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<document id="406FC19D738F21146E296FDE04A36423" ID-DOI="10.1080/0022293021000055441" ID-ISSN="1464-5262" ID-Zenodo-Dep="4653568" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617217246595" checkinUser="carolina" docAuthor="Peart, R. A." docDate="2004" docId="03CD6212D57EFFFEFD90FE12EB98FDF1" docLanguage="en" docName="JNATHIST.38.301-336.pdf" docOrigin="Journal of Natural History 38 (3)" docSource="http://www.tandfonline.com/doi/abs/10.1080/0022293021000055441" docStyle="DocumentStyle:443144A73DAE3E092C51A378E4CD0138.2:JNatHist.2003-2004.journal_article.1cover" docStyleId="443144A73DAE3E092C51A378E4CD0138" docStyleName="JNatHist.2003-2004.journal_article.1cover" docStyleVersion="2" docTitle="Cymadusa vadosa Imbach 1967" docType="treatment" docVersion="4" lastPageNumber="334" masterDocId="FFF41A6AD560FFDCFFF0FFFBEA3AFFBD" masterDocTitle="A revision of the Cymadusa filosa complex (Crustacea: Amphipoda: Corophioidea: Ampithoidae)" masterLastPageNumber="336" masterPageNumber="301" pageNumber="330" updateTime="1698898288008" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="199ED1A860CB6D1B963E4962C70540BA">A revision of the Cymadusa filosa complex (Crustacea: Amphipoda: Corophioidea: Ampithoidae)</mods:title>
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<mods:date id="4A79A12EC72D9C7BA8CEEA7466148F00">2004</mods:date>
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<mods:number id="9CE7EFA45E425C4538892D320A2A06E2">2004-02-10</mods:number>
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<mods:number id="7456D61711FC96EEE9BB69AA97C9FD7E">3</mods:number>
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<subSubSection id="C37E808FD57EFFC2FD90FE12E957FD9C" pageId="30" pageNumber="330" type="nomenclature">
<paragraph id="8BDBD304D57EFFC2FD90FE12E9F6FDBC" blockId="30.[313,1105,489,604]" box="[608,972,489,513]" pageId="30" pageNumber="330">
<heading id="D0936468D57EFFC2FD90FE12E9F6FDBC" box="[608,972,489,513]" centered="true" fontSize="10" level="2" pageId="30" pageNumber="330" reason="3">
<taxonomicName id="4C64A887D57EFFC2FD90FE12E9F6FDBC" ID-CoL="335R7" authority="Imbach, 1967" authorityName="Imbach" authorityYear="1967" box="[608,972,489,513]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="30" pageNumber="330" phylum="Arthropoda" rank="species" species="vadosa">
<emphasis id="B9100F16D57EFFC2FD90FE12E912FDBC" bold="true" box="[608,808,489,513]" italics="true" pageId="30" pageNumber="330">Cymadusa vadosa</emphasis>
<bibRefCitation id="EFF5AEF5D57EFFC2FCC1FE12E9F6FDBC" author="IMBACH, M. C." box="[817,972,489,513]" pageId="30" pageNumber="330" pagination="39 - 167" refId="ref11465" refString="IMBACH, M. C., 1967, Gammaridean Amphipoda from the South China Sea, Naga Report, 4 (1), 39 - 167." type="journal article" year="1967">Imbach, 1967</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDBD304D57EFFC2FD31FDF2E957FD9C" blockId="30.[313,1105,489,604]" box="[705,877,521,545]" pageId="30" pageNumber="330">(figures 2022)</paragraph>
</subSubSection>
<subSubSection id="C37E808FD57EFFC2FEC9FDD0EE08FDE6" pageId="30" pageNumber="330" type="reference_group">
<paragraph id="8BDBD304D57EFFC2FEC9FDD0EE6BFDFD" blockId="30.[313,1105,489,604]" box="[313,1105,555,577]" pageId="30" pageNumber="330">
<treatmentCitationGroup id="AB74F42AD57EFFC2FEC9FDD0EE6BFDFD" box="[313,1105,555,577]" pageId="30" pageNumber="330">
<treatmentCitation id="0AC5F515D57EFFC2FEC9FDD0E8F5FDFD" author="IMBACH, M. C." box="[313,719,555,577]" page="90" pageId="30" pageNumber="330" year="1967">
<taxonomicName id="4C64A887D57EFFC2FEC9FDD0E8F5FDFD" ID-CoL="335R7" authority="Imbach, 1967: 89 - 90" authorityName="Imbach" authorityPageNumber="89 - 90" authorityYear="1967" box="[313,719,555,577]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="30" pageNumber="330" phylum="Arthropoda" rank="species" species="vadosa">
<emphasis id="B9100F16D57EFFC2FEC9FDD0EBD6FDFD" box="[313,492,555,576]" italics="true" pageId="30" pageNumber="330">Cymadusa vadosa</emphasis>
<bibRefCitation id="EFF5AEF5D57EFFC2FE05FDD0E8F5FDFD" author="IMBACH, M. C." box="[501,719,555,577]" pageId="30" pageNumber="330" pagination="39 - 167" refId="ref11465" refString="IMBACH, M. C., 1967, Gammaridean Amphipoda from the South China Sea, Naga Report, 4 (1), 39 - 167." type="journal article" year="1967">Imbach, 1967: 8990</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 32;
<treatmentCitation id="0AC5F515D57EFFC2FCD3FDD0EE77FDFD" author="POORE, A. G. B. &amp; LOWRY, J. K." box="[803,1101,555,577]" page="900" pageId="30" pageNumber="330" year="1997">
<bibRefCitation id="EFF5AEF5D57EFFC2FCD3FDD0EE77FDFD" author="POORE, A. G. B. &amp; LOWRY, J. K." box="[803,1101,555,577]" pageId="30" pageNumber="330" pagination="897 - 941" refId="ref11888" refString="POORE, A. G. B. and LOWRY, J. K., 1997, New ampithoid amphipods from Port Jackson, New South Wales, Australia (Crustacea: Amphipoda: Ampithoidae), Invertebrate Taxonomy, 11, 897 - 941." type="journal article" year="1997">Poore and Lowry, 1997: 900</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDBD304D57EFFC2FEC9FDBEEE08FDE6" blockId="30.[313,1105,489,604]" box="[313,1074,581,604]" pageId="30" pageNumber="330">
<treatmentCitationGroup id="AB74F42AD57EFFC2FEC9FDBEEE08FDE6" box="[313,1074,581,604]" pageId="30" pageNumber="330">
<treatmentCitation id="0AC5F515D57EFFC2FEC9FDBEE8F4FDE1" author="LEDOYER, M." box="[313,718,581,604]" page="18" pageId="30" pageNumber="330" year="1984">
<taxonomicName id="4C64A887D57EFFC2FEC9FDBEE8F4FDE1" ID-CoL="335PS" authority=": Ledoyer, 1984: 15 - 18" authorityName=": Ledoyer" authorityPageNumber="15 - 18" authorityYear="1984" box="[313,718,581,604]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="30" pageNumber="330" phylum="Arthropoda" rank="species" species="filosa">
<emphasis id="B9100F16D57EFFC2FEC9FDBEEBE4FDE7" box="[313,478,581,602]" italics="true" pageId="30" pageNumber="330">Cymadusa filosa</emphasis>
:
<bibRefCitation id="EFF5AEF5D57EFFC2FE1DFDBEE8F4FDE1" author="LEDOYER, M." box="[493,718,581,604]" pageId="30" pageNumber="330" pagination="137 - 181" refId="ref11630" refString="LEDOYER, M., 1984, Les Gammariens (Crustacea, Amphipoda) des herbiers de phanerogrames marines de Nouvelle Caledonie (region de Noumea), Memoires du Museum National d'Historie Naturelle Paris, Series 4, 1, 137 - 181." type="journal article" year="1984">Ledoyer, 1984: 1518</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figures 5, 6 (as Group I smooth).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37E808FD57EFFFEFE91FD85EB98FDF1" lastPageId="34" lastPageNumber="334" pageId="30" pageNumber="330" type="materials_examined">
<paragraph id="8BDBD304D57EFFC2FE91FD85E977FD0B" blockId="30.[313,1268,638,885]" pageId="30" pageNumber="330">
<emphasis id="B9100F16D57EFFC2FE91FD85E805FD2B" box="[353,575,638,662]" italics="true" pageId="30" pageNumber="330">Material examined.</emphasis>
MNHN, male,
<quantity id="4C9C7EE1D57EFFC2FCF1FD85E905FD2B" box="[769,831,638,662]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="30" pageNumber="330" unit="mm" value="8.0">8 mm</quantity>
in length, Noumea,
<collectingCountry id="F3739394D57EFFC2FBC9FD84EECAFD2B" box="[1081,1264,638,662]" name="New Caledonia" pageId="30" pageNumber="330">New Caledonia</collectingCountry>
, 22°9.6∞S, 166°16.2∞E, collected by M. Ledoyer.
</paragraph>
<paragraph id="8BDBD304D57EFFC2FE91FD44EE18FD6B" blockId="30.[313,1268,638,885]" box="[353,1058,702,726]" pageId="30" pageNumber="330">
<materialsCitation id="3B0CD959D57EFFC2FE91FD44EE24FD6B" box="[353,1054,702,726]" country="Vietnam" county="Sea" location="Bay of Nha-Trang" municipality="South" pageId="30" pageNumber="330" specimenCount="1" typeStatus="holotype">
<emphasis id="B9100F16D57EFFC2FE91FD44EBC1FD6B" box="[353,507,702,726]" italics="true" pageId="30" pageNumber="330">
<typeStatus id="54DF6DA6D57EFFC2FE91FD44EBA3FD6B" box="[353,409,703,726]" pageId="30" pageNumber="330">Type</typeStatus>
locality.
</emphasis>
<location id="8EBB85DFD57EFFC2FDF3FD45E8E1FD6B" LSID="urn:lsid:plazi:treatment:03CD6212D57EFFFEFD90FE12EB98FDF1:8EBB85DFD57EFFC2FDF3FD45E8E1FD6B" box="[515,731,702,726]" country="Vietnam" county="Sea" municipality="South" name="Bay of Nha-Trang" pageId="30" pageNumber="330">Bay of Nha-Trang</location>
,
<collectingCountry id="F3739394D57EFFC2FD18FD45E976FD6B" box="[744,844,702,726]" name="Vietnam" pageId="30" pageNumber="330">Vietnam</collectingCountry>
,
<collectingMunicipality id="6BBF497ED57EFFC2FCA9FD45E9A6FD6B" box="[857,924,702,726]" pageId="30" pageNumber="330">South</collectingMunicipality>
<collectingCountry id="F3739394D57EFFC2FC55FD45E9D0FD6B" box="[933,1002,702,726]" name="China" pageId="30" pageNumber="330">China</collectingCountry>
<collectingCounty id="62BAAB88D57EFFC2FC03FD45EE24FD6B" box="[1011,1054,702,726]" pageId="30" pageNumber="330">Sea</collectingCounty>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDBD304D57EFFC2FE91FD25E832FCC8" blockId="30.[313,1268,638,885]" pageId="30" pageNumber="330">
<emphasis id="B9100F16D57EFFC2FE91FD25EBEFFD48" box="[353,469,734,757]" italics="true" pageId="30" pageNumber="330">Diagnosis.</emphasis>
Antenna 1 with a two-articulate accessory flagellum. Antenna 2 with weakly setose margins. Gnathopod 1 merus produced to form a small, rounded anteroventral lobe; carpus broad and short, subtriangular. Gnathopod 2 propodal palm not defined by posterodistal tooth, with no mid-palmar tooth; dactylus subequal to palm.
</paragraph>
<paragraph id="8BDBD304D57EFFC2FEC9FC48EB82FC77" blockId="30.[313,1269,947,1736]" box="[313,440,947,970]" pageId="30" pageNumber="330">
<emphasis id="B9100F16D57EFFC2FEC9FC48EB82FC77" box="[313,440,947,970]" italics="true" pageId="30" pageNumber="330">Description</emphasis>
</paragraph>
<paragraph id="8BDBD304D57EFFC2FE91FC29E8F6FC57" blockId="30.[313,1269,947,1736]" box="[353,716,978,1002]" pageId="30" pageNumber="330">
Based on male,
<quantity id="4C9C7EE1D57EFFC2FDE9FC29E86DFC57" box="[537,599,978,1002]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="30" pageNumber="330" unit="mm" value="8.0">8 mm</quantity>
in length.
</paragraph>
<paragraph id="8BDBD304D57EFFC2FE91FC09EEDCFB54" blockId="30.[313,1269,947,1736]" pageId="30" pageNumber="330">
<emphasis id="B9100F16D57EFFC2FE91FC09E824FBB7" box="[353,542,1010,1034]" italics="true" pageId="30" pageNumber="330">Head. Antenna 1</emphasis>
longer than antenna 2; peduncular article 1 shorter than article 2, article 2 longer than article 3, article 3 shorter than article 1; primary flagellum incomplete but with at least seven articles; accessory flagellum with two articles.
<emphasis id="B9100F16D57EFFC2FEC9FBA9EB91FBD7" box="[313,427,1106,1130]" italics="true" pageId="30" pageNumber="330">Antenna 2</emphasis>
weakly setose on ventral margin; peduncular article 4 subequal to article 5.
<emphasis id="B9100F16D57EFFC2FEA5FB89EB85FB37" box="[341,447,1138,1162]" italics="true" pageId="30" pageNumber="330">Lower lip</emphasis>
outer plate outer lobe longer than inner lobe.
<emphasis id="B9100F16D57EFFC2FC21FB89EE00FB37" box="[977,1082,1138,1162]" italics="true" pageId="30" pageNumber="330">Mandible</emphasis>
molar setal row with eight robust setae; palp article 1 shorter than article 2, article 2 shorter than article 3, article 3 longer than article 1; long plumose setae on posterior margin and apex of article 3.
<emphasis id="B9100F16D57EFFC2FDF3FB2AE848FB54" box="[515,626,1233,1257]" italics="true" pageId="30" pageNumber="330">Maxilla 1</emphasis>
inner plate with five plumose setae.
<emphasis id="B9100F16D57EFFC2FBE9FB2AEEA8FB54" box="[1049,1170,1233,1257]" italics="true" pageId="30" pageNumber="330">Maxilliped</emphasis>
setose.
</paragraph>
<paragraph id="8BDBD304D57EFFC2FE91FB09EBB3F935" blockId="30.[313,1269,947,1736]" pageId="30" pageNumber="330">
<emphasis id="B9100F16D57EFFC2FE91FB09E868FAB4" box="[353,594,1265,1289]" italics="true" pageId="30" pageNumber="330">Pereon. Gnathopod 1</emphasis>
subequal to gnathopod 2; poorly setose; coxa produced forward, anterior margin straight, ventral margin with just a tuft of a few simple setae on posterior corner; basis subequal to the coxa, without long plumose setal fringe, distal lobe present; merus produced to form a short, subacute anteroventral lobe; carpus longer than merus, subequal to propodus, divergent at distal end, subtriangular; carpal lobe rounded; propodus ovoid; palm entire, convex, defining posterodistal tooth absent, mid-palmar tooth absent; dactylus longer than palm.
<emphasis id="B9100F16D57EFFC2FEC9FA2BEBF6FA54" box="[313,460,1488,1513]" italics="true" pageId="30" pageNumber="330">Gnathopod 2</emphasis>
weakly setose; coxa ventral margin with a tuft of simple setae on posterior corner; basis subequal in length to coxa, no setal fringe, distal lobe present; merus produced to form a small, rounded anteroventral lobe; carpus subequal to merus, shorter than propodus; propodus subquadrate; palm entire, slightly concave, defining posterodistal tooth absent, mid-palmar tooth absent; dactylus tapering evenly.
</paragraph>
<paragraph id="8BDBD304D57EFFC3FE91F96BE861F995" blockId="30.[313,1269,947,1736]" lastBlockId="31.[313,1268,1519,1736]" lastPageId="31" lastPageNumber="331" pageId="30" pageNumber="330">
<emphasis id="B9100F16D57EFFC2FE91F96BEBDFF915" box="[353,485,1680,1704]" italics="true" pageId="30" pageNumber="330">Pereopods 3</emphasis>
and
<emphasis id="B9100F16D57EFFC2FDECF96BE813F915" box="[540,553,1680,1704]" italics="true" pageId="30" pageNumber="330">4</emphasis>
similar in shape; coxae without setae.
<emphasis id="B9100F16D57EFFC2FC26F96BEE75F91A" box="[982,1103,1680,1704]" italics="true" pageId="30" pageNumber="330">Pereopod 5</emphasis>
coxae without setal fringe and medial setae.
<emphasis id="B9100F16D57EFFC2FD7EF94BE930F975" box="[654,778,1712,1736]" italics="true" pageId="30" pageNumber="330">Pereopod 6</emphasis>
basis with sparse medial setae.
<emphasis id="B9100F16D57EFFC2FB88F94BEECEF97A" box="[1144,1268,1712,1736]" italics="true" pageId="30" pageNumber="330">Pereopod 7</emphasis>
basis subrectangular.
<emphasis id="B9100F16D57FFFC3FDC7FA0BE8FBF9B5" box="[567,705,1520,1544]" italics="true" pageId="31" pageNumber="331">Pereopods 6</emphasis>
and
<emphasis id="B9100F16D57FFFC3FCF3FA0AE92AF9B5" box="[771,784,1521,1544]" italics="true" pageId="31" pageNumber="331">7</emphasis>
much longer than pereopods 35; distal articles with sparse setae.
</paragraph>
<caption id="DF1B838CD57FFFC3FEC9FA8EE986FA19" ID-DOI="http://doi.org/10.5281/zenodo.4653616" ID-Zenodo-Dep="4653616" httpUri="https://zenodo.org/record/4653616/files/figure.png" pageId="31" pageNumber="331" startId="31.[313,329,1397,1418]" targetBox="[317,1263,182,1366]" targetPageId="31">
<paragraph id="8BDBD304D57FFFC3FEC9FA8EE986FA19" blockId="31.[313,1268,1396,1445]" pageId="31" pageNumber="331">
FIG. 20.
<taxonomicName id="4C64A887D57FFFC3FE51FA8FE8CDFA37" authority="Imbach, 1967" authorityName="Imbach" authorityYear="1967" box="[417,759,1396,1418]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="31" pageNumber="331" phylum="Arthropoda" rank="species" species="vadosa">
<emphasis id="B9100F16D57FFFC3FE51FA8FE862FA34" box="[417,600,1396,1417]" italics="true" pageId="31" pageNumber="331">Cymadusa vadosa</emphasis>
<bibRefCitation id="EFF5AEF5D57FFFC3FD94FA8FE8CDFA37" author="IMBACH, M. C." box="[612,759,1396,1418]" pageId="31" pageNumber="331" pagination="39 - 167" refId="ref11465" refString="IMBACH, M. C., 1967, Gammaridean Amphipoda from the South China Sea, Naga Report, 4 (1), 39 - 167." type="journal article" year="1967">Imbach, 1967</bibRefCitation>
</taxonomicName>
. Male, 8 mm, MNHN, New Caledonia, Pacific Ocean. Scales represent 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BDBD304D57FFFFCFE91F9CBE84FF9B5" blockId="31.[313,1268,1519,1736]" lastBlockId="32.[313,1269,1488,1736]" lastPageId="32" lastPageNumber="332" pageId="31" pageNumber="331">
<emphasis id="B9100F16D57FFFC3FE91F9CBE806F9F5" box="[353,572,1584,1608]" italics="true" pageId="31" pageNumber="331">Pleon. Epimeron 3</emphasis>
subquadrate, with a small, subacute posterodistal tooth.
<emphasis id="B9100F16D57FFFC3FEC9F9ABEB9FF9D5" box="[313,421,1616,1640]" italics="true" pageId="31" pageNumber="331">Uropod 1</emphasis>
peduncle with 10 marginal robust setae; outer ramus longer than inner ramus.
<emphasis id="B9100F16D57FFFC3FE60F98BEBC1F935" box="[400,507,1648,1672]" italics="true" pageId="31" pageNumber="331">Uropod 2</emphasis>
outer ramus longer than inner ramus.
<emphasis id="B9100F16D57FFFC3FC34F98BEE15F935" box="[964,1071,1648,1672]" italics="true" pageId="31" pageNumber="331">Uropod 3</emphasis>
with three distal peduncular robust setae; outer ramus shorter than inner ramus, hooks both strongly recurved, setal fringe absent on lateral margin; inner ramus with both robust and slender setae.
<taxonomicName id="4C64A887D540FFFCFE29FA2BE81BFA55" authorityName="Pearse" authorityYear="1952" box="[473,545,1488,1512]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="32" pageNumber="332" phylum="Arthropoda" rank="genus">
<emphasis id="B9100F16D540FFFCFE29FA2BE81BFA55" box="[473,545,1488,1512]" italics="true" pageId="32" pageNumber="332">Telson</emphasis>
</taxonomicName>
truncated, with lateral and apical slender setae (not in oblique rows); apical cusps present.
</paragraph>
<caption id="DF1B838CD540FFFCFEC9FAA9E986FA3F" ID-DOI="http://doi.org/10.5281/zenodo.4653618" ID-Zenodo-Dep="4653618" httpUri="https://zenodo.org/record/4653618/files/figure.png" pageId="32" pageNumber="332" startId="32.[313,329,1362,1383]" targetBox="[316,1266,181,1328]" targetPageId="32">
<paragraph id="8BDBD304D540FFFCFEC9FAA9E986FA3F" blockId="32.[313,1268,1362,1410]" pageId="32" pageNumber="332">
FIG. 21.
<taxonomicName id="4C64A887D540FFFCFE50FAA9E8CCFAD5" authority="Imbach, 1967" authorityName="Imbach" authorityYear="1967" box="[416,758,1362,1384]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="32" pageNumber="332" phylum="Arthropoda" rank="species" species="vadosa">
<emphasis id="B9100F16D540FFFCFE50FAA9E86DFADA" box="[416,599,1362,1383]" italics="true" pageId="32" pageNumber="332">Cymadusa vadosa</emphasis>
<bibRefCitation id="EFF5AEF5D540FFFCFD93FAA9E8CCFAD5" author="IMBACH, M. C." box="[611,758,1362,1384]" pageId="32" pageNumber="332" pagination="39 - 167" refId="ref11465" refString="IMBACH, M. C., 1967, Gammaridean Amphipoda from the South China Sea, Naga Report, 4 (1), 39 - 167." type="journal article" year="1967">Imbach, 1967</bibRefCitation>
</taxonomicName>
. Male, 8 mm, MNHN, New Caledonia, Pacific Ocean. Scales represent 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BDBD304D540FFFDFE91F9EAE8FCFA55" blockId="32.[313,1269,1488,1736]" lastBlockId="33.[313,1268,1456,1512]" lastPageId="33" lastPageNumber="333" pageId="32" pageNumber="332">
<emphasis id="B9100F16D540FFFCFE91F9EAEBEAF995" box="[353,464,1553,1576]" italics="true" pageId="32" pageNumber="332">Variation.</emphasis>
Based on the above-documented animal, however, there is some leftright dimorphism present. This is probably just an artefact, most likely the gnathopod is damaged. Left gnathopod 2 weakly setose; coxa ventral margin with a tuft of simple setae on posterior corner; basis subequal in length to coxa but no setal fringe; merus produced to form a small, subacute anteroventral lobe; carpus subequal to merus, shorter than propodus; propodus subquadrate; palm entire, convex, defined by a small, subacute posterodistal tooth, mid-palmar tooth absent; dactylus subequal in length to palm, tapering evenly.
</paragraph>
<caption id="DF1B838CD541FFFDFEC9FABBEEE0FAD2" ID-DOI="http://doi.org/10.5281/zenodo.4653620" ID-Zenodo-Dep="4653620" httpUri="https://zenodo.org/record/4653620/files/figure.png" pageId="33" pageNumber="333" startId="33.[313,329,1344,1365]" targetBox="[316,1266,181,1312]" targetPageId="33">
<paragraph id="8BDBD304D541FFFDFEC9FABBEEE0FAD2" blockId="33.[313,1268,1343,1392]" pageId="33" pageNumber="333">
FIG. 22.
<taxonomicName id="4C64A887D541FFFDFE51FAC4E8CDFAE8" authority="Imbach, 1967" authorityName="Imbach" authorityYear="1967" box="[417,759,1343,1365]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="33" pageNumber="333" phylum="Arthropoda" rank="species" species="vadosa">
<emphasis id="B9100F16D541FFFDFE51FAC4E862FAE9" box="[417,600,1343,1364]" italics="true" pageId="33" pageNumber="333">Cymadusa vadosa</emphasis>
<bibRefCitation id="EFF5AEF5D541FFFDFD94FAC4E8CDFAE8" author="IMBACH, M. C." box="[612,759,1343,1365]" pageId="33" pageNumber="333" pagination="39 - 167" refId="ref11465" refString="IMBACH, M. C., 1967, Gammaridean Amphipoda from the South China Sea, Naga Report, 4 (1), 39 - 167." type="journal article" year="1967">Imbach, 1967</bibRefCitation>
</taxonomicName>
. Male, 8 mm, MNHN, New Caledonia, Pacific Ocean. Scales represent 0.5 mm for P57 and EP; scales represent 0.2 mm for U13, T.
</paragraph>
</caption>
<paragraph id="8BDBD304D541FFFDFEC9F9EBEBA6F995" blockId="33.[313,1269,1552,1736]" box="[313,412,1552,1576]" pageId="33" pageNumber="333">
<emphasis id="B9100F16D541FFFDFEC9F9EBEBA6F995" box="[313,412,1552,1576]" italics="true" pageId="33" pageNumber="333">Remarks</emphasis>
</paragraph>
<paragraph id="8BDBD304D541FFFEFE91F9CBE853FEF0" blockId="33.[313,1269,1552,1736]" lastBlockId="34.[313,1269,181,588]" lastPageId="34" lastPageNumber="334" pageId="33" pageNumber="333">
<bibRefCitation id="EFF5AEF5D541FFFDFE91F9CBE83DF9F5" author="LEDOYER, M." box="[353,519,1584,1608]" pageId="33" pageNumber="333" pagination="137 - 181" refId="ref11630" refString="LEDOYER, M., 1984, Les Gammariens (Crustacea, Amphipoda) des herbiers de phanerogrames marines de Nouvelle Caledonie (region de Noumea), Memoires du Museum National d'Historie Naturelle Paris, Series 4, 1, 137 - 181." type="journal article" year="1984">Ledoyer, 1984</bibRefCitation>
recorded this species as a form of
<taxonomicName id="4C64A887D541FFFDFC4AF9CBEE4CF9F5" authorityName=": Ledoyer" authorityYear="1984" box="[954,1142,1584,1608]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="33" pageNumber="333" phylum="Arthropoda" rank="species" species="filosa">
<emphasis id="B9100F16D541FFFDFC4AF9CBEE4CF9F5" box="[954,1142,1584,1608]" italics="true" pageId="33" pageNumber="333">Cymadusa filosa</emphasis>
</taxonomicName>
that most closely resembles
<taxonomicName id="4C64A887D541FFFDFDE2F9ABE8D8F9D5" authorityName="Imbach" authorityYear="1967" box="[530,738,1616,1640]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="33" pageNumber="333" phylum="Arthropoda" rank="species" species="vadosa">
<emphasis id="B9100F16D541FFFDFDE2F9ABE8D8F9D5" box="[530,738,1616,1640]" italics="true" pageId="33" pageNumber="333">Cymadusa vadosa</emphasis>
</taxonomicName>
. Both records are around
<quantity id="4C9C7EE1D541FFFDFBCAF9ABEE42F9DA" box="[1082,1144,1616,1640]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="33" pageNumber="333" unit="mm" value="8.0">8 mm</quantity>
in length (Imbachs is 8.5 mm) and are therefore easy to contrast. When compared to Imbachs original description from Bay of Nha-Trang,
<collectingCountry id="F3739394D541FFFDFCC1F96BE9ABF915" box="[817,913,1680,1704]" name="Vietnam" pageId="33" pageNumber="333">Vietnam</collectingCountry>
there are only a few differences. These include article 1 of antenna 1 subequal in length to article
<quantity id="4C9C7EE1D541FFFDFBA1F94BEEBAF975" box="[1105,1152,1712,1736]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="33" pageNumber="333" unit="in" value="2.0">2 in</quantity>
Imbachs animals and shorter than article
<quantity id="4C9C7EE1D542FFFEFD3DFF4EE8C5FF70" box="[717,767,181,205]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="34" pageNumber="334" unit="in" value="2.0">2 in</quantity>
Ledoyers animal; accessory flagellum of antenna 1 with one long article in Imbachs animal compared to two articles (Ledoyers animal); uropod 3 with more robust setae on the peduncle and inner ramus than on Ledoyers animal; epimeron 3 more rounded on Imbachs animal than on Ledoyers animal.
</paragraph>
<paragraph id="8BDBD304D542FFFEFE91FEAFEBECFE51" blockId="34.[313,1269,181,588]" pageId="34" pageNumber="334">
These differences are slight enough to be variations of the same species. Interestingly,
<bibRefCitation id="EFF5AEF5D542FFFEFE26FE8FE8BDFE31" author="LEDOYER, M." box="[470,647,372,396]" pageId="34" pageNumber="334" pagination="137 - 181" refId="ref11630" refString="LEDOYER, M., 1984, Les Gammariens (Crustacea, Amphipoda) des herbiers de phanerogrames marines de Nouvelle Caledonie (region de Noumea), Memoires du Museum National d'Historie Naturelle Paris, Series 4, 1, 137 - 181." type="journal article" year="1984">Ledoyer (1984)</bibRefCitation>
reported that there was leftright side dimorphism on the gnathopod 2 (noted in detail in the description). This could be just an artefact or damage as this
<typeStatus id="54DF6DA6D542FFFEFDE2FE4EE878FE70" box="[530,578,437,461]" pageId="34" pageNumber="334">type</typeStatus>
of dimorphism has not been recorded before in the family
<taxonomicName id="4C64A887D542FFFEFEC9FE2FEBE8FE51" box="[313,466,468,492]" class="Malacostraca" family="Ampithoidae" kingdom="Animalia" order="Amphipoda" pageId="34" pageNumber="334" phylum="Arthropoda" rank="family">Ampithoidae</taxonomicName>
.
</paragraph>
<paragraph id="8BDBD304D542FFFEFE91FE0FE9B0FDB1" blockId="34.[313,1269,181,588]" box="[353,906,500,524]" pageId="34" pageNumber="334">
<emphasis id="B9100F16D542FFFEFE91FE0FE8B5FDB1" box="[353,655,500,524]" italics="true" pageId="34" pageNumber="334">
Habitat.
<taxonomicName id="4C64A887D542FFFEFE37FE0FE8B5FDB1" authorityName="Imbach" authorityYear="1967" box="[455,655,500,524]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="34" pageNumber="334" phylum="Arthropoda" rank="species" species="vadosa">Cymadusa vadosa</taxonomicName>
</emphasis>
occurs on seagrasses.
</paragraph>
<paragraph id="8BDBD304D542FFFEFE91FDEFEB98FDF1" blockId="34.[313,1269,181,588]" pageId="34" pageNumber="334">
<emphasis id="B9100F16D542FFFEFE91FDEFEBD4FD91" box="[353,494,532,556]" italics="true" pageId="34" pageNumber="334">Distribution.</emphasis>
Bay of Nha-Trang,
<collectingCountry id="F3739394D542FFFEFD1CFDEFE96AFD91" box="[748,848,532,556]" name="Vietnam" pageId="34" pageNumber="334">Vietnam</collectingCountry>
, South
<collectingCountry id="F3739394D542FFFEFC41FDEFE9CCFD91" box="[945,1014,532,556]" name="China" pageId="34" pageNumber="334">China</collectingCountry>
Sea;
<collectingCountry id="F3739394D542FFFEFBCFFDEFEECEFD91" box="[1087,1268,532,556]" name="New Caledonia" pageId="34" pageNumber="334">New Caledonia</collectingCountry>
(
<figureCitation id="135FCF81D542FFFEFEB1FDCFEBB5FDF1" box="[321,399,564,588]" captionStart="MAP" captionStartId="11.[313,337,861,882]" captionTargetBox="[345,1236,179,831]" captionTargetId="figure-8@11.[343,1239,177,834]" captionTargetPageId="11" captionText="MAP 1. Global distribution of Cymadusa filosa Savigny, 1816 (*), Cymadusa ledoyeri n. sp. (•), Cymadusa tattersalli n. sp. (*), Cymadusa imbroglio Rabindranath, 1972 (Δ), Cymadusa setosa (Haswell, 1879) (v), Cymadusa vadosa Imbach, 1967 (&amp;)." figureDoi="http://doi.org/10.5281/zenodo.4653578" httpUri="https://zenodo.org/record/4653578/files/figure.png" pageId="34" pageNumber="334">Map 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>