325 lines
60 KiB
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325 lines
60 KiB
XML
<document id="30601E6678B52B6AA2A90C495631749F" ID-CLB-Dataset="26633" ID-DOI="10.11646/zootaxa.4612.3.5" ID-GBIF-Dataset="8b8d3d31-4988-436c-854d-2ce9730d0429" ID-ISSN="1175-5326" ID-Zenodo-Dep="3235176" ID-ZooBank="0BFCCA1E-B279-408A-A466-6A66CB936788" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1559194398912" checkinUser="plazi" docAuthor="Garbelotto, Thereza De A., Grazia, Jocelia & Campos, Luiz Alexandre" docDate="2019" docId="03DA87C9F128AE10C9BAD440C5C1F45C" docLanguage="en" docName="zootaxa.4612.3.5.pdf" docOrigin="Zootaxa 4612 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alloeogyna Garbelotto & Grazia & Campos 2019, gen. nov." docType="treatment" docUuid="7438EAB0-D32A-4651-BF69-BF583FE568D4" docUuidSource="ZooBank" docVersion="6" lastPageNumber="390" masterDocId="FFE3FFB1F129AE13C92DD07EC05DF51E" masterDocTitle="Description of two new genera of Ochlerini (Hemiptera: Pentatomidae: Discocephalinae)" masterLastPageNumber="400" masterPageNumber="387" pageNumber="388" updateTime="1698732372034" updateUser="ExternalLinkService">
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<mods:mods id="A0D0964806419D1B3371D3FCD9D0BC4D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5928EEE54F2EB548BC142DA79AEDF623">
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<mods:title id="45AE37181D865CEE780C5A68672960C4">Description of two new genera of Ochlerini (Hemiptera: Pentatomidae: Discocephalinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="562BD499C09E57DB4694BC7E5F90407C" type="personal">
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<mods:role id="375E6F5E8395F6614FE967E9589E60D3">
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<mods:roleTerm id="4E83592D18D553BAE6331E320C2A8777">Author</mods:roleTerm>
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<mods:namePart id="D5DB96F88CABE47F4598532D94D02116">Garbelotto, Thereza De A.</mods:namePart>
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<mods:affiliation id="2C6103E0D62454F9FE470EB243C030F8">Universidade do Sul Catarinense, Departamento de Ciências Biológicas, Av. José Acácio Moreira, 787, Dehon, Tubarão - SC, 88704 - 900.</mods:affiliation>
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<mods:nameIdentifier id="C608600F0AA43B2DA2833D8B62F21475" type="email">therezagarbelotto@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="AD487FC28E01ECE4C9F54A1FDF3CFB81" type="personal">
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<mods:roleTerm id="333C21E87DBA484FD0DF3095C1B22588">Author</mods:roleTerm>
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<mods:namePart id="7364A79FF1AA807AA86288754C5973F1">Grazia, Jocelia</mods:namePart>
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<mods:affiliation id="7BE19074F4414B652D0C351E82F24E8F">Universidade Federal do Rio Grande do Sul, Dep. Zoologia, Av. Bento Gonçalves, 9500, prédio 43435, Agronomia, Porto Alegre - RS, 91501 - 970.</mods:affiliation>
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<mods:nameIdentifier id="F1CA7B9895DEB75B28982C9900C70322" type="email">jocelia@ufrgs.br</mods:nameIdentifier>
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<mods:roleTerm id="125D19311BF663426D14B2D88CA463B4">Author</mods:roleTerm>
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<mods:namePart id="61CC7B36A5061007E2CECA61A4DF733C">Campos, Luiz Alexandre</mods:namePart>
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<mods:affiliation id="4D2194F89E6750722AE88728C857BA89">Universidade Federal do Rio Grande do Sul, Dep. Zoologia, Av. Bento Gonçalves, 9500, prédio 43435, Agronomia, Porto Alegre - RS, 91501 - 970.</mods:affiliation>
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<mods:nameIdentifier id="E5B8C7285F08D562F46F1CD868CA9467" type="email">luiz.campos@ufrgs.br</mods:nameIdentifier>
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<mods:title id="2A0D1C8B9C84FF63C9B22A61DA3358C5">Zootaxa</mods:title>
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<mods:part id="3803CFF69A2E6DB6265B6A6AACACC0BF">
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<mods:date id="BE185288B60390BCFD6668C2A6859C5F">2019</mods:date>
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<mods:number id="3020336556B2DE1585F75D5A491E616B">2019-05-30</mods:number>
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<mods:detail id="E1CE992DC54F2241CE5C732FFED0C9B6" type="volume">
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<mods:number id="E29B21AD45889AD0F3BD0A6752DA3185">4612</mods:number>
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<mods:detail id="54DDC627B7691EC633F02172ACB684DF" type="issue">
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<mods:number id="3DB59928F9D5F50D7BEEBE4F8577C9EA">3</mods:number>
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<mods:identifier id="1D283F394A418A1E77002935F365BA8B" type="DOI">10.11646/zootaxa.4612.3.5</mods:identifier>
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<mods:identifier id="293EA089FEB3D8E152CF5E68891F19AB" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03DA87C9F128AE10C9BAD440C5C1F45C" ID-GBIF-Taxon="164613180" LSID="urn:lsid:zoobank.org:act:7438EAB0-D32A-4651-BF69-BF583FE568D4" httpUri="http://treatment.plazi.org/id/03DA87C9F128AE10C9BAD440C5C1F45C" lastPageId="3" lastPageNumber="390" pageId="1" pageNumber="388">
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<subSubSection id="C3696554F128AE12C9BAD440C2EEF147" box="[151,691,1086,1113]" pageId="1" pageNumber="388" type="nomenclature">
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<paragraph id="8BCC36DFF128AE12C9BAD440C2EEF147" blockId="1.[151,691,1086,1113]" box="[151,691,1086,1113]" pageId="1" pageNumber="388">
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<heading id="D08481B3F128AE12C9BAD440C2EEF147" bold="true" box="[151,691,1086,1113]" fontSize="11" level="1" pageId="1" pageNumber="388" reason="1">
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<emphasis id="B907EACDF128AE12C9BAD440C2EEF147" bold="true" box="[151,691,1086,1113]" pageId="1" pageNumber="388">
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<taxonomicName id="4C734D5CF128AE12C9BAD440C263F147" authority="Garbelotto & Campos" authorityName="Garbelotto & Grazia & Campos" authorityYear="2019" box="[151,574,1086,1113]" class="Insecta" family="Pentatomidae" genus="Alloeogyna" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="388" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B907EACDF128AE12C9BAD440C17DF147" bold="true" box="[151,288,1086,1113]" italics="true" pageId="1" pageNumber="388">Alloeogyna</emphasis>
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Garbelotto & Campos
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</taxonomicName>
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<taxonomicNameLabel id="A23457B6F128AE12CB68D441C2EEF147" box="[581,691,1087,1113]" pageId="1" pageNumber="388" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3696554F128AE12C9BAD4FDC375F185" box="[151,808,1154,1179]" pageId="1" pageNumber="388" type="type_taxon">
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<paragraph id="8BCC36DFF128AE12C9BAD4FDC375F185" blockId="1.[151,808,1154,1179]" box="[151,808,1154,1179]" pageId="1" pageNumber="388">
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<typeStatus id="54C8887DF128AE12C9BAD4FDC094F185" box="[151,201,1155,1179]" pageId="1" pageNumber="388">Type</typeStatus>
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species:
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<taxonomicName id="4C734D5CF128AE12C80BD4FCC225F185" authority="Sailer, 1950" authorityName="Sailer" authorityYear="1950" box="[294,632,1154,1179]" class="Insecta" family="Pentatomidae" genus="Alitocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="brunneus">
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<emphasis id="B907EACDF128AE12C80BD4FCC1ABF185" bold="true" box="[294,502,1154,1179]" italics="true" pageId="1" pageNumber="388">Alitocoris brunneus</emphasis>
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Sailer, 1950
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</taxonomicName>
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, here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3696554F128AE12C9BAD4B4C45AF019" pageId="1" pageNumber="388" type="etymology">
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<paragraph id="8BCC36DFF128AE12C9BAD4B4C45AF019" blockId="1.[151,1437,1225,2044]" pageId="1" pageNumber="388">
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<emphasis id="B907EACDF128AE12C9BAD4B4C143F1FA" bold="true" box="[151,286,1226,1252]" pageId="1" pageNumber="388">Etymology.</emphasis>
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The name refers to the presence and shape of callosities in the female genital plates, unique among Neotropical pentatomids. Greek:
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<emphasis id="B907EACDF128AE12CB21D493C205F019" box="[524,600,1261,1287]" italics="true" pageId="1" pageNumber="388">alloios</emphasis>
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, another kind, different;
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<emphasis id="B907EACDF128AE12CA5DD490C3F8F019" box="[880,933,1262,1287]" italics="true" pageId="1" pageNumber="388">gyne</emphasis>
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, female.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3696554F128AE11C9EAD56FC20BF011" lastPageId="2" lastPageNumber="389" pageId="1" pageNumber="388" type="description">
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<paragraph id="8BCC36DFF128AE12C9EAD56FC1EEF089" blockId="1.[151,1437,1225,2044]" pageId="1" pageNumber="388">
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<emphasis id="B907EACDF128AE12C9EAD56FC104F032" bold="true" box="[199,345,1297,1324]" pageId="1" pageNumber="388">Description.</emphasis>
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Body oval, general color brown sometimes with pale to yellowish maculae and with small maculae at apex of radial vein on corium (
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<figureCitation id="13482A5AF128AE12CB19D54BC2FCF051" box="[564,673,1333,1359]" captionStart="FIGURES 2–5" captionStartId="4.[151,264,1894,1919]" captionTargetBox="[177,1379,215,1820]" captionTargetId="figure@4.[151,1435,231,1869]" captionTargetPageId="4" captionText="FIGURES 2–5. 2, Alloeogyna brunnea (Sailer) comb. nov., male; 3, Alloeogyna poecila Garbelotto & Campos sp. nov., male holotype; 4, Alloeogyna ampla Garbelotto & Campos sp. nov., female holotype; 5, Sailerus maculosus (Sailer) comb. nov., female holotype. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235180/files/figure.png" pageId="1" pageNumber="388">Figs. 2–4</figureCitation>
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). Antennae four- or five-segmented. Gonocoxites VIII with posterolateral callus. Pygophore with open posterolateral angles; inferior layer of ventral rim with foliaceus projections directed dorso-anteriorly.
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</paragraph>
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<paragraph id="8BCC36DFF128AE12C9EAD5DFC517F36E" blockId="1.[151,1437,1225,2044]" pageId="1" pageNumber="388">Head wider than long, declivous; punctures sometimes confluent forming wrinkles from the base of clypeus to the posterior margin of head; vertex tumid or flat. Mandibular plates longer than clypeus, apart at apex, apex rounded; lateral margins sinuous and reflected. Anteocular processes truncated. Antennal tubercles with a lateral spine. Antennae four- or five-segmented. Ventrally 1+1 longitudinal carinae posterior to the eyes, concolor with the surrounding surface to yellowish. Bucculae elevated, with an anterior triangular tooth and evanescent posteriorly, not attaining posterior margin of head. Labium reaching urosternites IV to VI, first segment on prosternum.</paragraph>
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<paragraph id="8BCC36DFF128AE12C9EAD604C3ABF2E5" blockId="1.[151,1437,1225,2044]" pageId="1" pageNumber="388">Pronotum wider than long, declivous; punctures confluent forming wrinkles between humeral angles. Cicatrices slightly tumid, punctured. Anterior margin concave medially; anterior angles with a small tooth projected laterally.Anterolateral margins slightly sinuous and reflected; posterior margin convex. Scutellum longer than wide, reaching connexival segment V; punctures forming shallow wrinkles on disc. Foveae black, flat and with inconspicuous punctures. Corium longer than scutellum, reaching at least the connexival segment VI; punctures denser on exocorium; 1+1 small pale to yellowish maculae at apex of radial vein. Membrane with six to eight veins. Pro-, meso- and metapleura with sparse punctures; pro- and mesosternum punctured and metasternum without punctures. Prosternum furrowed, meso- and metasternum with median thin carinae along entire length. Mesopleural evaporatorium narrow following posterior margin of mesopleura; metapleural evaporatorium occupying half the width of metapleura, lateral margin subrectilinear; peritreme spout, reaching half the evaporatorium width. Tibiae sulcated dorsally; third segment of metatarsi depressed dorsally in males and females.</paragraph>
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<paragraph id="8BCC36DFF12BAE11C9EAD0E7C469F5E5" blockId="2.[151,1437,153,1835]" pageId="2" pageNumber="389">Connexivum exposed from segments III to VII with a narrow lateral strip without punctures. Abdomen sometimes punctured on lateral margins; longitudinal furrow shallow on median third. Trichobothria lateral to an imaginary line tangential to spiracles; spiracles black, visible from urosternites II to VII.</paragraph>
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<paragraph id="8BCC36DFF12BAE11C9EAD17BC4B0F45D" blockId="2.[151,1437,153,1835]" pageId="2" pageNumber="389">Male. Antennae: third to last segment the widest, about two times wider than the next segment; also wider than corresponding segment in females. Articular membrane on posterior margin of tergite VII concave.</paragraph>
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<paragraph id="8BCC36DFF12BAE11C9EAD133C39AF625" blockId="2.[151,1437,153,1835]" pageId="2" pageNumber="389">
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Genitalia. Pygophore globose (
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<figureCitation id="13482A5AF12BAE11CB07D133C2FDF476" box="[554,672,333,360]" captionStart="FIGURES 6–18" captionStartId="5.[151,264,1829,1854]" captionTargetBox="[202,1389,191,1781]" captionTargetId="figure@5.[199,1389,191,1793]" captionTargetPageId="5" captionText="FIGURES 6–18. Alloeogyna Garbelotto & Campos gen. nov. male genitalia. 6–8, All. brunnea (Sailer) comb. nov. pygophore dorsal, ventral and posterior; 9–11, All. poecila Garbelotto & Campos sp. nov. pygophore dorsal, ventral and posterior. 12–15, All. brunnea comb. nov. phallus dorsal, posterior, anterior and lateral; 16–18, All. brunnea comb. nov. right paramere dorsal, ventral and lateral. Abbreviations: cp, conical projection; dp, dorsal process of phallotheca; dr, dorsal rim; ds, ductus seminis distalis; lph, lateral projection of phallotheca; pa, posterolateral angle; par, paramere; ph, phallotheca; pil, projection of inferior layer of ventral rim; v, vesica; vp, vesica process; vr, ventral rim; X, segment X. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235182/files/figure.png" pageId="2" pageNumber="389">Figs 6–11</figureCitation>
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). Posterolateral angle opened (
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<figureCitation id="13482A5AF12BAE11CD2ED133C436F479" box="[1027,1131,333,359]" captionStart="FIGURES 6–18" captionStartId="5.[151,264,1829,1854]" captionTargetBox="[202,1389,191,1781]" captionTargetId="figure@5.[199,1389,191,1793]" captionTargetPageId="5" captionText="FIGURES 6–18. Alloeogyna Garbelotto & Campos gen. nov. male genitalia. 6–8, All. brunnea (Sailer) comb. nov. pygophore dorsal, ventral and posterior; 9–11, All. poecila Garbelotto & Campos sp. nov. pygophore dorsal, ventral and posterior. 12–15, All. brunnea comb. nov. phallus dorsal, posterior, anterior and lateral; 16–18, All. brunnea comb. nov. right paramere dorsal, ventral and lateral. Abbreviations: cp, conical projection; dp, dorsal process of phallotheca; dr, dorsal rim; ds, ductus seminis distalis; lph, lateral projection of phallotheca; pa, posterolateral angle; par, paramere; ph, phallotheca; pil, projection of inferior layer of ventral rim; v, vesica; vp, vesica process; vr, ventral rim; X, segment X. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235182/files/figure.png" pageId="2" pageNumber="389">Figs 6, 9</figureCitation>
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- pa). Dorsal rim concave (
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<figureCitation id="13482A5AF12BAE11C9B2D10FC15DF495" box="[159,256,369,395]" captionStart="FIGURES 6–18" captionStartId="5.[151,264,1829,1854]" captionTargetBox="[202,1389,191,1781]" captionTargetId="figure@5.[199,1389,191,1793]" captionTargetPageId="5" captionText="FIGURES 6–18. Alloeogyna Garbelotto & Campos gen. nov. male genitalia. 6–8, All. brunnea (Sailer) comb. nov. pygophore dorsal, ventral and posterior; 9–11, All. poecila Garbelotto & Campos sp. nov. pygophore dorsal, ventral and posterior. 12–15, All. brunnea comb. nov. phallus dorsal, posterior, anterior and lateral; 16–18, All. brunnea comb. nov. right paramere dorsal, ventral and lateral. Abbreviations: cp, conical projection; dp, dorsal process of phallotheca; dr, dorsal rim; ds, ductus seminis distalis; lph, lateral projection of phallotheca; pa, posterolateral angle; par, paramere; ph, phallotheca; pil, projection of inferior layer of ventral rim; v, vesica; vp, vesica process; vr, ventral rim; X, segment X. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235182/files/figure.png" pageId="2" pageNumber="389">Figs 6, 9</figureCitation>
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- dr), with enlarged margin with less sclerotized appearance at base of posterolateral angles. Inferior layer of ventral rim with 1+1 foliaceous projections with rounded apex over genital cup, almost parallel to frontal plane of pygophore (
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<figureCitation id="13482A5AF12BAE11C810D1C7C1FCF4CD" box="[317,417,441,467]" captionStart="FIGURES 6–18" captionStartId="5.[151,264,1829,1854]" captionTargetBox="[202,1389,191,1781]" captionTargetId="figure@5.[199,1389,191,1793]" captionTargetPageId="5" captionText="FIGURES 6–18. Alloeogyna Garbelotto & Campos gen. nov. male genitalia. 6–8, All. brunnea (Sailer) comb. nov. pygophore dorsal, ventral and posterior; 9–11, All. poecila Garbelotto & Campos sp. nov. pygophore dorsal, ventral and posterior. 12–15, All. brunnea comb. nov. phallus dorsal, posterior, anterior and lateral; 16–18, All. brunnea comb. nov. right paramere dorsal, ventral and lateral. Abbreviations: cp, conical projection; dp, dorsal process of phallotheca; dr, dorsal rim; ds, ductus seminis distalis; lph, lateral projection of phallotheca; pa, posterolateral angle; par, paramere; ph, phallotheca; pil, projection of inferior layer of ventral rim; v, vesica; vp, vesica process; vr, ventral rim; X, segment X. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235182/files/figure.png" pageId="2" pageNumber="389">Figs 6, 9</figureCitation>
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- pil). Surface beneath foliaceous projections with striated lobular expansions, these striae attaining lateral margins of the projections (
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<figureCitation id="13482A5AF12BAE11CBADD1A3C2B8F4E9" box="[640,741,477,503]" captionStart="FIGURES 6–18" captionStartId="5.[151,264,1829,1854]" captionTargetBox="[202,1389,191,1781]" captionTargetId="figure@5.[199,1389,191,1793]" captionTargetPageId="5" captionText="FIGURES 6–18. Alloeogyna Garbelotto & Campos gen. nov. male genitalia. 6–8, All. brunnea (Sailer) comb. nov. pygophore dorsal, ventral and posterior; 9–11, All. poecila Garbelotto & Campos sp. nov. pygophore dorsal, ventral and posterior. 12–15, All. brunnea comb. nov. phallus dorsal, posterior, anterior and lateral; 16–18, All. brunnea comb. nov. right paramere dorsal, ventral and lateral. Abbreviations: cp, conical projection; dp, dorsal process of phallotheca; dr, dorsal rim; ds, ductus seminis distalis; lph, lateral projection of phallotheca; pa, posterolateral angle; par, paramere; ph, phallotheca; pil, projection of inferior layer of ventral rim; v, vesica; vp, vesica process; vr, ventral rim; X, segment X. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235182/files/figure.png" pageId="2" pageNumber="389">Figs 6, 9</figureCitation>
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,
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<figureCitation id="13482A5AF12BAE11CBDCD1A3C366F4E9" box="[753,827,477,503]" captionStart="FIGURES 19–20" captionStartId="6.[151,264,1082,1107]" captionTargetBox="[153,1435,181,1031]" captionTargetId="figure@6.[153,1438,181,1031]" captionTargetPageId="6" captionText="FIGURES 19–20. Alloeogyna brunnea (Sailer) comb. nov. scanning electron microscopy of the surface beneath the projection of the inferior layer of ventral rim. 19, general view; 20, detail." httpUri="https://zenodo.org/record/3235184/files/figure.png" pageId="2" pageNumber="389">19–20</figureCitation>
|
||
). Inferior layer of ventral rim with a median rhomboid conical projection, continued laterally as a low carinae directed to posterolateral angles. Median third of ventral rim U-shaped, with subtriangular ventral opening (
|
||
<figureCitation id="13482A5AF12BAE11CB81D25BC37EF721" box="[684,803,549,575]" captionStart="FIGURES 6–18" captionStartId="5.[151,264,1829,1854]" captionTargetBox="[202,1389,191,1781]" captionTargetId="figure@5.[199,1389,191,1793]" captionTargetPageId="5" captionText="FIGURES 6–18. Alloeogyna Garbelotto & Campos gen. nov. male genitalia. 6–8, All. brunnea (Sailer) comb. nov. pygophore dorsal, ventral and posterior; 9–11, All. poecila Garbelotto & Campos sp. nov. pygophore dorsal, ventral and posterior. 12–15, All. brunnea comb. nov. phallus dorsal, posterior, anterior and lateral; 16–18, All. brunnea comb. nov. right paramere dorsal, ventral and lateral. Abbreviations: cp, conical projection; dp, dorsal process of phallotheca; dr, dorsal rim; ds, ductus seminis distalis; lph, lateral projection of phallotheca; pa, posterolateral angle; par, paramere; ph, phallotheca; pil, projection of inferior layer of ventral rim; v, vesica; vp, vesica process; vr, ventral rim; X, segment X. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235182/files/figure.png" pageId="2" pageNumber="389">Figs 7, 10</figureCitation>
|
||
). Segment X subquadrangular with basal half membranous; apex with setae and not exceeding the posterior limit of inferior layer of ventral rim; lateral margins of the sclerotized portion expanded, and carinated, bearing dense tufts of long setae directed posterolaterally; anal opening apical; genital opening ventral in a longitudinal slit (
|
||
<figureCitation id="13482A5AF12BAE11CBC6D2EFC302F7B2" box="[747,863,657,684]" captionStart="FIGURES 6–18" captionStartId="5.[151,264,1829,1854]" captionTargetBox="[202,1389,191,1781]" captionTargetId="figure@5.[199,1389,191,1793]" captionTargetPageId="5" captionText="FIGURES 6–18. Alloeogyna Garbelotto & Campos gen. nov. male genitalia. 6–8, All. brunnea (Sailer) comb. nov. pygophore dorsal, ventral and posterior; 9–11, All. poecila Garbelotto & Campos sp. nov. pygophore dorsal, ventral and posterior. 12–15, All. brunnea comb. nov. phallus dorsal, posterior, anterior and lateral; 16–18, All. brunnea comb. nov. right paramere dorsal, ventral and lateral. Abbreviations: cp, conical projection; dp, dorsal process of phallotheca; dr, dorsal rim; ds, ductus seminis distalis; lph, lateral projection of phallotheca; pa, posterolateral angle; par, paramere; ph, phallotheca; pil, projection of inferior layer of ventral rim; v, vesica; vp, vesica process; vr, ventral rim; X, segment X. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235182/files/figure.png" pageId="2" pageNumber="389">Figs 6–11</figureCitation>
|
||
). Parameres ventral to, and hidden by the segment X (
|
||
<figureCitation id="13482A5AF12BAE11C996D2CBC129F7D1" box="[187,372,693,719]" captionStart="FIGURES 6–18" captionStartId="5.[151,264,1829,1854]" captionTargetBox="[202,1389,191,1781]" captionTargetId="figure@5.[199,1389,191,1793]" captionTargetPageId="5" captionText="FIGURES 6–18. Alloeogyna Garbelotto & Campos gen. nov. male genitalia. 6–8, All. brunnea (Sailer) comb. nov. pygophore dorsal, ventral and posterior; 9–11, All. poecila Garbelotto & Campos sp. nov. pygophore dorsal, ventral and posterior. 12–15, All. brunnea comb. nov. phallus dorsal, posterior, anterior and lateral; 16–18, All. brunnea comb. nov. right paramere dorsal, ventral and lateral. Abbreviations: cp, conical projection; dp, dorsal process of phallotheca; dr, dorsal rim; ds, ductus seminis distalis; lph, lateral projection of phallotheca; pa, posterolateral angle; par, paramere; ph, phallotheca; pil, projection of inferior layer of ventral rim; v, vesica; vp, vesica process; vr, ventral rim; X, segment X. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235182/files/figure.png" pageId="2" pageNumber="389">Figs 7–8, 10–11</figureCitation>
|
||
), elongated; head bilobed, covered with long setae, ventral lobe projected through the opening of ventral rim (
|
||
<figureCitation id="13482A5AF12BAE11C868D2A7C205F7EA" box="[325,600,729,756]" captionStart="FIGURES 6–18" captionStartId="5.[151,264,1829,1854]" captionTargetBox="[202,1389,191,1781]" captionTargetId="figure@5.[199,1389,191,1793]" captionTargetPageId="5" captionText="FIGURES 6–18. Alloeogyna Garbelotto & Campos gen. nov. male genitalia. 6–8, All. brunnea (Sailer) comb. nov. pygophore dorsal, ventral and posterior; 9–11, All. poecila Garbelotto & Campos sp. nov. pygophore dorsal, ventral and posterior. 12–15, All. brunnea comb. nov. phallus dorsal, posterior, anterior and lateral; 16–18, All. brunnea comb. nov. right paramere dorsal, ventral and lateral. Abbreviations: cp, conical projection; dp, dorsal process of phallotheca; dr, dorsal rim; ds, ductus seminis distalis; lph, lateral projection of phallotheca; pa, posterolateral angle; par, paramere; ph, phallotheca; pil, projection of inferior layer of ventral rim; v, vesica; vp, vesica process; vr, ventral rim; X, segment X. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235182/files/figure.png" pageId="2" pageNumber="389">Figs 7–8, 10–11, 16–18</figureCitation>
|
||
). Phallus (
|
||
<figureCitation id="13482A5AF12BAE11CBFCD2A7C308F7ED" box="[721,853,728,755]" captionStart="FIGURES 6–18" captionStartId="5.[151,264,1829,1854]" captionTargetBox="[202,1389,191,1781]" captionTargetId="figure@5.[199,1389,191,1793]" captionTargetPageId="5" captionText="FIGURES 6–18. Alloeogyna Garbelotto & Campos gen. nov. male genitalia. 6–8, All. brunnea (Sailer) comb. nov. pygophore dorsal, ventral and posterior; 9–11, All. poecila Garbelotto & Campos sp. nov. pygophore dorsal, ventral and posterior. 12–15, All. brunnea comb. nov. phallus dorsal, posterior, anterior and lateral; 16–18, All. brunnea comb. nov. right paramere dorsal, ventral and lateral. Abbreviations: cp, conical projection; dp, dorsal process of phallotheca; dr, dorsal rim; ds, ductus seminis distalis; lph, lateral projection of phallotheca; pa, posterolateral angle; par, paramere; ph, phallotheca; pil, projection of inferior layer of ventral rim; v, vesica; vp, vesica process; vr, ventral rim; X, segment X. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235182/files/figure.png" pageId="2" pageNumber="389">Figs 12–15</figureCitation>
|
||
). Phallotheca bulb shaped; basal lateral projections paired, broad and spatulated, with truncated apex (
|
||
<figureCitation id="13482A5AF12BAE11CBFCD283C379F609" box="[721,804,764,791]" captionStart="FIGURES 6–18" captionStartId="5.[151,264,1829,1854]" captionTargetBox="[202,1389,191,1781]" captionTargetId="figure@5.[199,1389,191,1793]" captionTargetPageId="5" captionText="FIGURES 6–18. Alloeogyna Garbelotto & Campos gen. nov. male genitalia. 6–8, All. brunnea (Sailer) comb. nov. pygophore dorsal, ventral and posterior; 9–11, All. poecila Garbelotto & Campos sp. nov. pygophore dorsal, ventral and posterior. 12–15, All. brunnea comb. nov. phallus dorsal, posterior, anterior and lateral; 16–18, All. brunnea comb. nov. right paramere dorsal, ventral and lateral. Abbreviations: cp, conical projection; dp, dorsal process of phallotheca; dr, dorsal rim; ds, ductus seminis distalis; lph, lateral projection of phallotheca; pa, posterolateral angle; par, paramere; ph, phallotheca; pil, projection of inferior layer of ventral rim; v, vesica; vp, vesica process; vr, ventral rim; X, segment X. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235182/files/figure.png" pageId="2" pageNumber="389">Fig. 15</figureCitation>
|
||
, lph); ventral and dorsal processes sinuous (
|
||
<figureCitation id="13482A5AF12BAE11CC38D283C5CAF609" box="[1301,1431,764,791]" captionStart="FIGURES 6–18" captionStartId="5.[151,264,1829,1854]" captionTargetBox="[202,1389,191,1781]" captionTargetId="figure@5.[199,1389,191,1793]" captionTargetPageId="5" captionText="FIGURES 6–18. Alloeogyna Garbelotto & Campos gen. nov. male genitalia. 6–8, All. brunnea (Sailer) comb. nov. pygophore dorsal, ventral and posterior; 9–11, All. poecila Garbelotto & Campos sp. nov. pygophore dorsal, ventral and posterior. 12–15, All. brunnea comb. nov. phallus dorsal, posterior, anterior and lateral; 16–18, All. brunnea comb. nov. right paramere dorsal, ventral and lateral. Abbreviations: cp, conical projection; dp, dorsal process of phallotheca; dr, dorsal rim; ds, ductus seminis distalis; lph, lateral projection of phallotheca; pa, posterolateral angle; par, paramere; ph, phallotheca; pil, projection of inferior layer of ventral rim; v, vesica; vp, vesica process; vr, ventral rim; X, segment X. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235182/files/figure.png" pageId="2" pageNumber="389">Figs 13, 15</figureCitation>
|
||
, dp). Ductus seminis distalis surpassing base of phallotheca (
|
||
<figureCitation id="13482A5AF12BAE11CA15D35FC3E6F625" box="[824,955,800,827]" captionStart="FIGURES 6–18" captionStartId="5.[151,264,1829,1854]" captionTargetBox="[202,1389,191,1781]" captionTargetId="figure@5.[199,1389,191,1793]" captionTargetPageId="5" captionText="FIGURES 6–18. Alloeogyna Garbelotto & Campos gen. nov. male genitalia. 6–8, All. brunnea (Sailer) comb. nov. pygophore dorsal, ventral and posterior; 9–11, All. poecila Garbelotto & Campos sp. nov. pygophore dorsal, ventral and posterior. 12–15, All. brunnea comb. nov. phallus dorsal, posterior, anterior and lateral; 16–18, All. brunnea comb. nov. right paramere dorsal, ventral and lateral. Abbreviations: cp, conical projection; dp, dorsal process of phallotheca; dr, dorsal rim; ds, ductus seminis distalis; lph, lateral projection of phallotheca; pa, posterolateral angle; par, paramere; ph, phallotheca; pil, projection of inferior layer of ventral rim; v, vesica; vp, vesica process; vr, ventral rim; X, segment X. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235182/files/figure.png" pageId="2" pageNumber="389">Figs 12–15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC36DFF12BAE11C9EAD33BC2E5F641" blockId="2.[151,1437,153,1835]" box="[199,696,837,863]" pageId="2" pageNumber="389">Female. Antennal segments uniformly wide.</paragraph>
|
||
<paragraph id="8BCC36DFF12BAE11C9EAD317C20BF011" blockId="2.[151,1437,153,1835]" pageId="2" pageNumber="389">
|
||
Genitalia. Gonocoxites VIII longer than wide; posterior margin subrectilinear to sinuous over laterotergites IX; disc with posterolateral callus and sutural margins elevated, both varying in size; surface between sutural margins and calli with oblique wrinkles (
|
||
<figureCitation id="13482A5AF12BAE11CB29D3CFC2D5F6D5" box="[516,648,945,971]" captionStart="FIGURES 21–27" captionStartId="8.[151,264,1817,1842]" captionTargetBox="[197,1391,179,1771]" captionTargetId="figure@8.[197,1393,179,1773]" captionTargetPageId="8" captionText="FIGURES 21–27. Female genital plates (ventroposterior view) and internal ducts. 21, 25 All. brunnea (Sailer) comb. nov.; 22, 26, All. poecila Garbelotto & Campos sp. nov.; 23, All. ampla Garbelotto & Campos sp. nov.; 24, 27 Sailerus maculosus (Sailer) comb. nov. Abbreviations: cm, conical microsculptures; cs, capsula seminalis; dr, ductus receptaculi; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpIX, gonapophyses IX; id, inner duct of vesicular area; laVIII, laterotergites VIII; laIX, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pi, pars intermedialis; rs, ring sclerites; tgIX, thickening of gonapophyses IX; vi, thickening of vaginal intima; VII, urosternite VII. Scale bars: 1mm." httpUri="https://zenodo.org/record/3235186/files/figure.png" pageId="2" pageNumber="389">Figs 21–23</figureCitation>
|
||
). Laterotergites VIII triangular, wider than long; spiracles present in a callus and posterior to lateral angle of gonocoxites VIII (
|
||
<figureCitation id="13482A5AF12BAE11CA38D3ABC3C5F6F1" box="[789,920,981,1007]" captionStart="FIGURES 21–27" captionStartId="8.[151,264,1817,1842]" captionTargetBox="[197,1391,179,1771]" captionTargetId="figure@8.[197,1393,179,1773]" captionTargetPageId="8" captionText="FIGURES 21–27. Female genital plates (ventroposterior view) and internal ducts. 21, 25 All. brunnea (Sailer) comb. nov.; 22, 26, All. poecila Garbelotto & Campos sp. nov.; 23, All. ampla Garbelotto & Campos sp. nov.; 24, 27 Sailerus maculosus (Sailer) comb. nov. Abbreviations: cm, conical microsculptures; cs, capsula seminalis; dr, ductus receptaculi; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpIX, gonapophyses IX; id, inner duct of vesicular area; laVIII, laterotergites VIII; laIX, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pi, pars intermedialis; rs, ring sclerites; tgIX, thickening of gonapophyses IX; vi, thickening of vaginal intima; VII, urosternite VII. Scale bars: 1mm." httpUri="https://zenodo.org/record/3235186/files/figure.png" pageId="2" pageNumber="389">Figs 21–23</figureCitation>
|
||
). Gonocoxites IX reduced to thin longitudinal carina externally (
|
||
<figureCitation id="13482A5AF12BAE11C870D387C183F10D" box="[349,478,1017,1043]" captionStart="FIGURES 21–27" captionStartId="8.[151,264,1817,1842]" captionTargetBox="[197,1391,179,1771]" captionTargetId="figure@8.[197,1393,179,1773]" captionTargetPageId="8" captionText="FIGURES 21–27. Female genital plates (ventroposterior view) and internal ducts. 21, 25 All. brunnea (Sailer) comb. nov.; 22, 26, All. poecila Garbelotto & Campos sp. nov.; 23, All. ampla Garbelotto & Campos sp. nov.; 24, 27 Sailerus maculosus (Sailer) comb. nov. Abbreviations: cm, conical microsculptures; cs, capsula seminalis; dr, ductus receptaculi; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpIX, gonapophyses IX; id, inner duct of vesicular area; laVIII, laterotergites VIII; laIX, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pi, pars intermedialis; rs, ring sclerites; tgIX, thickening of gonapophyses IX; vi, thickening of vaginal intima; VII, urosternite VII. Scale bars: 1mm." httpUri="https://zenodo.org/record/3235186/files/figure.png" pageId="2" pageNumber="389">Figs 21–23</figureCitation>
|
||
), and inner portion medially narrowed (
|
||
<figureCitation id="13482A5AF12BAE11CABAD387C447F10D" box="[919,1050,1016,1043]" captionStart="FIGURES 21–27" captionStartId="8.[151,264,1817,1842]" captionTargetBox="[197,1391,179,1771]" captionTargetId="figure@8.[197,1393,179,1773]" captionTargetPageId="8" captionText="FIGURES 21–27. Female genital plates (ventroposterior view) and internal ducts. 21, 25 All. brunnea (Sailer) comb. nov.; 22, 26, All. poecila Garbelotto & Campos sp. nov.; 23, All. ampla Garbelotto & Campos sp. nov.; 24, 27 Sailerus maculosus (Sailer) comb. nov. Abbreviations: cm, conical microsculptures; cs, capsula seminalis; dr, ductus receptaculi; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpIX, gonapophyses IX; id, inner duct of vesicular area; laVIII, laterotergites VIII; laIX, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pi, pars intermedialis; rs, ring sclerites; tgIX, thickening of gonapophyses IX; vi, thickening of vaginal intima; VII, urosternite VII. Scale bars: 1mm." httpUri="https://zenodo.org/record/3235186/files/figure.png" pageId="2" pageNumber="389">Figs 25–26</figureCitation>
|
||
). Laterotergites IX triangular, with rounded apex not surpassing the posterior margins of laterotergites VIII; lateral and mesial margins levelled (
|
||
<figureCitation id="13482A5AF12BAE11CC46D463C0BFF145" captionStart="FIGURES 21–27" captionStartId="8.[151,264,1817,1842]" captionTargetBox="[197,1391,179,1771]" captionTargetId="figure@8.[197,1393,179,1773]" captionTargetPageId="8" captionText="FIGURES 21–27. Female genital plates (ventroposterior view) and internal ducts. 21, 25 All. brunnea (Sailer) comb. nov.; 22, 26, All. poecila Garbelotto & Campos sp. nov.; 23, All. ampla Garbelotto & Campos sp. nov.; 24, 27 Sailerus maculosus (Sailer) comb. nov. Abbreviations: cm, conical microsculptures; cs, capsula seminalis; dr, ductus receptaculi; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpIX, gonapophyses IX; id, inner duct of vesicular area; laVIII, laterotergites VIII; laIX, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pi, pars intermedialis; rs, ring sclerites; tgIX, thickening of gonapophyses IX; vi, thickening of vaginal intima; VII, urosternite VII. Scale bars: 1mm." httpUri="https://zenodo.org/record/3235186/files/figure.png" pageId="2" pageNumber="389">Figs 21–23</figureCitation>
|
||
); in inner view anterior margin sinuous (
|
||
<figureCitation id="13482A5AF12BAE11CB81D43FC36DF145" box="[684,816,1088,1115]" captionStart="FIGURES 21–27" captionStartId="8.[151,264,1817,1842]" captionTargetBox="[197,1391,179,1771]" captionTargetId="figure@8.[197,1393,179,1773]" captionTargetPageId="8" captionText="FIGURES 21–27. Female genital plates (ventroposterior view) and internal ducts. 21, 25 All. brunnea (Sailer) comb. nov.; 22, 26, All. poecila Garbelotto & Campos sp. nov.; 23, All. ampla Garbelotto & Campos sp. nov.; 24, 27 Sailerus maculosus (Sailer) comb. nov. Abbreviations: cm, conical microsculptures; cs, capsula seminalis; dr, ductus receptaculi; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpIX, gonapophyses IX; id, inner duct of vesicular area; laVIII, laterotergites VIII; laIX, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pi, pars intermedialis; rs, ring sclerites; tgIX, thickening of gonapophyses IX; vi, thickening of vaginal intima; VII, urosternite VII. Scale bars: 1mm." httpUri="https://zenodo.org/record/3235186/files/figure.png" pageId="2" pageNumber="389">Figs 25–26</figureCitation>
|
||
). Segment X hidden by laterotergites IX. Pars comunis with conical microsculptures. Gonapophyses VIII and IX with folds bearing conical microsculptures; folds of gonapophyses IX following anterior margin of gonocoxites IX (
|
||
<figureCitation id="13482A5AF12BAE11CA67D4F7C393F1BD" box="[842,974,1160,1187]" captionStart="FIGURES 21–27" captionStartId="8.[151,264,1817,1842]" captionTargetBox="[197,1391,179,1771]" captionTargetId="figure@8.[197,1393,179,1773]" captionTargetPageId="8" captionText="FIGURES 21–27. Female genital plates (ventroposterior view) and internal ducts. 21, 25 All. brunnea (Sailer) comb. nov.; 22, 26, All. poecila Garbelotto & Campos sp. nov.; 23, All. ampla Garbelotto & Campos sp. nov.; 24, 27 Sailerus maculosus (Sailer) comb. nov. Abbreviations: cm, conical microsculptures; cs, capsula seminalis; dr, ductus receptaculi; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpIX, gonapophyses IX; id, inner duct of vesicular area; laVIII, laterotergites VIII; laIX, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pi, pars intermedialis; rs, ring sclerites; tgIX, thickening of gonapophyses IX; vi, thickening of vaginal intima; VII, urosternite VII. Scale bars: 1mm." httpUri="https://zenodo.org/record/3235186/files/figure.png" pageId="2" pageNumber="389">Figs 25–26</figureCitation>
|
||
). Thickening of gonapophyses IX as 1+1 circles, each one lateral to the margins of thickening of vaginal intima (
|
||
<figureCitation id="13482A5AF12BAE11CAE4D4D3C413F1D9" box="[969,1102,1196,1223]" captionStart="FIGURES 21–27" captionStartId="8.[151,264,1817,1842]" captionTargetBox="[197,1391,179,1771]" captionTargetId="figure@8.[197,1393,179,1773]" captionTargetPageId="8" captionText="FIGURES 21–27. Female genital plates (ventroposterior view) and internal ducts. 21, 25 All. brunnea (Sailer) comb. nov.; 22, 26, All. poecila Garbelotto & Campos sp. nov.; 23, All. ampla Garbelotto & Campos sp. nov.; 24, 27 Sailerus maculosus (Sailer) comb. nov. Abbreviations: cm, conical microsculptures; cs, capsula seminalis; dr, ductus receptaculi; gcVIII, gonocoxites VIII; gcIX, gonocoxites IX; gpIX, gonapophyses IX; id, inner duct of vesicular area; laVIII, laterotergites VIII; laIX, laterotergites IX; md, median duct of vesicular area; od, outer duct of vesicular area; pi, pars intermedialis; rs, ring sclerites; tgIX, thickening of gonapophyses IX; vi, thickening of vaginal intima; VII, urosternite VII. Scale bars: 1mm." httpUri="https://zenodo.org/record/3235186/files/figure.png" pageId="2" pageNumber="389">Figs 25–26</figureCitation>
|
||
). Ring sclerites laterally constricted. Median duct of vesicular area enlarged at base; pars intermedialis longer than wide; processes of capsula seminalis thin; annular crests narrowed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3696554F12BAE11C9EAD567C1FAF216" pageId="2" pageNumber="389" type="discussion">
|
||
<paragraph id="8BCC36DFF12BAE11C9EAD567C3CAF369" blockId="2.[151,1437,153,1835]" pageId="2" pageNumber="389">
|
||
<emphasis id="B907EACDF12BAE11C9EAD567C166F02D" bold="true" box="[199,315,1305,1331]" pageId="2" pageNumber="389">Remarks.</emphasis>
|
||
<taxonomicName id="4C734D5CF12BAE11C86FD567C19FF02D" authority="Garbelotto & Grazia & Campos, 2019" authorityName="Garbelotto & Grazia & Campos" authorityYear="2019" box="[322,450,1305,1331]" class="Insecta" family="Pentatomidae" genus="Alloeogyna" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="389" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B907EACDF12BAE11C86FD567C19FF02D" box="[322,450,1305,1331]" italics="true" pageId="2" pageNumber="389">Alloeogyna</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457B6F12BAE11C8E5D567C26DF02D" box="[456,560,1305,1331]" pageId="2" pageNumber="389" rank="genus">
|
||
<emphasis id="B907EACDF12BAE11C8E5D567C26DF02D" bold="true" box="[456,560,1305,1331]" pageId="2" pageNumber="389">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is proposed based on the phylogeny of the
|
||
<taxonomicName id="4C734D5CF12BAE11CD3ED567C4CBF02D" authorityName="Distant" authorityYear="1911" box="[1043,1174,1305,1331]" class="Insecta" family="Pentatomidae" genus="Herrichella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="389" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B907EACDF12BAE11CD3ED567C4CBF02D" box="[1043,1174,1305,1331]" italics="true" pageId="2" pageNumber="389">Herrichella</emphasis>
|
||
</taxonomicName>
|
||
clade (Ochlerini) to include the species in clade I (
|
||
<figureCitation id="13482A5AF12BAE11C8FCD543C24CF049" box="[465,529,1341,1367]" captionStart="FIGURE 1" captionStartId="3.[151,250,1096,1121]" captionTargetBox="[153,1435,392,1020]" captionTargetId="figure@3.[151,1436,392,1020]" captionTargetPageId="3" captionText="FIGURE 1. Relationships of the ‘clade D’ of the Herrichela clade of Ochlerini, showing the relationships between Sailerus gen. nov., Alloeogyna gen. nov. and close related genera, as well relationship between Alloeogyna gen. nov. species. Modified from Garbelotto et al. (2013)." httpUri="https://zenodo.org/record/3235178/files/figure.png" pageId="2" pageNumber="389">Fig. 1</figureCitation>
|
||
) (
|
||
<bibRefCitation id="EFE24B2EF12BAE11CB07D543C378F049" author="Garbelotto, T. A. & Campos, L. A. & Grazia, J." box="[554,805,1341,1367]" pageId="2" pageNumber="389" pagination="452 - 472" refId="ref9489" refString="Garbelotto, T. A., Campos, L. A. & Grazia, J. (2013) Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society, 168, 452 - 472. https: // doi. org / 10.1111 / zoj. 12032" type="journal article" year="2013">
|
||
Garbelotto
|
||
<emphasis id="B907EACDF12BAE11CB87D540C2BFF049" box="[682,738,1341,1367]" italics="true" pageId="2" pageNumber="389">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
). The two synapomorphies supporting clade I are female traits: the sutural margins of gonocoxites VIII swollen in the apical half (character 51(1) of
|
||
<bibRefCitation id="EFE24B2EF12BAE11CDBBD51FC5CDF065" author="Garbelotto, T. A. & Campos, L. A. & Grazia, J." box="[1174,1424,1377,1403]" pageId="2" pageNumber="389" pagination="452 - 472" refId="ref9489" refString="Garbelotto, T. A., Campos, L. A. & Grazia, J. (2013) Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society, 168, 452 - 472. https: // doi. org / 10.1111 / zoj. 12032" type="journal article" year="2013">
|
||
Garbelotto
|
||
<emphasis id="B907EACDF12BAE11CC38D51CC510F065" box="[1301,1357,1377,1403]" italics="true" pageId="2" pageNumber="389">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
), and the callus on the disc projecting to the lateral angles of gonocoxites VIII (character 52(2) of
|
||
<bibRefCitation id="EFE24B2EF12BAE11CDCED5FBC089F0DD" author="Garbelotto, T. A. & Campos, L. A. & Grazia, J." pageId="2" pageNumber="389" pagination="452 - 472" refId="ref9489" refString="Garbelotto, T. A., Campos, L. A. & Grazia, J. (2013) Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society, 168, 452 - 472. https: // doi. org / 10.1111 / zoj. 12032" type="journal article" year="2013">
|
||
Garbelotto
|
||
<emphasis id="B907EACDF12BAE11CC4ED5F8C5C0F081" box="[1379,1437,1413,1439]" italics="true" pageId="2" pageNumber="389">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
). Other characters are recognized as diagnostic for the genus, as follows. For females: gonocoxites IX reduced to thin longitudinal carinae externally, and laterally narrowed internally (character 62(1) of
|
||
<bibRefCitation id="EFE24B2EF12BAE11CDBBD5B3C5CDF0F9" author="Garbelotto, T. A. & Campos, L. A. & Grazia, J." box="[1174,1424,1485,1511]" pageId="2" pageNumber="389" pagination="452 - 472" refId="ref9489" refString="Garbelotto, T. A., Campos, L. A. & Grazia, J. (2013) Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society, 168, 452 - 472. https: // doi. org / 10.1111 / zoj. 12032" type="journal article" year="2013">
|
||
Garbelotto
|
||
<emphasis id="B907EACDF12BAE11CC38D5B0C510F0F9" box="[1301,1357,1485,1511]" italics="true" pageId="2" pageNumber="389">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
), laterotergites IX with a basal constriction internally (character 64(1) of
|
||
<bibRefCitation id="EFE24B2EF12BAE11CA82D58FC4F9F315" author="Garbelotto, T. A. & Campos, L. A. & Grazia, J." box="[943,1188,1521,1547]" pageId="2" pageNumber="389" pagination="452 - 472" refId="ref9489" refString="Garbelotto, T. A., Campos, L. A. & Grazia, J. (2013) Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society, 168, 452 - 472. https: // doi. org / 10.1111 / zoj. 12032" type="journal article" year="2013">
|
||
Garbelotto
|
||
<emphasis id="B907EACDF12BAE11CD00D58CC439F315" box="[1069,1124,1521,1547]" italics="true" pageId="2" pageNumber="389">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
) and the thickening of gonapophyses IX composed of a pair of circles (character 67(4) of
|
||
<bibRefCitation id="EFE24B2EF12BAE11CAA6D66BC4DAF331" author="Garbelotto, T. A. & Campos, L. A. & Grazia, J." box="[907,1159,1557,1583]" pageId="2" pageNumber="389" pagination="452 - 472" refId="ref9489" refString="Garbelotto, T. A., Campos, L. A. & Grazia, J. (2013) Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society, 168, 452 - 472. https: // doi. org / 10.1111 / zoj. 12032" type="journal article" year="2013">
|
||
Garbelotto
|
||
<emphasis id="B907EACDF12BAE11CD27D668C41EF331" box="[1034,1091,1557,1583]" italics="true" pageId="2" pageNumber="389">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
). For males, the subrectangular projections of the inferior layer of ventral rim, posterolateral angles opened, ventral opening of pygophore triangular, 1+1 lateral carinae on segment X, and bilobed parameres.
|
||
</paragraph>
|
||
<paragraph id="8BCC36DFF12BAE11C9EAD6FFC1FAF216" blockId="2.[151,1437,153,1835]" pageId="2" pageNumber="389">
|
||
<taxonomicName id="4C734D5CF12BAE11C9EAD6FFC164F385" authority="Garbelotto & Grazia & Campos, 2019" authorityName="Garbelotto & Grazia & Campos" authorityYear="2019" box="[199,313,1665,1691]" class="Insecta" family="Pentatomidae" genus="Alloegyna" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="389" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B907EACDF12BAE11C9EAD6FFC164F385" box="[199,313,1665,1691]" italics="true" pageId="2" pageNumber="389">Alloegyna</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457B6F12BAE11C86CD6FFC1F1F385" box="[321,428,1665,1691]" pageId="2" pageNumber="389" rank="genus">
|
||
<emphasis id="B907EACDF12BAE11C86CD6FFC1F1F385" bold="true" box="[321,428,1665,1691]" pageId="2" pageNumber="389">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is the sister group of
|
||
<taxonomicName id="4C734D5CF12BAE11CB8AD6FFC441F385" authority="Garbelotto & Campos" authorityName="Garbelotto & Campos" authorityYear="2016" box="[679,1052,1665,1691]" class="Insecta" family="Pentatomidae" genus="Stapecolis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="389" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B907EACDF12BAE11CB8AD6FFC347F385" box="[679,794,1665,1691]" italics="true" pageId="2" pageNumber="389">Stapecolis</emphasis>
|
||
Garbelotto & Campos
|
||
</taxonomicName>
|
||
and this relationship is supported by the second antennal segment shorter than the first, and the gonocoxites IX as a carina externally (
|
||
<bibRefCitation id="EFE24B2EF12BAE11CC2BD6DBC0A5F3FD" author="Garbelotto, T. A. & Campos, L. A. & Grazia, J." pageId="2" pageNumber="389" pagination="452 - 472" refId="ref9489" refString="Garbelotto, T. A., Campos, L. A. & Grazia, J. (2013) Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society, 168, 452 - 472. https: // doi. org / 10.1111 / zoj. 12032" type="journal article" year="2013">
|
||
Garbelotto
|
||
<emphasis id="B907EACDF12BAE11CCA5D6D8C0E9F3FD" italics="true" pageId="2" pageNumber="389">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE24B2EF12BAE11C829D6B7C11CF3FA" author="Garbelotto, T. A. & Campos, L. A. & Grazia, J." box="[260,321,1737,1764]" pageId="2" pageNumber="389" pagination="545 - 552" refId="ref9622" refString="Garbelotto, T. A., Campos, L. A. & Grazia, J. (2016) Stapecolis, new genus of Ochlerini (Hemiptera: Heteroptera: Discocephalinae). Zootaxa, 4137 (4), 545 - 552. http: // doi. org / 10.11646 / zootaxa. 4137.4.6" type="journal article" year="2016">2016</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C734D5CF12BAE11C878D6B7C195F3FD" authorityName="Garbelotto & Campos" authorityYear="2016" box="[341,456,1737,1763]" class="Insecta" family="Pentatomidae" genus="Stapecolis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="389" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B907EACDF12BAE11C878D6B7C195F3FD" box="[341,456,1737,1763]" italics="true" pageId="2" pageNumber="389">Stapecolis</emphasis>
|
||
</taxonomicName>
|
||
is supported by other synapomorphies and shows some unique pygophoral characters (
|
||
<bibRefCitation id="EFE24B2EF12BAE11C9B2D693C1C6F216" author="Garbelotto, T. A. & Campos, L. A. & Grazia, J." box="[159,411,1773,1800]" pageId="2" pageNumber="389" pagination="545 - 552" refId="ref9622" refString="Garbelotto, T. A., Campos, L. A. & Grazia, J. (2016) Stapecolis, new genus of Ochlerini (Hemiptera: Heteroptera: Discocephalinae). Zootaxa, 4137 (4), 545 - 552. http: // doi. org / 10.11646 / zootaxa. 4137.4.6" type="journal article" year="2016">
|
||
Garbelotto
|
||
<emphasis id="B907EACDF12BAE11C832D690C105F219" box="[287,344,1773,1799]" italics="true" pageId="2" pageNumber="389">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3696554F12BAE11C9EAD76EC45FF235" box="[199,1026,1808,1835]" pageId="2" pageNumber="389" type="distribution">
|
||
<paragraph id="8BCC36DFF12BAE11C9EAD76EC45FF235" blockId="2.[151,1437,153,1835]" box="[199,1026,1808,1835]" pageId="2" pageNumber="389">
|
||
<emphasis id="B907EACDF12BAE11C9EAD76EC13CF235" bold="true" box="[199,353,1808,1835]" pageId="2" pageNumber="389">Distribution.</emphasis>
|
||
Mexico, Guatemala, Honduras, El Salvador and Nicaragua.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSection id="E2FC2F34F12BAE10C9BAD703C5C1F45C" lastPageId="3" lastPageNumber="390" pageId="2" pageNumber="389" type="key">
|
||
<subSubSection id="C3696554F12BAE11C9BAD703C3E4F289" box="[151,953,1917,1944]" pageId="2" pageNumber="389" type="nomenclature">
|
||
<paragraph id="8BCC36DFF12BAE11C9BAD703C3E4F289" blockId="2.[151,953,1917,1944]" box="[151,953,1917,1944]" pageId="2" pageNumber="389">
|
||
<heading id="D08481B3F12BAE11C9BAD703C3E4F289" bold="true" box="[151,953,1917,1944]" fontSize="11" level="1" pageId="2" pageNumber="389" reason="1">
|
||
<emphasis id="B907EACDF12BAE11C9BAD703C3E4F289" bold="true" box="[151,953,1917,1944]" pageId="2" pageNumber="389">
|
||
Key to the species of
|
||
<taxonomicName id="4C734D5CF12BAE11C8B0D703C319F289" authority="Garbelotto & Campos" authorityName="Garbelotto & Grazia & Campos" authorityYear="2019" box="[413,836,1917,1944]" class="Insecta" family="Pentatomidae" genus="Alloeogyna" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="389" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B907EACDF12BAE11C8B0D703C27BF286" bold="true" box="[413,550,1917,1944]" italics="true" pageId="2" pageNumber="389">Alloeogyna</emphasis>
|
||
Garbelotto & Campos
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457B6F12BAE11CA66D703C3E4F289" box="[843,953,1917,1943]" pageId="2" pageNumber="389" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3696554F12BAE10C9BAD7BCC5C1F45C" lastPageId="3" lastPageNumber="390" pageId="2" pageNumber="389" type="multiple">
|
||
<key id="FFE261EEF12BAE10C9BAD7BCC5C1F45C" lastPageId="3" lastPageNumber="390" pageId="2" pageNumber="389">
|
||
<footnote id="E8682AD1F12BAE11C9BAD7BCC5C1F2EB" pageId="2" pageNumber="389">
|
||
<keyStep id="30872E7AF12BAE11C9BAD7BCC5C1F2EB" pageId="2" pageNumber="389">
|
||
<paragraph id="8BCC36DFF12BAE11C9BAD7BCC5C1F2C9" blockId="2.[151,1436,1985,2037]" box="[151,1436,1985,2007]" pageId="2" pageNumber="389">
|
||
<keyLead id="308295EAF12BAE11C9BAD7BCC5C1F2C9" box="[151,1436,1985,2007]" pageId="2" pageNumber="389">1. Antennae 4-segmented................................................................................ 2</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BCC36DFF12BAE11C9BAD7A1C5C1F2EB" blockId="2.[151,1436,1985,2037]" box="[151,1436,2015,2037]" pageId="2" pageNumber="389">
|
||
<keyLead id="308295EAF12BAE11C9BAD7A1C5C1F2EB" box="[151,1436,2015,2037]" pageId="2" pageNumber="389">
|
||
1’. Antennae 5-segmented..........................................
|
||
<taxonomicName id="4C734D5CF12BAE11CAE4D7A1C486F2EB" authority="(Sailer)" baseAuthorityName="Sailer" box="[969,1243,2015,2037]" class="Insecta" family="Pentatomidae" genus="Alloeogyna" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="brunnea" status="comb. nov.">
|
||
<emphasis id="B907EACDF12BAE11CAE4D7A1C4D1F2EA" box="[969,1164,2015,2036]" italics="true" pageId="2" pageNumber="389">Alloeogyna brunnea</emphasis>
|
||
(Sailer)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457B6F12BAE11CDCDD7A1C513F2EA" box="[1248,1358,2015,2036]" pageId="2" pageNumber="389" rank="species">
|
||
<emphasis id="B907EACDF12BAE11CDCDD7A1C513F2EA" bold="true" box="[1248,1358,2015,2036]" pageId="2" pageNumber="389">comb. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(Fig. 2)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</footnote>
|
||
<keyStep id="30872E7AF12AAE10C9BAD0E4C5C1F45C" pageId="3" pageNumber="390">
|
||
<paragraph id="8BCC36DFF12AAE10C9BAD0E4C5C1F5F4" blockId="3.[151,1437,153,322]" pageId="3" pageNumber="390">
|
||
<keyLead id="308295EAF12AAE10C9BAD0E4C5C1F5F4" pageId="3" pageNumber="390">
|
||
2. Total length less than
|
||
<quantity id="4C8B9B3AF12AAE10C894D0E4C1A4F5B1" box="[441,505,154,175]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="390" unit="mm" value="10.0">10mm</quantity>
|
||
; labium reaching fifth urosternite; corium reaching half of connexivum VI; body dorsal surface light brown with 1+1 pale spots at lateral angle of the cicatrices of pronotum, and 1+1 pale spots on margin of foveae; ventral surface of head with yellowish longitudinal maculae posterior to the eyes.............
|
||
<taxonomicName id="4C734D5CF12AAE10CD6FD0AAC4A1F5F7" authority="Garbelotto & Grazia & Campos, 2019" authorityName="Garbelotto & Grazia & Campos" authorityYear="2019" box="[1090,1276,212,233]" class="Insecta" family="Pentatomidae" genus="Alloeogyna" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="poecila" status="sp. nov.">
|
||
<emphasis id="B907EACDF12AAE10CD6FD0AAC4A1F5F7" box="[1090,1276,212,233]" italics="true" pageId="3" pageNumber="390">Alloeogyna poecila</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457B6F12AAE10CC2FD0AAC513F5F7" box="[1282,1358,212,233]" pageId="3" pageNumber="390" rank="species">
|
||
<emphasis id="B907EACDF12AAE10CC2FD0AAC513F5F7" bold="true" box="[1282,1358,212,233]" pageId="3" pageNumber="390">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13482A5AF12AAE10CC76D0AAC5C9F5F4" box="[1371,1428,212,234]" captionStart="FIGURES 2–5" captionStartId="4.[151,264,1894,1919]" captionTargetBox="[177,1379,215,1820]" captionTargetId="figure@4.[151,1435,231,1869]" captionTargetPageId="4" captionText="FIGURES 2–5. 2, Alloeogyna brunnea (Sailer) comb. nov., male; 3, Alloeogyna poecila Garbelotto & Campos sp. nov., male holotype; 4, Alloeogyna ampla Garbelotto & Campos sp. nov., female holotype; 5, Sailerus maculosus (Sailer) comb. nov., female holotype. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235180/files/figure.png" pageId="3" pageNumber="390">Fig. 3</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BCC36DFF12AAE10C9BAD08CC5C1F45C" blockId="3.[151,1437,153,322]" pageId="3" pageNumber="390">
|
||
<keyLead id="308295EAF12AAE10C9BAD08CC5C1F45C" pageId="3" pageNumber="390">
|
||
2’. Total length at least
|
||
<quantity id="4C8B9B3AF12AAE10C883D08CC1B0F419" box="[430,493,242,263]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="390" unit="mm" value="11.0">11mm</quantity>
|
||
; labium reaching fourth urosternite; corium reaching posterior margin of connexivum VI; body dorsal surface brown, without spots at lateral angle of the cicatrices of pronotum, and on margin of foveae; ventral surface of head without yellowish maculae..............................................
|
||
<taxonomicName id="4C734D5CF12AAE10CD61D152C4A1F45F" authority="Garbelotto & Grazia & Campos, 2019" authorityName="Garbelotto & Grazia & Campos" authorityYear="2019" box="[1100,1276,300,321]" class="Insecta" family="Pentatomidae" genus="Alloeogyna" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="ampla" status="sp. nov.">
|
||
<emphasis id="B907EACDF12AAE10CD61D152C4A1F45F" box="[1100,1276,300,321]" italics="true" pageId="3" pageNumber="390">Alloeogyna ampla</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23457B6F12AAE10CC2FD152C513F45F" box="[1282,1358,300,321]" pageId="3" pageNumber="390" rank="species">
|
||
<emphasis id="B907EACDF12AAE10CC2FD152C513F45F" bold="true" box="[1282,1358,300,321]" pageId="3" pageNumber="390">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13482A5AF12AAE10CC76D152C5C8F45C" box="[1371,1429,300,322]" captionStart="FIGURES 2–5" captionStartId="4.[151,264,1894,1919]" captionTargetBox="[177,1379,215,1820]" captionTargetId="figure@4.[151,1435,231,1869]" captionTargetPageId="4" captionText="FIGURES 2–5. 2, Alloeogyna brunnea (Sailer) comb. nov., male; 3, Alloeogyna poecila Garbelotto & Campos sp. nov., male holotype; 4, Alloeogyna ampla Garbelotto & Campos sp. nov., female holotype; 5, Sailerus maculosus (Sailer) comb. nov., female holotype. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3235180/files/figure.png" pageId="3" pageNumber="390">Fig. 4</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |