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<document id="AEA55F51805EB213877AAE287B7EB2CA" ID-CLB-Dataset="6281" ID-DOI="10.11646/phytotaxa.15.1.3" ID-GBIF-Dataset="9d402fa7-54cf-4147-b17a-44d15ac482a4" ID-ISSN="1179-3163" ID-Zenodo-Dep="4906379" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622644583337" checkinUser="felipe" docAuthor="Porter, J. Mark" docDate="2011" docId="03DA87C9FFFC936AE0B1FF70FA13F644" docLanguage="en" docName="Phytotaxa.15.15-25.pdf.imf" docOrigin="Phytotaxa 15" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Aliciella cliffordii J. M. Porter 2011, sp. nov." docType="treatment" docVersion="6" lastPageNumber="20" masterDocId="FFE3FFB1FFF8936FE026FFE7FF8FF177" masterDocTitle="Two new Aliciella species and a new subspecies in Ipomopsis (Polemoniaceae) from the western United States of America" masterLastPageNumber="25" masterPageNumber="15" pageNumber="19" updateTime="1698996633771" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5E53B562CF4DA5DDA6A021CA48D17116">Two new Aliciella species and a new subspecies in Ipomopsis (Polemoniaceae) from the western United States of America</mods:title>
</mods:titleInfo>
<mods:name id="A1E73D97BBF2C46A35E6821E6B981FDA" type="personal">
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<mods:namePart id="CA74D3D205A74C884815471613F65122">Porter, J. Mark</mods:namePart>
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<mods:title id="70766C8B0A26AD56630658AE6C315582">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="07960828F279EDE76AF86327FB6E99E7">
<mods:date id="93106ACFFE265FE5870D9C5FEFF5D980">2011</mods:date>
<mods:detail id="AA8635FEAB2A3B19CFBBEAC7C0FE5164" type="pubDate">
<mods:number id="ECD00032F3B4D58563E3D5B3619ADA53">2011-12-31</mods:number>
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<mods:classification id="884FC45A78CA9211F4D704EA8ACB7DA3">journal article</mods:classification>
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<mods:identifier id="F7ECF13B7D8A8FDE96810D69A2731965" type="DOI">10.11646/phytotaxa.15.1.3</mods:identifier>
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<treatment id="03DA87C9FFFC936AE0B1FF70FA13F644" ID-DOI="http://doi.org/10.5281/zenodo.4906389" ID-GBIF-Taxon="182597800" ID-Zenodo-Dep="4906389" LSID="urn:lsid:plazi:treatment:03DA87C9FFFC936AE0B1FF70FA13F644" httpUri="http://treatment.plazi.org/id/03DA87C9FFFC936AE0B1FF70FA13F644" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
<subSubSection id="C3696554FFFC936BE0B1FF70FD4BF1C5" box="[151,708,151,178]" pageId="4" pageNumber="19" type="nomenclature">
<paragraph id="8BCC36DFFFFC936BE0B1FF70FD4BF1C5" blockId="4.[151,708,151,178]" box="[151,708,151,178]" pageId="4" pageNumber="19">
<heading id="D08481B3FFFC936BE0B1FF70FD4BF1C5" box="[151,708,151,178]" fontSize="11" level="2" pageId="4" pageNumber="19" reason="3">
<emphasis id="B907EACDFFFC936BE0B1FF70FDEAF1C6" bold="true" box="[151,613,151,177]" pageId="4" pageNumber="19">
<taxonomicName id="4C734D5CFFFC936BE0B1FF70FE73F1C6" ID-CoL="BS4D" authority="J.M.Porter" authorityName="J. M. Porter" authorityYear="2011" box="[151,508,151,177]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="cliffordii" status="sp. nov.">
<emphasis id="B907EACDFFFC936BE0B1FF70FEE3F1C6" bold="true" box="[151,364,151,177]" italics="true" pageId="4" pageNumber="19">Aliciella cliffordii</emphasis>
J.M.Porter
</taxonomicName>
,
<emphasis id="B907EACDFFFC936BE22CFF70FDEAF1C6" bold="true" box="[522,613,151,177]" italics="true" pageId="4" pageNumber="19">
<taxonomicNameLabel id="A23457B6FFFC936BE22CFF70FDEAF1C6" box="[522,613,151,177]" pageId="4" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</emphasis>
(
<figureCitation id="13482A5AFFFC936BE253FF7FFD33F1C5" box="[629,700,152,178]" captionStart="FIGURE 2" captionStartId="4.[151,255,1074,1098]" captionTargetBox="[151,1436,458,1050]" captionTargetId="figure-447@4.[151,1436,458,1050]" captionTargetPageId="4" captionText="FIGURE 2. A. Habit and architecture of Aliciella cliffordii, scale= 3.0 cm, J.M.Porter, K.D.Heil, &amp; A.Clifford 13528 (Photo J.M.Porter). B. Cymose inflorescence and flowers, scale= 5.0 mm, J.M.Porter, K.D.Heil, &amp; A.Clifford 13528 (Photo J.M.Porter). C. Bombilius sp. visiting flowers of Aliciella cliffordii, scale= 5.0 mm, J.M.Porter, K.D.Heil, &amp; A.Clifford 13528 (Photo J.M.Porter)." figureDoi="http://doi.org/10.5281/zenodo.4906385" httpUri="https://zenodo.org/record/4906385/files/figure.png" pageId="4" pageNumber="19">Fig. 2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3696554FFFC936BE0B1FF02FABCF18B" box="[151,1331,229,252]" pageId="4" pageNumber="19" type="diagnosis">
<paragraph id="8BCC36DFFFFC936BE0B1FF02FABCF18B" blockId="4.[151,1331,229,252]" box="[151,1331,229,252]" pageId="4" pageNumber="19">
<emphasis id="B907EACDFFFC936BE0B1FF02FABCF18B" box="[151,1331,229,252]" italics="true" pageId="4" pageNumber="19">
Species
<taxonomicName id="4C734D5CFFFC936BE0CBFF02FE77F18B" box="[237,504,229,252]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="subSpecies" species="cliffordii" subSpecies="similis">Aliciella cliffordii similis</taxonomicName>
<taxonomicName id="4C734D5CFFFC936BE1D8FF02FCA7F18B" authority="(A. Gray)" authorityName="J. M. Porter" authorityYear="1998" baseAuthorityName="A. Gray" box="[510,808,229,252]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="haydenii">Aliciella haydenii (A. Gray)</taxonomicName>
J.M.
<taxonomicName id="4C734D5CFFFC936BE37BFF02FC2CF18B" authority="Porter, 2011" authorityName="Porter" authorityYear="2011" box="[861,931,229,252]" pageId="4" pageNumber="21" rank="subSpecies" status="subsp. nov." subSpecies="porter">Porter</taxonomicName>
, sed corolla caerulea vel
<taxonomicName id="4C734D5CFFFC936BE492FF02FB6CF18B" authorityName="Moench" authorityYear="1802" box="[1204,1251,229,252]" class="Magnoliopsida" family="Polemoniaceae" genus="Phlox" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="alba">alba</taxonomicName>
differt.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3696554FFFC936BE0B1FECBFDFAF6F8" pageId="4" pageNumber="19" type="materials_examined">
<paragraph id="8BCC36DFFFFC936BE0B1FECBFE39F0E4" blockId="4.[151,1436,300,403]" pageId="4" pageNumber="19">
<materialsCitation id="3B1B3C82FFFC936BE0B1FECBFE3EF0E4" ID-GBIF-Occurrence="3127432301" collectingDate="2003-05-06" collectionCode="ARIZ, SJNM" collectorName="J. Mark Porter &amp; K. D. Heil &amp; A. Clifford" country="United States of America" county="Apache County" elevation="2225" latitude="36.53361" location="Apache County" longLatPrecision="19" longitude="-109.21778" pageId="4" pageNumber="19" specimenCount="1" stateProvince="Arizona" typeStatus="holotype">
<collectorName id="26865309FFFC936BE0B1FECBFF57F031" box="[151,216,300,326]" pageId="4" pageNumber="19">Type</collectorName>
:—
<emphasis id="B907EACDFFFC936BE0D8FECBFC94F032" box="[254,795,300,325]" italics="true" pageId="4" pageNumber="19">
<collectorName id="26865309FFFC936BE0D8FECBFEB9F032" box="[254,310,300,325]" name="J. Mark Porter" pageId="4" pageNumber="19">J.M.</collectorName>
<taxonomicName id="4C734D5CFFFC936BE110FECBFE05F032" authority="Porter, 2011" authorityName="Porter" authorityYear="2011" box="[310,394,300,325]" pageId="4" pageNumber="21" rank="subSpecies" status="subsp. nov." subSpecies="porter">Porter</taxonomicName>
,
<collectorName id="26865309FFFC936BE1BCFECBFD9EF032" box="[410,529,300,325]" pageId="4" pageNumber="19">K.D.Heil</collectorName>
, &amp;
<collectorName id="26865309FFFC936BE264FECBFD4DF032" box="[578,706,300,325]" pageId="4" pageNumber="19">A.Clifford</collectorName>
13528
</emphasis>
;
<collectingCountry id="F364764FFFFC936BE309FECBFC09F031" box="[815,902,300,326]" name="United States of America" pageId="4" pageNumber="19">U.S.A.</collectingCountry>
<collectingRegion id="49B7F83DFFFC936BE3B7FECBFC74F031" box="[913,1019,300,326]" country="United States of America" name="Arizona" pageId="4" pageNumber="19">Arizona</collectingRegion>
.
<collectingCounty id="62AD4E53FFFC936BE42CFECBFB59F031" box="[1034,1238,300,326]" pageId="4" pageNumber="19">Apache County</collectingCounty>
:
<collectorName id="26865309FFFC936BE4C3FECBFEA9F01B" pageId="4" pageNumber="19">Navajo Indian Reservation</collectorName>
, ca.
<locationDeviation id="9A7E2FFDFFFC936BE17CFEB5FD9BF01B" box="[346,532,338,364]" location="Cove" pageId="4" pageNumber="19">
<quantity id="4C8B9B3AFFFC936BE17CFEB5FE1BF01B" box="[346,404,338,364]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="19" unit="km" value="5.0">5 km</quantity>
S of Cove
</locationDeviation>
;
<geoCoordinate id="EE475018FFFC936BE206FEB5FD47F01B" box="[544,712,338,364]" degrees="36" direction="north" minutes="32" orientation="latitude" pageId="4" pageNumber="19" precision="15" seconds="01" value="36.53361">36° 32’ 01” N</geoCoordinate>
,
<geoCoordinate id="EE475018FFFC936BE2F2FEB5FC00F01B" box="[724,911,338,364]" degrees="109" direction="west" minutes="13" orientation="longitude" pageId="4" pageNumber="19" precision="15" seconds="04" value="-109.21778">109° 13’ 04” W</geoCoordinate>
,
<quantity id="4C8B9B3AFFFC936BE3BCFEB5FC7CF01B" box="[922,1011,338,364]" metricMagnitude="3" metricUnit="m" metricValue="2.225" pageId="4" pageNumber="19" unit="m" value="2225.0">
<elevation id="005ED1ECFFFC936BE3BCFEB5FC7CF01B" box="[922,1011,338,364]" metricMagnitude="3" metricUnit="m" metricValue="2.225" pageId="4" pageNumber="19" unit="m" value="2225.0">2225 m</elevation>
</quantity>
elev.,
<date id="FFCD101FFFFC936BE419FEB5FB5FF01B" box="[1087,1232,338,364]" pageId="4" pageNumber="19" value="2003-05-06">
<collectingDate id="EF89E9F7FFFC936BE419FEB5FB5FF01B" box="[1087,1232,338,364]" pageId="4" pageNumber="19" value="2003-05-06">6 May 2003</collectingDate>
</date>
. (
<typeStatus id="54C8887DFFFC936BE4C5FEB5FAC0F01B" box="[1251,1359,338,364]" pageId="4" pageNumber="19" type="holotype">holotype</typeStatus>
:
<collectingCountry id="F364764FFFFC936BE57DFEB5FA17F01B" box="[1371,1432,338,364]" name="South Africa" pageId="4" pageNumber="19">RSA</collectingCountry>
; isotypes:
<collectionCode id="ED62AE1AFFFC936BE120FE9EFEC2F0E4" box="[262,333,377,403]" country="USA" lsid="urn:lsid:biocol.org:col:15814" name="University of Arizona" pageId="4" pageNumber="19" type="Herbarium">ARIZ</collectionCode>
,
<collectionCode id="ED62AE1AFFFC936BE17FFE9EFE26F0E4" box="[345,425,377,403]" country="USA" lsid="urn:lsid:biocol.org:col:14919" name="San Juan College" pageId="4" pageNumber="19" type="Herbarium">SJNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="DF0C6657FFFC936BE0B1FBD5FDAFF5C6" ID-DOI="http://doi.org/10.5281/zenodo.4906385" ID-Zenodo-Dep="4906385" httpUri="https://zenodo.org/record/4906385/files/figure.png" pageId="4" pageNumber="19" startId="4.[151,255,1074,1098]" targetBox="[151,1436,458,1050]" targetPageId="4">
<paragraph id="8BCC36DFFFFC936BE0B1FBD5FDAFF5C6" blockId="4.[151,1436,1074,1201]" pageId="4" pageNumber="19">
<emphasis id="B907EACDFFFC936BE0B1FBD5FE95F53D" bold="true" box="[151,282,1074,1098]" pageId="4" pageNumber="19">FIGURE 2.</emphasis>
A. Habit and architecture of
<taxonomicName id="4C734D5CFFFC936BE27BFBD5FC90F53E" authorityName="J. M. Porter" authorityYear="2011" box="[605,799,1074,1097]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="cliffordii">
<emphasis id="B907EACDFFFC936BE27BFBD5FC90F53E" box="[605,799,1074,1097]" italics="true" pageId="4" pageNumber="19">Aliciella cliffordii</emphasis>
</taxonomicName>
, scale= 3.0 cm,
<emphasis id="B907EACDFFFC936BE3F2FBD5FA14F53E" box="[980,1435,1074,1097]" italics="true" pageId="4" pageNumber="19">
J.M.
<taxonomicName id="4C734D5CFFFC936BE422FBD5FBC4F53E" authority="Porter, 2011" authorityName="Porter" authorityYear="2011" box="[1028,1099,1074,1097]" pageId="4" pageNumber="21" rank="subSpecies" status="subsp. nov." subSpecies="porter">Porter</taxonomicName>
, K.D.Heil, &amp; A.Clifford 13528
</emphasis>
(Photo J.M.Porter). B. Cymose inflorescence and flowers, scale= 5.0 mm,
<emphasis id="B907EACDFFFC936BE3F6FBB2FA13F51B" box="[976,1436,1109,1132]" italics="true" pageId="4" pageNumber="19">
J.M.
<taxonomicName id="4C734D5CFFFC936BE426FBB2FBC7F51B" authority="Porter, 2011" authorityName="Porter" authorityYear="2011" box="[1024,1096,1109,1132]" pageId="4" pageNumber="21" rank="subSpecies" status="subsp. nov." subSpecies="porter">Porter</taxonomicName>
, K.D.Heil, &amp; A.Clifford 13528
</emphasis>
(Photo J.M.Porter). C.
<emphasis id="B907EACDFFFC936BE1B0FB9FFD8AF5F8" box="[406,517,1144,1167]" italics="true" pageId="4" pageNumber="19">Bombilius</emphasis>
sp. visiting flowers of
<taxonomicName id="4C734D5CFFFC936BE32FFB9FFC40F5F8" authorityName="J. M. Porter" authorityYear="2011" box="[777,975,1144,1167]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="cliffordii">
<emphasis id="B907EACDFFFC936BE32FFB9FFC40F5F8" box="[777,975,1144,1167]" italics="true" pageId="4" pageNumber="19">Aliciella cliffordii</emphasis>
</taxonomicName>
, scale= 5.0 mm,
<emphasis id="B907EACDFFFC936BE4B6FB9FFEC5F5C6" italics="true" pageId="4" pageNumber="19">
J.M.
<taxonomicName id="4C734D5CFFFC936BE4E6FB9FFA86F5F8" authority="Porter, 2011" authorityName="Porter" authorityYear="2011" box="[1216,1289,1144,1167]" pageId="4" pageNumber="21" rank="subSpecies" status="subsp. nov." subSpecies="porter">Porter</taxonomicName>
, K.D.Heil, &amp; A.Clifford 13528
</emphasis>
(Photo J.M.Porter).
</paragraph>
</caption>
<paragraph id="8BCC36DFFFFC936BE0E0FB03FDFAF6F8" blockId="4.[151,1436,1252,2012]" pageId="4" pageNumber="19">
Taprooted, short-lived perennial, with a well-developed basal rosette of leaves, thus appearing biennial,
<quantity id="4C8B9B3AFFFC936BE0B1FAEDFE83F453" box="[151,268,1290,1316]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="9.0" metricValueMin="1.0" pageId="4" pageNumber="19" unit="cm" value="50.0" valueMax="90.0" valueMin="10.0">10–90 cm</quantity>
tall, stems sparsely and coarsely glandular pubescent with uniseriate glandular trichomes, erect but freely and divaricately branching, sometimes to the base, the lateral branches overtopping the primary axis. Basal leaves forming a rosette, entire, coarsely toothed to once-pinnatifid,
<quantity id="4C8B9B3AFFFC936BE3D8FABFFBE3F405" box="[1022,1132,1368,1394]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="5.0" metricValueMin="1.2" pageId="4" pageNumber="19" unit="cm" value="3.1" valueMax="5.0" valueMin="1.2">1.2–5 cm</quantity>
long, the rachis broad,
<quantity id="4C8B9B3AFFFC936BE559FABFFF7FF4EF" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="4.5" metricValueMin="1.0" pageId="4" pageNumber="19" unit="mm" value="2.75" valueMax="4.5" valueMin="1.0">1– 4.5 mm</quantity>
wide, the segments 6–15, entire to rarely lobed, glandular and crisp puberulent with white, uniseriate eglandular trichomes, lobes cuspidate or mucronate. Lower cauline leaves pinnatifid to more commonly entire and linear, gradually to abruptly reduced in size upward, ultimately upper cauline leaves entire,
<quantity id="4C8B9B3AFFFC936BE50FFA2BFA13F491" box="[1321,1436,1484,1510]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="18.0" metricValueMin="1.0" pageId="4" pageNumber="19" unit="mm" value="9.5" valueMax="18.0" valueMin="1.0">1–18 mm</quantity>
long, glandular puberulent. Inflorescence loosely open cymose-panicle, the flowers mostly crowded at the tips of the branches. Calyx cylindrical to campanulate,
<quantity id="4C8B9B3AFFFC936BE339F9FEFC12F744" box="[799,925,1561,1587]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" metricValueMax="4.7" metricValueMin="3.0" pageId="4" pageNumber="19" unit="mm" value="3.85" valueMax="4.7" valueMin="3.0">3–4.7 mm</quantity>
long, tube
<quantity id="4C8B9B3AFFFC936BE409F9FEFB4BF744" box="[1071,1220,1561,1587]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="3.5" metricValueMin="1.8" pageId="4" pageNumber="19" unit="mm" value="2.65" valueMax="3.5" valueMin="1.8">1.8–3.5 mm</quantity>
long at anthesis, glandular. Corolla
<quantity id="4C8B9B3AFFFC936BE157F9A7FE6FF72D" box="[369,480,1600,1626]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.8" metricValueMin="0.6" pageId="4" pageNumber="19" unit="mm" value="12.0" valueMax="18.0" valueMin="6.0">6–18 mm</quantity>
long, blue, paling to nearly white, corolla externally glabrous but bearing a patch of glandular trichomes centrally, at the proximal end (base) of the corolla lobe, narrowly funnelformsalverform; the tube,
<quantity id="4C8B9B3AFFFC936BE1B4F96AFD96F7D0" box="[402,537,1677,1703]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="12.0" metricValueMin="5.5" pageId="4" pageNumber="19" unit="mm" value="8.75" valueMax="12.0" valueMin="5.5">5.5–12 mm</quantity>
long, lobes narrowly oblanceolate, 3–5.5(–6.0) mm long,
<quantity id="4C8B9B3AFFFC936BE4E0F96AFADCF7D0" box="[1222,1363,1677,1703]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="3.5" metricValueMin="1.9" pageId="4" pageNumber="19" unit="mm" value="2.7" valueMax="3.5" valueMin="1.9">1.9–3.5 mm</quantity>
wide. Stamens equally inserted in the upper tube (at the sinuses of the corolla lobes), the free portion ca.
<quantity id="4C8B9B3AFFFC936BE530F953FAD6F7B9" box="[1302,1369,1716,1742]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="19" unit="mm" value="1.0">1 mm</quantity>
long, anthers
<quantity id="4C8B9B3AFFFC936BE0D4F93DFEF0F783" box="[242,383,1754,1780]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="4" pageNumber="19" unit="mm" value="1.85" valueMax="2.2" valueMin="1.5">1.5–2.2 mm</quantity>
long, slightly exserted. Style well exserted and approaching herkogamous to included and reverse herkogamous. Fruit a capsule,
<quantity id="4C8B9B3AFFFC936BE27EF8E6FD3BF66C" box="[600,692,1793,1819]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="4" pageNumber="19" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3-5 mm</quantity>
long,
<quantity id="4C8B9B3AFFFC936BE2DBF8E6FC06F66C" box="[765,905,1793,1819]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.9" metricValueMin="2.3" pageId="4" pageNumber="19" unit="mm" value="2.5999999999999996" valueMax="2.9" valueMin="2.3">2.3–2.9 mm</quantity>
wide, ovoid, dehiscing at the apex but valves remaining attached at the base, locules 3. Seed
<quantity id="4C8B9B3AFFFC936BE29BF8CFFCC6F635" box="[701,841,1832,1858]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="4" pageNumber="19" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
long,
<quantity id="4C8B9B3AFFFC936BE3B5F8CFFB86F635" box="[915,1033,1832,1858]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="4" pageNumber="19" unit="mm" value="0.85" valueMax="1.0" valueMin="0.7">0.7–1 mm</quantity>
wide, narrowly ovoid to fusiform, or angled-truncate and lung-shaped, wingless or with a narrow partial wing at one end or along the margin, less than
<quantity id="4C8B9B3AFFFC936BE122F892FED2F6F8" box="[260,349,1909,1935]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="19" unit="mm" value="0.2">0.2 mm</quantity>
wide, pale gray-brown.
</paragraph>
</subSubSection>
<subSubSection id="C3696554FFFC936AE0E0F87CFDA0F03B" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19" type="distribution">
<paragraph id="8BCC36DFFFFC936AE0E0F87CFD55F188" blockId="4.[151,1436,1252,2012]" lastBlockId="5.[151,1436,152,1608]" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
<emphasis id="B907EACDFFFC936BE0E0F87CFED0F6C2" bold="true" box="[198,351,1947,1973]" pageId="4" pageNumber="19">Distribution</emphasis>
:—
<taxonomicName id="4C734D5CFFFC936BE1A2F87BFDD8F6C2" authorityName="J. M. Porter" authorityYear="2011" box="[388,599,1948,1973]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="cliffordii">
<emphasis id="B907EACDFFFC936BE1A2F87BFDD8F6C2" box="[388,599,1948,1973]" italics="true" pageId="4" pageNumber="19">Aliciella cliffordii</emphasis>
</taxonomicName>
is apparently restricted to Apache County,
<collectingRegion id="49B7F83DFFFC936BE441F87BFB4AF6C1" box="[1127,1221,1948,1974]" country="United States of America" name="Arizona" pageId="4" pageNumber="19">Arizona</collectingRegion>
and adjacent San Juan County,
<collectingRegion id="49B7F83DFFFC936BE110F825FE5EF6AB" box="[310,465,1986,2012]" country="United States of America" name="New Mexico" pageId="4" pageNumber="19">New Mexico</collectingRegion>
, occurring in sandy or clay badlands, associated with pinyon-juniper woodland and ponderosa pine forests. Soils are generally red. Occurrences range in elev. from
<quantity id="4C8B9B3AFFFD936AE41AFF7FFB51F1C5" box="[1084,1246,152,178]" metricMagnitude="3" metricUnit="m" metricValue="1.7525" metricValueMax="1.98" metricValueMin="1.525" pageId="5" pageNumber="20" unit="m" value="1752.5" valueMax="1980.0" valueMin="1525.0">1525–1980 m</quantity>
(
<quantity id="4C8B9B3AFFFD936AE4CAFF7FFA00F1C5" box="[1260,1423,152,178]" metricMagnitude="3" metricUnit="m" metricValue="1.7526" metricValueMax="1.9812" metricValueMin="1.524" pageId="5" pageNumber="20" unit="ft" value="5750.0" valueMax="6500.0" valueMin="5000.0">5000–6500 ft</quantity>
). A very localized species, it is endemic to the Navajo Nation on Beautiful Mountain and the adjacent eastern slopes of the Lukachukai and Chuska Mountains.
</paragraph>
<paragraph id="8BCC36DFFFFD936AE0E0FEECFDA0F03B" blockId="5.[151,1436,152,1608]" pageId="5" pageNumber="20">
<emphasis id="B907EACDFFFD936AE0E0FEECFEC7F052" bold="true" box="[198,328,267,293]" pageId="5" pageNumber="20">Phenology</emphasis>
:—Flowering begins in May and continues through June or rarely to early July. Fruiting begins in mid-May and lasts through July.
</paragraph>
</subSubSection>
<subSubSection id="C3696554FFFD936AE0E0FEBFFB6EF3F5" pageId="5" pageNumber="20" type="etymology">
<paragraph id="8BCC36DFFFFD936AE0E0FEBFFC2EF0ED" blockId="5.[151,1436,152,1608]" pageId="5" pageNumber="20">
<emphasis id="B907EACDFFFD936AE0E0FEBFFEC2F005" bold="true" box="[198,333,344,370]" pageId="5" pageNumber="20">Etymology</emphasis>
:—The specific epithet,
<taxonomicName id="4C734D5CFFFD936AE25BFEBEFD6DF005" authorityName="J. M. Porter" authorityYear="2011" box="[637,738,345,370]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="cliffordii">
<emphasis id="B907EACDFFFD936AE25BFEBEFD6DF005" box="[637,738,345,370]" italics="true" pageId="5" pageNumber="20">cliffordii</emphasis>
</taxonomicName>
, honors Arnold Clifford, enthusiastic field botanist and important collector in the Four Corners region and Navajo Nation.
</paragraph>
<paragraph id="8BCC36DFFFFD936AE0E0FE41FB6EF3F5" blockId="5.[151,1436,152,1608]" pageId="5" pageNumber="20">
<materialsCitation id="3B1B3C82FFFD936AE0E0FE41FF40F379" ID-GBIF-Occurrence="3127432313" collectingDate="1974-06-08" collectionCode="BRY, NY" collectorName="D. Atwood" county="San Juan Co." location="Chuska Mtns." pageId="5" pageNumber="20" specimenCount="2" stateProvince="Arizona" typeStatus="paratype">
<emphasis id="B907EACDFFFD936AE0E0FE41FD64F0B7" bold="true" box="[198,747,422,448]" pageId="5" pageNumber="20">
Additional specimens examined (
<typeStatus id="54C8887DFFFD936AE242FE41FD6CF0B7" box="[612,739,422,448]" pageId="5" pageNumber="20" type="paratype">paratypes</typeStatus>
)
</emphasis>
:— U.S.A.
<emphasis id="B907EACDFFFD936AE354FE41FC73F0B7" bold="true" box="[882,1020,422,448]" pageId="5" pageNumber="20">
<collectingRegion id="49B7F83DFFFD936AE354FE41FC73F0B7" box="[882,1020,422,448]" country="United States of America" name="Arizona" pageId="5" pageNumber="20">ARIZONA</collectingRegion>
</emphasis>
.
<emphasis id="B907EACDFFFD936AE42AFE41FB69F0C8" bold="true" box="[1036,1254,422,448]" pageId="5" pageNumber="20">APACHE COUNTY</emphasis>
:
<locationDeviation id="9A7E2FFDFFFD936AE4D0FE41FF57F090" location="Cove" pageId="5" pageNumber="20">
<quantity id="4C8B9B3AFFFD936AE4D0FE41FAEDF0B7" box="[1270,1378,422,448]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" metricValueMax="4.828032" metricValueMin="1.609344" pageId="5" pageNumber="20" unit="mi" value="2.0" valueMax="3.0" valueMin="1.0">1 to 3 mi</quantity>
S of Cove
</locationDeviation>
, in the
<location id="8EAC6004FFFD936AE109FE2AFE5BF090" LSID="urn:lsid:plazi:treatment:03DA87C9FFFC936AE0B1FF70FA13F644:8EAC6004FFFD936AE109FE2AFE5BF090" box="[303,468,461,487]" county="San Juan Co." name="Chuska Mtns." pageId="5" pageNumber="20" stateProvince="Arizona">Chuska Mtns.</location>
(label states
<collectingCounty id="62AD4E53FFFD936AE256FE2AFC81F090" box="[624,782,461,487]" pageId="5" pageNumber="20">San Juan Co.</collectingCounty>
, NM, incorrectly),
<date id="FFCD101FFFFD936AE3D5FE2AFB0AF090" box="[1011,1157,461,487]" pageId="5" pageNumber="20" value="1974-06-08">
<collectingDate id="EF89E9F7FFFD936AE3D5FE2AFB0AF090" box="[1011,1157,461,487]" pageId="5" pageNumber="20" value="1974-06-08">8 June 1974</collectingDate>
</date>
,
<emphasis id="B907EACDFFFD936AE4B4FE2AFAC4F091" box="[1170,1355,461,486]" italics="true" pageId="5" pageNumber="20">
<collectorName id="26865309FFFD936AE4B4FE2AFA87F091" box="[1170,1288,461,486]" pageId="5" pageNumber="20">D.Atwood</collectorName>
6361
</emphasis>
(
<collectionCode id="ED62AE1AFFFD936AE57AFE2AFA17F090" box="[1372,1432,461,487]" country="USA" lsid="urn:lsid:biocol.org:col:15673" name="Brigham Young University - S.L. Welsh Herbarium" pageId="5" pageNumber="20" type="Herbarium">BRY</collectionCode>
,
<collectionCode id="ED62AE1AFFFD936AE0B1FE13FF49F379" box="[151,198,500,526]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="5" pageNumber="20" type="Herbarium">NY</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1B3C82FFFD936AE0FDFE13FE37F343" ID-GBIF-Occurrence="3127432304" collectingDate="1991-05-14" collectionCode="RSA" collectorName="San Juan Co., NM" location="Lukachukai Mtns" municipality="Navajo Nation" pageId="5" pageNumber="20" specimenCount="1" stateProvince="ARIZONA">
<collectingMunicipality id="6BA8ACA5FFFD936AE0FDFE13FE04F379" box="[219,395,500,526]" pageId="5" pageNumber="20">Navajo Nation</collectingMunicipality>
,
<locationDeviation id="9A7E2FFDFFFD936AE1B1FE13FDDBF379" box="[407,596,500,526]" location="Red Rock" pageId="5" pageNumber="20">W of Red Rock</locationDeviation>
, on the eastern foot of
<location id="8EAC6004FFFD936AE343FE13FBB8F379" LSID="urn:lsid:plazi:treatment:03DA87C9FFFC936AE0B1FF70FA13F644:8EAC6004FFFD936AE343FE13FBB8F379" box="[869,1079,500,526]" municipality="Navajo Nation" name="Lukachukai Mtns" pageId="5" pageNumber="20" stateProvince="ARIZONA">Lukachukai Mtns</location>
, red silty loam,
<date id="FFCD101FFFFD936AE4D0FE13FA18F379" box="[1270,1431,500,526]" pageId="5" pageNumber="20" value="1991-05-14">
<collectingDate id="EF89E9F7FFFD936AE4D0FE13FA18F379" box="[1270,1431,500,526]" pageId="5" pageNumber="20" value="1991-05-14">14 May 1991</collectingDate>
</date>
,
<emphasis id="B907EACDFFFD936AE0B1FDFDFED2F344" box="[151,349,538,563]" italics="true" pageId="5" pageNumber="20">
<collectorName id="26865309FFFD936AE0B1FDFDFF44F344" box="[151,203,538,563]" name="San Juan Co., NM" pageId="5" pageNumber="20">J.M.</collectorName>
<taxonomicName id="4C734D5CFFFD936AE0EDFDFDFE95F344" authority="Porter, 2011" authorityName="Porter" authorityYear="2011" box="[203,282,538,563]" pageId="5" pageNumber="21" rank="subSpecies" status="subsp. nov." subSpecies="porter">Porter</taxonomicName>
9559
</emphasis>
(
<collectionCode id="ED62AE1AFFFD936AE149FDFDFE20F343" box="[367,431,538,564]" country="USA" lsid="urn:lsid:biocol.org:col:15601" name="Rancho Santa Ana Botanic Garden" pageId="5" pageNumber="20" type="Herbarium">RSA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1B3C82FFFD936AE1E3FDFDFBB8F32C" ID-GBIF-Occurrence="3127432334" collectingDate="2001-09-27" collectionCode="RSA, SJNM" collectorName="K. Heil &amp; A. Clifford" county="Navajo Nation" location="Wingate Formation" municipality="Mexican Cry Mesa" pageId="5" pageNumber="20" specimenCount="1" stateProvince="ARIZONA">
<collectingCounty id="62AD4E53FFFD936AE1E3FDFDFDF8F343" box="[453,631,538,564]" pageId="5" pageNumber="20">Navajo Nation</collectingCounty>
, drainage and road to
<collectingMunicipality id="6BA8ACA5FFFD936AE3A2FDFDFBE2F343" box="[900,1133,538,564]" pageId="5" pageNumber="20">Mexican Cry Mesa</collectingMunicipality>
, on lower member of the
<location id="8EAC6004FFFD936AE0B1FDA6FE0EF32C" LSID="urn:lsid:plazi:treatment:03DA87C9FFFC936AE0B1FF70FA13F644:8EAC6004FFFD936AE0B1FDA6FE0EF32C" box="[151,385,577,603]" county="Navajo Nation" municipality="Mexican Cry Mesa" name="Wingate Formation" pageId="5" pageNumber="20" stateProvince="ARIZONA">Wingate Formation</location>
,
<date id="FFCD101FFFFD936AE1A8FDA6FDB9F32C" box="[398,566,577,603]" pageId="5" pageNumber="20" value="2001-09-27">
<collectingDate id="EF89E9F7FFFD936AE1A8FDA6FDB9F32C" box="[398,566,577,603]" pageId="5" pageNumber="20" value="2001-09-27">27 Sept. 2001</collectingDate>
</date>
,
<emphasis id="B907EACDFFFD936AE262FDA6FC0CF32D" box="[580,899,577,602]" italics="true" pageId="5" pageNumber="20">
<collectorName id="26865309FFFD936AE262FDA6FD1DF32D" box="[580,658,577,602]" pageId="5" pageNumber="20">K.Heil</collectorName>
&amp;
<collectorName id="26865309FFFD936AE29CFDA6FCBFF32D" box="[698,816,577,602]" pageId="5" pageNumber="20">A.Clifford</collectorName>
18375
</emphasis>
(
<collectionCode id="ED62AE1AFFFD936AE3B5FDA6FC5EF32C" box="[915,977,577,603]" country="USA" lsid="urn:lsid:biocol.org:col:15601" name="Rancho Santa Ana Botanic Garden" pageId="5" pageNumber="20" type="Herbarium">RSA</collectionCode>
,
<collectionCode id="ED62AE1AFFFD936AE3F8FDA6FBA0F32C" box="[990,1071,577,603]" country="USA" lsid="urn:lsid:biocol.org:col:14919" name="San Juan College" pageId="5" pageNumber="20" type="Herbarium">SJNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1B3C82FFFD936AE462FDA7FB51F3F5" ID-GBIF-Occurrence="3127432319" collectingDate="1970-05-15" collectionCode="BRY, NY" collectorName="D. Atwood" location="Hwy" municipality="Shiprock" pageId="5" pageNumber="20" specimenCount="1" stateProvince="New Mexico">
<emphasis id="B907EACDFFFD936AE462FDA7FE8EF3F7" bold="true" pageId="5" pageNumber="20">
<collectingRegion id="49B7F83DFFFD936AE462FDA7FA9EF32D" box="[1092,1297,576,602]" country="United States of America" name="New Mexico" pageId="5" pageNumber="20">NEW MEXICO</collectingRegion>
. SAN JUAN COUNTY
</emphasis>
:
<quantity id="4C8B9B3AFFFD936AE136FD8FFEDBF3F5" box="[272,340,616,642]" metricMagnitude="4" metricUnit="m" metricValue="1.9312128" pageId="5" pageNumber="20" unit="mi" value="12.0">12 mi</quantity>
west of
<collectingMunicipality id="6BA8ACA5FFFD936AE19EFD8FFDADF3F5" box="[440,546,616,642]" pageId="5" pageNumber="20">Shiprock</collectingMunicipality>
along
<location id="8EAC6004FFFD936AE257FD8FFD25F3F5" LSID="urn:lsid:plazi:treatment:03DA87C9FFFC936AE0B1FF70FA13F644:8EAC6004FFFD936AE257FD8FFD25F3F5" box="[625,682,616,642]" municipality="Shiprock" name="Hwy" pageId="5" pageNumber="20" stateProvince="New Mexico">Hwy</location>
504,
<date id="FFCD101FFFFD936AE2CAFD8FFC04F3F5" box="[748,907,616,642]" pageId="5" pageNumber="20" value="1970-05-15">
<collectingDate id="EF89E9F7FFFD936AE2CAFD8FFC04F3F5" box="[748,907,616,642]" pageId="5" pageNumber="20" value="1970-05-15">15 May 1970</collectingDate>
</date>
,
<emphasis id="B907EACDFFFD936AE3B1FD8FFBC0F3F6" box="[919,1103,616,641]" italics="true" pageId="5" pageNumber="20">
<collectorName id="26865309FFFD936AE3B1FD8FFB82F3F6" box="[919,1037,616,641]" pageId="5" pageNumber="20">D.Atwood</collectorName>
2525
</emphasis>
(
<collectionCode id="ED62AE1AFFFD936AE478FD8FFB15F3F5" box="[1118,1178,616,642]" country="USA" lsid="urn:lsid:biocol.org:col:15673" name="Brigham Young University - S.L. Welsh Herbarium" pageId="5" pageNumber="20" type="Herbarium">BRY</collectionCode>
,
<collectionCode id="ED62AE1AFFFD936AE483FD8FFB5BF3F5" box="[1189,1236,616,642]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="5" pageNumber="20" type="Herbarium">NY</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3696554FFFD936AE0E0FD69FD24F73F" pageId="5" pageNumber="20" type="discussion">
<paragraph id="8BCC36DFFFFD936AE0E0FD69FD8EF573" blockId="5.[151,1436,152,1608]" pageId="5" pageNumber="20">
<emphasis id="B907EACDFFFD936AE0E0FD69FEC5F3DF" bold="true" box="[198,330,654,680]" pageId="5" pageNumber="20">Discussion</emphasis>
:—
<taxonomicName id="4C734D5CFFFD936AE149FD68FBC2F3DF" authority="J. M. Porter" authorityName="J. M. Porter" box="[367,1101,654,680]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="subSection" section="Aliciella" subSection="Subnudae">
<emphasis id="B907EACDFFFD936AE149FD68FE5DF3DF" box="[367,466,655,680]" italics="true" pageId="5" pageNumber="20">Aliciella</emphasis>
Brand sect.
<emphasis id="B907EACDFFFD936AE250FD68FD56F3DF" box="[630,729,655,680]" italics="true" pageId="5" pageNumber="20">Aliciella</emphasis>
subsect.
<emphasis id="B907EACDFFFD936AE377FD68FC4BF3DF" box="[849,964,655,680]" italics="true" pageId="5" pageNumber="20">Subnudae</emphasis>
J.M.Porter
</taxonomicName>
(
<bibRefCitation id="EFE24B2EFFFD936AE444FD69FAA7F3DF" author="Porter, J. M." box="[1122,1320,654,680]" pageId="5" pageNumber="20" pagination="23 - 46" refId="ref9524" refString="Porter, J. M. (1998) Aliciella, a recircumscribed genus of Polemoniaceae. Aliso 17: 23 - 46." type="journal article" year="1998">Porter 1998: 31</bibRefCitation>
; see also
<bibRefCitation id="EFE24B2EFFFD936AE0B1FD52FE35F3B8" author="Porter, J. M. &amp; Johnson, L. A." box="[151,442,693,719]" pageId="5" pageNumber="20" pagination="55 - 91" refId="ref9636" refString="Porter, J. M. &amp; Johnson, L. A. (2000) A phylogenetic classification of Polemoniaceae. Aliso 19: 55 - 91." type="journal article" year="2000">Porter and Johnson 2000</bibRefCitation>
) is largely of the
<collectingRegion id="49B7F83DFFFD936AE2A0FD52FD7CF3B8" box="[646,755,693,719]" country="United States of America" name="Colorado" pageId="5" pageNumber="20">Colorado</collectingRegion>
Plateau. It boasts some of the more colorful and attractive members of the
<taxonomicName id="4C734D5CFFFD936AE171FD3BFE13F382" authorityName="Linnaeus" authorityYear="1753" box="[343,412,732,757]" class="Magnoliopsida" family="Polemoniaceae" genus="Phlox" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis id="B907EACDFFFD936AE171FD3BFE13F382" box="[343,412,732,757]" italics="true" pageId="5" pageNumber="20">Phlox</emphasis>
</taxonomicName>
family in the Four Corners region, including
<taxonomicName id="4C734D5CFFFD936AE39FFD3BFE2AF26B" authority="(Greene ex Brand 1907: 119) J. M. Porter (1998: 33)" authorityName="J. M. Porter" authorityPageNumber="33" authorityYear="1998" baseAuthorityName="Greene ex Brand" baseAuthorityPageNumber="119" baseAuthorityYear="1907" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="formosa">
<emphasis id="B907EACDFFFD936AE39FFD3BFC44F382" box="[953,971,732,757]" italics="true" pageId="5" pageNumber="20">A</emphasis>
.
<emphasis id="B907EACDFFFD936AE3FCFD3BFBB5F382" box="[986,1082,732,757]" italics="true" pageId="5" pageNumber="20">formosa</emphasis>
(Greene ex
<bibRefCitation id="EFE24B2EFFFD936AE4EAFD3BFA1CF381" author="Brand, A." box="[1228,1427,732,758]" pageId="5" pageNumber="20" pagination="250" refId="ref8765" refString="Brand, A. (1907) Polemoniaceae. Pp. 1 - 203. In: Das Pflanzenreich vol. 4: 250, ed A. Engler. Leipzig: W. Engelmann." type="journal article" year="1907">Brand 1907: 119</bibRefCitation>
) J.M.
<bibRefCitation id="EFE24B2EFFFD936AE0E0FCE5FE2AF26B" author="Porter, J. M." box="[198,421,770,796]" pageId="5" pageNumber="20" pagination="23 - 46" refId="ref9524" refString="Porter, J. M. (1998) Aliciella, a recircumscribed genus of Polemoniaceae. Aliso 17: 23 - 46." type="journal article" year="1998">Porter (1998: 33)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C734D5CFFFD936AE192FCE5FBBAF26B" authority="(Gray 1870: 276) J. M. Porter (1998: 33)" authorityName="J. M. Porter" authorityPageNumber="33" authorityYear="1998" baseAuthorityName="Gray" baseAuthorityPageNumber="276" baseAuthorityYear="1870" box="[436,1077,770,796]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="subnuda">
<emphasis id="B907EACDFFFD936AE192FCE5FE49F26C" box="[436,454,770,795]" italics="true" pageId="5" pageNumber="20">A</emphasis>
.
<emphasis id="B907EACDFFFD936AE1F0FCE5FDB2F26C" box="[470,573,770,795]" italics="true" pageId="5" pageNumber="20">subnuda</emphasis>
(
<bibRefCitation id="EFE24B2EFFFD936AE269FCE5FC9BF26B" author="Gray, A." box="[591,788,770,796]" pageId="5" pageNumber="20" pagination="247 - 282" refId="ref9235" refString="Gray, A. (1870) Revision of North American Polemoniaceae. Proceedings of the American Academy of Arts and Sciences 8: 247 - 282." type="journal article" year="1870">Gray 1870: 276</bibRefCitation>
) J.M.
<bibRefCitation id="EFE24B2EFFFD936AE370FCE5FBBAF26B" author="Porter, J. M." box="[854,1077,770,796]" pageId="5" pageNumber="20" pagination="23 - 46" refId="ref9524" refString="Porter, J. M. (1998) Aliciella, a recircumscribed genus of Polemoniaceae. Aliso 17: 23 - 46." type="journal article" year="1998">Porter (1998: 33)</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C734D5CFFFD936AE451FCE5FA8BF26C" authorityName="J. M. Porter" authorityYear="1998" baseAuthorityName="A. Gray" box="[1143,1284,770,796]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="haydenii">
<emphasis id="B907EACDFFFD936AE451FCE5FB06F26C" box="[1143,1161,770,795]" italics="true" pageId="5" pageNumber="20">A</emphasis>
.
<emphasis id="B907EACDFFFD936AE4BDFCE5FA8BF26C" box="[1179,1284,770,795]" italics="true" pageId="5" pageNumber="20">haydenii</emphasis>
</taxonomicName>
. The showy magenta flowers of
<taxonomicName id="4C734D5CFFFD936AE159FCCEFE70F235" authorityName="J. M. Porter" authorityYear="1998" baseAuthorityName="Greene ex Brand" baseAuthorityYear="1907" box="[383,511,809,835]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="formosa">
<emphasis id="B907EACDFFFD936AE159FCCEFE1EF235" box="[383,401,809,834]" italics="true" pageId="5" pageNumber="20">A</emphasis>
.
<emphasis id="B907EACDFFFD936AE1B9FCCEFE70F235" box="[415,511,809,834]" italics="true" pageId="5" pageNumber="20">formosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C734D5CFFFD936AE211FCCEFD33F235" authorityName="J. M. Porter" authorityYear="1998" baseAuthorityName="A. Gray" box="[567,700,809,835]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="haydenii">
<emphasis id="B907EACDFFFD936AE211FCCEFDC6F235" box="[567,585,809,834]" italics="true" pageId="5" pageNumber="20">A</emphasis>
.
<emphasis id="B907EACDFFFD936AE271FCCEFD33F235" box="[599,700,809,834]" italics="true" pageId="5" pageNumber="20">haydenii</emphasis>
</taxonomicName>
are hawkmoth pollinated, with over 90% of insect visits due to hawkmoths (Porter 1993;
<bibRefCitation id="EFE24B2EFFFD936AE1E1FCB7FCA7F21D" author="Porter, J. M. &amp; Floyd-Hanna, L." box="[455,808,848,874]" pageId="5" pageNumber="20" pagination="236 - 248" refId="ref9548" refString="Porter, J. M. &amp; Floyd-Hanna, L. (1993) A phylogenetic approach to natural historical and demographic explanations of rarity of Gilia formosa. Pp. 236 - 248. In: R. Sivinski and K. Lightfoot eds., Proceedings of the Southwestern rare and endangered plant conference. Miscellaneous publication No. 2. New Mexico Forestry and Resources Conservation Division, Energy, Minerals and Natural Resources Department, Santa Fe, New Mexico. 390 pp." type="book chapter" year="1993">Porter and Floyd-Hanna 1993</bibRefCitation>
). There are two subspecies of
<taxonomicName id="4C734D5CFFFD936AE4AEFCB7FA82F21E" authorityName="J. M. Porter" authorityYear="1998" baseAuthorityName="A. Gray" box="[1160,1293,848,874]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="haydenii">
<emphasis id="B907EACDFFFD936AE4AEFCB7FB15F21E" box="[1160,1178,848,873]" italics="true" pageId="5" pageNumber="20">A</emphasis>
.
<emphasis id="B907EACDFFFD936AE48EFCB7FA82F21E" box="[1192,1293,848,873]" italics="true" pageId="5" pageNumber="20">haydenii</emphasis>
</taxonomicName>
recognized, differing in corolla size, density of glands on the external corolla, and in part corolla color upon drying. Regardless of these differences, populations and the two races of
<taxonomicName id="4C734D5CFFFD936AE38FFC7AFBBEF2C1" authorityName="J. M. Porter" authorityYear="1998" baseAuthorityName="A. Gray" box="[937,1073,925,951]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="haydenii">
<emphasis id="B907EACDFFFD936AE38FFC7AFC34F2C1" box="[937,955,925,950]" italics="true" pageId="5" pageNumber="20">A</emphasis>
.
<emphasis id="B907EACDFFFD936AE3EDFC7AFBBEF2C1" box="[971,1073,925,950]" italics="true" pageId="5" pageNumber="20">haydenii</emphasis>
</taxonomicName>
share a common reproductive and pollination system (Porter 1993) and show some intergradation. This is consistent with the hypothesis that they represent a single species.
</paragraph>
<paragraph id="8BCC36DFFFFD936AE0E0FBF6FD24F73F" blockId="5.[151,1436,152,1608]" pageId="5" pageNumber="20">
The series of isolated and disjunct populations near Beautiful Mountain,
<collectingRegion id="49B7F83DFFFD936AE46BFBF6FB24F55C" box="[1101,1195,1041,1067]" country="United States of America" name="Arizona" pageId="5" pageNumber="20">Arizona</collectingRegion>
are very similar to
<taxonomicName id="4C734D5CFFFD936AE0B1FBDFFEE1F526" authorityName="J. M. Porter" authorityYear="1998" baseAuthorityName="A. Gray" box="[151,366,1080,1105]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="haydenii">
<emphasis id="B907EACDFFFD936AE0B1FBDFFEE1F526" box="[151,366,1080,1105]" italics="true" pageId="5" pageNumber="20">Aliciella haydenii</emphasis>
</taxonomicName>
, but differ in their narrower corolla tube and lobes and pale floral color, having blue to nearly white corollas (
<figureCitation id="13482A5AFFFD936AE1BAFBB9FD94F50F" box="[412,539,1118,1144]" captionStart="FIGURE 2" captionStartId="4.[151,255,1074,1098]" captionTargetBox="[151,1436,458,1050]" captionTargetId="figure-447@4.[151,1436,458,1050]" captionTargetPageId="4" captionText="FIGURE 2. A. Habit and architecture of Aliciella cliffordii, scale= 3.0 cm, J.M.Porter, K.D.Heil, &amp; A.Clifford 13528 (Photo J.M.Porter). B. Cymose inflorescence and flowers, scale= 5.0 mm, J.M.Porter, K.D.Heil, &amp; A.Clifford 13528 (Photo J.M.Porter). C. Bombilius sp. visiting flowers of Aliciella cliffordii, scale= 5.0 mm, J.M.Porter, K.D.Heil, &amp; A.Clifford 13528 (Photo J.M.Porter)." figureDoi="http://doi.org/10.5281/zenodo.4906385" httpUri="https://zenodo.org/record/4906385/files/figure.png" pageId="5" pageNumber="20">Fig. 2A, B</figureCitation>
). The populations in the Beautiful Mountain area are pollinated primarily by beeflies (
<emphasis id="B907EACDFFFD936AE136FB62FE70F5E9" box="[272,511,1157,1182]" italics="true" pageId="5" pageNumber="20">Bombilius lancifer</emphasis>
<bibRefCitation id="EFE24B2EFFFD936AE22FFB62FCD3F5E8" author="Osten Sacken, C. R." box="[521,860,1157,1183]" pageId="5" pageNumber="20" pagination="189 - 354" refId="ref9409" refString="Osten Sacken, C. R. (1877) Western Diptera: Descriptions of new genera and species of Diptera from the region west of the Mississippi and especially from California. U. S. Geological and Geographical Survey of the Territories Bulletin 8: 189 - 354" type="journal article" year="1877">Osten Sacken [1877: 251]</bibRefCitation>
;
<figureCitation id="13482A5AFFFD936AE34AFB62FC5FF5E8" box="[876,976,1157,1183]" captionStart="FIGURE 2" captionStartId="4.[151,255,1074,1098]" captionTargetBox="[151,1436,458,1050]" captionTargetId="figure-447@4.[151,1436,458,1050]" captionTargetPageId="4" captionText="FIGURE 2. A. Habit and architecture of Aliciella cliffordii, scale= 3.0 cm, J.M.Porter, K.D.Heil, &amp; A.Clifford 13528 (Photo J.M.Porter). B. Cymose inflorescence and flowers, scale= 5.0 mm, J.M.Porter, K.D.Heil, &amp; A.Clifford 13528 (Photo J.M.Porter). C. Bombilius sp. visiting flowers of Aliciella cliffordii, scale= 5.0 mm, J.M.Porter, K.D.Heil, &amp; A.Clifford 13528 (Photo J.M.Porter)." figureDoi="http://doi.org/10.5281/zenodo.4906385" httpUri="https://zenodo.org/record/4906385/files/figure.png" pageId="5" pageNumber="20">Fig. 2C</figureCitation>
), and visited, to a lesser degree, by anthophorid bees, representing an ethological distinction from
<taxonomicName id="4C734D5CFFFD936AE3BEFB4BFBACF5B2" authorityName="J. M. Porter" authorityYear="1998" baseAuthorityName="A. Gray" box="[920,1059,1196,1222]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="haydenii">
<emphasis id="B907EACDFFFD936AE3BEFB4BFC25F5B2" box="[920,938,1196,1221]" italics="true" pageId="5" pageNumber="20">A</emphasis>
.
<emphasis id="B907EACDFFFD936AE39DFB4BFBACF5B2" box="[955,1059,1196,1221]" italics="true" pageId="5" pageNumber="20">haydenii</emphasis>
</taxonomicName>
. Field studies over a four-year period, summing to nearly 200 hours of observation time, have verified that these populations are not visited by hawkmoths, rather
<emphasis id="B907EACDFFFD936AE18EFB1EFE35F465" box="[424,442,1273,1298]" italics="true" pageId="5" pageNumber="20">B</emphasis>
.
<emphasis id="B907EACDFFFD936AE1EDFB1EFDA6F465" box="[459,553,1273,1298]" italics="true" pageId="5" pageNumber="20">lancifer</emphasis>
(73% of visits) and several species of anthophorid bees (18% of visits). Nonpollinating visitors (e.g., syrphid flies) account for the remaining visits. I hypothesize that the populations near Beautiful Mountain correspond to a different reproductive and pollination system from
<taxonomicName id="4C734D5CFFFD936AE4FAFAA0FAEDF417" authorityName="J. M. Porter" authorityYear="1998" baseAuthorityName="A. Gray" box="[1244,1378,1350,1376]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="haydenii">
<emphasis id="B907EACDFFFD936AE4FAFAA0FB61F417" box="[1244,1262,1351,1376]" italics="true" pageId="5" pageNumber="20">A</emphasis>
.
<emphasis id="B907EACDFFFD936AE4DBFAA0FAEDF417" box="[1277,1378,1351,1376]" italics="true" pageId="5" pageNumber="20">haydenii</emphasis>
</taxonomicName>
, and therefore comprise a genetically independent species. In accordance, I treat it at the rank of species. While
<taxonomicName id="4C734D5CFFFD936AE5A5FA8AFE8FF4DA" authorityName="J. M. Porter" authorityYear="2011" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="cliffordii">
<emphasis id="B907EACDFFFD936AE5A5FA8AFA1AF4F1" box="[1411,1429,1389,1414]" italics="true" pageId="5" pageNumber="20">A</emphasis>
.
<emphasis id="B907EACDFFFD936AE0B1FA73FE8FF4DA" box="[151,256,1428,1453]" italics="true" pageId="5" pageNumber="20">cliffordii</emphasis>
</taxonomicName>
seems technically weak if considering the morphological features alone, the different pollination system and geographic disjunction is consistent with a hypothesis of peripatric speciation, involving a change in pollination mechanisms. This provides an explicit hypothesis that can be experimentally tested. It is also important to note that, in the field, it is readily distinguishable from
<taxonomicName id="4C734D5CFFFD936AE392F9EFFBB5F756" authorityName="J. M. Porter" authorityYear="1998" baseAuthorityName="A. Gray" box="[948,1082,1544,1570]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="haydenii">
<emphasis id="B907EACDFFFD936AE392F9EFFC49F756" box="[948,966,1544,1569]" italics="true" pageId="5" pageNumber="20">A</emphasis>
.
<emphasis id="B907EACDFFFD936AE3F3F9EFFBB5F756" box="[981,1082,1544,1569]" italics="true" pageId="5" pageNumber="20">haydenii</emphasis>
</taxonomicName>
(
<figureCitation id="13482A5AFFFD936AE46CF9EFFB46F755" box="[1098,1225,1544,1570]" captionStart="FIGURE 3" captionStartId="6.[151,257,862,886]" captionTargetBox="[154,1436,153,834]" captionTargetId="figure-184@6.[154,1440,153,835]" captionTargetPageId="6" captionText="FIGURE 3. A. Habit and architecture of Aliciella haydenii, scale= 3.0 cm, J.M.Porter &amp; K.D.Heil 12455, (Photo J.M.Porter). B. Cymose inflorescence and flowers, scale= 3.0 mm, J.M.Porter &amp; K.D.Heil 12455, (Photo J.M.Porter)." figureDoi="http://doi.org/10.5281/zenodo.4906387" httpUri="https://zenodo.org/record/4906387/files/figure.png" pageId="5" pageNumber="20">Fig. 3A, B</figureCitation>
). The two species can be distinguished using the following key:
</paragraph>
</subSubSection>
<subSubSection id="C3696554FFFD936AE0B1F99BFA13F644" pageId="5" pageNumber="20" type="key">
<keyStep id="30872E7AFFFD936AE0B1F99BFA13F644" pageId="5" pageNumber="20">
<key id="FFE261EEFFFD936AE0B1F99BFA13F644" pageId="5" pageNumber="20">
<paragraph id="8BCC36DFFFFD936AE0B1F99BFA13F7A4" blockId="5.[151,1436,1660,1843]" pageId="5" pageNumber="20">
<keyLead id="308295EAFFFD936AE0B1F99BFA13F7A4" pageId="5" pageNumber="20">
1 Corolla blue, paling to nearly white when fresh; corolla lobes narrowly lanceolate,
<quantity id="4C8B9B3AFFFD936AE418F99BFB18F7E4" box="[1086,1175,1660,1683]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="5" pageNumber="20" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
long,
<quantity id="4C8B9B3AFFFD936AE4FCF99BFAD4F7E4" box="[1242,1371,1660,1683]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="3.5" metricValueMin="1.9" pageId="5" pageNumber="20" unit="mm" value="2.7" valueMax="3.5" valueMin="1.9">1.9–3.5 mm</quantity>
wide; free portion of filaments
<quantity id="4C8B9B3AFFFD936AE1F5F97BFDDCF7C4" box="[467,595,1692,1715]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="15.0" metricValueMin="2.0" pageId="5" pageNumber="20" unit="mm" value="0.85" valueMax="1.5" valueMin="0.2">0.2–1.5 mm</quantity>
(mean
<quantity id="4C8B9B3AFFFD936AE283F97BFD74F7C4" box="[677,763,1692,1715]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="20" unit="mm" value="0.9">0.9 mm</quantity>
); restricted to the Beautiful Mountain area and near Red Rock,
<collectingRegion id="49B7F83DFFFD936AE0E1F95BFE92F7A4" box="[199,285,1724,1747]" country="United States of America" name="Arizona" pageId="5" pageNumber="20">Arizona</collectingRegion>
<emphasis id="B907EACDFFFD936AE103F95BFA13F7A4" box="[293,1436,1724,1747]" italics="true" pageId="5" pageNumber="20">
.........................................................................................................................................................
<taxonomicName id="4C734D5CFFFD936AE503F95BFA13F7A4" ID-CoL="BS4D" authorityName="J. M. Porter" authorityYear="2011" box="[1317,1436,1724,1747]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="cliffordii">A. cliffordii</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCC36DFFFFD936AE0B1F93BFA13F644" blockId="5.[151,1436,1660,1843]" pageId="5" pageNumber="20">
<keyLead id="308295EAFFFD936AE0B1F93BFA13F644" pageId="5" pageNumber="20">
- Corolla magenta when fresh; corolla lobes oval to oblanceolate, slightly wider than above,
<quantity id="4C8B9B3AFFFD936AE48DF93BFA96F784" box="[1195,1305,1756,1779]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="9.0" metricValueMin="3.5" pageId="5" pageNumber="20" unit="mm" value="6.25" valueMax="9.0" valueMin="3.5">3.5–9 mm</quantity>
long,
<quantity id="4C8B9B3AFFFD936AE546F93BFF7FF664" metricMagnitude="-3" metricUnit="m" metricValue="3.1" metricValueMax="4.2" metricValueMin="2.0" pageId="5" pageNumber="20" unit="mm" value="3.1" valueMax="4.2" valueMin="2.0">2–4.2 mm</quantity>
wide; free portion of filaments
<quantity id="4C8B9B3AFFFD936AE26CF91BFD45F664" box="[586,714,1788,1811]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.7" metricValueMin="0.8" pageId="5" pageNumber="20" unit="mm" value="1.75" valueMax="2.7" valueMin="0.8">0.8–2.7 mm</quantity>
(mean
<quantity id="4C8B9B3AFFFD936AE33BF91BFCFBF664" box="[797,884,1788,1811]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="20" unit="mm" value="1.5">1.5 mm</quantity>
); scattered in the Four Corners region of
<collectingRegion id="49B7F83DFFFD936AE514F91BFE36F644" isEnumeration="true" pageId="5" pageNumber="20">Colorado, New Mexico and Utah</collectingRegion>
<emphasis id="B907EACDFFFD936AE1E3F8FBFA13F644" box="[453,1436,1820,1843]" italics="true" pageId="5" pageNumber="20">
.................................................................................................................................
<taxonomicName id="4C734D5CFFFD936AE505F8FBFA13F644" ID-CoL="BS4F" authorityName="J. M. Porter" authorityYear="1998" baseAuthorityName="A. Gray" box="[1315,1436,1820,1843]" class="Magnoliopsida" family="Polemoniaceae" genus="Aliciella" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="haydenii">A. haydenii</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</key>
</keyStep>
</subSubSection>
</treatment>
</document>
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