1727]" country="Papua New Guinea" name="Southern Highlands" pageId="17" pageNumber="933">Southern Highlands</collectingRegion>
|
||
and
|
||
<collectingRegion id="49B7F815D404457C07B0E3FA86009FCA" box="[709,798,1703,1727]" country="Papua New Guinea" name="Chimbu" pageId="17" pageNumber="933">Chimbu</collectingRegion>
|
||
Provinces by having two rather than (usually) one preocular scale. The overall dark coloration of
|
||
<taxonomicName id="4C734D74D407457F06E8E5D3810199D3" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[925,1055,142,166]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="934" phylum="Chordata" rank="species" species="derooijae" status="sp. nov.">
|
||
<emphasis id="B907EAE5D407457F06E8E5D3810199D3" box="[925,1055,142,166]" italics="true" pageId="18" pageNumber="934">S. derooijae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED407457F015CE5D3819A99D3" box="[1065,1156,142,166]" pageId="18" pageNumber="934" rank="species">sp. nov.</taxonomicNameLabel>
|
||
distinguishes it from the sympatric
|
||
<taxonomicName id="4C734D74D407457F0740E5EC879D99BC" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[565,643,177,201]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="934" phylum="Chordata" rank="species" species="iridis" status="sp. nov.">
|
||
<emphasis id="B907EAE5D407457F0740E5EC879D99BC" box="[565,643,177,201]" italics="true" pageId="18" pageNumber="934">S. iridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED407457F07F9E5EC87FA99BC" box="[652,740,177,201]" pageId="18" pageNumber="934" rank="species">sp. nov.</taxonomicNameLabel>
|
||
described above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |