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<document id="1BCD57999A07511F7F5EED50F2E6F619" ID-CLB-Dataset="22090" ID-DOI="10.5852/ejt.2017.331" ID-GBIF-Dataset="acf565ee-29ba-45e7-9ef3-d1e85dec9ae7" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829776" ID-ZooBank="3FD5D922-B91F-42BB-9D7B-7DFB98841732" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589572797790" checkinUser="carolina" docAuthor="Ikeda, Hiroshi &amp; Yoshitomi, Hiroyuki" docDate="2017" docId="4A5D70554515FFFAFDACBB4CFB1CFB5F" docLanguage="en" docName="ejt-2017-331.pdf.imf" docOrigin="European Journal of Taxonomy 331" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Intybia takaraensis" docType="treatment" docVersion="7" lastPageNumber="25" masterDocId="B664082D4503FFE2FFE6BC65FF8CFFBE" masterDocTitle="Revision of the genus Intybia (Coleoptera, Malachiidae) from Japan" masterLastPageNumber="31" masterPageNumber="1" pageNumber="23" updateTime="1698839968837" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6B545B89F9D9EC690851FCF91FFC8643">Revision of the genus Intybia (Coleoptera, Malachiidae) from Japan</mods:title>
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<mods:namePart id="892179C283AE7F5295BB331B325D4C95">Ikeda, Hiroshi</mods:namePart>
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<mods:namePart id="26A3D4547A852999C8D0F20112495028">Yoshitomi, Hiroyuki</mods:namePart>
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<mods:affiliation id="2A6F0FAC6C02DDFCF0BFD60D45D94419">Email: hymushi @ agr. ehime-u. ac. jp &amp; urn: lsid: zoobank. org: author: 744 FA 6 FC- 2 D 84 - 41 B 7 - 8 A 6 C-C 1611 CC 68 EE 1</mods:affiliation>
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<treatment id="4A5D70554515FFFAFDACBB4CFB1CFB5F" ID-DOI="http://doi.org/10.5281/zenodo.3848100" ID-GBIF-Taxon="164161042" ID-Zenodo-Dep="3848100" LSID="urn:lsid:plazi:treatment:4A5D70554515FFFAFDACBB4CFB1CFB5F" httpUri="http://treatment.plazi.org/id/4A5D70554515FFFAFDACBB4CFB1CFB5F" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
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<taxonomicName id="05F4BAC04515FFF4FDACBB4CFC65F8FD" authority="(Nakane, 1955)" baseAuthorityName="Nakane" baseAuthorityYear="1955" box="[586,1001,1832,1859]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="takaraensis">
<emphasis id="F0801D514515FFF4FDACBB4CFCA7F8FD" box="[586,811,1833,1859]" italics="true" pageId="22" pageNumber="23">Intybia takaraensis</emphasis>
(Nakane, 1955)
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<paragraph id="C24BC1434515FFF4FDF8BB29FB90F8D8" blockId="22.[542,1052,1832,1930]" box="[542,1052,1867,1894]" pageId="22" pageNumber="23">[Japanese name: Ruri-ki-obi-jôkai-modoki]</paragraph>
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<subSubSection id="8AEE92C84515FFF4FDC8BB0AFB8AF837" box="[558,1030,1903,1930]" pageId="22" pageNumber="23" type="description">
<paragraph id="C24BC1434515FFF4FDC8BB0AFB8AF837" blockId="22.[542,1052,1832,1930]" box="[558,1030,1903,1930]" pageId="22" pageNumber="23">
<figureCitation id="5ACFDDC64515FFF4FDC8BB0AFD0AF837" box="[558,646,1903,1929]" captionStart="Fig" captionStartId="4.[189,232,1893,1919]" captionTargetBox="[198,1389,740,1831]" captionTargetId="figure@4.[189,1398,723,1853]" captionTargetPageId="4" captionText="Fig. 1. Dorsal habitus Intybia spp., ♂. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov., holotype. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956), holotype. Scales = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3829778" httpUri="https://zenodo.org/record/3829778/files/figure.png" pageId="22" pageNumber="23">Figs 1F</figureCitation>
,
<figureCitation id="5ACFDDC64515FFF4FD75BB0AFD3EF837" box="[659,690,1903,1929]" captionStart="Fig" captionStartId="5.[190,233,1894,1920]" captionTargetBox="[195,1390,781,1842]" captionTargetId="figure@5.[190,1399,771,1854]" captionTargetPageId="5" captionText="Fig. 2. Dosal habitus of Intybia spp., ♀. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov., paratype. D. I. pelegrini pelegrini (Pic, 1910). E. I.niponica (Lewis, 1895). F. I.takaraensis (Nakane, 1955).G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956), paratype. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3829782" httpUri="https://zenodo.org/record/3829782/files/figure.png" pageId="22" pageNumber="23">2F</figureCitation>
,
<figureCitation id="5ACFDDC64515FFF4FD58BB0AFD52F837" box="[702,734,1903,1929]" captionStart="Fig" captionStartId="7.[190,233,1895,1921]" captionTargetBox="[265,1321,271,1836]" captionTargetId="figure@7.[246,1331,265,1859]" captionTargetPageId="7" captionText="Fig. 4. Right antennae of Intybia spp., ♂. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829786" httpUri="https://zenodo.org/record/3829786/files/figure.png" pageId="22" pageNumber="23">4F</figureCitation>
,
<figureCitation id="5ACFDDC64515FFF4FD0FBB0AFC85F834" box="[745,777,1903,1930]" captionStart="Fig" captionStartId="8.[190,233,1894,1920]" captionTargetBox="[285,1298,276,1852]" captionTargetId="figure@8.[273,1302,265,1859]" captionTargetPageId="8" captionText="Fig. 5. Right antennae of Intybia spp., ♀. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829788" httpUri="https://zenodo.org/record/3829788/files/figure.png" pageId="22" pageNumber="23">5F</figureCitation>
,
<figureCitation id="5ACFDDC64515FFF4FCF3BB0AFCB8F837" box="[789,820,1903,1929]" captionStart="Fig" captionStartId="9.[190,233,1895,1921]" captionTargetBox="[263,1324,265,1858]" captionTargetId="figure@9.[263,1324,265,1859]" captionTargetPageId="9" captionText="Fig. 6. SEM photographs of right male antennomere III of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C, H. I. donan sp. nov., showing projection (H). D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955)." figureDoi="http://doi.org/10.5281/zenodo.3829790" httpUri="https://zenodo.org/record/3829790/files/figure.png" pageId="22" pageNumber="23">6F</figureCitation>
,
<figureCitation id="5ACFDDC64515FFF4FCA6BB0AFCE8F837" box="[832,868,1903,1929]" captionStart="Fig" captionStartId="11.[190,233,1928,1954]" captionTargetBox="[192,1397,1222,1886]" captionTargetId="figure@11.[192,1400,1222,1886]" captionTargetPageId="11" captionText="Fig. 8. Endophallus of Intybia spp., A. Intybia histrio (Kiesenwetter, 1874). B. I. pelegrini pelegrini (Pic, 1910). C. I. takaraensis (Nakane, 1955). D. I. kishiii (Nakane, 1955). Abbreviations: gp = gonoporal piece; l = ligula; sgp = semi-gonoporal piece; sp = spinous plate. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829794" httpUri="https://zenodo.org/record/3829794/files/figure.png" pageId="22" pageNumber="23">8C</figureCitation>
,
<figureCitation id="5ACFDDC64515FFF4FC97BB0AFC1CF837" box="[881,912,1903,1929]" captionStart="Fig" captionStartId="12.[190,233,1893,1919]" captionTargetBox="[191,1394,543,1850]" captionTargetId="figure@12.[189,1398,538,1851]" captionTargetPageId="12" captionText="Fig. 9. Spinous area and spinous plate of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica(Lewis,1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955).Abbreviations: sa = spinous area; sp = spinous plate. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829796" httpUri="https://zenodo.org/record/3829796/files/figure.png" pageId="22" pageNumber="23">9F</figureCitation>
,
<figureCitation id="5ACFDDC64515FFF4FC7ABB0AFC47F837" box="[924,971,1903,1929]" captionStart="Fig" captionStartId="13.[190,233,1893,1919]" captionTargetBox="[216,1374,635,1827]" captionTargetId="figure@13.[188,1397,614,1853]" captionTargetPageId="13" captionText="Fig. 10. Endophallic sclerites of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). Abbreviations: gp = gonoporal piece; l = ligula; sgp = semi-gonoporal piece. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829798" httpUri="https://zenodo.org/record/3829798/files/figure.png" pageId="22" pageNumber="23">10F</figureCitation>
,
<figureCitation id="5ACFDDC64515FFF4FC30BB0AFB8AF837" box="[982,1030,1903,1929]" captionStart="Fig" captionStartId="15.[190,233,1895,1921]" captionTargetBox="[196,1385,1174,1842]" captionTargetId="figure@15.[192,1400,1167,1854]" captionTargetPageId="15" captionText="Fig. 11.Map showing thedistribution of Intybia spp.A.The histriogroup:●:Intybia histrio(Kiesenwetter, 1874), ▲: I. tsushimensis (Satô &amp; Ohbayashi, 1968), ■: I. donan sp. nov.). B. The pelegrini group: ○: I. pelegrini (Pic, 1910), △: I. niponica (Lewis, 1895), □: I. takaraensis (Nakane, 1955), ×: I. kishiii (Nakane, 1955), +: I. kawasakii (Nakane, 1956)." figureDoi="http://doi.org/10.5281/zenodo.3829800" httpUri="https://zenodo.org/record/3829800/files/figure.png" pageId="22" pageNumber="23">11B</figureCitation>
</paragraph>
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<treatmentCitation id="4355E7524515FFF4FF5BBBD4FD0DF874" author="Nakane T." box="[189,641,1968,1995]" page="376" pageId="22" pageNumber="23" year="1955">
<taxonomicName id="05F4BAC04515FFF4FF5BBBD4FD0DF874" authority="Nakane, 1955 b: 376" authorityName="Nakane" authorityPageNumber="376" authorityYear="1955" box="[189,641,1968,1995]" class="Insecta" family="Melyridae" genus="Laius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="takaraensis">
<emphasis id="F0801D514515FFF4FF5BBBD4FE01F874" box="[189,397,1968,1994]" italics="true" pageId="22" pageNumber="23">Laius takaraensis</emphasis>
<bibRefCitation id="A665BCB24515FFF4FE72BBD5FD0DF874" author="Nakane T." box="[404,641,1968,1995]" pageId="22" pageNumber="23" pagination="373 - 378" refId="ref17772" refString="Nakane T. 1955 b. Marine insects of the Tokara Islands V. Three new species of the genus Laius Guerin from Kyusyu and the Tokara Islands, with notes on a species from Mariana. Publication of the Seto Mariana Biological Laboratory 4 (2 - 3): 373 - 378." type="journal article" year="1955">Nakane, 1955b: 376</bibRefCitation>
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[
<typeStatus id="1D4F7FE14515FFF4FD69BBD4FD48F875" box="[655,708,1969,1995]" pageId="22" pageNumber="23">type</typeStatus>
locality: Takarajima Is., Tokara Isls.].
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<taxonomicName id="05F4BAC04514FFF5FF5BBD68FE00FE99" authorityName="Nakane" authorityYear="1955" box="[189,396,269,295]" class="Insecta" family="Melyridae" genus="Laius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="takaraensis">
<emphasis id="F0801D514514FFF5FF5BBD68FE00FE99" box="[189,396,269,295]" italics="true" pageId="23" pageNumber="24">Laius takaraensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="4355E7524514FFF5FE4EBD69FDFCFE99" author="Nakane T." box="[424,624,268,295]" page="13" pageId="23" pageNumber="24" year="1957">
<bibRefCitation id="A665BCB24514FFF5FE4EBD69FDFCFE99" author="Nakane T." box="[424,624,268,295]" pageId="23" pageNumber="24" pagination="13 - 14" refId="ref17872" refString="Nakane T. 1957. On Japanese Malachiinae (Coleoptera, Melyrdae) (2). Akitsu 6: 13 - 14." type="journal article" year="1957">Nakane 1957: 13</bibRefCitation>
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, fig. 4. —
<treatmentCitation id="4355E7524514FFF5FD0CBD69FC4FFE98" author="Nakane T." box="[746,963,268,294]" page="186" pageId="23" pageNumber="24" year="1963">
<bibRefCitation id="A665BCB24514FFF5FD0CBD69FC4FFE98" author="Nakane T." box="[746,963,268,294]" pageId="23" pageNumber="24" pagination="186" refId="ref17896" refString="Nakane T. 1963. Melyridae. In: Nakane T. Ohbayashi K. &amp; Kurosawa Y. (eds), Iconographia Insectorum Japonicorum Colore Naturali Edita 2: 186, pl. 93. Hokuryu-kan. [In Japanese.]" type="journal article" year="1963">Nakane 1963: 186</bibRefCitation>
</treatmentCitation>
, pl. 93, fig. 13. —
<treatmentCitation id="4355E7524514FFF5FB7FBD69FAFEFE98" author="Nakane T." box="[1177,1394,268,295]" page="164" pageId="23" pageNumber="24" year="1983">
<bibRefCitation id="A665BCB24514FFF5FB7FBD69FAFEFE98" author="Nakane T." box="[1177,1394,268,295]" pageId="23" pageNumber="24" pagination="161 - 166" refId="ref17940" refString="Nakane T. 1983. Notes on Japanese Melyridae (Coleoptera). Kitakyushu no Kontyu 30 (3): 161 - 166." type="journal article" year="1983">Nakane 1983: 164</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="4355E7524514FFF5FEE1BD55FE30FEF4" author="Sato M." box="[263,444,304,331]" page="164" pageId="23" pageNumber="24" year="1985">
<bibRefCitation id="A665BCB24514FFF5FEE1BD55FE30FEF4" author="Sato M." box="[263,444,304,331]" pageId="23" pageNumber="24" pagination="164 - 165" refId="ref18206" refString="Sato M. 1985. Melyridae. In: Ueno S. Kurosawa Y. &amp; Sato M. (eds), The Coleoptera of Japan in Color, 3: 164 - 165, pl. 26. Hoikusha, Osaka. [In Japanese.]" type="journal article" year="1985">Satô 1985: 164</bibRefCitation>
</treatmentCitation>
, pl. 26, fig. 7. —
<treatmentCitation id="4355E7524514FFF5FD6FBD55FCB3FEF4" author="Sato M." box="[649,831,304,330]" page="366" pageId="23" pageNumber="24" year="1989">
<bibRefCitation id="A665BCB24514FFF5FD6FBD55FCB3FEF4" author="Sato M." box="[649,831,304,330]" pageId="23" pageNumber="24" pagination="366" refId="ref18255" refString="Sato M. 1989. Melyridae. In: Hirahima Y. (et al.) A Check List of Japanese Insects 1: 366. [In Japanese.]" type="journal article" year="1989">Satô 1989: 366</bibRefCitation>
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.
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</paragraph>
<paragraph id="C24BC1434514FFF5FF5BBD36FB03FE2E" blockId="23.[189,1399,268,436]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="E2E4E66D4514FFF5FF5BBD36FB03FE2E" pageId="23" pageNumber="24">
<taxonomicName id="05F4BAC04514FFF5FF5BBD36FE11FED3" baseAuthorityName="Nakane" baseAuthorityYear="1955" box="[189,413,339,365]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="takaraensis">
<emphasis id="F0801D514514FFF5FF5BBD36FE11FED3" box="[189,413,339,365]" italics="true" pageId="23" pageNumber="24">Intybia takaraensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="4355E7524514FFF5FE51BD36FD18FED3" author="Wittmer W." box="[439,660,339,365]" page="181" pageId="23" pageNumber="24" year="1997">
<bibRefCitation id="A665BCB24514FFF5FE51BD36FD18FED3" author="Wittmer W." box="[439,660,339,365]" pageId="23" pageNumber="24" pagination="181 - 211" refId="ref18419" refString="Wittmer W. 1997. Zur Kenntnis der Gattungen Intybia Pascoe und Stenolaius Wittmer (Coleoptera, Malachiidae). Japanese Journal of Systematic Entomology 3 (2): 181 - 211." type="journal article" year="1997">Wittmer 1997: 181</bibRefCitation>
</treatmentCitation>
, abb. 39–46. —
<treatmentCitation id="4355E7524514FFF5FCB7BD31FB95FED3" author="Mayor A. J." box="[849,1049,339,366]" page="417" pageId="23" pageNumber="24" year="2007">
<bibRefCitation id="A665BCB24514FFF5FCB7BD31FB95FED3" author="Mayor A. J." box="[849,1049,339,366]" pageId="23" pageNumber="24" pagination="415 - 454" refId="ref17633" refString="Mayor A. J. 2007. Malachiidae. In: Lobl I. &amp; Smetana A. (ed.) Catalogue of Palaeartic Coleoptera, Vol. 4, pp. 415 - 454. Apollo Books, Vester Skerninge. https: // doi. org / 10.1163 / 9789004260894" type="journal article" year="2007">Mayor 2007: 417</bibRefCitation>
</treatmentCitation>
[catalogued]. —
<treatmentCitation id="4355E7524514FFF5FB07BD36FE3FFE2E" author="Yoshitomi H. &amp; Hayashi N." page="19" pageId="23" pageNumber="24" year="2011">
<bibRefCitation id="A665BCB24514FFF5FB07BD36FE3FFE2E" author="Yoshitomi H. &amp; Hayashi N." pageId="23" pageNumber="24" pagination="18 - 24" refId="ref18569" refString="Yoshitomi H. &amp; Hayashi N. 2011. List of the family Melyridae and its related families in Japan. Sayabane, New Series, Tokyo (2): 18 - 24. [In Japanese.]" type="journal article" year="2011">Yoshitomi &amp; Hayashi 2011: 19</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="4355E7524514FFF5FE05BD13FCBEFE2F" author="Tshernyshev S. E." box="[483,818,374,401]" page="575" pageId="23" pageNumber="24" year="2012">
<bibRefCitation id="A665BCB24514FFF5FE05BD13FCBEFE2F" author="Tshernyshev S. E." box="[483,818,374,401]" pageId="23" pageNumber="24" pagination="575 - 587" refId="ref18358" refString="Tshernyshev S. E. 2012. A review of soft-winged flower beetle fauna (Coleoptera, Malachiidae) of North Asia. Eurosian Entomological Journal 11 (6): 575 - 587." type="journal article" year="2012">Tshernyshev 2012: 575–587</bibRefCitation>
</treatmentCitation>
, table 1. —
<treatmentCitation id="4355E7524514FFF5FC26BD13FB07FE2E" author="Plonski I. S." box="[960,1163,374,400]" page="66" pageId="23" pageNumber="24" year="2013">
<bibRefCitation id="A665BCB24514FFF5FC26BD13FB07FE2E" author="Plonski I. S." box="[960,1163,374,400]" pageId="23" pageNumber="24" pagination="61 - 68" refId="ref18033" refString="Plonski I. S. 2013. Studies on the genus Intybia Pascoe (Coleoptera: Malachiidae) I. Some nomenclatorial acts and faunistic records. Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen 65: 61 - 68." type="journal article" year="2013">Plonski 2013: 66</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C24BC1434514FFF5FF5BBDFFFC8AFE0A" blockId="23.[189,1399,268,436]" box="[189,774,409,436]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="E2E4E66D4514FFF5FF5BBDFFFC8AFE0A" box="[189,774,409,436]" pageId="23" pageNumber="24">
<treatmentCitation id="4355E7524514FFF5FF5BBDFFFDD0FE0A" author="Nakane T." box="[189,604,409,436]" page="33" pageId="23" pageNumber="24" year="1956">
<taxonomicName id="05F4BAC04514FFF5FF5BBDFFFDD0FE0A" authority="Nakane 1956: 33" authorityName="Nakane" authorityPageNumber="33" authorityYear="1956" box="[189,604,409,436]" class="Insecta" family="Melyridae" genus="Laius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="tokaraensis">
<emphasis id="F0801D514514FFF5FF5BBDFFFE01FE0D" box="[189,397,409,435]" italics="true" pageId="23" pageNumber="24">Laius tokaraensis</emphasis>
<bibRefCitation id="A665BCB24514FFF5FE72BDFCFDD0FE0A" author="Nakane T." box="[404,604,409,436]" pageId="23" pageNumber="24" pagination="159 - 174" refId="ref17827" refString="Nakane T. 1956. New or little-known Coleoptera from Japan and its abject regions, XIII. The scientific Reports of the Saikyo University (Natural Science and Living Science) (A) 2: 29 - 44, 159 - 174." type="journal article" year="1956">Nakane 1956: 33</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[misspelling].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="8AEE92C84514FFF5FF5BBDBEFED3FD33" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="C24BC1434514FFF5FF5BBDBEFEBAFE4B" blockId="23.[189,310,475,501]" box="[189,310,475,501]" pageId="23" pageNumber="24">
<heading id="9903762F4514FFF5FF5BBDBEFEBAFE4B" bold="true" box="[189,310,475,501]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="F0801D514514FFF5FF5BBDBEFEBAFE4B" bold="true" box="[189,310,475,501]" pageId="23" pageNumber="24">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC1434514FFF5FF5BBE6CFED3FD33" blockId="23.[189,1399,521,653]" pageId="23" pageNumber="24">
Coloration of body black with bluish luster on head, pronotum and elytra; elytra with a wide yellow fascia; antennomere III heart-like shape; gonoporal piece straight, slender, slightly curved basally; semigonoporal piece and ligula absent; spinous area (
<figureCitation id="5ACFDDC64514FFF5FD09BE35FCC9FDD4" box="[751,837,592,618]" captionStart="Fig" captionStartId="12.[190,233,1893,1919]" captionTargetBox="[191,1394,543,1850]" captionTargetId="figure@12.[189,1398,538,1851]" captionTargetPageId="12" captionText="Fig. 9. Spinous area and spinous plate of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica(Lewis,1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955).Abbreviations: sa = spinous area; sp = spinous plate. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829796" httpUri="https://zenodo.org/record/3829796/files/figure.png" pageId="23" pageNumber="24">Fig. 9F</figureCitation>
) densely covered with long spines and with oval spinous plate.
</paragraph>
</subSubSection>
<subSubSection id="8AEE92C84514FFF5FF5BBED1FB95FA10" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="C24BC1434514FFF5FF5BBED1FE26FD70" blockId="23.[189,426,692,718]" box="[189,426,692,718]" pageId="23" pageNumber="24">
<heading id="9903762F4514FFF5FF5BBED1FE26FD70" bold="true" box="[189,426,692,718]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="F0801D514514FFF5FF5BBED1FE26FD70" bold="true" box="[189,426,692,718]" pageId="23" pageNumber="24">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC1434514FFF5FF5BBE86FEDBFC13" blockId="23.[189,1399,739,941]" pageId="23" pageNumber="24">
<collectingCountry id="BAE381D34514FFF5FF5BBE86FE95FD43" box="[189,281,739,765]" name="Japan" pageId="23" pageNumber="24">JAPAN</collectingCountry>
, Honshu:
<collectingRegion id="00300FA14514FFF5FE74BE86FD82FD43" box="[402,526,739,765]" country="Japan" name="Gifu" pageId="23" pageNumber="24">Gifu Pref.</collectingRegion>
:
<specimenCount id="D4F20ACA4514FFF5FDFCBE86FDC6FD43" box="[538,586,739,765]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
,
<collectingRegion id="00300FA14514FFF5FDBEBE86FD1DFD43" box="[600,657,739,765]" country="Japan" name="Gifu" pageId="23" pageNumber="24">Gifu</collectingRegion>
, Japan,
<date id="B64AE7834514FFF5FD14BE86FC02FD43" box="[754,910,739,766]" pageId="23" pageNumber="24" value="1947-06-25">25 Jun. 1947</date>
, D. Matusita leg. (
<collectionCode id="A4E559864514FFF5FB8BBE86FB31FD43" box="[1133,1213,739,765]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
). –
<collectingRegion id="00300FA14514FFF5FB0ABE86FAFFFD43" box="[1260,1395,739,765]" country="Japan" name="Aichi" pageId="23" pageNumber="24">Aichi Pref.</collectingRegion>
:
<specimenCount id="D4F20ACA4514FFF5FF5BBF63FE8CFC9F" box="[189,256,774,801]" pageId="23" pageNumber="24" type="female">2 ♀♀</specimenCount>
, Yahagi-gawa, Murataka, Anjô-shi,
<date id="B64AE7834514FFF5FD4FBF63FCC8FC9E" box="[681,836,774,800]" pageId="23" pageNumber="24" value="1998-09-23">23 Sep. 1998</date>
and
<date id="B64AE7834514FFF5FC9BBF63FB9CFC9E" box="[893,1040,774,800]" pageId="23" pageNumber="24" value="1999-07-17">17 Jul. 1999</date>
, M. Hasegawa leg. (
<collectionCode id="A4E559864514FFF5FAE7BF63FAD8FC9E" box="[1281,1364,774,800]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
). –
<collectingRegion id="00300FA14514FFF5FF5BBF4CFEBEFCFD" box="[189,306,809,835]" country="Japan" name="Mie" pageId="23" pageNumber="24">Mie Pref.</collectingRegion>
:
<specimenCount id="D4F20ACA4514FFF5FED9BF4CFEE0FCFA" box="[319,364,809,836]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
, Yokkaichi,
<date id="B64AE7834514FFF5FDE5BF4CFD1DFCFD" box="[515,657,809,836]" pageId="23" pageNumber="24" value="1979-06-05">5 Jun. 1979</date>
, H. Nagai leg. (
<collectionCode id="A4E559864514FFF5FCB7BF4CFC28FCFD" box="[849,932,809,835]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA4514FFF5FC5CBF4CFC65FCFA" box="[954,1001,809,836]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
, same data,
<date id="B64AE7834514FFF5FB9BBF4FFB71FCFD" box="[1149,1277,809,836]" pageId="23" pageNumber="24" value="1978-08">Aug. 1978</date>
, H. Nagai leg. (
<collectionCode id="A4E559864514FFF5FF1EBF28FEC7FCD9" box="[248,331,845,871]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA4514FFF5FE86BF28FE00FCD9" box="[352,396,845,871]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
, Yokkaichi, Saburô-chô,
<date id="B64AE7834514FFF5FD50BF28FCC6FCD9" box="[694,842,844,871]" pageId="23" pageNumber="24" value="1982-07-12">12 Jul. 1982</date>
, B. Tanaka leg. (
<collectionCode id="A4E559864514FFF5FBE9BF28FBEEFCD9" box="[1039,1122,845,871]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
). –
<collectingRegion id="00300FA14514FFF5FB6ABF29FAB4FCD8" box="[1164,1336,844,870]" country="Japan" name="Shimane" pageId="23" pageNumber="24">Shimane Pref.</collectingRegion>
:
<specimenCount id="D4F20ACA4514FFF5FAA5BF28FAFDFCD9" box="[1347,1393,845,871]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
,
<specimenCount id="D4F20ACA4514FFF5FF5BBF15FF73FC34" box="[189,255,880,907]" pageId="23" pageNumber="24" type="female">5 ♀♀</specimenCount>
, Tsunozu, Gotsu,
<date id="B64AE7834514FFF5FE29BF15FDFCFC35" box="[463,624,880,907]" pageId="23" pageNumber="24" value="2015-08-19">19 Aug. 2015</date>
, H. Yoshitomi leg. (
<collectionCode id="A4E559864514FFF5FCB3BF15FC24FC34" box="[853,936,880,906]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA4514FFF5FC5DBF15FC6AFC34" box="[955,998,880,906]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
, st. 24,
<date id="B64AE7834514FFF5FBA6BF15FB5DFC34" box="[1088,1233,880,906]" pageId="23" pageNumber="24" value="2012-07-12">12 Jul. 2012</date>
, H. Yoshitomi leg. (
<collectionCode id="A4E559864514FFF5FF1EBFF6FEC7FC13" box="[248,331,915,941]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
).
</paragraph>
<paragraph id="C24BC1434514FFF5FF5BBFB0FEDBFBC5" blockId="23.[189,1399,980,1147]" pageId="23" pageNumber="24">
<collectingCountry id="BAE381D34514FFF5FF5BBFB0FE95FC51" box="[189,281,981,1007]" name="Japan" pageId="23" pageNumber="24">JAPAN</collectingCountry>
, Shikoku:
<collectingRegion id="00300FA14514FFF5FE74BFB1FDACFC50" box="[402,544,980,1006]" country="Japan" name="Ehime" pageId="23" pageNumber="24">Ehime Pref.</collectingRegion>
:
<specimenCount id="D4F20ACA4514FFF5FDCFBFB1FDD9FC51" box="[553,597,980,1007]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
, Nakajima,
<date id="B64AE7834514FFF5FD39BFB1FCF9FC51" box="[735,885,980,1007]" pageId="23" pageNumber="24" value="1957-10-13">13 Oct. 1957</date>
, F. Takechi leg. (
<collectionCode id="A4E559864514FFF5FBD2BFB1FB08FC50" box="[1076,1156,980,1006]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA4514FFF5FB7CBFB1FB64FC51" box="[1178,1256,980,1007]" pageId="23" pageNumber="24" type="female">11 ♀♀</specimenCount>
, Nishihabumachi, Matsuyama-shi, riverside of Shigenobu-gawa,
<geoCoordinate id="A7C0A7844514FFF5FCA7BF93FC71FBAC" box="[833,1021,1014,1042]" degrees="33" direction="north" minutes="48" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="22.61" value="33.80628">33°48′22.61′′ N</geoCoordinate>
,
<geoCoordinate id="A7C0A7844514FFF5FBEDBF93FB5EFBAC" box="[1035,1234,1014,1042]" degrees="132" direction="east" minutes="41" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="59.42" value="132.69984">132°41′59.42′′ E</geoCoordinate>
,
<date id="B64AE7834514FFF5FB06BF9DFAFEFBAC" box="[1248,1394,1015,1042]" pageId="23" pageNumber="24" value="2015-09-04">4 Sep. 2015</date>
, H. Ikeda leg. (
<collectionCode id="A4E559864514FFF5FE8CB87EFE31FB8B" box="[362,445,1051,1077]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA4514FFF5FE35B87EFD9BFB8B" box="[467,535,1051,1077]" pageId="23" pageNumber="24" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="D4F20ACA4514FFF5FDCEB87EFDF7FB8B" box="[552,635,1051,1077]" pageId="23" pageNumber="24" type="female">10 ♀♀</specimenCount>
, same data,
<date id="B64AE7834514FFF5FCF4B87EFC3FFB88" box="[786,947,1051,1078]" pageId="23" pageNumber="24" value="2015-09-10">10 Sep. 2015</date>
, H. Ikeda leg. (
<collectionCode id="A4E559864514FFF5FB8BB87EFB4CFB8B" box="[1133,1216,1051,1077]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA4514FFF5FB31B87EFA97FB8B" box="[1239,1307,1051,1077]" pageId="23" pageNumber="24" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="D4F20ACA4514FFF5FACDB87EFAFCFB8B" box="[1323,1392,1051,1077]" pageId="23" pageNumber="24" type="female">3 ♀♀</specimenCount>
, Shioya-kaigan, Kitagawara, Masaki-chô, Iyo-gun, estuary of Shigenobu-gawa,
<date id="B64AE7834514FFF5FB80B85BFA88FBE7" box="[1126,1284,1086,1113]" pageId="23" pageNumber="24" value="2015-04-20">20 Apr. 2015</date>
, H. Ikeda leg. (
<collectionCode id="A4E559864514FFF5FF1EB804FEC7FBC5" box="[248,331,1121,1147]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
).
</paragraph>
<paragraph id="C24BC1434514FFF5FF5BB8C6FEA9FAF4" blockId="23.[189,1399,1186,1354]" pageId="23" pageNumber="24">
<collectingCountry id="BAE381D34514FFF5FF5BB8C6FE95FB03" box="[189,281,1187,1213]" name="Japan" pageId="23" pageNumber="24">JAPAN</collectingCountry>
, Tokara Isls.: Takarajima:
<specimenCount id="D4F20ACA4514FFF5FDADB8C7FD01FB03" box="[587,653,1186,1213]" pageId="23" pageNumber="24" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="D4F20ACA4514FFF5FD7DB8C6FD4AFB03" box="[667,710,1187,1213]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
, Takara-jima,
<date id="B64AE7834514FFF5FC89B8C6FC7DFB02" box="[879,1009,1186,1213]" pageId="23" pageNumber="24" value="1960-07-03">3 Jul. 1960</date>
, M. Satô leg. (
<collectionCode id="A4E559864514FFF5FB7CB8C6FB61FB03" box="[1178,1261,1187,1213]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA4514FFF5FAE7B8C6FAA0FB03" box="[1281,1324,1187,1213]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
, same data,
<date id="B64AE7834514FFF5FF1DB8A3FE08FB5E" box="[251,388,1222,1248]" pageId="23" pageNumber="24" value="1962-06-03">3 Jun. 1962</date>
, M. Satô leg. (
<collectionCode id="A4E559864514FFF5FDCDB8A3FDF2FB5E" box="[555,638,1222,1248]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA4514FFF5FD77B8A3FD30FB5E" box="[657,700,1222,1248]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
, same data,
<date id="B64AE7834514FFF5FCACB8A3FC5FFB5E" box="[842,979,1222,1248]" pageId="23" pageNumber="24" value="1962-06-04">4 Jun. 1962</date>
, M. Satô leg. (
<collectionCode id="A4E559864514FFF5FB9CB8A3FB41FB5E" box="[1146,1229,1222,1248]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA4514FFF5FB06B8A3FAAFFB5E" box="[1248,1315,1222,1248]" pageId="23" pageNumber="24" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="D4F20ACA4514FFF5FAC8B8A3FAFCFB5E" box="[1326,1392,1222,1248]" pageId="23" pageNumber="24" type="female">2 ♀♀</specimenCount>
, same data,
<date id="B64AE7834514FFF5FEA7B88CFE41FABD" box="[321,461,1257,1284]" pageId="23" pageNumber="24" value="1962-06-05">5 Jun. 1962</date>
, M. Satô leg. (
<collectionCode id="A4E559864514FFF5FD9CB88CFD41FABD" box="[634,717,1257,1283]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA4514FFF5FD04B88CFC82FABA" box="[738,782,1257,1284]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
, same data,
<date id="B64AE7834514FFF5FC44B88CFBA1FABD" box="[930,1069,1257,1284]" pageId="23" pageNumber="24" value="1962-06-06">6 Jun. 1962</date>
, M. Satô leg. (
<collectionCode id="A4E559864514FFF5FB3DB88CFAA2FABD" box="[1243,1326,1257,1283]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA4514FFF5FAA5B88CFAFDFABA" box="[1347,1393,1257,1284]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
,
<specimenCount id="D4F20ACA4514FFF5FF5BB969FE8DFA99" box="[189,257,1292,1319]" pageId="23" pageNumber="24" type="female">4 ♀♀</specimenCount>
, same data,
<date id="B64AE7834514FFF5FE72B968FDADFA98" box="[404,545,1292,1319]" pageId="23" pageNumber="24" value="1962-06-07">7 Jun. 1962</date>
, M. Satô leg. (
<collectionCode id="A4E559864514FFF5FD36B969FCAFFA98" box="[720,803,1292,1318]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
);
<specimenCount id="D4F20ACA4514FFF5FCDEB969FCEBFA99" box="[824,871,1292,1319]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
,
<specimenCount id="D4F20ACA4514FFF5FC92B969FC34FA99" box="[884,952,1292,1319]" pageId="23" pageNumber="24" type="female">2 ♀♀</specimenCount>
, same data,
<date id="B64AE7834514FFF5FBAAB969FB54FA98" box="[1100,1240,1292,1319]" pageId="23" pageNumber="24" value="1962-06-08">8 Jun. 1962</date>
, M. Satô leg. (
<collectionCode id="A4E559864514FFF5FF23B955FE94FAF4" box="[197,280,1328,1354]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
).
</paragraph>
<paragraph id="C24BC1434514FFF5FF5BB914FB95FA10" blockId="23.[189,1399,1391,1455]" pageId="23" pageNumber="24">
<collectingCountry id="BAE381D34514FFF5FF5BB914FE95FA35" box="[189,281,1393,1419]" name="Japan" pageId="23" pageNumber="24">JAPAN</collectingCountry>
, Amami-Ôshima: Amami Isls.: Tokunoshima:
<specimenCount id="D4F20ACA4514FFF5FCB5B914FC10FA35" box="[851,924,1393,1419]" pageId="23" pageNumber="24" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="D4F20ACA4514FFF5FC4BB914FB8FFA35" box="[941,1027,1393,1419]" pageId="23" pageNumber="24" type="female">10 ♀♀</specimenCount>
, Senma Beach,
<date id="B64AE7834514FFF5FB29B914FAFDFA35" box="[1231,1393,1393,1419]" pageId="23" pageNumber="24" value="2001-06-20">20 Jun. 2001</date>
, M. Satô leg. (
<collectionCode id="A4E559864514FFF5FEBAB9F1FE23FA10" box="[348,431,1428,1454]" country="Japan" name="Ehime University" pageId="23" pageNumber="24">EUMJ</collectionCode>
).
<specimenCount id="D4F20ACA4514FFF5FE25B9F1FE7DFA11" box="[451,497,1428,1455]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
, Koniya,
<date id="B64AE7834514FFF5FD84B9F1FD71FA11" box="[610,765,1428,1455]" pageId="23" pageNumber="24" value="1954-04-20">20 Apr. 1954</date>
, S. Takagi leg. (
<collectionCode id="A4E559864514FFF5FC5FB9F1FB80FA10" box="[953,1036,1428,1454]" pageId="23" pageNumber="24">SEHU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="8AEE92C84514FFFAFF5BB9B0FAD0FCDC" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
<paragraph id="C24BC1434514FFF5FF5BB9B0FEE7FA51" blockId="23.[189,363,1493,1519]" box="[189,363,1493,1519]" pageId="23" pageNumber="24">
<heading id="9903762F4514FFF5FF5BB9B0FEE7FA51" bold="true" box="[189,363,1493,1519]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="F0801D514514FFF5FF5BB9B0FEE7FA51" bold="true" box="[189,363,1493,1519]" pageId="23" pageNumber="24">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC1434514FFF5FF0CBA61FEA6F9A0" blockId="23.[189,1398,1540,1707]" box="[234,298,1540,1566]" pageId="23" pageNumber="24">
<emphasis id="F0801D514514FFF5FF0CBA61FEA6F9A0" bold="true" box="[234,298,1540,1566]" pageId="23" pageNumber="24">Male</emphasis>
</paragraph>
<paragraph id="C24BC1434514FFF5FF5BBA42FE4AF915" blockId="23.[189,1398,1540,1707]" pageId="23" pageNumber="24">
Body (
<figureCitation id="5ACFDDC64514FFF5FEE9BA42FEEBF9FF" box="[271,359,1575,1601]" captionStart="Fig" captionStartId="4.[189,232,1893,1919]" captionTargetBox="[198,1389,740,1831]" captionTargetId="figure@4.[189,1398,723,1853]" captionTargetPageId="4" captionText="Fig. 1. Dorsal habitus Intybia spp., ♂. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov., holotype. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956), holotype. Scales = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3829778" httpUri="https://zenodo.org/record/3829778/files/figure.png" pageId="23" pageNumber="24">Fig. 1F</figureCitation>
) oval, shining, sparsely covered with fulvous hairs. Coloration of head, pronotum and elytra black with bluish luster; antennae yellow, with apical 8 or 9 segments black; elytra with a wide yellow fascia just before the middle; tibiae, trochanters, apex of coxae, fore femora; apical portion of middle femora yellow.
</paragraph>
<paragraph id="C24BC1434514FFFAFF5BBAB7FE34FE31" blockId="23.[189,1399,1746,2020]" lastBlockId="24.[189,1399,267,399]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
Head moderately densely punctate; eyes oval and prominent laterally; HL/HW 0.62–0.82 (0.69). Antennae (
<figureCitation id="5ACFDDC64514FFF5FEDBBA90FE18F8AE" box="[317,404,1781,1808]" captionStart="Fig" captionStartId="7.[190,233,1895,1921]" captionTargetBox="[265,1321,271,1836]" captionTargetId="figure@7.[246,1331,265,1859]" captionTargetPageId="7" captionText="Fig. 4. Right antennae of Intybia spp., ♂. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829786" httpUri="https://zenodo.org/record/3829786/files/figure.png" pageId="23" pageNumber="24">Fig. 4F</figureCitation>
) short, reaching base of elytra; antennomere I stout; antennomere III (
<figureCitation id="5ACFDDC64514FFF5FB2DBA90FAAFF8AE" box="[1227,1315,1781,1808]" captionStart="Fig" captionStartId="9.[190,233,1895,1921]" captionTargetBox="[263,1324,265,1858]" captionTargetId="figure@9.[263,1324,265,1859]" captionTargetPageId="9" captionText="Fig. 6. SEM photographs of right male antennomere III of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C, H. I. donan sp. nov., showing projection (H). D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955)." figureDoi="http://doi.org/10.5281/zenodo.3829790" httpUri="https://zenodo.org/record/3829790/files/figure.png" pageId="23" pageNumber="24">Fig. 6F</figureCitation>
) heartlike shape, deeply concave in dorso-mesal portion; L/W = 1.12; approximate ratio of each antennomere (n = 1) as 9.7: 1.0: 12.7: 2.3: 2.3: 2.3: 3.0:2.7:2.7: 2.7: 4.3. Pronotum transverse, broader than head; sides subparallel in the anterior half, then sinuately and strongly curved posteriorly; base slightly but broadly emarginated; surface rather densely punctate at the sides, rugosely so before hind angles, more or less sparsely and finely punctate in middle; PW/PL 1.13–1.40 (1.22). Elytra oval, moderately convex, broadened posteriorly, densely punctate at lateral parts; EL/EW 1.15–1.48 (1.36); EL/PL 2.39–3.07 (2.68); EW/PW 1.48–1.83 (1.60); TL/EW 1.92–2.42 (2.28). Aedeagus about
<quantity id="050C6CA6451BFFFAFBDDBD6EFB2EFE98" box="[1083,1186,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="24" pageNumber="25" unit="mm" value="0.73">0.73 mm</quantity>
in AL. Gonoporal piece (
<figureCitation id="5ACFDDC6451BFFFAFEEABD4BFEE1FEF7" box="[268,365,302,329]" captionStart="Fig" captionStartId="11.[190,233,1928,1954]" captionTargetBox="[192,1397,1222,1886]" captionTargetId="figure@11.[192,1400,1222,1886]" captionTargetPageId="11" captionText="Fig. 8. Endophallus of Intybia spp., A. Intybia histrio (Kiesenwetter, 1874). B. I. pelegrini pelegrini (Pic, 1910). C. I. takaraensis (Nakane, 1955). D. I. kishiii (Nakane, 1955). Abbreviations: gp = gonoporal piece; l = ligula; sgp = semi-gonoporal piece; sp = spinous plate. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829794" httpUri="https://zenodo.org/record/3829794/files/figure.png" pageId="24" pageNumber="25">Figs 8C</figureCitation>
,
<figureCitation id="5ACFDDC6451BFFFAFE9ABD4AFE27FEF7" box="[380,427,303,329]" captionStart="Fig" captionStartId="13.[190,233,1893,1919]" captionTargetBox="[216,1374,635,1827]" captionTargetId="figure@13.[188,1397,614,1853]" captionTargetPageId="13" captionText="Fig. 10. Endophallic sclerites of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). Abbreviations: gp = gonoporal piece; l = ligula; sgp = semi-gonoporal piece. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829798" httpUri="https://zenodo.org/record/3829798/files/figure.png" pageId="24" pageNumber="25">10F</figureCitation>
) straight, slender, slightly curved in basally,
<quantity id="050C6CA6451BFFFAFC25BD4AFBA2FEF7" box="[963,1070,303,329]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="24" pageNumber="25" unit="mm" value="0.44">0.44 mm</quantity>
in GL; GL/AL 0.60. Semigonoporal piece and ligula absent. Spinous area (
<figureCitation id="5ACFDDC6451BFFFAFCE9BD37FCE5FED2" box="[783,873,338,364]" captionStart="Fig" captionStartId="12.[190,233,1893,1919]" captionTargetBox="[191,1394,543,1850]" captionTargetId="figure@12.[189,1398,538,1851]" captionTargetPageId="12" captionText="Fig. 9. Spinous area and spinous plate of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica(Lewis,1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955).Abbreviations: sa = spinous area; sp = spinous plate. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829796" httpUri="https://zenodo.org/record/3829796/files/figure.png" pageId="24" pageNumber="25">Fig. 9F</figureCitation>
) densely covered with long spines; spinous plate large and ovate.
</paragraph>
<paragraph id="C24BC143451BFFFAFF0CBDCAFEC9FE77" blockId="24.[189,1399,431,634]" box="[234,325,431,457]" pageId="24" pageNumber="25">
<emphasis id="F0801D51451BFFFAFF0CBDCAFEC9FE77" bold="true" box="[234,325,431,457]" pageId="24" pageNumber="25">Female</emphasis>
</paragraph>
<paragraph id="C24BC143451BFFFAFF5BBDB7FB2FFDC4" blockId="24.[189,1399,431,634]" pageId="24" pageNumber="25">
Similar to male in general appearance (
<figureCitation id="5ACFDDC6451BFFFAFD42BDB7FC8EFE52" box="[676,770,466,492]" captionStart="Fig" captionStartId="5.[190,233,1894,1920]" captionTargetBox="[195,1390,781,1842]" captionTargetId="figure@5.[190,1399,771,1854]" captionTargetPageId="5" captionText="Fig. 2. Dosal habitus of Intybia spp., ♀. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov., paratype. D. I. pelegrini pelegrini (Pic, 1910). E. I.niponica (Lewis, 1895). F. I.takaraensis (Nakane, 1955).G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956), paratype. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3829782" httpUri="https://zenodo.org/record/3829782/files/figure.png" pageId="24" pageNumber="25">Fig. 2F</figureCitation>
); coloration of antennae yellow, with apical 7 or 8 segments black; HL/HW 0.54–0.68 (0.61); PW/PL 1.14–1.37 (1.24); EL/EW 1.22–1.53 (1.38); EL/ PL 2.60–3.03 (2.79); EW/PW 1.5–1.81 (1.63); TL/EW 1.99–2.41 (2.22). Antennae (
<figureCitation id="5ACFDDC6451BFFFAFB2FBE7DFAA9FD8D" box="[1225,1317,536,563]" captionStart="Fig" captionStartId="8.[190,233,1894,1920]" captionTargetBox="[285,1298,276,1852]" captionTargetId="figure@8.[273,1302,265,1859]" captionTargetPageId="8" captionText="Fig. 5. Right antennae of Intybia spp., ♀. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956). Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3829788" httpUri="https://zenodo.org/record/3829788/files/figure.png" pageId="24" pageNumber="25">Fig. 5F</figureCitation>
) long; antennomere III long, expanded laterally, shallowly concave in dorsal part; L/W = 2.00; approximate ratio of each antennomere (n = 1) as 9.7: 1.0: 6.7:3.0:3.0: 3.3: 3.3: 3.3: 3.3: 2.7:5.0.
</paragraph>
<paragraph id="C24BC143451BFFFAFF5BBEFCFEF9FD0D" blockId="24.[189,373,665,691]" box="[189,373,665,691]" pageId="24" pageNumber="25">
<heading id="9903762F451BFFFAFF5BBEFCFEF9FD0D" bold="true" box="[189,373,665,691]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="F0801D51451BFFFAFF5BBEFCFEF9FD0D" bold="true" box="[189,373,665,691]" pageId="24" pageNumber="25">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC143451BFFFAFF5BBEA2FAD0FCBB" blockId="24.[189,1398,711,773]" pageId="24" pageNumber="25">Male (n = 18): TL 2.97–3.54 (3.18) mm; HL 0.50–0.62 (0.56) mm; HW 0.73–0.91 (0.81) mm; PW 0.77–1.03 (0.87) mm; PL 0.60–0.82 (0.72) mm; EL 1.67–2.14 (1.90) mm; EW 1.27–1.70 (1.40) mm.</paragraph>
<paragraph id="C24BC143451BFFFAFF5BBF41FAD0FCDC" blockId="24.[189,1398,804,866]" pageId="24" pageNumber="25">Female (n = 20): TL 2.78–3.26 (3.01) mm; HL 0.39–0.52 (0.47) mm; HW 0.71–0.85 (0.77) mm; PW 0.75–0.95 (0.83) mm; PL 0.62–0.75 (0.67) mm; EL 1.73–2.04 (1.87) mm; EW 1.17–1.58 (1.36) mm.</paragraph>
</subSubSection>
<subSubSection id="8AEE92C8451BFFFAFF5BBFE4FC48FC50" pageId="24" pageNumber="25" type="distribution">
<paragraph id="C24BC143451BFFFAFF5BBFE4FE5BFC25" blockId="24.[189,471,897,923]" box="[189,471,897,923]" pageId="24" pageNumber="25">
<emphasis id="F0801D51451BFFFAFF5BBFE4FEDAFC25" bold="true" box="[189,342,897,923]" pageId="24" pageNumber="25">Distribution</emphasis>
(
<figureCitation id="5ACFDDC6451BFFFAFE80BFE4FE42FC25" box="[358,462,897,923]" captionStart="Fig" captionStartId="15.[190,233,1895,1921]" captionTargetBox="[196,1385,1174,1842]" captionTargetId="figure@15.[192,1400,1167,1854]" captionTargetPageId="15" captionText="Fig. 11.Map showing thedistribution of Intybia spp.A.The histriogroup:●:Intybia histrio(Kiesenwetter, 1874), ▲: I. tsushimensis (Satô &amp; Ohbayashi, 1968), ■: I. donan sp. nov.). B. The pelegrini group: ○: I. pelegrini (Pic, 1910), △: I. niponica (Lewis, 1895), □: I. takaraensis (Nakane, 1955), ×: I. kishiii (Nakane, 1955), +: I. kawasakii (Nakane, 1956)." figureDoi="http://doi.org/10.5281/zenodo.3829800" httpUri="https://zenodo.org/record/3829800/files/figure.png" pageId="24" pageNumber="25">Fig. 11B</figureCitation>
)
</paragraph>
<paragraph id="C24BC143451BFFFAFF5BBFD5FC48FC50" blockId="24.[189,1398,942,1006]" pageId="24" pageNumber="25">
<collectingCountry id="BAE381D3451BFFFAFF5BBFD5FE8CFC74" box="[189,256,944,970]" name="Japan" pageId="24" pageNumber="25">Japan</collectingCountry>
(Honshu, Shikoku, Kyushu, Shôdo-shima, Takara-jima, Tokuno-shima [NR], Amami-Ôshima [NR]);
<collectingCountry id="BAE381D3451BFFFAFEF5BFB6FED5FC53" box="[275,345,979,1005]" name="China" pageId="24" pageNumber="25">China</collectingCountry>
(
<collectingRegion id="00300FA1451BFFFAFE8EBFB6FE34FC53" box="[360,440,979,1005]" country="China" name="Fujian" pageId="24" pageNumber="25">Fujian</collectingRegion>
, NE
<collectingCountry id="BAE381D3451BFFFAFE14BFB6FDB1FC50" box="[498,573,979,1006]" name="China" pageId="24" pageNumber="25">China</collectingCountry>
),
<collectingCountry id="BAE381D3451BFFFAFDB6BFB6FD12FC53" box="[592,670,979,1005]" name="Russia" pageId="24" pageNumber="25">Russia</collectingCountry>
(Primorye, Wrangel Is.).
</paragraph>
</subSubSection>
<subSubSection id="8AEE92C8451BFFFAFF5BB868FB84FBC7" pageId="24" pageNumber="25" type="discussion">
<paragraph id="C24BC143451BFFFAFF5BB868FEA3FB99" blockId="24.[189,303,1037,1063]" box="[189,303,1037,1063]" pageId="24" pageNumber="25">
<heading id="9903762F451BFFFAFF5BB868FEA3FB99" bold="true" box="[189,303,1037,1063]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="F0801D51451BFFFAFF5BB868FEA3FB99" bold="true" box="[189,303,1037,1063]" pageId="24" pageNumber="25">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC143451BFFFAFF5BB85EFB84FBC7" blockId="24.[189,1398,1083,1145]" pageId="24" pageNumber="25">
This species is closely similar to
<taxonomicName id="05F4BAC0451BFFFAFDA6B859FC23FBE8" authority="(Lewis, 1895)" baseAuthorityName="Lewis" baseAuthorityYear="1895" box="[576,943,1083,1110]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="niponica">
<emphasis id="F0801D51451BFFFAFDA6B859FC8DFBEB" box="[576,769,1084,1110]" italics="true" pageId="24" pageNumber="25">Intybia niponica</emphasis>
(
<bibRefCitation id="A665BCB2451BFFFAFCF6B85EFC2AFBE8" author="Lewis G." box="[784,934,1083,1110]" pageId="24" pageNumber="25" pagination="98 - 122" refId="ref17589" refString="Lewis G. 1895. On the Daschillidae and malacoderm Coleoptera of Japan. The Annals and Magazine of Natural History (6) 16: 98 - 122, pl. 6. https: // doi. org / 10.1080 / 00222939508680233" type="journal article" year="1895">Lewis, 1895</bibRefCitation>
)
</taxonomicName>
, and differs from it in the coloration of the body with a bluish luster and the shape of the male antennomere III.
</paragraph>
</subSubSection>
<subSubSection id="8AEE92C8451BFFFAFF5BB8FDFB1CFB5F" pageId="24" pageNumber="25" type="biology_ecology">
<paragraph id="C24BC143451BFFFAFF5BB8FDFE0DFB0D" blockId="24.[189,385,1176,1203]" box="[189,385,1176,1203]" pageId="24" pageNumber="25">
<heading id="9903762F451BFFFAFF5BB8FDFE0DFB0D" bold="true" box="[189,385,1176,1203]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="F0801D51451BFFFAFF5BB8FDFE0DFB0D" bold="true" box="[189,385,1176,1203]" pageId="24" pageNumber="25">Biological notes</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC143451BFFFAFF5BB8A2FB1CFB5F" blockId="24.[189,1168,1223,1250]" box="[189,1168,1223,1250]" pageId="24" pageNumber="25">This species inhabits beach plant communities or grasslands areas near large rivers.</paragraph>
</subSubSection>
</treatment>
</document>
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