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<mods:title id="04491400C5CA7B36D2925C6D3634A2E9">Revision of the status of some genus-level water mite taxa in the families Pionidae Thor, 1900, Aturidae Thor, 1900, and Nudomideopsidae Smith, 1990 (Acari: Hydrachnidiae)</mods:title>
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<treatment id="03DA87C4FFA2174CFF4C6C9CAF4FFD08" ID-DOI="http://doi.org/10.5281/zenodo.6109944" ID-GBIF-Taxon="119596500" ID-Zenodo-Dep="6109944" LSID="urn:lsid:plazi:treatment:03DA87C4FFA2174CFF4C6C9CAF4FFD08" httpUri="http://treatment.plazi.org/id/03DA87C4FFA2174CFF4C6C9CAF4FFD08" lastPageId="43" lastPageNumber="154" pageId="42" pageNumber="153">
<subSubSection id="C3696559FFA2174DFF4C6C9CA827F89A" box="[151,571,1789,1815]" pageId="42" pageNumber="153" type="nomenclature">
<paragraph id="8BCC36D2FFA2174DFF4C6C9CA827F89A" blockId="42.[151,571,1789,1815]" box="[151,571,1789,1815]" pageId="42" pageNumber="153">
<heading id="D08481BEFFA2174DFF4C6C9CA827F89A" bold="true" box="[151,571,1789,1815]" fontSize="11" level="1" pageId="42" pageNumber="153" reason="1">
<emphasis id="B907EAC0FFA2174DFF4C6C9CA827F89A" bold="true" box="[151,571,1789,1815]" pageId="42" pageNumber="153">
Genus
<taxonomicName id="4C734D51FFA2174DFF356C9CA827F89A" authority="Smith, 1983" authorityName="Smith" authorityYear="1983" box="[238,571,1789,1815]" class="Arachnida" family="Mideopsidae" genus="Paramideopsis" kingdom="Animalia" order="Trombidiformes" pageId="42" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FFA2174DFF356C9CABBCF89A" bold="true" box="[238,416,1789,1815]" italics="true" pageId="42" pageNumber="153">Paramideopsis</emphasis>
Smith, 1983
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3696559FFA2174DFF4C6D25A9B1F87E" pageId="42" pageNumber="153" type="reference_group">
<paragraph id="8BCC36D2FFA2174DFF4C6D25A8DDF8D7" blockId="42.[151,977,1860,2035]" box="[151,705,1860,1882]" pageId="42" pageNumber="153">
<taxonomicName id="4C734D51FFA2174DFF4C6D25ABA3F8D7" authority="Smith, 1983" authorityName="Smith" authorityYear="1983" box="[151,447,1860,1882]" class="Arachnida" family="Mideopsidae" genus="Paramideopsis" kingdom="Animalia" order="Trombidiformes" pageId="42" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FFA2174DFF4C6D25AB33F8D7" box="[151,303,1860,1882]" italics="true" pageId="42" pageNumber="153">Paramideopsis</emphasis>
<bibRefCitation id="EFE24B23FFA2174DFEEE6D25ABA3F8D7" author="Smith" box="[309,447,1860,1882]" pageId="42" pageNumber="153" refString="Smith, I. M. (1983 b) Description of larvae and adults of Paramideopsis susanae n. gen., n. sp., with remarks on phylogeny and classification of Mideopsidae (Acari: Parasitengona: Arrenuroidea). Canadian Entomologist, 115, 529 - 538. http: // dx. doi. org / 10.4039 / Ent 115529 - 5" type="journal article" year="1983" yearSuffix="b">Smith, 1983b</bibRefCitation>
</taxonomicName>
, pp. 529538, figs. 113.
</paragraph>
<paragraph id="8BCC36D2FFA2174DFF4C6D03A8D5F8F5" blockId="42.[151,977,1860,2035]" box="[151,713,1890,1912]" pageId="42" pageNumber="153">
<taxonomicName id="4C734D51FFA2174DFF4C6D03ABA6F8F5" authority="Smith, 1990" authorityName="Smith" authorityYear="1990" box="[151,442,1890,1912]" class="Arachnida" family="Mideopsidae" genus="Paramideopsis" kingdom="Animalia" order="Trombidiformes" pageId="42" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FFA2174DFF4C6D03AB33F8F5" box="[151,303,1890,1912]" italics="true" pageId="42" pageNumber="153">Paramideopsis</emphasis>
:
<bibRefCitation id="EFE24B23FFA2174DFEE76D03ABA6F8F5" author="Smith" box="[316,442,1890,1912]" pageId="42" pageNumber="153" refString="Smith, I. M. (1990) Proposal of Nudomideopsidae fam. nov. (Acari: Arrenuroidea) with a review of North American taxa and description of a new subgenus and species of Nudomideopsis Szalay, 1945. Canadian Entomologist, 122, 229 - 252. http: // dx. doi. org / 10.4039 / Ent 122229 - 3" type="journal article" year="1990">Smith, 1990</bibRefCitation>
</taxonomicName>
, pp. 245249, figs. 1822.
</paragraph>
<paragraph id="8BCC36D2FFA2174DFF4C6DE0A8B6F83B" blockId="42.[151,977,1860,2035]" pageId="42" pageNumber="153">
<taxonomicName id="4C734D51FFA2174DFF4C6DE0A80DF81A" authority="Smith &amp; Cook, 1991" authorityName="Smith &amp; Cook" authorityYear="1991" box="[151,529,1921,1943]" class="Arachnida" family="Mideopsidae" genus="Paramideopsis" kingdom="Animalia" order="Trombidiformes" pageId="42" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FFA2174DFF4C6DE0AB33F81A" box="[151,303,1921,1943]" italics="true" pageId="42" pageNumber="153">Paramideopsis</emphasis>
:
<bibRefCitation id="EFE24B23FFA2174DFEE76DE0A80DF81A" author="Smith" box="[316,529,1921,1943]" pageId="42" pageNumber="153" refString="Smith, I. M. &amp; Cook, D. R. (1991) Water mites. Chapter 16. In: Thorp, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. Academic Press, San Diego, CA, pp. 523 - 592. [total page number 911 pp.]" type="book chapter" year="1991">Smith &amp; Cook, 1991</bibRefCitation>
</taxonomicName>
, pp. 551, 577, figs. 16.82, 16.90, 16.16.259.
<taxonomicName id="4C734D51FFA2174DFF4C6DC1ABA8F83B" authority="Cook, 1992" authorityName="Cook" authorityYear="1992" box="[151,436,1952,1974]" class="Arachnida" family="Mideopsidae" genus="Paramideopsis" kingdom="Animalia" order="Trombidiformes" pageId="42" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FFA2174DFF4C6DC1AB33F83B" box="[151,303,1952,1974]" italics="true" pageId="42" pageNumber="153">Paramideopsis</emphasis>
:
<bibRefCitation id="EFE24B23FFA2174DFEE76DC1ABA8F83B" author="Cook" box="[316,436,1952,1974]" pageId="42" pageNumber="153" refString="Cook, D. R. (1992) Water mites (Hydracarina) from driven wells in New Zealand: taxa other than Notoaturinae Besch. Stygologia, 7, 43 - 62." type="journal article" year="1992">Cook, 1992</bibRefCitation>
</taxonomicName>
, pp. 5658, figs. 5257.
</paragraph>
<paragraph id="8BCC36D2FFA2174DFF4C6DDFA9B1F87E" blockId="42.[151,977,1860,2035]" pageId="42" pageNumber="153">
<taxonomicName id="4C734D51FFA2174DFF4C6DDFABEFF859" authority="Smith et al., 2001" authorityName="Smith et al." authorityYear="2001" box="[151,499,1982,2005]" class="Arachnida" family="Mideopsidae" genus="Paramideopsis" kingdom="Animalia" order="Trombidiformes" pageId="42" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FFA2174DFF4C6DDFAB33F859" box="[151,303,1982,2004]" italics="true" pageId="42" pageNumber="153">Paramideopsis</emphasis>
:
<bibRefCitation id="EFE24B23FFA2174DFEE76DDFABEFF859" author="Smith" box="[316,499,1982,2005]" pageId="42" pageNumber="153" refString="Smith, I. M., Cook, D. R. &amp; Smith, B. P. (2001) Water mites and other arachnids. Chapter 16. In: Thorpe, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. 2 nd Edition. Academic Press, Waltham, Massachusetts, pp. 551 - 659. [1056 pp.] http: // dx. doi. org / 10.1016 / B 978 - 012690647 - 9 / 50017 - X" type="book chapter" year="2001">
Smith
<emphasis id="B907EAC0FFA2174DFEA46DA1ABB7F859" box="[383,427,1982,2005]" italics="true" pageId="42" pageNumber="153">et al</emphasis>
., 2001
</bibRefCitation>
</taxonomicName>
, pp. 579, 611, figs. 174, 177, 386.
<taxonomicName id="4C734D51FFA2174DFF4C6DBCABEFF87E" authority="Smith et al., 2010" authorityName="Smith et al." authorityYear="2010" box="[151,499,2013,2035]" class="Arachnida" family="Mideopsidae" genus="Paramideopsis" kingdom="Animalia" order="Trombidiformes" pageId="42" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FFA2174DFF4C6DBCAB33F87E" box="[151,303,2013,2035]" italics="true" pageId="42" pageNumber="153">Paramideopsis</emphasis>
:
<bibRefCitation id="EFE24B23FFA2174DFEE76DBCABEFF87E" author="Smith" box="[316,499,2013,2035]" pageId="42" pageNumber="153" refString="Smith, I. M., Cook, D. R. &amp; Smith, B. P. (2010) Water mites and other arachnids. Chapter 15. In: Thorpe, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. 3 rd Edition. Academic Press, Waltham, Massachusetts, pp. 485 - 586. [total page number: 1021 pp.] http: // dx. doi. org / 10.1016 / B 978 - 0 - 12 - 374855 - 3.00015 - 7" type="book chapter" year="2010">
Smith
<emphasis id="B907EAC0FFA2174DFEA46DBFABB7F87E" box="[383,427,2013,2035]" italics="true" pageId="42" pageNumber="153">et al</emphasis>
., 2010
</bibRefCitation>
</taxonomicName>
, pp. 522, 553, figs. 15.174, 15.177, 15.386.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FFA3174CFF4C6AF6A863FE5D" pageId="43" pageNumber="154" type="diagnosis">
<paragraph id="8BCC36D2FFA3174CFF4C6AF6AECFFF58" blockId="43.[151,1436,151,645]" pageId="43" pageNumber="154">
<emphasis id="B907EAC0FFA3174CFF4C6AF6AB42FF22" bold="true" box="[151,350,151,176]" pageId="43" pageNumber="154">
Diagnosis.
<emphasis id="B907EAC0FFA3174CFEC16AF9AB42FF22" bold="true" box="[282,350,152,175]" italics="true" pageId="43" pageNumber="154">Larva</emphasis>
</emphasis>
(modified from
<bibRefCitation id="EFE24B23FFA3174CFDC46AF6A8A8FF3D" author="Smith" box="[543,692,151,176]" pageId="43" pageNumber="154" refString="Smith, I. M. (1983 b) Description of larvae and adults of Paramideopsis susanae n. gen., n. sp., with remarks on phylogeny and classification of Mideopsidae (Acari: Parasitengona: Arrenuroidea). Canadian Entomologist, 115, 529 - 538. http: // dx. doi. org / 10.4039 / Ent 115529 - 5" type="journal article" year="1983" yearSuffix="b">Smith 1983b</bibRefCitation>
,
<bibRefCitation id="EFE24B23FFA3174CFD196AF9A8E7FF3D" author="Smith" box="[706,763,152,176]" pageId="43" pageNumber="154" refString="Smith, I. M. (1990) Proposal of Nudomideopsidae fam. nov. (Acari: Arrenuroidea) with a review of North American taxa and description of a new subgenus and species of Nudomideopsis Szalay, 1945. Canadian Entomologist, 122, 229 - 252. http: // dx. doi. org / 10.4039 / Ent 122229 - 3" type="journal article" year="1990">1990</bibRefCitation>
): Character states of family
<taxonomicName id="4C734D51FFA3174CFB9D6AF6AF0AFF3D" box="[1094,1302,151,176]" class="Arachnida" family="Nudomideopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acarina" pageId="43" pageNumber="154" phylum="Arthropoda" rank="family">Nudomideopsidae</taxonomicName>
. First coxal plate with setae
<emphasis id="B907EAC0FFA3174CFE976ADDAB74FF59" box="[332,360,188,212]" italics="true" pageId="43" pageNumber="154">1a</emphasis>
long, third coxal plate with transverse muscle attachment scar relatively long.
</paragraph>
<paragraph id="8BCC36D2FFA3174CFF1C6ABEA863FE5D" blockId="43.[151,1436,151,645]" pageId="43" pageNumber="154">
<emphasis id="B907EAC0FFA3174CFF1C6ABEAB0EFF75" bold="true" box="[199,274,223,248]" italics="true" pageId="43" pageNumber="154">Adults</emphasis>
(modified from
<bibRefCitation id="EFE24B23FFA3174CFE0A6ABEA84AFF75" author="Smith" box="[465,598,223,248]" pageId="43" pageNumber="154" refString="Smith, I. M. (1990) Proposal of Nudomideopsidae fam. nov. (Acari: Arrenuroidea) with a review of North American taxa and description of a new subgenus and species of Nudomideopsis Szalay, 1945. Canadian Entomologist, 122, 229 - 252. http: // dx. doi. org / 10.4039 / Ent 122229 - 3" type="journal article" year="1990">Smith 1990</bibRefCitation>
): Character states of family
<taxonomicName id="4C734D51FFA3174CFC7B6ABEAE73FF75" box="[928,1135,223,248]" class="Arachnida" family="Nudomideopsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acarina" pageId="43" pageNumber="154" phylum="Arthropoda" rank="family">Nudomideopsidae</taxonomicName>
. Idiosoma nearly round in shape; convex dorsally. First coxal plate not extending to anterior edge of shield; suture line between third and fourth coxal plate ending medially near posteromedial angle on posterior coxal group so that fourth plate has a short medial edge. Genital field bearing five to seven pairs of acetabula that are free in gonopore in both sexes; fourth coxal plate bearing small projection that partially covers opening for insertion of fourth leg, with suture line at posterior edge moderately distinct. Pedipalp tibia stocky and slightly expanded ventrally; bearing two slender setae ventrally in proximal half of segment.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FFA3174CFF1C6BBDAF63FD95" pageId="43" pageNumber="154" type="materials_examined">
<paragraph id="8BCC36D2FFA3174CFF1C6BBDA8D8FE78" blockId="43.[151,1436,151,645]" box="[199,708,476,501]" pageId="43" pageNumber="154">
<emphasis id="B907EAC0FFA3174CFF1C6BBDAB7EFE78" bold="true" box="[199,354,476,501]" pageId="43" pageNumber="154">
<typeStatus id="54C88870FFA3174CFF1C6BBDAB1EFE78" box="[199,258,476,501]" pageId="43" pageNumber="154">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C734D51FFA3174CFEB26BBDA8D8FE78" authority="Smith." authorityName="Smith." box="[361,708,476,501]" class="Arachnida" family="Mideopsidae" genus="Paramideopsis" kingdom="Animalia" order="Trombidiformes" pageId="43" pageNumber="154" phylum="Arthropoda" rank="species" species="susanae">
<emphasis id="B907EAC0FFA3174CFEB26BBDA86DFE79" box="[361,625,476,501]" italics="true" pageId="43" pageNumber="154">Paramideopsis susanae</emphasis>
Smith.
</taxonomicName>
</paragraph>
<paragraph id="8BCC36D2FFA3174CFF1C6B9EAF63FD95" blockId="43.[151,1436,151,645]" box="[199,1407,511,536]" pageId="43" pageNumber="154">
<emphasis id="B907EAC0FFA3174CFF1C6B9EAB8FFD95" bold="true" box="[199,403,511,536]" pageId="43" pageNumber="154">Species included.</emphasis>
<taxonomicName id="4C734D51FFA3174CFE416B9EA8F0FD95" authority="Smith" authorityName="Smith" box="[410,748,511,536]" class="Arachnida" family="Mideopsidae" genus="Paramideopsis" kingdom="Animalia" order="Trombidiformes" pageId="43" pageNumber="154" phylum="Arthropoda" rank="species" species="susanae">
<emphasis id="B907EAC0FFA3174CFE416B9EA8BFFD95" box="[410,675,511,536]" italics="true" pageId="43" pageNumber="154">Paramideopsis susanae</emphasis>
Smith
</taxonomicName>
(western North
<collectingCountry id="F3647642FFA3174CFC786B9EAE14FD95" box="[931,1032,511,536]" name="United States of America" pageId="43" pageNumber="154">America</collectingCountry>
),
<taxonomicName id="4C734D51FFA3174CFBC06860AED4FD95" authority="Cook" authorityName="Cook" box="[1051,1224,511,536]" class="Arachnida" family="Mideopsidae" genus="Paramideopsis" kingdom="Animalia" order="Trombidiformes" pageId="43" pageNumber="154" phylum="Arthropoda" rank="species" species="kyphus">
<emphasis id="B907EAC0FFA3174CFBC06860AE9FFD95" box="[1051,1155,511,536]" italics="true" pageId="43" pageNumber="154">P. kyphus</emphasis>
Cook
</taxonomicName>
(
<collectingCountry id="F3647642FFA3174CFB0C6861AF6FFD95" box="[1239,1395,511,536]" name="New Zealand" pageId="43" pageNumber="154">New Zealand</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C3696559FFA3174CFF1C6845A95AFDB0" box="[199,838,548,573]" pageId="43" pageNumber="154" type="distribution">
<paragraph id="8BCC36D2FFA3174CFF1C6845A95AFDB0" blockId="43.[151,1436,151,645]" box="[199,838,548,573]" pageId="43" pageNumber="154">
<emphasis id="B907EAC0FFA3174CFF1C6845AB7CFDB0" bold="true" box="[199,352,548,573]" pageId="43" pageNumber="154">Distribution.</emphasis>
Western North
<collectingCountry id="F3647642FFA3174CFDCA6845A86DFDB0" box="[529,625,548,573]" name="United States of America" pageId="43" pageNumber="154">America</collectingCountry>
and
<collectingCountry id="F3647642FFA3174CFD736844A95EFDB0" box="[680,834,548,573]" name="New Zealand" pageId="43" pageNumber="154">New Zealand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FFA3174CFF1C6826AF4FFD08" pageId="43" pageNumber="154" type="discussion">
<paragraph id="8BCC36D2FFA3174CFF1C6826AF4FFD08" blockId="43.[151,1436,151,645]" pageId="43" pageNumber="154">
<emphasis id="B907EAC0FFA3174CFF1C6826AB50FDED" bold="true" box="[199,332,583,608]" pageId="43" pageNumber="154">Discussion.</emphasis>
<bibRefCitation id="EFE24B23FFA3174CFE886826ABE9FDED" author="Smith" box="[339,501,583,608]" pageId="43" pageNumber="154" refString="Smith, I. M. (1983 b) Description of larvae and adults of Paramideopsis susanae n. gen., n. sp., with remarks on phylogeny and classification of Mideopsidae (Acari: Parasitengona: Arrenuroidea). Canadian Entomologist, 115, 529 - 538. http: // dx. doi. org / 10.4039 / Ent 115529 - 5" type="journal article" year="1983" yearSuffix="b">Smith (1983b)</bibRefCitation>
proposed
<taxonomicName id="4C734D51FFA3174CFDB26826A90DFDED" box="[617,785,583,608]" class="Arachnida" family="Mideopsidae" genus="Paramideopsis" kingdom="Animalia" order="Trombidiformes" pageId="43" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FFA3174CFDB26826A90DFDED" box="[617,785,583,608]" italics="true" pageId="43" pageNumber="154">Paramideopsis</emphasis>
</taxonomicName>
as a distinct genus. This treatment was followed by
<bibRefCitation id="EFE24B23FFA3174CFA826826AAFFFD09" author="Smith" pageId="43" pageNumber="154" refString="Smith, I. M. (1990) Proposal of Nudomideopsidae fam. nov. (Acari: Arrenuroidea) with a review of North American taxa and description of a new subgenus and species of Nudomideopsis Szalay, 1945. Canadian Entomologist, 122, 229 - 252. http: // dx. doi. org / 10.4039 / Ent 122229 - 3" type="journal article" year="1990">Smith (1990)</bibRefCitation>
,
<bibRefCitation id="EFE24B23FFA3174CFF34680DAB9CFD09" author="Cook" box="[239,384,620,645]" pageId="43" pageNumber="154" refString="Cook, D. R. (1992) Water mites (Hydracarina) from driven wells in New Zealand: taxa other than Notoaturinae Besch. Stygologia, 7, 43 - 62." type="journal article" year="1992">Cook (1992)</bibRefCitation>
,
<bibRefCitation id="EFE24B23FFA3174CFE57680DA866FD09" author="Smith" box="[396,634,620,645]" pageId="43" pageNumber="154" refString="Smith, I. M. &amp; Cook, D. R. (1991) Water mites. Chapter 16. In: Thorp, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. Academic Press, San Diego, CA, pp. 523 - 592. [total page number 911 pp.]" type="book chapter" year="1991">Smith &amp; Cook (1991</bibRefCitation>
,
<bibRefCitation id="EFE24B23FFA3174CFD5E680DA8A2FD09" author="Smith" box="[645,702,620,644]" pageId="43" pageNumber="154" refString="Smith, I. M. &amp; Cook, D. R. (1999) An assessment of global distribution patterns in water mites (Acari: Hydrachnida). In: Needham, G. R., Mitchell, R. D., Horn, D. J. &amp; Welbourn, W. C. (Eds.), Acarology IX. Vol. 2. Symposia. Ohio Biological Survey, Columbus, pp. 109 - 124." type="book" year="1999">1999</bibRefCitation>
) and
<bibRefCitation id="EFE24B23FFA3174CFD26680DA9D6FD09" author="Smith" box="[765,970,620,645]" pageId="43" pageNumber="154" refString="Smith, I. M., Cook, D. R. &amp; Smith, B. P. (2001) Water mites and other arachnids. Chapter 16. In: Thorpe, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. 2 nd Edition. Academic Press, Waltham, Massachusetts, pp. 551 - 659. [1056 pp.] http: // dx. doi. org / 10.1016 / B 978 - 012690647 - 9 / 50017 - X" type="book chapter" year="2001">
Smith
<emphasis id="B907EAC0FFA3174CFC9C680CA964FD08" box="[839,888,620,645]" italics="true" pageId="43" pageNumber="154">et al</emphasis>
. (2001
</bibRefCitation>
, 2010), and is followed by us here.
</paragraph>
</subSubSection>
</treatment>
</document>