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<document id="5959325D4D92CA1434538F58ABE5123C" ID-DOI="10.5281/zenodo.10881992" ID-ISSN="0936-9902" ID-Zenodo-Dep="10881992" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1711474576113" checkinUser="juliana" docAuthor="Weitzman, Stanley H. &amp; Malabarba, Luiz R." docDate="1999" docId="03DA87B22A11FFCDFC47FBBE4AF3FA20" docLanguage="en" docName="IchthyolExplorFres.10.1.1-43.pdf.imf" docOrigin="Ichthyol. Explor. Freshwaters 10 (1)" docTitle="Spintherobolus leptoura Weitzman &amp; Malabarba 1999, new species" docType="treatment" docUuid="0F30D6DE-EB03-42AC-8CA5-F13DF31DD414" docUuidSource="ZooBank" docVersion="3" lastPageNumber="39" masterDocId="FFE3FFCA2A33FFEBFFB0FF824B09FF9B" masterDocTitle="Systematics of Spintherobolus (Teleostei: Characidae: Cheirodontinae) from eastern Brazil" masterLastPageNumber="43" masterPageNumber="1" pageNumber="35" updateTime="1711475764197" updateUser="ExternalLinkService">
<mods:mods id="297615C1FEF3222EAD6B85B13865A85A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="215BF53F01329DBB380440E7702F524F">
<mods:title id="B13007E03C10BFA1C8BDE907BCF9C435">Systematics of Spintherobolus (Teleostei: Characidae: Cheirodontinae) from eastern Brazil</mods:title>
</mods:titleInfo>
<mods:name id="BA28A02EA77F98F8F287EFB41BA966C6" type="personal">
<mods:role id="4DE54F157C8BEE6461BE37743AFC1E85">
<mods:roleTerm id="F999A288E566B2CAAC6B21CC22A2592D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B5D08AEADBD4C290EDC2F5EC843AA42B">Weitzman, Stanley H.</mods:namePart>
<mods:affiliation id="0010098FCB7653A49328DCC2E1C3EC09">Division of Fishes, Department of Vertebrate Zoology, MRC- 159, Smithsonian Institution, Washington D. C. 20560, USA.</mods:affiliation>
</mods:name>
<mods:name id="57CE09D4FBC3FFFF795BA2AFC771CA19" type="personal">
<mods:role id="54632DA3BB97A974F8918A3C70F8F2C6">
<mods:roleTerm id="CDEDE568C6C0B6DDC68EBE7854C6EC5C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1F5A62B763E1540AA61A5EF839B3B32B">Malabarba, Luiz R.</mods:namePart>
<mods:affiliation id="5C06D445478E6F0FB357B293AE22D54A">Laboratório de Ictiologia, Museu de Ciências e Tecnologia, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga, 6681, CEP 90619 - 900, Porto Alegre, RS, Brazil, and Departamento de Zoologia, IB, Universidade Federal do Rio Grande do Sul, Porto Alegre, Brazil.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6FA2E0E95F9626C0BCD6C945F1A0DD74">text</mods:typeOfResource>
<mods:relatedItem id="835686AF47B63F78816FB8F4A47299B6" type="host">
<mods:titleInfo id="AF4D5A604F46FC8BDB3EABEADAB0E7EB">
<mods:title id="F421A2DFA59A6DFED84F2B3D82C7373F">Ichthyol. Explor. Freshwaters</mods:title>
</mods:titleInfo>
<mods:part id="B3D4F3547913DB4F81C06A8A6993DB2A">
<mods:date id="6652B607932F757A9A84DDB18861A1B3">1999</mods:date>
<mods:detail id="119C914D4213E38FEF050C73B38EDBDD" type="pubDate">
<mods:number id="2225AF0C2E4F8E69E73A6B2732962C85">1999-01-01</mods:number>
</mods:detail>
<mods:detail id="9BFFB69411E2C973ED0CFAE744B9FF29" type="volume">
<mods:number id="41D3B7EF23C0177EE11396E817A27D55">10</mods:number>
</mods:detail>
<mods:detail id="3ADB5ED97153D41B47FD1FDB3FA0369F" type="issue">
<mods:number id="F54082F44DCF7AFE133D298A535F748F">1</mods:number>
</mods:detail>
<mods:extent id="7EA2920E753EA04DE7C26A1FE74860C0" unit="page">
<mods:start id="50AD606D7F9C6DF98DD5EDAFC6D5B91E">1</mods:start>
<mods:end id="9BF0A279C67D2370C569FC07F498CB1B">43</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6F73B24BEDFEA650127DB1CEABB9D42E">journal article</mods:classification>
<mods:identifier id="B7A74AEFD806ABE4354489815827B2FE" type="DOI">10.5281/zenodo.10881992</mods:identifier>
<mods:identifier id="F3BE9255E7166496EA4C73DA3CCB0A0B" type="ISSN">0936-9902</mods:identifier>
<mods:identifier id="AC8722501C2CDBA43ED1AA9F496B91B9" type="Zenodo-Dep">10881992</mods:identifier>
<mods:identifier id="2C4AC019C97E6EBA3DEE1BF174195D0A" type="ZooBank">DB264AEA-C4EC-48B7-BE35-56761CA13225</mods:identifier>
</mods:mods>
<treatment id="03DA87B22A11FFCDFC47FBBE4AF3FA20" ID-DOI="http://doi.org/10.5281/zenodo.10881819" ID-Zenodo-Dep="10881819" LSID="urn:lsid:zoobank.org:act:0F30D6DE-EB03-42AC-8CA5-F13DF31DD414" httpUri="http://treatment.plazi.org/id/03DA87B22A11FFCDFC47FBBE4AF3FA20" lastPageId="38" lastPageNumber="39" pageId="34" pageNumber="35">
<subSubSection id="C369652F2A11FFC9FC47FBBE4D7CFBF9" box="[1015,1653,1084,1122]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="8BCC36A42A11FFC9FC47FBBE4D7CFBF9" blockId="34.[1015,1653,1084,1168]" box="[1015,1653,1084,1122]" pageId="34" pageNumber="35">
<heading id="D08481C82A11FFC9FC47FBBE4D7CFBF9" box="[1015,1653,1084,1122]" centered="true" fontSize="9" level="4" pageId="34" pageNumber="35" reason="4">
<taxonomicName id="4C734D272A11FFC9FC47FBBE4E98FBF9" ID-CoL="4Z63R" authority="Weitzman &amp; Malabarba, 1999" authorityName="Weitzman &amp; Malabarba" authorityYear="1999" box="[1015,1425,1084,1122]" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="leptoura" status="sp. nov.">
<emphasis id="B907EAB62A11FFC9FC47FBBE4FFBFBF9" box="[1015,1266,1084,1122]" italics="true" pageId="34" pageNumber="35">Spintherobolus</emphasis>
<emphasis id="B907EAB62A11FFC9FAB4FBBE4E9EFBF9" box="[1284,1431,1084,1122]" italics="true" pageId="34" pageNumber="35">leptoura,</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23457CD2A11FFC9FA18FBBE4D7CFBF9" box="[1448,1653,1084,1122]" pageId="34" pageNumber="35" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C369652F2A11FFC9FB66FBE84E95FB0B" box="[1238,1436,1130,1168]" pageId="34" pageNumber="35" type="description">
<paragraph id="8BCC36A42A11FFC9FB66FBE84E95FB0B" blockId="34.[1015,1653,1084,1168]" box="[1238,1436,1130,1168]" pageId="34" pageNumber="35">
<heading id="D08481C82A11FFC9FB66FBE84E95FB0B" box="[1238,1436,1130,1168]" centered="true" fontSize="9" level="4" pageId="34" pageNumber="35" reason="1">
(
<figureCitation id="13482A212A11FFC9FB50FBE84E99FB0B" box="[1248,1424,1130,1168]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="35.[207,272,2036,2082]" captionStartId-1="35.[205,267,2144,2188]" captionTargetBox-0="[210,2446,290,1130]" captionTargetBox-1="[206,2443,1144,1985]" captionTargetPageId-0="35" captionTargetPageId-1="35" captionText-0="Fig. 36. Spintherobolus leptoura, MZUSP 41854, holotype, adult male 19.1 mm SL, Brazil, São Paulo, fazenda Dalila, Registro." captionText-1="Fig. 37. Spintherobolus leptoura, MZUSP 41854, paratype, adult female 25.0 mm SL, Brazil, São Paulo, fazenda Dalila, Registro." figureDoi-0="http://doi.org/10.5281/zenodo.10882080" figureDoi-1="http://doi.org/10.5281/zenodo.10882082" httpUri-0="https://zenodo.org/record/10882080/files/figure.png" httpUri-1="https://zenodo.org/record/10882082/files/figure.png" pageId="34" pageNumber="35">Figs. 36-37</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C369652F2A11FFC9FC15FB414D07F934" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8BCC36A42A11FFC9FC15FB414DC9FADF" blockId="34.[931,1736,1219,1348]" pageId="34" pageNumber="35">
<materialsCitation id="3B1B3CF92A11FFC9FC15FB414DB1FADF" collectingDate="1990-02-16" collectionCode="MZUSP" collectorName="M. Damato &amp; O. Oyakawa" country="Brazil" location="rio Quilombo" municipality="Registro" pageId="34" pageNumber="35" specimenCode="MZUSP 41854" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="holotype">
<typeStatus id="54C888062A11FFC9FC15FB414F4EFB73" box="[933,1095,1219,1256]" pageId="34" pageNumber="35">Holotype</typeStatus>
.
<specimenCode id="DBD59EDF2A11FFC9FBD1FB414E46FB73" box="[1121,1359,1219,1256]" pageId="34" pageNumber="35">MZUSP 41854</specimenCode>
,
<specimenCount id="9D75FD2D2A11FFC9FAD2FB414EBAFB73" box="[1378,1459,1219,1256]" pageId="34" pageNumber="35" type="male">male</specimenCount>
,
<quantity id="4C8B9B412A11FFC9FA78FB414D59FB73" box="[1480,1616,1219,1256]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="34" pageNumber="35" unit="mm" value="19.1">19.1 mm</quantity>
SL;
<location id="8EAC607F2A11FFC9F92CFB414F44FA8D" LSID="urn:lsid:plazi:treatment:03DA87B22A11FFCDFC47FBBE4AF3FA20:8EAC607F2A11FFC9F92CFB414F44FA8D" country="Brazil" municipality="Registro" name="rio Quilombo" pageId="34" pageNumber="35" stateProvince="Sao Paulo">rio Quilombo</location>
,
<location id="8EAC607F2A11FFC9FBD6FB724E54FA8D" LSID="urn:lsid:plazi:treatment:03DA87B22A11FFCDFC47FBBE4AF3FA20:8EAC607F2A11FFC9FBD6FB724E54FA8D" box="[1126,1373,1264,1302]" country="Brazil" municipality="Registro" name="fazenda Dalila" pageId="34" pageNumber="35" stateProvince="Sao Paulo">fazenda Dalila</location>
,
<collectingMunicipality id="6BA8ACDE2A11FFC9FAC5FB724EF6FA8D" box="[1397,1535,1264,1302]" pageId="34" pageNumber="35">Registro</collectingMunicipality>
,
<collectingRegion id="49B7F8462A11FFC9F9A8FB724DB7FA8D" box="[1560,1726,1264,1302]" country="Brazil" name="Sao Paulo" pageId="34" pageNumber="35">São Paulo</collectingRegion>
,
<collectingCountry id="F36476342A11FFC9FC14FA9C4F0EFADF" box="[932,1031,1310,1348]" name="Brazil" pageId="34" pageNumber="35" value="Brazil">Brazil</collectingCountry>
;
<date id="FFCD10642A11FFC9FBACFA9C4FEBFADF" box="[1052,1250,1310,1348]" pageId="34" pageNumber="35" value="1990-02-16">
<collectingDate id="EF89E98C2A11FFC9FBACFA9C4FEBFADF" box="[1052,1250,1310,1348]" pageId="34" pageNumber="35" value="1990-02-16">16 Feb 1990</collectingDate>
</date>
,
<collectorName id="268653722A11FFC9FB47FA9C4EB8FADF" box="[1271,1457,1310,1348]" pageId="34" pageNumber="35">M. Damato</collectorName>
&amp;
<collectorName id="268653722A11FFC9FA5AFA9C4DB1FADF" box="[1514,1720,1310,1348]" pageId="34" pageNumber="35">O. Oyakawa</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BCC36A42A11FFC9FC14FAF84D07F934" blockId="34.[930,1734,1402,1711]" pageId="34" pageNumber="35">
<materialsCitation id="3B1B3CF92A11FFC9FC14FAF848D9F9BE" collectingDate="1990-02-16" collectionCode="MZUSP" collectorName="M. Damato &amp; O. Oyakawa" country="Brazil" location="rio Quilombo" municipality="Registro" pageId="34" pageNumber="35" specimenCode="MZUSP 51023" specimenCount="11" specimenCount-female="1" specimenCount-male="3" stateProvince="Sao Paulo" typeStatus="paratype">
<typeStatus id="54C888062A11FFC9FC14FAF84F45FA3B" box="[932,1100,1402,1440]" pageId="34" pageNumber="35">Paratypes</typeStatus>
.
<specimenCode id="DBD59EDF2A11FFC9FBD6FAF84E5AFA3B" box="[1126,1363,1402,1440]" pageId="34" pageNumber="35">MZUSP 51023</specimenCode>
,
<specimenCount id="9D75FD2D2A11FFC9FAD3FAF84ED7FA3B" box="[1379,1502,1402,1440]" count="2" pageId="34" pageNumber="35" type="male">2 males</specimenCount>
,
<quantity id="4C8B9B412A11FFC9FA42FAF84DCFFA3B" box="[1522,1734,1402,1440]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" metricValueMax="1.96" metricValueMin="1.72" pageId="34" pageNumber="35" unit="mm" value="18.4" valueMax="19.6" valueMin="17.2">17.2-19.6 mm</quantity>
SL,
<specimenCount id="9D75FD2D2A11FFC9FC5BFA2548F2FA50" box="[1003,1019,1447,1483]" pageId="34" pageNumber="35">7</specimenCount>
immature - females,
<quantity id="4C8B9B412A11FFC9FA37FA254D55FA50" box="[1415,1628,1447,1483]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" metricValueMax="2.7199999999999998" metricValueMin="1.6199999999999999" pageId="34" pageNumber="35" unit="mm" value="21.7" valueMax="27.2" valueMin="16.2">16.2-27.2 mm</quantity>
SL,
<specimenCount id="9D75FD2D2A11FFC9F908FA2548FCFA63" count="1" pageId="34" pageNumber="35" type="male">1 male</specimenCount>
, c&amp;s,
<quantity id="4C8B9B412A11FFC9FBD4FA564FE4FA63" box="[1124,1261,1492,1528]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" pageId="34" pageNumber="35" unit="mm" value="19.9">19.9 mm</quantity>
SL,
<specimenCount id="9D75FD2D2A11FFC9FAF2FA564ED8FA63" box="[1346,1489,1492,1528]" count="1" pageId="34" pageNumber="35" type="female">1 female</specimenCount>
, c&amp;s,
<quantity id="4C8B9B412A11FFC9F98DFA564DCFFA63" box="[1597,1734,1492,1528]" metricMagnitude="-2" metricUnit="m" metricValue="2.52" pageId="34" pageNumber="35" unit="mm" value="25.2">25.2 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="3B1B3CF92A11FFC9FC52F9834EC1F9C9" collectingDate="1990-02-16" collectionCode="USNM" collectorName="M. Damato &amp; O. Oyakawa" country="Brazil" location="rio Quilombo" municipality="Registro" pageId="34" pageNumber="35" specimenCode="USNM 346386" specimenCount="3" specimenCount-immature="2" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="holotype">
<specimenCode id="DBD59EDF2A11FFC9FC52F9834FDDF9BE" box="[994,1236,1537,1573]" pageId="34" pageNumber="35">USNM 346386</specimenCode>
,
<specimenCount id="9D75FD2D2A11FFC9FB58F9834E5CF9BE" box="[1256,1365,1537,1573]" count="1" pageId="34" pageNumber="35" type="male">1 male</specimenCount>
,
<quantity id="4C8B9B412A11FFC9FAD9F9834EFDF9BE" box="[1385,1524,1537,1573]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="34" pageNumber="35" unit="mm" value="21.8">21.8 mm</quantity>
SL,
<specimenCount id="9D75FD2D2A11FFC9F9F0F98348EAF9C9" count="2" pageId="34" pageNumber="35" type="immature">2 immature</specimenCount>
- females, 16.2-25.0 mm SL
</materialsCitation>
;
<materialsCitation id="3B1B3CF92A11FFC9FA6EF9AD4D01F934" collectingDate="1990-02-16" collectionCode="MCP" collectorName="M. Damato &amp; O. Oyakawa" country="Brazil" location="rio Quilombo" municipality="Registro" pageId="34" pageNumber="35" specimenCode="MCP 19254" specimenCount="4" specimenCount-female="1" specimenCount-immature="2" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="holotype">
<specimenCode id="DBD59EDF2A11FFC9FA6EF9AD4D96F9C9" box="[1502,1695,1583,1618]" pageId="34" pageNumber="35">MCP 19254</specimenCode>
,
<specimenCount id="9D75FD2D2A11FFC9F907F9AD48FDF9E4" count="1" pageId="34" pageNumber="35" type="male">1 male</specimenCount>
,
<quantity id="4C8B9B412A11FFC9FBA1F9DE4F91F9E4" box="[1041,1176,1628,1663]" metricMagnitude="-2" metricUnit="m" metricValue="1.89" pageId="34" pageNumber="35" unit="mm" value="18.9">18.9 mm</quantity>
SL;
<specimenCount id="9D75FD2D2A11FFC9FB5EF9DE4EB8F9E4" box="[1262,1457,1628,1663]" count="2" pageId="34" pageNumber="35" type="immature">2 immature</specimenCount>
-
<specimenCount id="9D75FD2D2A11FFC9FA56F9DE4D68F9E4" box="[1510,1633,1628,1663]" pageId="34" pageNumber="35" type="female">females</specimenCount>
,
<quantity id="4C8B9B412A11FFC9F9CDF9DE4F26F934" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.54" metricValueMin="1.66" pageId="34" pageNumber="35" unit="mm" value="21.0" valueMax="25.4" valueMin="16.6">16.6- 25.4 mm</quantity>
SL; collected with holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C369652F2A11FFC9FC14F9614E02F761" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="8BCC36A42A11FFC9FC14F9614E02F761" blockId="34.[929,1735,1763,2298]" pageId="34" pageNumber="35">
Diagnosis. Most elongate and slender species, adult males differing from other small species by caudal peduncle depth and body depth (
<figureCitation id="13482A212A11FFC9F9F4F8B94DB4F8FF" box="[1604,1725,1851,1892]" captionStart="Fig" captionStartId="18.[949,999,879,915]" captionTargetBox="[948,1754,211,843]" captionTargetPageId="18" captionText="Fig. 24. Graph of body depth as a function of standard length for Spintherobolus leptoura (black symbols) and S. ankoseion (empty symbols). Axes are logarithmic. Note that the 95 % confidence intervals in lengths of about 21 to 28 mm SL do not overlap with the means of the of the compared species indicating a significant difference in these size ranges. These growth pattern differences are used to separate these species in the key and text." figureDoi="http://doi.org/10.5281/zenodo.10882050" httpUri="https://zenodo.org/record/10882050/files/figure.png" pageId="34" pageNumber="35">Figs. 24</figureCitation>
,
<figureCitation id="13482A212A11FFC9FC12F8EA48CFF80A" box="[930,966,1896,1937]" captionStart="Fig" captionStartId="18.[948,998,1937,1973]" captionTargetBox="[947,1750,1260,1892]" captionTargetPageId="18" captionText="Fig. 25. Graph of caudal peduncle depth as a function of standard length for Spintherobolus leptoura (black symbols) and S. ankoseion (empty symbols). Axes are logarithmic. Note that the 95 % confidence intervals in lengths of about 21 to 27 mm SL do not overlap with the means of the compared species indicating a significant difference in these size ranges. These growth pattern differences are used to separate these species in the key and text." figureDoi="http://doi.org/10.5281/zenodo.10882052" httpUri="https://zenodo.org/record/10882052/files/figure.png" pageId="34" pageNumber="35">25</figureCitation>
,
<figureCitation id="13482A212A11FFC9FC51F8EA4F0CF80A" box="[993,1029,1896,1937]" captionStart="Fig" captionStartId="36.[1226,1294,1149,1196]" captionTargetBox="[1225,2300,308,1096]" captionTargetPageId="36" captionText="Fig. 38. Graph of caudal peduncle depth as a function of standard length of Spintherobolus leptoura (black symbols) and S. broccae (empty symbols). Axes are logarithmic. Note that the 95 % confidence intervals in lengths of about 18 to 24 mm SL do not overlap with the means of these of the compared species indicating a significant difference in these size ranges. Comparison of these growth patterns are discussed under the diagnosis of S. leptoura." figureDoi="http://doi.org/10.5281/zenodo.10882084" httpUri="https://zenodo.org/record/10882084/files/figure.png" pageId="34" pageNumber="35">38</figureCitation>
). Body depth in
<taxonomicName id="4C734D272A11FFC9FA87F8EA4EDFF80A" authorityName="Weitzman &amp; Malabarba" authorityYear="1999" box="[1335,1494,1896,1937]" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="leptoura">
<emphasis id="B907EAB62A11FFC9FA87F8EA4E59F80A" box="[1335,1360,1896,1937]" italics="true" pageId="34" pageNumber="35">S.</emphasis>
<emphasis id="B907EAB62A11FFC9FAEEF8EA4EDFF80A" box="[1374,1494,1896,1937]" italics="true" pageId="34" pageNumber="35">leptoura</emphasis>
</taxonomicName>
26.6-29.7, x = 28.3 % SL, versus 29.3-37.9, x =
<quantity id="4C8B9B412A11FFC9FA2EF81A4D0FF827" box="[1438,1542,1944,1980]" metricMagnitude="-1" metricUnit="m" metricValue="8.6868" pageId="34" pageNumber="35" unit="in" value="34.2">34.2 in</quantity>
<taxonomicName id="4C734D272A11FFC9F9A1F81A4DCDF827" box="[1553,1732,1944,1980]" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="ankoseion">
<emphasis id="B907EAB62A11FFC9F9A1F81A4DCDF827" box="[1553,1732,1944,1980]" italics="true" pageId="34" pageNumber="35">S. ankoseion</emphasis>
</taxonomicName>
(
<figureCitation id="13482A212A11FFC9FC1FF8474F13F877" box="[943,1050,1989,2028]" captionStart="Fig" captionStartId="18.[949,999,879,915]" captionTargetBox="[948,1754,211,843]" captionTargetPageId="18" captionText="Fig. 24. Graph of body depth as a function of standard length for Spintherobolus leptoura (black symbols) and S. ankoseion (empty symbols). Axes are logarithmic. Note that the 95 % confidence intervals in lengths of about 21 to 28 mm SL do not overlap with the means of the of the compared species indicating a significant difference in these size ranges. These growth pattern differences are used to separate these species in the key and text." figureDoi="http://doi.org/10.5281/zenodo.10882050" httpUri="https://zenodo.org/record/10882050/files/figure.png" pageId="34" pageNumber="35">Fig. 24</figureCitation>
), and 29.2-32.8, x =
<quantity id="4C8B9B412A11FFC9FAD0F8474EC0F877" box="[1376,1481,1989,2028]" metricMagnitude="-1" metricUnit="m" metricValue="7.747" pageId="34" pageNumber="35" unit="in" value="30.5">30.5 in</quantity>
<taxonomicName id="4C734D272A11FFC9FA64F8474D6FF877" box="[1492,1638,1989,2028]" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="broccae">
<emphasis id="B907EAB62A11FFC9FA64F8474EE4F877" box="[1492,1517,1989,2028]" italics="true" pageId="34" pageNumber="35">S.</emphasis>
<emphasis id="B907EAB62A11FFC9FA4AF8474D64F877" box="[1530,1645,1989,2028]" italics="true" pageId="34" pageNumber="35">broccae.</emphasis>
</taxonomicName>
Caudal peduncle depth in
<taxonomicName id="4C734D272A11FFC9FA84F8734EDCF782" authorityName="Weitzman &amp; Malabarba" authorityYear="1999" box="[1332,1493,2033,2073]" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="leptoura">
<emphasis id="B907EAB62A11FFC9FA84F8734EDCF782" box="[1332,1493,2033,2073]" italics="true" pageId="34" pageNumber="35">S. leptoura</emphasis>
</taxonomicName>
11.0-13.1, x = 12.3 % SL, versus 12.8-15.6, x =
<quantity id="4C8B9B412A11FFC9FA61F7A24D37F7D8" box="[1489,1598,2080,2115]" metricMagnitude="-1" metricUnit="m" metricValue="3.7592" pageId="34" pageNumber="35" unit="in" value="14.8">14.8 in</quantity>
<emphasis id="B907EAB62A11FFC9F9E0F7A248FCF7EB" italics="true" pageId="34" pageNumber="35">
<taxonomicName id="4C734D272A11FFC9F9E0F7A248E6F7EB" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="ankoseion">S. ankoseion</taxonomicName>
,
</emphasis>
and 12.6-15.1, x =
<quantity id="4C8B9B412A11FFC9FAEDF7CE4EC7F7EB" box="[1373,1486,2124,2160]" metricMagnitude="-1" metricUnit="m" metricValue="3.5052" pageId="34" pageNumber="35" unit="in" value="13.8">13.8 in</quantity>
<taxonomicName id="4C734D272A11FFC9FA53F7CE4D7FF7EB" box="[1507,1654,2124,2160]" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="broccae">
<emphasis id="B907EAB62A11FFC9FA53F7CE4EF5F7EB" box="[1507,1532,2124,2160]" italics="true" pageId="34" pageNumber="35">S.</emphasis>
<emphasis id="B907EAB62A11FFC9F9BAF7CE4D74F7EB" box="[1546,1661,2124,2160]" italics="true" pageId="34" pageNumber="35">broccae.</emphasis>
</taxonomicName>
Although these ratios overlap in juveniles, they do not in adult males and females as demonstrated by
<figureCitation id="13482A212A11FFC9FC69F7514F8CF761" box="[985,1157,2259,2298]" captionStart="Fig" captionStartId="18.[949,999,879,915]" captionTargetBox="[948,1754,211,843]" captionTargetPageId="18" captionText="Fig. 24. Graph of body depth as a function of standard length for Spintherobolus leptoura (black symbols) and S. ankoseion (empty symbols). Axes are logarithmic. Note that the 95 % confidence intervals in lengths of about 21 to 28 mm SL do not overlap with the means of the of the compared species indicating a significant difference in these size ranges. These growth pattern differences are used to separate these species in the key and text." figureDoi="http://doi.org/10.5281/zenodo.10882050" httpUri="https://zenodo.org/record/10882050/files/figure.png" pageId="34" pageNumber="35">Figures 24</figureCitation>
and
<figureCitation id="13482A212A11FFC9FB50F7514E0AF761" box="[1248,1283,2259,2298]" captionStart="Fig" captionStartId="18.[948,998,1937,1973]" captionTargetBox="[947,1750,1260,1892]" captionTargetPageId="18" captionText="Fig. 25. Graph of caudal peduncle depth as a function of standard length for Spintherobolus leptoura (black symbols) and S. ankoseion (empty symbols). Axes are logarithmic. Note that the 95 % confidence intervals in lengths of about 21 to 27 mm SL do not overlap with the means of the compared species indicating a significant difference in these size ranges. These growth pattern differences are used to separate these species in the key and text." figureDoi="http://doi.org/10.5281/zenodo.10882052" httpUri="https://zenodo.org/record/10882052/files/figure.png" pageId="34" pageNumber="35">25</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C369652F2A11FFCDFC13F6AE4859FBBF" lastPageId="38" lastPageNumber="39" pageId="34" pageNumber="35" type="description">
<paragraph id="8BCC36A42A11FFC8FC13F6AE4876F3D4" blockId="34.[930,1735,2348,2615]" lastBlockId="35.[201,1280,2377,3394]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
Description. Morphometric data are summarized in
<tableCitation id="C6F1031F2A11FFC9FB9BF6DE4FA8F619" box="[1067,1185,2396,2434]" captionStart="Table 5" captionStartId="36.[72,183,2072,2118]" captionTargetBox="[71,2289,2244,3457]" captionTargetPageId="36" captionText="Table 5. Morphometries of Spintherobolus leptoura. Table includes measurements of all type specimens. Values for the holotype are given separately." httpUri="http://table.plazi.org/id/DF0C662C2A17FFCFFFF8F79A49E6F7E7" pageId="34" pageNumber="35" tableUuid="DF0C662C2A17FFCFFFF8F79A49E6F7E7">Table 5</tableCitation>
. Body compressed and relatively elongate. Predorsal body profile slightly convex with a slight concavity at nape. Body profile along dorsal-fin base posteroventrally inclined becoming straight or slightly concave between dorsal and caudal fins in females and nearly straight to slightly convex in males. Ventral profile from tip of mandible to pelvic-fin origin convex and slightly concave between pelvic- and anal-fin origins. Entire body profile along anal-fin base concave in females. This same profile convex in anterior anal-fin lobe of males. Remaining posterior anal-fin body profile nearly straight and continuous with straight or nearly straight caudal peduncle profile. Ventral profile of caudal peduncle somewhat concave in females, but straight and slightly serrated in males because of prominent ventral procurrent caudal-fin rays.
</paragraph>
<caption id="DF0C662C2A10FFC8FF7FF8764A7EF7C3" ID-DOI="http://doi.org/10.5281/zenodo.10882080" ID-Zenodo-Dep="10882080" httpUri="https://zenodo.org/record/10882080/files/figure.png" pageId="35" pageNumber="36" startId="35.[207,272,2036,2082]" targetBox="[210,2446,290,1130]" targetPageId="35" targetType="figure">
<paragraph id="8BCC36A42A10FFC8FF7FF8764A7EF7C3" blockId="35.[205,2437,2036,2136]" pageId="35" pageNumber="36">
Fig. 36.
<taxonomicName id="4C734D272A10FFC8FEC1F876482AF7B9" authority=", MZUSP" authorityName="MZUSP" box="[369,803,2036,2082]" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="leptoura">
<emphasis id="B907EAB62A10FFC8FEC1F876482AF7B9" box="[369,803,2036,2082]" italics="true" pageId="35" pageNumber="36">Spintherobolus leptoura,</emphasis>
</taxonomicName>
MZUSP 41854, holotype, adult male 19.1 mm SL, Brazil, São Paulo, fazenda Dalila, Registro.
</paragraph>
</caption>
<caption id="DF0C662C2A10FFC8FF7DF7E2490EF75A" ID-DOI="http://doi.org/10.5281/zenodo.10882082" ID-Zenodo-Dep="10882082" httpUri="https://zenodo.org/record/10882082/files/figure.png" pageId="35" pageNumber="36" startId="35.[205,267,2144,2188]" targetBox="[206,2443,1144,1985]" targetPageId="35">
<paragraph id="8BCC36A42A10FFC8FF7DF7E2490EF75A" blockId="35.[205,2438,2144,2241]" pageId="35" pageNumber="36">
Fig. 37.
<taxonomicName id="4C734D272A10FFC8FEC9F7E24838F717" authority=", MZUSP" authorityName="MZUSP" box="[377,817,2144,2188]" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="leptoura">
<emphasis id="B907EAB62A10FFC8FEC9F7E24838F717" box="[377,817,2144,2188]" italics="true" pageId="35" pageNumber="36">Spintherobolus leptoura,</emphasis>
</taxonomicName>
MZUSP 41854, paratype, adult female 25.0 mm SL, Brazil, São Paulo, fazenda Dalila, Registro.
</paragraph>
</caption>
<paragraph id="8BCC36A42A10FFC8FEA8F3D54AA6F2D9" blockId="35.[201,1280,2377,3394]" pageId="35" pageNumber="36">Snout short, shorter than eye diameter. Mouth terminal, angled posteroventrally; maxilla reaching line drawn vertically through anterior mar ­ gin of eye.</paragraph>
<paragraph id="8BCC36A42A10FFC8FA2EF6CE4D3DF668" blockId="35.[1354,2438,2380,3454]" pageId="35" pageNumber="36">
Dorsal-fin rays, ii,
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all specimens. Dorsal-fin origin slightly posterior to midbody. Adipose fin absent.
</paragraph>
<paragraph id="8BCC36A42A10FFCFFA2CF5824F57FC6F" blockId="35.[1354,2438,2380,3454]" lastBlockId="36.[72,1150,299,1977]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
Anal-fin rays iii, 14 (anterior unbranched rays iii-iv, x = 3.1, n = 16; branched anal-fin rays 12-14, x = 12.6, n = 16). Anterior lobe of anal fin of males strongly developed, rounded profile. Distal tip of longest (and largest) rays reaching posteriorly beyond distal tip of most posterior anal-fin rays. Distal profile of posterior anal-fin lobe concave. Distal profile of entire anal fin in females concave, with length of first 6 branched rays decreasing in size and posterior rays smaller, near ­ ly of equal size. Branched anal-fin rays 1 -5 of males very large, and rays 2 to 4 most enlarged and best developed, becoming more than 5 times wider than following posterior rays. Rays 2-4 with a process on anteroproximal face of each lepidotrich. Processes inserted between lepidotrichia of next anterior ray. Each bony ray segment of posterior branch of branched rays 2 to 4, and sometimes also first and fifth branched rays, not rectangular as other branches of same ray, but all with a posteroventral acute and elongate angle extending towards distal tip of that fin ray. Segments of branched fin rays 2 to 4 progressively fused to each other.
<typeStatus id="54C888062A17FFCFFDE5FC89482DFCDA" box="[597,804,779,833]" pageId="36" pageNumber="37">Holotype</typeStatus>
only specimen with hooks. These on two segments of first branched ray. Hooks extremely reduced, with 2 or 3 hooks on posterior face of both segments.
</paragraph>
<paragraph id="8BCC36A42A17FFCFFF27FC784BD2F911" blockId="36.[72,1150,299,1977]" pageId="36" pageNumber="37">
Pectoral-fin rays i,9 (branched rays 8-11, x = 8.9, n = 14). Posterior tip of longest pectoral-fin ray extended beyond pelvic-fin origin in both males and females. Pelvic-fin rays, i,
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all counted specimens (n = 18). Tip of longest ray reaching to or beyond anal-fin origin in both sexes. Pelvic fin of males with short, wide-based hooks on ventromedial face of unbranched ray and first two branched rays (
<specimenCount id="9D75FD2D2A17FFCFFE4BFA5F49C9F989" box="[507,704,1501,1554]" count="1" pageId="36" pageNumber="37" type="male">one male</specimenCount>
had them on first to third branched rays) along almost their total length.
</paragraph>
<caption id="DF0C662C2A17FFCFFB7AFBFF4D04F9C3" ID-DOI="http://doi.org/10.5281/zenodo.10882084" ID-Zenodo-Dep="10882084" httpUri="https://zenodo.org/record/10882084/files/figure.png" pageId="36" pageNumber="37" startId="36.[1226,1294,1149,1196]" targetBox="[1225,2300,308,1096]" targetPageId="36" targetType="figure">
<paragraph id="8BCC36A42A17FFCFFB7AFBFF4D04F9C3" blockId="36.[1225,2298,1149,1624]" pageId="36" pageNumber="37">
Fig. 38. Graph of caudal peduncle depth as a function of standard length of
<taxonomicName id="4C734D272A17FFCFF9DCFB304318FB79" box="[1644,2065,1202,1250]" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="leptoura">
<emphasis id="B907EAB62A17FFCFF9DCFB304C7CFB79" box="[1644,1909,1202,1250]" italics="true" pageId="36" pageNumber="37">Spintherobolus</emphasis>
<emphasis id="B907EAB62A17FFCFF831FB304318FB79" box="[1921,2065,1202,1250]" italics="true" pageId="36" pageNumber="37">leptoura</emphasis>
</taxonomicName>
(black symbols) and
<taxonomicName id="4C734D272A17FFCFFA3AFB6A4D3FFA8D" box="[1418,1590,1256,1302]" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="broccae">
<emphasis id="B907EAB62A17FFCFFA3AFB6A4EA3FA8D" box="[1418,1450,1256,1302]" italics="true" pageId="36" pageNumber="37">S.</emphasis>
<emphasis id="B907EAB62A17FFCFFA06FB6A4D3FFA8D" box="[1462,1590,1256,1302]" italics="true" pageId="36" pageNumber="37">broccae</emphasis>
</taxonomicName>
(empty symbols). Axes are logarithmic. Note that the 95 % confidence intervals in lengths of about 18 to 24 mm SL do not overlap with the means of these of the compared species indicating a significant difference in these size ranges. Comparison of these growth patterns are discussed under the diagnosis of
<taxonomicName id="4C734D272A17FFCFFAF1F9AE4D0CF9C3" box="[1345,1541,1580,1624]" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="leptoura">S. leptoura</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BCC36A42A17FFCEFF27F9104A86FE7F" blockId="36.[72,1150,299,1977]" lastBlockId="37.[171,981,218,1341]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">Principal caudal-fin ray count 10/ 9. Dorsal procurrent caudal-fin rays 9 (7- 10, x = 8.8, n = 9). Ventral procurrent caudal-fin rays, 15 (13-15, x = 14.1, n = 10). Ventral procurrent caudal-fin rays of males strongly-developed, with their proximal tips fused, slab shaped, and expanded in a sagittal plane. Four or five posteriormost caudal (preural) vertebrae (including posterior half centrum) supporting caudal fin and procurrent rays and bearing large, modified hemal spines. Those elements anterior to hemal spine of antepenultimate vertebra fused to each other and having lateral bony processes between muscles and skin. Proximal tips of anterior ventral procurrent rays undeveloped.</paragraph>
<caption id="DF0C662C2A17FFCFFFF8F79A49E6F7E7" ID-Table-UUID="DF0C662C2A17FFCFFFF8F79A49E6F7E7" httpUri="http://table.plazi.org/id/DF0C662C2A17FFCFFFF8F79A49E6F7E7" pageId="36" pageNumber="37" startId="36.[72,183,2072,2118]" targetBox="[71,2289,2244,3457]" targetIsTable="true" targetPageId="36" targetType="table">
<paragraph id="8BCC36A42A17FFCFFFF8F79A49E6F7E7" blockId="36.[72,2297,2072,2172]" pageId="36" pageNumber="37">
Table 5. Morphometries of
<taxonomicName id="4C734D272A17FFCFFDDDF79A4F15F7DD" box="[621,1052,2072,2118]" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="leptoura">
<emphasis id="B907EAB62A17FFCFFDDDF79A487FF7DD" box="[621,886,2072,2118]" italics="true" pageId="36" pageNumber="37">Spintherobolus</emphasis>
<emphasis id="B907EAB62A17FFCFFC36F79A4F29F7DD" box="[902,1056,2072,2118]" italics="true" pageId="36" pageNumber="37">leptoura.</emphasis>
</taxonomicName>
Table includes measurements of all type specimens. Values for the holotype are given separately.
</paragraph>
</caption>
<paragraph id="8BCC36A42A17FFCFFC82F74643E8F21A" blockId="36.[67,2299,2227,3470]" pageId="36" pageNumber="37">
<table id="F973C4042A170014FFF7F74643F8F21A" box="[71,2289,2244,3457]" gridcols="10" gridrows="21" pageId="36" pageNumber="37">
<tr id="354334E62A170014FFF7F74643F8F76A" box="[71,2289,2244,2289]" gridrow="0" pageId="36" pageNumber="37" rowspan-0="1">
<th id="76925D9A2A170014FC82F74648E8F76A" box="[818,993,2244,2289]" gridcol="1" gridrow="0" pageId="36" pageNumber="37">holotype</th>
<th id="76925D9A2A170014FBF6F7464D41F76A" box="[1094,1608,2244,2289]" colspan="4" colspanRight="3" gridcol="2" gridrow="0" pageId="36" pageNumber="37">males</th>
<th id="76925D9A2A170014F96DF74643F8F76A" box="[1757,2289,2244,2289]" colspan="4" colspanRight="3" gridcol="6" gridrow="0" pageId="36" pageNumber="37">females</th>
</tr>
<tr id="354334E62A170014FFF7F68F43F8F6A7" box="[71,2289,2317,2364]" gridrow="1" pageId="36" pageNumber="37" rowspan-0="1" rowspan-1="1">
<td id="76925D9A2A170014FBF6F68F4F54F6A7" box="[1094,1117,2317,2364]" gridcol="2" gridrow="1" pageId="36" pageNumber="37">n</td>
<td id="76925D9A2A170014FB1AF68F4FF1F6A7" box="[1194,1272,2317,2364]" gridcol="3" gridrow="1" pageId="36" pageNumber="37">low</td>
<td id="76925D9A2A170014FAF6F68F4E97F6A7" box="[1350,1438,2317,2364]" gridcol="4" gridrow="1" pageId="36" pageNumber="37">high</td>
<td id="76925D9A2A170014FA6CF68F4D41F6A7" box="[1500,1608,2317,2364]" gridcol="5" gridrow="1" pageId="36" pageNumber="37">mean</td>
<td id="76925D9A2A170014F96DF68F4C0FF6A7" box="[1757,1798,2317,2364]" gridcol="6" gridrow="1" pageId="36" pageNumber="37">n</td>
<td id="76925D9A2A170014F8E9F68F4CAFF6A7" box="[1881,1958,2317,2364]" gridcol="7" gridrow="1" pageId="36" pageNumber="37">low</td>
<td id="76925D9A2A170014F842F68F4340F6A7" box="[2034,2121,2317,2364]" gridcol="8" gridrow="1" pageId="36" pageNumber="37">high</td>
<td id="76925D9A2A170014F738F68F43F8F6A7" box="[2184,2289,2317,2364]" gridcol="9" gridrow="1" pageId="36" pageNumber="37">mean</td>
</tr>
<tr id="354334E62A170014FFF7F6E343F8F614" box="[71,2289,2401,2447]" gridrow="2" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F6E34971F614" box="[71,632,2401,2447]" gridcol="0" gridrow="2" pageId="36" pageNumber="37">Standard length [mm]</th>
<td id="76925D9A2A170014FC82F6E348E8F614" box="[818,993,2401,2447]" gridcol="1" gridrow="2" pageId="36" pageNumber="37">21.8</td>
<td id="76925D9A2A170014FBF6F6E34F54F614" box="[1094,1117,2401,2447]" gridcol="2" gridrow="2" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF6E34FF1F614" box="[1194,1272,2401,2447]" gridcol="3" gridrow="2" pageId="36" pageNumber="37">17.3</td>
<td id="76925D9A2A170014FAF6F6E34E97F614" box="[1350,1438,2401,2447]" gridcol="4" gridrow="2" pageId="36" pageNumber="37">21.8</td>
<td id="76925D9A2A170014FA6CF6E34D41F614" box="[1500,1608,2401,2447]" gridcol="5" gridrow="2" pageId="36" pageNumber="37">19.5</td>
<td id="76925D9A2A170014F96DF6E34C0FF614" box="[1757,1798,2401,2447]" gridcol="6" gridrow="2" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F6E34CAFF614" box="[1881,1958,2401,2447]" gridcol="7" gridrow="2" pageId="36" pageNumber="37">16.2</td>
<td id="76925D9A2A170014F842F6E34340F614" box="[2034,2121,2401,2447]" gridcol="8" gridrow="2" pageId="36" pageNumber="37">27.2</td>
<td id="76925D9A2A170014F738F6E343F8F614" box="[2184,2289,2401,2447]" gridcol="9" gridrow="2" pageId="36" pageNumber="37">20.9</td>
</tr>
<tr id="354334E62A170014FFF7F62C43F8F640" box="[71,2289,2478,2523]" gridrow="3" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F62C43F8F640" box="[71,2289,2478,2523]" colspan="10" colspanRight="9" gridcol="0" gridrow="3" pageId="36" pageNumber="37">Percentage of standard length</th>
</tr>
<tr id="354334E62A170014FFF7F66143F8F594" box="[71,2289,2531,2575]" gridrow="4" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F6614971F594" box="[71,632,2531,2575]" gridcol="0" gridrow="4" pageId="36" pageNumber="37">Snout to anal-fin origin</th>
<td id="76925D9A2A170014FC82F66148E8F594" box="[818,993,2531,2575]" gridcol="1" gridrow="4" pageId="36" pageNumber="37">58.7</td>
<td id="76925D9A2A170014FBF6F6614F54F594" box="[1094,1117,2531,2575]" gridcol="2" gridrow="4" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF6614FF1F594" box="[1194,1272,2531,2575]" gridcol="3" gridrow="4" pageId="36" pageNumber="37">57.3</td>
<td id="76925D9A2A170014FAF6F6614E97F594" box="[1350,1438,2531,2575]" gridcol="4" gridrow="4" pageId="36" pageNumber="37">61.9</td>
<td id="76925D9A2A170014FA6CF6614D41F594" box="[1500,1608,2531,2575]" gridcol="5" gridrow="4" pageId="36" pageNumber="37">58.9</td>
<td id="76925D9A2A170014F96DF6614C0FF594" box="[1757,1798,2531,2575]" gridcol="6" gridrow="4" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F6614CAFF594" box="[1881,1958,2531,2575]" gridcol="7" gridrow="4" pageId="36" pageNumber="37">58.6</td>
<td id="76925D9A2A170014F842F6614340F594" box="[2034,2121,2531,2575]" gridcol="8" gridrow="4" pageId="36" pageNumber="37">62.2</td>
<td id="76925D9A2A170014F738F66143F8F594" box="[2184,2289,2531,2575]" gridcol="9" gridrow="4" pageId="36" pageNumber="37">60.5</td>
</tr>
<tr id="354334E62A170014FFF7F59B43F8F5DD" box="[71,2289,2585,2630]" gridrow="5" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F59B4971F5DD" box="[71,632,2585,2630]" gridcol="0" gridrow="5" pageId="36" pageNumber="37">Snout to dorsal-fin origin</th>
<td id="76925D9A2A170014FC82F59B48E8F5DD" box="[818,993,2585,2630]" gridcol="1" gridrow="5" pageId="36" pageNumber="37">56.0</td>
<td id="76925D9A2A170014FBF6F59B4F54F5DD" box="[1094,1117,2585,2630]" gridcol="2" gridrow="5" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF59B4FF1F5DD" box="[1194,1272,2585,2630]" gridcol="3" gridrow="5" pageId="36" pageNumber="37">53.8</td>
<td id="76925D9A2A170014FAF6F59B4E97F5DD" box="[1350,1438,2585,2630]" gridcol="4" gridrow="5" pageId="36" pageNumber="37">56.0</td>
<td id="76925D9A2A170014FA6CF59B4D41F5DD" box="[1500,1608,2585,2630]" gridcol="5" gridrow="5" pageId="36" pageNumber="37">55.3</td>
<td id="76925D9A2A170014F96DF59B4C0FF5DD" box="[1757,1798,2585,2630]" gridcol="6" gridrow="5" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F59B4CAFF5DD" box="[1881,1958,2585,2630]" gridcol="7" gridrow="5" pageId="36" pageNumber="37">53.5</td>
<td id="76925D9A2A170014F842F59B4340F5DD" box="[2034,2121,2585,2630]" gridcol="8" gridrow="5" pageId="36" pageNumber="37">57.0</td>
<td id="76925D9A2A170014F738F59B43F8F5DD" box="[2184,2289,2585,2630]" gridcol="9" gridrow="5" pageId="36" pageNumber="37">55.5</td>
</tr>
<tr id="354334E62A170014FFF7F5CC43F8F5E0" box="[71,2289,2638,2683]" gridrow="6" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F5CC4971F5E0" box="[71,632,2638,2683]" gridcol="0" gridrow="6" pageId="36" pageNumber="37">Snout to pelvic-fin origin</th>
<td id="76925D9A2A170014FC82F5CC48E8F5E0" box="[818,993,2638,2683]" gridcol="1" gridrow="6" pageId="36" pageNumber="37">44.5</td>
<td id="76925D9A2A170014FBF6F5CC4F54F5E0" box="[1094,1117,2638,2683]" gridcol="2" gridrow="6" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF5CC4FF1F5E0" box="[1194,1272,2638,2683]" gridcol="3" gridrow="6" pageId="36" pageNumber="37">42.2</td>
<td id="76925D9A2A170014FAF6F5CC4E97F5E0" box="[1350,1438,2638,2683]" gridcol="4" gridrow="6" pageId="36" pageNumber="37">44.5</td>
<td id="76925D9A2A170014FA6CF5CC4D41F5E0" box="[1500,1608,2638,2683]" gridcol="5" gridrow="6" pageId="36" pageNumber="37">43.4</td>
<td id="76925D9A2A170014F96DF5CC4C0FF5E0" box="[1757,1798,2638,2683]" gridcol="6" gridrow="6" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F5CC4CAFF5E0" box="[1881,1958,2638,2683]" gridcol="7" gridrow="6" pageId="36" pageNumber="37">37.4</td>
<td id="76925D9A2A170014F842F5CC4340F5E0" box="[2034,2121,2638,2683]" gridcol="8" gridrow="6" pageId="36" pageNumber="37">45.1</td>
<td id="76925D9A2A170014F738F5CC43F8F5E0" box="[2184,2289,2638,2683]" gridcol="9" gridrow="6" pageId="36" pageNumber="37">41.6</td>
</tr>
<tr id="354334E62A170014FFF7F50143F8F52A" box="[71,2289,2691,2737]" gridrow="7" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F5014971F52A" box="[71,632,2691,2737]" gridcol="0" gridrow="7" pageId="36" pageNumber="37">Dorsal-fin base length</th>
<td id="76925D9A2A170014FC82F50148E8F52A" box="[818,993,2691,2737]" gridcol="1" gridrow="7" pageId="36" pageNumber="37">11.3</td>
<td id="76925D9A2A170014FBF6F5014F54F52A" box="[1094,1117,2691,2737]" gridcol="2" gridrow="7" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF5014FF1F52A" box="[1194,1272,2691,2737]" gridcol="3" gridrow="7" pageId="36" pageNumber="37">11.3</td>
<td id="76925D9A2A170014FAF6F5014E97F52A" box="[1350,1438,2691,2737]" gridcol="4" gridrow="7" pageId="36" pageNumber="37">12.5</td>
<td id="76925D9A2A170014FA6CF5014D41F52A" box="[1500,1608,2691,2737]" gridcol="5" gridrow="7" pageId="36" pageNumber="37">11.7</td>
<td id="76925D9A2A170014F96DF5014C0FF52A" box="[1757,1798,2691,2737]" gridcol="6" gridrow="7" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F5014CAFF52A" box="[1881,1958,2691,2737]" gridcol="7" gridrow="7" pageId="36" pageNumber="37">9.6</td>
<td id="76925D9A2A170014F842F5014340F52A" box="[2034,2121,2691,2737]" gridcol="8" gridrow="7" pageId="36" pageNumber="37">12.9</td>
<td id="76925D9A2A170014F738F50143F8F52A" box="[2184,2289,2691,2737]" gridcol="9" gridrow="7" pageId="36" pageNumber="37">11.5</td>
</tr>
<tr id="354334E62A170014FFF7F53543F8F57D" box="[71,2289,2743,2790]" gridrow="8" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F5354971F57D" box="[71,632,2743,2790]" gridcol="0" gridrow="8" pageId="36" pageNumber="37">Anal-fin base length</th>
<td id="76925D9A2A170014FC82F53548E8F57D" box="[818,993,2743,2790]" gridcol="1" gridrow="8" pageId="36" pageNumber="37">21.1</td>
<td id="76925D9A2A170014FBF6F5354F54F57D" box="[1094,1117,2743,2790]" gridcol="2" gridrow="8" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF5354FF1F57D" box="[1194,1272,2743,2790]" gridcol="3" gridrow="8" pageId="36" pageNumber="37">21.1</td>
<td id="76925D9A2A170014FAF6F5354E97F57D" box="[1350,1438,2743,2790]" gridcol="4" gridrow="8" pageId="36" pageNumber="37">22.9</td>
<td id="76925D9A2A170014FA6CF5354D41F57D" box="[1500,1608,2743,2790]" gridcol="5" gridrow="8" pageId="36" pageNumber="37">21.9</td>
<td id="76925D9A2A170014F96DF5354C0FF57D" box="[1757,1798,2743,2790]" gridcol="6" gridrow="8" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F5354CAFF57D" box="[1881,1958,2743,2790]" gridcol="7" gridrow="8" pageId="36" pageNumber="37">21.0</td>
<td id="76925D9A2A170014F842F5354340F57D" box="[2034,2121,2743,2790]" gridcol="8" gridrow="8" pageId="36" pageNumber="37">24.0</td>
<td id="76925D9A2A170014F738F53543F8F57D" box="[2184,2289,2743,2790]" gridcol="9" gridrow="8" pageId="36" pageNumber="37">22.8</td>
</tr>
<tr id="354334E62A170014FFF7F56F43F8F480" box="[71,2289,2797,2843]" gridrow="9" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F56F4971F480" box="[71,632,2797,2843]" gridcol="0" gridrow="9" pageId="36" pageNumber="37">Caudal peduncle length</th>
<td id="76925D9A2A170014FC82F56F48E8F480" box="[818,993,2797,2843]" gridcol="1" gridrow="9" pageId="36" pageNumber="37">18.4</td>
<td id="76925D9A2A170014FBF6F56F4F54F480" box="[1094,1117,2797,2843]" gridcol="2" gridrow="9" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF56F4FF1F480" box="[1194,1272,2797,2843]" gridcol="3" gridrow="9" pageId="36" pageNumber="37">16.7</td>
<td id="76925D9A2A170014FAF6F56F4E97F480" box="[1350,1438,2797,2843]" gridcol="4" gridrow="9" pageId="36" pageNumber="37">21.5</td>
<td id="76925D9A2A170014FA6CF56F4D41F480" box="[1500,1608,2797,2843]" gridcol="5" gridrow="9" pageId="36" pageNumber="37">18.8</td>
<td id="76925D9A2A170014F96DF56F4C0FF480" box="[1757,1798,2797,2843]" gridcol="6" gridrow="9" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F56F4CAFF480" box="[1881,1958,2797,2843]" gridcol="7" gridrow="9" pageId="36" pageNumber="37">15.2</td>
<td id="76925D9A2A170014F842F56F4340F480" box="[2034,2121,2797,2843]" gridcol="8" gridrow="9" pageId="36" pageNumber="37">18.1</td>
<td id="76925D9A2A170014F738F56F43F8F480" box="[2184,2289,2797,2843]" gridcol="9" gridrow="9" pageId="36" pageNumber="37">163</td>
</tr>
<tr id="354334E62A170014FFF7F4A043F8F4CA" box="[71,2289,2850,2897]" gridrow="10" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F4A04971F4CA" box="[71,632,2850,2897]" gridcol="0" gridrow="10" pageId="36" pageNumber="37">Caudal peduncle depth</th>
<td id="76925D9A2A170014FC82F4A048E8F4CA" box="[818,993,2850,2897]" gridcol="1" gridrow="10" pageId="36" pageNumber="37">12.4</td>
<td id="76925D9A2A170014FBF6F4A04F54F4CA" box="[1094,1117,2850,2897]" gridcol="2" gridrow="10" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF4A04FF1F4CA" box="[1194,1272,2850,2897]" gridcol="3" gridrow="10" pageId="36" pageNumber="37">11.0</td>
<td id="76925D9A2A170014FAF6F4A04E97F4CA" box="[1350,1438,2850,2897]" gridcol="4" gridrow="10" pageId="36" pageNumber="37">13.1</td>
<td id="76925D9A2A170014FA6CF4A04D41F4CA" box="[1500,1608,2850,2897]" gridcol="5" gridrow="10" pageId="36" pageNumber="37">12.3</td>
<td id="76925D9A2A170014F96DF4A04C0FF4CA" box="[1757,1798,2850,2897]" gridcol="6" gridrow="10" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F4A04CAFF4CA" box="[1881,1958,2850,2897]" gridcol="7" gridrow="10" pageId="36" pageNumber="37">9.5</td>
<td id="76925D9A2A170014F842F4A04340F4CA" box="[2034,2121,2850,2897]" gridcol="8" gridrow="10" pageId="36" pageNumber="37">13.5</td>
<td id="76925D9A2A170014F738F4A043F8F4CA" box="[2184,2289,2850,2897]" gridcol="9" gridrow="10" pageId="36" pageNumber="37">11.8</td>
</tr>
<tr id="354334E62A170014FFF7F4D543F8F41D" box="[71,2289,2903,2950]" gridrow="11" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F4D54971F41D" box="[71,632,2903,2950]" gridcol="0" gridrow="11" pageId="36" pageNumber="37">Depth at dorsal-fin origin</th>
<td id="76925D9A2A170014FC82F4D548E8F41D" box="[818,993,2903,2950]" gridcol="1" gridrow="11" pageId="36" pageNumber="37">27.5</td>
<td id="76925D9A2A170014FBF6F4D54F54F41D" box="[1094,1117,2903,2950]" gridcol="2" gridrow="11" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF4D54FF1F41D" box="[1194,1272,2903,2950]" gridcol="3" gridrow="11" pageId="36" pageNumber="37">26.6</td>
<td id="76925D9A2A170014FAF6F4D54E97F41D" box="[1350,1438,2903,2950]" gridcol="4" gridrow="11" pageId="36" pageNumber="37">29.7</td>
<td id="76925D9A2A170014FA6CF4D54D41F41D" box="[1500,1608,2903,2950]" gridcol="5" gridrow="11" pageId="36" pageNumber="37">28.3</td>
<td id="76925D9A2A170014F96DF4D54C0FF41D" box="[1757,1798,2903,2950]" gridcol="6" gridrow="11" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F4D54CAFF41D" box="[1881,1958,2903,2950]" gridcol="7" gridrow="11" pageId="36" pageNumber="37">26.6</td>
<td id="76925D9A2A170014F842F4D54340F41D" box="[2034,2121,2903,2950]" gridcol="8" gridrow="11" pageId="36" pageNumber="37">33.7</td>
<td id="76925D9A2A170014F738F4D543F8F41D" box="[2184,2289,2903,2950]" gridcol="9" gridrow="11" pageId="36" pageNumber="37">29.9</td>
</tr>
<tr id="354334E62A170014FFF7F40F43F8F420" box="[71,2289,2957,3003]" gridrow="12" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F40F4971F420" box="[71,632,2957,3003]" gridcol="0" gridrow="12" pageId="36" pageNumber="37">Dorsal-fin length</th>
<td id="76925D9A2A170014FC82F40F48E8F420" box="[818,993,2957,3003]" gridcol="1" gridrow="12" pageId="36" pageNumber="37">25.7</td>
<td id="76925D9A2A170014FBF6F40F4F54F420" box="[1094,1117,2957,3003]" gridcol="2" gridrow="12" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF40F4FF1F420" box="[1194,1272,2957,3003]" gridcol="3" gridrow="12" pageId="36" pageNumber="37">22.5</td>
<td id="76925D9A2A170014FAF6F40F4E97F420" box="[1350,1438,2957,3003]" gridcol="4" gridrow="12" pageId="36" pageNumber="37">28.6</td>
<td id="76925D9A2A170014FA6CF40F4D41F420" box="[1500,1608,2957,3003]" gridcol="5" gridrow="12" pageId="36" pageNumber="37">26.4</td>
<td id="76925D9A2A170014F96DF40F4C0FF420" box="[1757,1798,2957,3003]" gridcol="6" gridrow="12" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F40F4CAFF420" box="[1881,1958,2957,3003]" gridcol="7" gridrow="12" pageId="36" pageNumber="37">23.0</td>
<td id="76925D9A2A170014F842F40F4340F420" box="[2034,2121,2957,3003]" gridcol="8" gridrow="12" pageId="36" pageNumber="37">29.4</td>
<td id="76925D9A2A170014F738F40F43F8F420" box="[2184,2289,2957,3003]" gridcol="9" gridrow="12" pageId="36" pageNumber="37">26.2</td>
</tr>
<tr id="354334E62A170014FFF7F44143F8F46A" box="[71,2289,3011,3057]" gridrow="13" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F4414971F46A" box="[71,632,3011,3057]" gridcol="0" gridrow="13" pageId="36" pageNumber="37">Pelvic-fin length</th>
<td id="76925D9A2A170014FC82F44148E8F46A" box="[818,993,3011,3057]" gridcol="1" gridrow="13" pageId="36" pageNumber="37">20.2</td>
<td id="76925D9A2A170014FBF6F4414F54F46A" box="[1094,1117,3011,3057]" gridcol="2" gridrow="13" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF4414FF1F46A" box="[1194,1272,3011,3057]" gridcol="3" gridrow="13" pageId="36" pageNumber="37">17.8</td>
<td id="76925D9A2A170014FAF6F4414E97F46A" box="[1350,1438,3011,3057]" gridcol="4" gridrow="13" pageId="36" pageNumber="37">20.2</td>
<td id="76925D9A2A170014FA6CF4414D41F46A" box="[1500,1608,3011,3057]" gridcol="5" gridrow="13" pageId="36" pageNumber="37">18.8</td>
<td id="76925D9A2A170014F96DF4414C0FF46A" box="[1757,1798,3011,3057]" gridcol="6" gridrow="13" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F4414CAFF46A" box="[1881,1958,3011,3057]" gridcol="7" gridrow="13" pageId="36" pageNumber="37">14.2</td>
<td id="76925D9A2A170014F842F4414340F46A" box="[2034,2121,3011,3057]" gridcol="8" gridrow="13" pageId="36" pageNumber="37">19.0</td>
<td id="76925D9A2A170014F738F44143F8F46A" box="[2184,2289,3011,3057]" gridcol="9" gridrow="13" pageId="36" pageNumber="37">17.0</td>
</tr>
<tr id="354334E62A170014FFF7F47543F8F3BC" box="[71,2289,3063,3111]" gridrow="14" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F4754971F3BC" box="[71,632,3063,3111]" gridcol="0" gridrow="14" pageId="36" pageNumber="37">Pectoral-fin length</th>
<td id="76925D9A2A170014FC82F47548E8F3BC" box="[818,993,3063,3111]" gridcol="1" gridrow="14" pageId="36" pageNumber="37">17.9</td>
<td id="76925D9A2A170014FBF6F4754F54F3BC" box="[1094,1117,3063,3111]" gridcol="2" gridrow="14" pageId="36" pageNumber="37">5</td>
<td id="76925D9A2A170014FB1AF4754FF1F3BC" box="[1194,1272,3063,3111]" gridcol="3" gridrow="14" pageId="36" pageNumber="37">15.6</td>
<td id="76925D9A2A170014FAF6F4754E97F3BC" box="[1350,1438,3063,3111]" gridcol="4" gridrow="14" pageId="36" pageNumber="37">17.9</td>
<td id="76925D9A2A170014FA6CF4754D41F3BC" box="[1500,1608,3063,3111]" gridcol="5" gridrow="14" pageId="36" pageNumber="37">16.9</td>
<td id="76925D9A2A170014F96DF4754C0FF3BC" box="[1757,1798,3063,3111]" gridcol="6" gridrow="14" pageId="36" pageNumber="37">10</td>
<td id="76925D9A2A170014F8E9F4754CAFF3BC" box="[1881,1958,3063,3111]" gridcol="7" gridrow="14" pageId="36" pageNumber="37">14.6</td>
<td id="76925D9A2A170014F842F4754340F3BC" box="[2034,2121,3063,3111]" gridcol="8" gridrow="14" pageId="36" pageNumber="37">19.0</td>
<td id="76925D9A2A170014F738F47543F8F3BC" box="[2184,2289,3063,3111]" gridcol="9" gridrow="14" pageId="36" pageNumber="37">16.3</td>
</tr>
<tr id="354334E62A170014FFF7F3AD43F8F3C6" box="[71,2289,3119,3165]" gridrow="15" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F3AD4971F3C6" box="[71,632,3119,3165]" gridcol="0" gridrow="15" pageId="36" pageNumber="37">Bony head length</th>
<td id="76925D9A2A170014FC82F3AD48E8F3C6" box="[818,993,3119,3165]" gridcol="1" gridrow="15" pageId="36" pageNumber="37">28.9</td>
<td id="76925D9A2A170014FBF6F3AD4F54F3C6" box="[1094,1117,3119,3165]" gridcol="2" gridrow="15" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF3AD4FF1F3C6" box="[1194,1272,3119,3165]" gridcol="3" gridrow="15" pageId="36" pageNumber="37">28.9</td>
<td id="76925D9A2A170014FAF6F3AD4E97F3C6" box="[1350,1438,3119,3165]" gridcol="4" gridrow="15" pageId="36" pageNumber="37">30.4</td>
<td id="76925D9A2A170014FA6CF3AD4D41F3C6" box="[1500,1608,3119,3165]" gridcol="5" gridrow="15" pageId="36" pageNumber="37">29.3</td>
<td id="76925D9A2A170014F96DF3AD4C0FF3C6" box="[1757,1798,3119,3165]" gridcol="6" gridrow="15" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F3AD4CAFF3C6" box="[1881,1958,3119,3165]" gridcol="7" gridrow="15" pageId="36" pageNumber="37">26.1</td>
<td id="76925D9A2A170014F842F3AD4340F3C6" box="[2034,2121,3119,3165]" gridcol="8" gridrow="15" pageId="36" pageNumber="37">30.2</td>
<td id="76925D9A2A170014F738F3AD43F8F3C6" box="[2184,2289,3119,3165]" gridcol="9" gridrow="15" pageId="36" pageNumber="37">28.2</td>
</tr>
<tr id="354334E62A170014FFF7F3FF43F8F332" box="[71,2289,3197,3241]" gridrow="16" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F3FF43F8F332" box="[71,2289,3197,3241]" colspan="10" colspanRight="9" gridcol="0" gridrow="16" pageId="36" pageNumber="37">Percentage of head length</th>
</tr>
<tr id="354334E62A170014FFF7F33343F8F344" box="[71,2289,3249,3295]" gridrow="17" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F3334971F344" box="[71,632,3249,3295]" gridcol="0" gridrow="17" pageId="36" pageNumber="37">Snout length</th>
<td id="76925D9A2A170014FC82F33348E8F344" box="[818,993,3249,3295]" gridcol="1" gridrow="17" pageId="36" pageNumber="37">19.5</td>
<td id="76925D9A2A170014FBF6F3334F54F344" box="[1094,1117,3249,3295]" gridcol="2" gridrow="17" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF3334FF1F344" box="[1194,1272,3249,3295]" gridcol="3" gridrow="17" pageId="36" pageNumber="37">19.0</td>
<td id="76925D9A2A170014FAF6F3334E97F344" box="[1350,1438,3249,3295]" gridcol="4" gridrow="17" pageId="36" pageNumber="37">22.4</td>
<td id="76925D9A2A170014FA6CF3334D41F344" box="[1500,1608,3249,3295]" gridcol="5" gridrow="17" pageId="36" pageNumber="37">19.9</td>
<td id="76925D9A2A170014F96DF3334C0FF344" box="[1757,1798,3249,3295]" gridcol="6" gridrow="17" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F3334CAFF344" box="[1881,1958,3249,3295]" gridcol="7" gridrow="17" pageId="36" pageNumber="37">18.9</td>
<td id="76925D9A2A170014F842F3334340F344" box="[2034,2121,3249,3295]" gridcol="8" gridrow="17" pageId="36" pageNumber="37">23.2</td>
<td id="76925D9A2A170014F738F33343F8F344" box="[2184,2289,3249,3295]" gridcol="9" gridrow="17" pageId="36" pageNumber="37">20.9</td>
</tr>
<tr id="354334E62A170014FFF7F36443F8F28E" box="[71,2289,3302,3349]" gridrow="18" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F3644971F28E" box="[71,632,3302,3349]" gridcol="0" gridrow="18" pageId="36" pageNumber="37">Upper jaw length</th>
<td id="76925D9A2A170014FC82F36448E8F28E" box="[818,993,3302,3349]" gridcol="1" gridrow="18" pageId="36" pageNumber="37">19.5</td>
<td id="76925D9A2A170014FBF6F3644F54F28E" box="[1094,1117,3302,3349]" gridcol="2" gridrow="18" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF3644FF1F28E" box="[1194,1272,3302,3349]" gridcol="3" gridrow="18" pageId="36" pageNumber="37">19.0</td>
<td id="76925D9A2A170014FAF6F3644E97F28E" box="[1350,1438,3302,3349]" gridcol="4" gridrow="18" pageId="36" pageNumber="37">21.4</td>
<td id="76925D9A2A170014FA6CF3644D41F28E" box="[1500,1608,3302,3349]" gridcol="5" gridrow="18" pageId="36" pageNumber="37">19.9</td>
<td id="76925D9A2A170014F96DF3644C0FF28E" box="[1757,1798,3302,3349]" gridcol="6" gridrow="18" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F3644CAFF28E" box="[1881,1958,3302,3349]" gridcol="7" gridrow="18" pageId="36" pageNumber="37">18.4</td>
<td id="76925D9A2A170014F842F3644340F28E" box="[2034,2121,3302,3349]" gridcol="8" gridrow="18" pageId="36" pageNumber="37">22.9</td>
<td id="76925D9A2A170014F738F36443F8F28E" box="[2184,2289,3302,3349]" gridcol="9" gridrow="18" pageId="36" pageNumber="37">21.0</td>
</tr>
<tr id="354334E62A170014FFF7F29F43F8F2DE" box="[71,2289,3357,3397]" gridrow="19" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F29F4971F2DE" box="[71,632,3357,3397]" gridcol="0" gridrow="19" pageId="36" pageNumber="37">Horizontal eye diameter</th>
<td id="76925D9A2A170014FC82F29F48E8F2DE" box="[818,993,3357,3397]" gridcol="1" gridrow="19" pageId="36" pageNumber="37">27.0</td>
<td id="76925D9A2A170014FBF6F29F4F54F2DE" box="[1094,1117,3357,3397]" gridcol="2" gridrow="19" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF29F4FF1F2DE" box="[1194,1272,3357,3397]" gridcol="3" gridrow="19" pageId="36" pageNumber="37">25.9</td>
<td id="76925D9A2A170014FAF6F29F4E97F2DE" box="[1350,1438,3357,3397]" gridcol="4" gridrow="19" pageId="36" pageNumber="37">28.1</td>
<td id="76925D9A2A170014FA6CF29F4D41F2DE" box="[1500,1608,3357,3397]" gridcol="5" gridrow="19" pageId="36" pageNumber="37">26.9</td>
<td id="76925D9A2A170014F96DF29F4C0FF2DE" box="[1757,1798,3357,3397]" gridcol="6" gridrow="19" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F29F4CAFF2DE" box="[1881,1958,3357,3397]" gridcol="7" gridrow="19" pageId="36" pageNumber="37">23.2</td>
<td id="76925D9A2A170014F842F29F4340F2DE" box="[2034,2121,3357,3397]" gridcol="8" gridrow="19" pageId="36" pageNumber="37">30.3</td>
<td id="76925D9A2A170014F738F29F43F8F2DE" box="[2184,2289,3357,3397]" gridcol="9" gridrow="19" pageId="36" pageNumber="37">27.4</td>
</tr>
<tr id="354334E62A170014FFF7F2C443F8F21A" box="[71,2289,3398,3457]" gridrow="20" pageId="36" pageNumber="37">
<th id="76925D9A2A170014FFF7F2C44971F21A" box="[71,632,3398,3457]" gridcol="0" gridrow="20" pageId="36" pageNumber="37">Least interorbital width</th>
<td id="76925D9A2A170014FC82F2C448E8F21A" box="[818,993,3398,3457]" gridcol="1" gridrow="20" pageId="36" pageNumber="37">27.0</td>
<td id="76925D9A2A170014FBF6F2C44F54F21A" box="[1094,1117,3398,3457]" gridcol="2" gridrow="20" pageId="36" pageNumber="37">6</td>
<td id="76925D9A2A170014FB1AF2C44FF1F21A" box="[1194,1272,3398,3457]" gridcol="3" gridrow="20" pageId="36" pageNumber="37">26.3</td>
<td id="76925D9A2A170014FAF6F2C44E97F21A" box="[1350,1438,3398,3457]" gridcol="4" gridrow="20" pageId="36" pageNumber="37">29.3</td>
<td id="76925D9A2A170014FA6CF2C44D41F21A" box="[1500,1608,3398,3457]" gridcol="5" gridrow="20" pageId="36" pageNumber="37">27.5</td>
<td id="76925D9A2A170014F96DF2C44C0FF21A" box="[1757,1798,3398,3457]" gridcol="6" gridrow="20" pageId="36" pageNumber="37">12</td>
<td id="76925D9A2A170014F8E9F2C44CAFF21A" box="[1881,1958,3398,3457]" gridcol="7" gridrow="20" pageId="36" pageNumber="37">25.5</td>
<td id="76925D9A2A170014F842F2C44340F21A" box="[2034,2121,3398,3457]" gridcol="8" gridrow="20" pageId="36" pageNumber="37">30.3</td>
<td id="76925D9A2A170014F738F2C443F8F21A" box="[2184,2289,3398,3457]" gridcol="9" gridrow="20" pageId="36" pageNumber="37">27.9</td>
</tr>
</table>
</paragraph>
<paragraph id="8BCC36A42A16FFCEFF58FE684A7DFCE3" blockId="37.[171,981,218,1341]" pageId="37" pageNumber="38">
Scales cycloid. Lateral line with 4 perforated scales (n = 10). Scales in longitudinal series 33 (n = 3). Scale rows between dorsal-fin origin and lateral-line series 5 (n = 6), and between lateralline series and pelvic-fin origin 5 (4-5, x = 4.5, n = 6). Scale rows around caudal peduncle
<quantity id="4C8B9B412A16FFCEFC35FD4E48DCFD6A" box="[901,981,716,753]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="37" pageNumber="38" unit="in" value="14.0">14 in</quantity>
all countable specimens (n = 2). No modified scales on caudal fin. Scale sheath along anal-fin base absent.
</paragraph>
<paragraph id="8BCC36A42A16FFCEFF57FC024953FB2E" blockId="37.[171,981,218,1341]" pageId="37" pageNumber="38">
Teeth. All jaw teeth with basal pedicle and somewhat laterally expanded distal conical cusp. Anterior dentary teeth tricuspid but each with single cusp laterally and posteriorly. Premaxillary and maxillary teeth all conical. Premaxillary teeth 7-8, maxillary teeth 5-6 and dentary teeth
<quantity id="4C8B9B412A16FFCEFF1DFB134A27FB2E" box="[173,302,1169,1205]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="37" pageNumber="38" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">10-11 in</quantity>
2 c&amp;s specimens.
</paragraph>
<paragraph id="8BCC36A42A16FFCEFF57FB3F4858FAA6" blockId="37.[171,981,218,1341]" pageId="37" pageNumber="38">
Vertebrae 33 (range = 32-34, x = 32.9, n = 16). Branchiostegal rays
<quantity id="4C8B9B412A16FFCEFE4AFB6A493EFA94" box="[506,567,1256,1295]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="37" pageNumber="38" unit="in" value="4.0">4 in</quantity>
two cleared and stained specimens, arranged as in other species.
</paragraph>
<paragraph id="8BCC36A42A16FFCEFF1DFAED49EFF6D5" blockId="37.[167,979,1391,2605]" pageId="37" pageNumber="38">
Color in alcohol. See
<figureCitation id="13482A212A16FFCEFD9AFAED4804FA02" box="[554,781,1391,1433]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="35.[207,272,2036,2082]" captionStartId-1="35.[205,267,2144,2188]" captionTargetBox-0="[210,2446,290,1130]" captionTargetBox-1="[206,2443,1144,1985]" captionTargetPageId-0="35" captionTargetPageId-1="35" captionText-0="Fig. 36. Spintherobolus leptoura, MZUSP 41854, holotype, adult male 19.1 mm SL, Brazil, São Paulo, fazenda Dalila, Registro." captionText-1="Fig. 37. Spintherobolus leptoura, MZUSP 41854, paratype, adult female 25.0 mm SL, Brazil, São Paulo, fazenda Dalila, Registro." figureDoi-0="http://doi.org/10.5281/zenodo.10882080" figureDoi-1="http://doi.org/10.5281/zenodo.10882082" httpUri-0="https://zenodo.org/record/10882080/files/figure.png" httpUri-1="https://zenodo.org/record/10882082/files/figure.png" pageId="37" pageNumber="38">Figures 36-37</figureCitation>
for the approximate preserved and similar color patterns of adult males and females. A discrete lateral body stripe extends from just dorsal to the dorsalmost part of the gill opening to the caudal peduncle where it continues onto the caudal fin as a very solid dark wedge with its apex covering the two middle caudal-fin rays. The dark stripe consists of a dusting of dark chromatophores anterior to the pseudotympanum area and this continues posterior to this, merging with the dark lateral stripe that becomes dense with dark chromatophores external to the dorsal region of the abdomen. This stripe increases in width, and density posteriorly to the base of the caudal fin. In the photographs of a male and female (
<figureCitation id="13482A212A16FFCEFCEAF7954BC6F7F0" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="35.[207,272,2036,2082]" captionStartId-1="35.[205,267,2144,2188]" captionTargetBox-0="[210,2446,290,1130]" captionTargetBox-1="[206,2443,1144,1985]" captionTargetPageId-0="35" captionTargetPageId-1="35" captionText-0="Fig. 36. Spintherobolus leptoura, MZUSP 41854, holotype, adult male 19.1 mm SL, Brazil, São Paulo, fazenda Dalila, Registro." captionText-1="Fig. 37. Spintherobolus leptoura, MZUSP 41854, paratype, adult female 25.0 mm SL, Brazil, São Paulo, fazenda Dalila, Registro." figureDoi-0="http://doi.org/10.5281/zenodo.10882080" figureDoi-1="http://doi.org/10.5281/zenodo.10882082" httpUri-0="https://zenodo.org/record/10882080/files/figure.png" httpUri-1="https://zenodo.org/record/10882082/files/figure.png" pageId="37" pageNumber="38">Fig. 36- 37</figureCitation>
), there appears to be a dark humeral spot just posterior to the more dorsal region of the opercular flap. This is the area of the anterior and posterior pseudotympanums and the black pigment coating the swimbladder can be seen through the skin, appearing black in the print.
</paragraph>
<paragraph id="8BCC36A42A16FFCEFF54F6D14E21FD58" blockId="37.[167,979,1391,2605]" lastBlockId="37.[1034,1849,218,2560]" pageId="37" pageNumber="38">The scale borders above the lateral dark stripe are covered by a single row of dark chromatophores, giving the dorsal part of the body a retic ­ ulate pattern. Just ventral to the stripe the scale borders continue to be dark for about one to one half the width of a scale row. The belly and sides ventral to the lateral stripe are pale cream color or yellowish (the color of preserved muscle tissue) and the ventrolateral surface of the abdomen may be covered with dark chromatophores of varying density. The posterior termination of the caudal peduncle, both dorsal and ventral to the lateral stripe, is white, without dark pigment of any kind, except for the procurrent caudal-fin rays in this region which are covered with dark chromatophores.</paragraph>
<paragraph id="8BCC36A42A16FFCEFBF9FD4B4FD5F94F" blockId="37.[1034,1849,218,2560]" pageId="37" pageNumber="38">The pectoral and pelvic fins of both sexes are hyaline with scattered dark chromatophores, especially on the anterior undivided ray of both fins. The dorsal fin is more or less hyaline except for its anterior dorsal border where the anterior undivided rays and the first and second branched rays have numerous dark chromatophores. The rays posterior to these dark rays progressively have fewer dark chromatophores distally. The caudal fin is hyaline, especially for the proximal one third of its length. There is a thin scattering of small dark chromatophores over the rays and membranes, especially along their distal length. A second longitudinal dark, but relatively broad stripe covers the muscular anterior base of the fin and extends posteriorly to almost completely cover the short anal-fin rays that follow the large, elongate, anterior anal-fin lobe. The anterior lobe of the anal fin, distal to its muscular base is hyaline except for the basal one third the distal tips of the anterior unbranched and the first two branched rays. These are covered with dark chromatophores.</paragraph>
<paragraph id="8BCC36A42A16FFCEFBF8F95B4EF3F59B" blockId="37.[1034,1849,218,2560]" pageId="37" pageNumber="38">The head is pale brown around the mouth with a scattering of dark chromatophores over the ventral surface of the snout, jaws, and oper ­ cular region. The snout region, especially the jaws possess a scattering of dark chromatophores. The dorsal surface of the snout and the area between the eyes are pale brown, while the dorsum of the cranium and the nape are dark brown. The head area posterior to the eye and extending ventrally from the parietal region to the preopercle and branchiostegal rays is white except for some dark chromatophores in the dorsal region of the oper ­ cle. The circumorbital region ventral to the eye is dark, with a series of dark chromatophores, giving the impression of a spot ventral to the eye when these chromatophores are expanded. The opercular region is dark dorsally and pale brown to white below in both sexes.</paragraph>
<paragraph id="8BCC36A42A15FFCDFF9FFF4D483BFE42" blockId="38.[45,851,207,473]" pageId="38" pageNumber="39">Color in life. Life colors have not been recorded, but we expect that life colors are very much like those in preservative except that the dark stripes are usually black, the abdominal region white and the two white areas at the posterior termination of the caudal peduncle are intense white.</paragraph>
<paragraph id="8BCC36A42A15FFCDFF9EFD8E4859FBBF" blockId="38.[43,851,524,1060]" pageId="38" pageNumber="39">
Sexual dimorphism. The sexes are easily identified externally by the shape of anal fins (
<figureCitation id="13482A212A15FFCDFD7CFDBB4B5AFD17" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="35.[207,272,2036,2082]" captionStartId-1="35.[205,267,2144,2188]" captionTargetBox-0="[210,2446,290,1130]" captionTargetBox-1="[206,2443,1144,1985]" captionTargetPageId-0="35" captionTargetPageId-1="35" captionText-0="Fig. 36. Spintherobolus leptoura, MZUSP 41854, holotype, adult male 19.1 mm SL, Brazil, São Paulo, fazenda Dalila, Registro." captionText-1="Fig. 37. Spintherobolus leptoura, MZUSP 41854, paratype, adult female 25.0 mm SL, Brazil, São Paulo, fazenda Dalila, Registro." figureDoi-0="http://doi.org/10.5281/zenodo.10882080" figureDoi-1="http://doi.org/10.5281/zenodo.10882082" httpUri-0="https://zenodo.org/record/10882080/files/figure.png" httpUri-1="https://zenodo.org/record/10882082/files/figure.png" pageId="38" pageNumber="39">Figs. 36- 37</figureCitation>
). The distal border of the anal fin of females is concave and the anterior lobe has an acute distal tip. When this lobe is depressed it does not reach the posterior termination of the fin. The anterior lobe of the anal fin of males is largely rounded, with the tip of largest rays reaching beyond the tip of last anal-fin rays. The shape of first to fifth branched anal-fin rays, the modified ventral procurrent caudal-fin rays and the presence of hooks in the pelvic fins are also characteristic for males.
</paragraph>
</subSubSection>
<subSubSection id="C369652F2A15FFCDFF9CFBD44921FAA8" pageId="38" pageNumber="39" type="etymology">
<paragraph id="8BCC36A42A15FFCDFF9CFBD44921FAA8" blockId="38.[42,850,1110,1331]" pageId="38" pageNumber="39">
Etymology. The word
<taxonomicName id="4C734D272A15FFCDFE1BFBD4492CFBE5" box="[427,549,1110,1150]" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="leptoura">
<emphasis id="B907EAB62A15FFCDFE1BFBD4492CFBE5" box="[427,549,1110,1150]" italics="true" pageId="38" pageNumber="39">leptoura</emphasis>
</taxonomicName>
is from the Greek leptos meaning small, thin or delicate and oura, Greek for tail. The name is given in reference to the relatively slender caudal peduncle of this species. A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C369652F2A15FFCDFF9BFAE44AF3FA20" pageId="38" pageNumber="39" type="biology_ecology">
<paragraph id="8BCC36A42A15FFCDFF9BFAE44AF3FA20" blockId="38.[41,849,1382,1467]" pageId="38" pageNumber="39">Ecological notes. We have no ecological infor ­ mation about this species.</paragraph>
</subSubSection>
</treatment>
</document>
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