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<document id="FF6FA5EBD3341FBBE90057D586186E74" ID-CLB-Dataset="22090" ID-DOI="10.5852/ejt.2017.331" ID-GBIF-Dataset="acf565ee-29ba-45e7-9ef3-d1e85dec9ae7" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829776" ID-ZooBank="3FD5D922-B91F-42BB-9D7B-7DFB98841732" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589572797790" checkinUser="carolina" docAuthor="Ikeda, Hiroshi &amp; Yoshitomi, Hiroyuki" docDate="2017" docId="4A5D70554501FFE1FD92B869FAFAFC11" docLanguage="en" docName="ejt-2017-331.pdf.imf" docOrigin="European Journal of Taxonomy 331" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Intybia Pascoe 1866" docType="key" docVersion="7" lastPageNumber="4" masterDocId="B664082D4503FFE2FFE6BC65FF8CFFBE" masterDocTitle="Revision of the genus Intybia (Coleoptera, Malachiidae) from Japan" masterLastPageNumber="31" masterPageNumber="1" pageNumber="3" updateTime="1698839968837" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="82A9019458E353198674A164FFB78BCC">Revision of the genus Intybia (Coleoptera, Malachiidae) from Japan</mods:title>
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<mods:name id="60FDF337C28E89A75673F6031B71292E" type="personal">
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<mods:namePart id="B508DB70B3BE22F5DF989117E1F15E17">Ikeda, Hiroshi</mods:namePart>
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<mods:namePart id="90A4FB8D0D8C09484908CB44F0B330E3">Yoshitomi, Hiroyuki</mods:namePart>
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<mods:affiliation id="12DCAA138A065667ABF225D591045F34">Email: hymushi @ agr. ehime-u. ac. jp &amp; urn: lsid: zoobank. org: author: 744 FA 6 FC- 2 D 84 - 41 B 7 - 8 A 6 C-C 1611 CC 68 EE 1</mods:affiliation>
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<mods:title id="00C6D3D974DFAC68EB24290AC991CDE3">European Journal of Taxonomy</mods:title>
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<treatment id="4A5D70554501FFE1FD92B869FAFAFC11" ID-DOI="http://doi.org/10.5281/zenodo.3848094" ID-GBIF-Taxon="164161038" ID-Zenodo-Dep="3848094" LSID="urn:lsid:plazi:treatment:4A5D70554501FFE1FD92B869FAFAFC11" httpUri="http://treatment.plazi.org/id/4A5D70554501FFE1FD92B869FAFAFC11" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection id="8AEE92C84501FFE0FD92B869FC33FB98" box="[628,959,1035,1062]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="C24BC1434501FFE0FD92B869FC33FB98" blockId="2.[628,959,1035,1062]" box="[628,959,1035,1062]" pageId="2" pageNumber="3">
<heading id="9903762F4501FFE0FD92B869FC33FB98" box="[628,959,1035,1062]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
Genus
<taxonomicName id="05F4BAC04501FFE0FD21B86EFC33FB98" authority="Pascoe, 1866" authorityName="Pascoe" authorityYear="1866" box="[711,959,1035,1062]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="F0801D514501FFE0FD21B86EFC91FB98" bold="true" box="[711,797,1035,1062]" italics="true" pageId="2" pageNumber="3">Intybia</emphasis>
<bibRefCitation id="A665BCB24501FFE0FCC2B869FC33FB98" author="Pascoe F. P." box="[804,959,1036,1062]" pageId="2" pageNumber="3" pagination="443 - 493" refId="ref17965" refString="Pascoe F. P. 1866. Notices of new or little-known genera and species of Coleoptera. Journal of Entomology 2: 443 - 493." type="journal article" year="1866">Pascoe, 1866</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<treatmentCitationGroup id="E2E4E66D4501FFE0FF5BB828FC72FBD6" box="[189,1022,1101,1128]" pageId="2" pageNumber="3">
<subSubSection id="8AEE92C84501FFE0FF5BB828FE70FBD6" box="[189,508,1101,1128]" pageId="2" pageNumber="3" type="reference_group">
<paragraph id="C24BC1434501FFE0FF5BB828FE70FBD6" blockId="2.[189,1022,1101,1128]" box="[189,508,1101,1128]" pageId="2" pageNumber="3">
<treatmentCitation id="4355E7524501FFE0FF5BB828FE62FBD9" author="Pascoe F. P." box="[189,494,1101,1128]" page="448" pageId="2" pageNumber="3" year="1866">
<taxonomicName id="05F4BAC04501FFE0FF5BB828FE62FBD9" authority="Pascoe, 1866: 448" authorityName="Pascoe" authorityPageNumber="448" authorityYear="1866" box="[189,494,1101,1128]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="F0801D514501FFE0FF5BB828FE9CFBD9" box="[189,272,1101,1127]" italics="true" pageId="2" pageNumber="3">Intybia</emphasis>
<bibRefCitation id="A665BCB24501FFE0FEF1B82BFE62FBD9" author="Pascoe F. P." box="[279,494,1101,1128]" pageId="2" pageNumber="3" pagination="443 - 493" refId="ref17965" refString="Pascoe F. P. 1866. Notices of new or little-known genera and species of Coleoptera. Journal of Entomology 2: 443 - 493." type="journal article" year="1866">Pascoe, 1866: 448</bibRefCitation>
</taxonomicName>
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[
</paragraph>
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<subSubSection id="8AEE92C84501FFE0FE1AB82BFC72FBD6" box="[508,1022,1101,1128]" pageId="2" pageNumber="3" type="type_taxon">
<paragraph id="C24BC1434501FFE0FE1AB82BFC72FBD6" blockId="2.[189,1022,1101,1128]" box="[508,1022,1101,1128]" pageId="2" pageNumber="3">
type species:
<taxonomicName id="05F4BAC04501FFE0FD7BB828FC7FFBD6" authority="Pascoe, 1866" authorityName="Pascoe" authorityYear="1866" box="[669,1011,1101,1128]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="guttata">
<emphasis id="F0801D514501FFE0FD7BB828FCC6FBD9" box="[669,842,1101,1127]" italics="true" pageId="2" pageNumber="3">Intybia guttata</emphasis>
<bibRefCitation id="A665BCB24501FFE0FCB4B82BFC7FFBD6" author="Pascoe F. P." box="[850,1011,1101,1128]" pageId="2" pageNumber="3" pagination="443 - 493" refId="ref17965" refString="Pascoe F. P. 1866. Notices of new or little-known genera and species of Coleoptera. Journal of Entomology 2: 443 - 493." type="journal article" year="1866">Pascoe, 1866</bibRefCitation>
</taxonomicName>
].
</paragraph>
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</treatmentCitationGroup>
<subSubSection id="8AEE92C84501FFE0FF5BB8EAFEB8F9DF" pageId="2" pageNumber="3" type="discussion">
<paragraph id="C24BC1434501FFE0FF5BB8EAFE8FFB17" blockId="2.[189,259,1167,1193]" box="[189,259,1167,1193]" pageId="2" pageNumber="3">
<heading id="9903762F4501FFE0FF5BB8EAFE8FFB17" bold="true" box="[189,259,1167,1193]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="F0801D514501FFE0FF5BB8EAFE8FFB17" bold="true" box="[189,259,1167,1193]" pageId="2" pageNumber="3">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC1434501FFE0FF5BB8D8FE85FA4C" blockId="2.[189,1399,1213,1522]" pageId="2" pageNumber="3">
The genus
<taxonomicName id="05F4BAC04501FFE0FEA0B8D8FE15FB69" authorityName="Pascoe" authorityYear="1866" box="[326,409,1213,1239]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="F0801D514501FFE0FEA0B8D8FE15FB69" box="[326,409,1213,1239]" italics="true" pageId="2" pageNumber="3">Intybia</emphasis>
</taxonomicName>
belongs to the tribe
<taxonomicName id="05F4BAC04501FFE0FD78B8D8FCA6FB69" authorityName="Muslant &amp; Rey" authorityYear="1867" box="[670,810,1213,1239]" class="Insecta" family="Melyridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Apalochrini">Apalochrini</taxonomicName>
of the subfamily
<taxonomicName id="05F4BAC04501FFE0FBECB8D8FB17FB69" authorityName="Fleming" authorityYear="1821" box="[1034,1179,1213,1239]" class="Insecta" family="Melyridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Malachiinae">Malachiinae</taxonomicName>
together with the related genera
<taxonomicName id="05F4BAC04501FFE0FE81B884FE24FB44" authorityName="Guerin-Meneville" authorityYear="1830" box="[359,424,1249,1274]" class="Insecta" family="Melyridae" genus="Laius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="F0801D514501FFE0FE81B884FE24FB44" box="[359,424,1249,1274]" italics="true" pageId="2" pageNumber="3">Laius</emphasis>
</taxonomicName>
and
<taxonomicName id="05F4BAC04501FFE0FE39B885FCD6FB45" authority="(Champion, 1921)" baseAuthorityName="Champion" baseAuthorityYear="1921" box="[479,858,1248,1275]" class="Insecta" family="Melyridae" genus="Dicranolaius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="F0801D514501FFE0FE39B885FDF6FB44" box="[479,634,1248,1274]" italics="true" pageId="2" pageNumber="3">Dicranolaius</emphasis>
(
<bibRefCitation id="A665BCB24501FFE0FD6EB885FCDDFB45" author="Champion G. C." box="[648,849,1248,1275]" pageId="2" pageNumber="3" pagination="322 - 343" refId="ref17367" refString="Champion G. C. 1921. Notes on various african and asiatic species of Laius, Guerin, with an account at their accessory ♂ - characters [Coleoptera]. The Annals and Magazine of Natural History 9 (7): 322 - 343. https: // doi. org / 10.1080 / 00222932108632526" type="journal article" year="1921">Champion, 1921</bibRefCitation>
)
</taxonomicName>
. The genus
<taxonomicName id="05F4BAC04501FFE0FC04B885FBB9FB44" authorityName="Pascoe" authorityYear="1866" box="[994,1077,1248,1274]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="F0801D514501FFE0FC04B885FBB9FB44" box="[994,1077,1248,1274]" italics="true" pageId="2" pageNumber="3">Intybia</emphasis>
</taxonomicName>
is characterized as follows: elytra with yellowish to reddish fascia or maculation in most species (vs fully black with bluish luster in
<taxonomicName id="05F4BAC04501FFE0FF39B94DFEACFAFF" authorityName="Guerin-Meneville" authorityYear="1830" box="[223,288,1320,1345]" class="Insecta" family="Melyridae" genus="Laius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="F0801D514501FFE0FF39B94DFEACFAFF" box="[223,288,1320,1345]" italics="true" pageId="2" pageNumber="3">Laius</emphasis>
</taxonomicName>
); antennomere II short; antennomere III in male modified; simple protarsus and protibia in male (vs two tarsal combs in protarsi V in
<taxonomicName id="05F4BAC04501FFE0FD5EB92FFCDFFADA" baseAuthorityName="Champion" baseAuthorityYear="1921" box="[696,851,1354,1380]" class="Insecta" family="Melyridae" genus="Dicranolaius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="F0801D514501FFE0FD5EB92FFCDFFADA" box="[696,851,1354,1380]" italics="true" pageId="2" pageNumber="3">Dicranolaius</emphasis>
</taxonomicName>
); front legs slender (vs male protibia of
<taxonomicName id="05F4BAC04501FFE0FAD3B92EFAFAFADA" authorityName="Guerin-Meneville" authorityYear="1830" box="[1333,1398,1355,1380]" class="Insecta" family="Melyridae" genus="Laius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="F0801D514501FFE0FAD3B92EFAFAFADA" box="[1333,1398,1355,1380]" italics="true" pageId="2" pageNumber="3">Laius</emphasis>
</taxonomicName>
thickened basally and inside of basal area distinctly hollow); aedeagal apex round in Japanese species (vs concave in most
<taxonomicName id="05F4BAC04501FFE0FE54B9F4FE7FFA14" authorityName="Guerin-Meneville" authorityYear="1830" box="[434,499,1425,1450]" class="Insecta" family="Melyridae" genus="Laius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="F0801D514501FFE0FE54B9F4FE7FFA14" box="[434,499,1425,1450]" italics="true" pageId="2" pageNumber="3">Laius</emphasis>
</taxonomicName>
species); endophallic structure composed of 1–3 sclerites (gonoporal piece, semi-gonoporal piece, ligula, and 0–2 spinous plate(s)) (Evers 1994;
<bibRefCitation id="A665BCB24501FFE0FC04B9D1FAADFA70" author="Asano M. &amp; Kawashima I." box="[994,1313,1460,1486]" pageId="2" pageNumber="3" pagination="261 - 266" refId="ref17325" refString="Asano M. &amp; Kawashima I. 2010. Discovery of the genus Dicranolaius (Coleoptera, Malachiidae) from Japan with description of a new species. Japanese Journal of Systematic Entomology 16 (2): 261 - 266." type="journal article" year="2010">Asano &amp; Kawashima 2010</bibRefCitation>
;
<bibRefCitation id="A665BCB24501FFE0FACDB9D1FF71FA4C" author="Asano M." pageId="2" pageNumber="3" pagination="77 - 82" refId="ref17280" refString="Asano M. 2015. Taxonomic notes on some malachiid beetles of Taiwan, with description of a new species of the genus Intybia Pascoe (Coleoptera: Malachiidae). Japanese Journal of Systematic Entomology 21 (1): 77 - 82." type="journal article" year="2015">Asano 2015</bibRefCitation>
).
</paragraph>
<paragraph id="C24BC1434501FFE0FF5BBA7DFE90F98C" blockId="2.[189,284,1560,1586]" box="[189,284,1560,1586]" pageId="2" pageNumber="3">
<heading id="9903762F4501FFE0FF5BBA7DFE90F98C" bold="true" box="[189,284,1560,1586]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="F0801D514501FFE0FF5BBA7DFE90F98C" bold="true" box="[189,284,1560,1586]" pageId="2" pageNumber="3">Gender</emphasis>
</heading>
</paragraph>
<paragraph id="C24BC1434501FFE0FF5BBA22FEB8F9DF" blockId="2.[189,308,1607,1633]" box="[189,308,1607,1633]" pageId="2" pageNumber="3">Feminine.</paragraph>
</subSubSection>
<subSubSection id="8AEE92C84501FFE0FF5BBAEDFC83F91D" box="[189,783,1672,1699]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="C24BC1434501FFE0FF5BBAEDFC83F91D" blockId="2.[189,783,1672,1699]" box="[189,783,1672,1699]" pageId="2" pageNumber="3">
<heading id="9903762F4501FFE0FF5BBAEDFC83F91D" bold="true" box="[189,783,1672,1699]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="F0801D514501FFE0FF5BBAEDFC83F91D" bold="true" box="[189,783,1672,1699]" pageId="2" pageNumber="3">
Key to the Japanese species of the genus
<taxonomicName id="05F4BAC04501FFE0FD5FBAEDFC83F91D" authorityName="Pascoe" authorityYear="1866" box="[697,783,1672,1699]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="F0801D514501FFE0FD5FBAEDFC83F91D" bold="true" box="[697,783,1672,1699]" italics="true" pageId="2" pageNumber="3">Intybia</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8AEE92C84501FFE1FF5BBAD2FAFAFC11" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="key">
<key id="B66596724501FFE1FF5BBAD2FAFAFC11" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<keyStep id="7900D9E64501FFE0FF5BBAD2FAFBF8A6" pageId="2" pageNumber="3">
<paragraph id="C24BC1434501FFE0FF5BBAD2FAFAF94A" blockId="2.[189,1399,1719,1816]" pageId="2" pageNumber="3">
<keyLead id="790562764501FFE0FF5BBAD2FAFAF94A" pageId="2" pageNumber="3">
1. Elytra with fascia, but sometimes fascia divided into two spots by suture; distributed in
<collectingRegion id="00300FA14501FFE0FB03BAD2FAD5F96F" box="[1253,1369,1719,1745]" country="Japan" name="Hokkaido" pageId="2" pageNumber="3">Hokkaido</collectingRegion>
to Ryukyu ……………………………………………………………………………………………2
</keyLead>
</paragraph>
<paragraph id="C24BC1434501FFE0FF27BA9BFAFBF8A6" blockId="2.[189,1399,1719,1816]" box="[193,1399,1789,1816]" pageId="2" pageNumber="3">
<keyLead id="790562764501FFE0FF27BA9BFAFBF8A6" box="[193,1399,1789,1816]" pageId="2" pageNumber="3">
– Elytra with four spots;distributed in Ryukyu (Yonaguni-jima)………………………
<taxonomicName id="05F4BAC04501FFE0FB5EBA9BFA97F8A6" authority="Ikeda &amp; Yoshitomi, 2017" authorityName="Ikeda &amp; Yoshitomi" authorityYear="2017" box="[1208,1307,1789,1816]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="donan" status="sp. nov.">
<emphasis id="F0801D514501FFE0FB5EBA9BFA97F8A6" bold="true" box="[1208,1307,1789,1816]" italics="true" pageId="2" pageNumber="3">I.donan</emphasis>
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<taxonomicNameLabel id="EBB3A02A4501FFE0FAF9BA9BFAFBF8A6" box="[1311,1399,1790,1816]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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<keyStep id="7900D9E64501FFE0FF5BBB5AFAFAF8C3" pageId="2" pageNumber="3">
<paragraph id="C24BC1434501FFE0FF5BBB5AFAFBF8E7" blockId="2.[189,1399,1855,1917]" box="[189,1399,1855,1881]" pageId="2" pageNumber="3">
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2. Elytral fascia widened at suture (
<figureCitation id="5ACFDDC64501FFE0FD8EBB5AFD21F8E7" box="[616,685,1855,1881]" captionStart="Fig" captionStartId="4.[189,232,1893,1919]" captionTargetBox="[198,1389,740,1831]" captionTargetId="figure@4.[189,1398,723,1853]" captionTargetPageId="4" captionText="Fig. 1. Dorsal habitus Intybia spp., ♂. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov., holotype. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956), holotype. Scales = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3829778" httpUri="https://zenodo.org/record/3829778/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
D–G; 2D–G) ……………………………………………… 3
</keyLead>
</paragraph>
<paragraph id="C24BC1434501FFE0FF27BB07FAFAF8C3" blockId="2.[189,1399,1855,1917]" box="[193,1398,1890,1917]" pageId="2" pageNumber="3">
<keyLead id="790562764501FFE0FF27BB07FAFAF8C3" box="[193,1398,1890,1917]" pageId="2" pageNumber="3">
– Elytral fascia narrowed at suture, or divided into two spots by suture (
<figureCitation id="5ACFDDC64501FFE0FBCDBB07FB5AF8C2" box="[1067,1238,1890,1916]" captionStart="Fig" captionStartId="4.[189,232,1893,1919]" captionTargetBox="[198,1389,740,1831]" captionTargetId="figure@4.[189,1398,723,1853]" captionTargetPageId="4" captionText="Fig. 1. Dorsal habitus Intybia spp., ♂. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov., holotype. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica (Lewis, 1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956), holotype. Scales = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3829778" httpUri="https://zenodo.org/record/3829778/files/figure.png" pageId="2" pageNumber="3">Figs 1A, B, H</figureCitation>
;
<figureCitation id="5ACFDDC64501FFE0FB02BB07FB7EF8C2" box="[1252,1266,1890,1916]" captionStart="Fig" captionStartId="5.[190,233,1894,1920]" captionTargetBox="[195,1390,781,1842]" captionTargetId="figure@5.[190,1399,771,1854]" captionTargetPageId="5" captionText="Fig. 2. Dosal habitus of Intybia spp., ♀. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov., paratype. D. I. pelegrini pelegrini (Pic, 1910). E. I.niponica (Lewis, 1895). F. I.takaraensis (Nakane, 1955).G. I. kishiii (Nakane, 1955). H. I. kawasakii (Nakane, 1956), paratype. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3829782" httpUri="https://zenodo.org/record/3829782/files/figure.png" pageId="2" pageNumber="3">2</figureCitation>
A–B, H)..6
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<paragraph id="C24BC1434501FFE0FF5BBBC6FAFBF85F" blockId="2.[189,1399,1955,2017]" pageId="2" pageNumber="3">
<keyLead id="790562764501FFE0FF5BBBC6FAFBF85F" pageId="2" pageNumber="3">
3. Fore legs fully yellow; head wide, about
<quantity id="050C6CA64501FFE0FD52BBC6FCABF803" box="[692,807,1955,1982]" metricMagnitude="-2" metricUnit="m" metricValue="1.7779999999999998" metricValueMax="2.286" metricValueMin="1.27" pageId="2" pageNumber="3" unit="in" value="0.7" valueMax="0.9" valueMin="0.5">0.5–0.9 in</quantity>
HL/HW; eyes prominent; endophallus with spinous plate (
<figureCitation id="5ACFDDC64501FFE0FED7BBA3FEF4F85F" box="[305,376,1990,2017]" captionStart="Fig" captionStartId="11.[190,233,1928,1954]" captionTargetBox="[192,1397,1222,1886]" captionTargetId="figure@11.[192,1400,1222,1886]" captionTargetPageId="11" captionText="Fig. 8. Endophallus of Intybia spp., A. Intybia histrio (Kiesenwetter, 1874). B. I. pelegrini pelegrini (Pic, 1910). C. I. takaraensis (Nakane, 1955). D. I. kishiii (Nakane, 1955). Abbreviations: gp = gonoporal piece; l = ligula; sgp = semi-gonoporal piece; sp = spinous plate. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829794" httpUri="https://zenodo.org/record/3829794/files/figure.png" pageId="2" pageNumber="3">Figs 8</figureCitation>
B–C, 9D–F) …………………………………………………………………………… 4
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</paragraph>
<paragraph id="C24BC1434500FFE1FF27BD69FAFAFEF7" blockId="3.[193,1398,267,329]" pageId="3" pageNumber="4">
<keyLead id="790562764500FFE1FF27BD69FAFAFEF7" pageId="3" pageNumber="4">
– Fore legs fully black; head narrow, about
<quantity id="050C6CA64500FFE1FD24BD6EFCB9FE9B" box="[706,821,267,293]" metricMagnitude="-2" metricUnit="m" metricValue="2.667" metricValueMax="3.048" metricValueMin="2.286" pageId="3" pageNumber="4" unit="in" value="1.05" valueMax="1.2" valueMin="0.9">0.9–1.2 in</quantity>
HL/HW; eyes not prominent; spinous plate absent (
<figureCitation id="5ACFDDC64500FFE1FF15BD4BFEC2FEF7" box="[243,334,302,329]" captionStart="Fig" captionStartId="11.[190,233,1928,1954]" captionTargetBox="[192,1397,1222,1886]" captionTargetId="figure@11.[192,1400,1222,1886]" captionTargetPageId="11" captionText="Fig. 8. Endophallus of Intybia spp., A. Intybia histrio (Kiesenwetter, 1874). B. I. pelegrini pelegrini (Pic, 1910). C. I. takaraensis (Nakane, 1955). D. I. kishiii (Nakane, 1955). Abbreviations: gp = gonoporal piece; l = ligula; sgp = semi-gonoporal piece; sp = spinous plate. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829794" httpUri="https://zenodo.org/record/3829794/files/figure.png" pageId="3" pageNumber="4">Figs8D</figureCitation>
,
<figureCitation id="5ACFDDC64500FFE1FEB3BD4AFEF7FEF7" box="[341,379,303,329]" captionStart="Fig" captionStartId="12.[190,233,1893,1919]" captionTargetBox="[191,1394,543,1850]" captionTargetId="figure@12.[189,1398,538,1851]" captionTargetPageId="12" captionText="Fig. 9. Spinous area and spinous plate of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica(Lewis,1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955).Abbreviations: sa = spinous area; sp = spinous plate. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829796" httpUri="https://zenodo.org/record/3829796/files/figure.png" pageId="3" pageNumber="4">9G</figureCitation>
)…………………………………………………………………
<taxonomicName id="05F4BAC04500FFE1FB82BD4AFAFAFEF7" authority="(Nakane, 1955)" baseAuthorityName="Nakane" baseAuthorityYear="1955" box="[1124,1398,302,329]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="kishiii">
<emphasis id="F0801D514500FFE1FB82BD4AFB48FEF7" bold="true" box="[1124,1220,302,329]" italics="true" pageId="3" pageNumber="4">I.kishiii</emphasis>
(Nakane,1955)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7900D9E64500FFE1FF5BBD15FAFAFE6E" pageId="3" pageNumber="4">
<paragraph id="C24BC1434500FFE1FF5BBD15FAFBFE35" blockId="3.[189,1399,368,465]" box="[189,1399,368,395]" pageId="3" pageNumber="4">
<keyLead id="790562764500FFE1FF5BBD15FAFBFE35" box="[189,1399,368,395]" pageId="3" pageNumber="4">
4. Body oval; femur black; endophallus with one spinous plate (
<figureCitation id="5ACFDDC64500FFE1FBE7BD15FBE4FE34" box="[1025,1128,368,394]" captionStart="Fig" captionStartId="11.[190,233,1928,1954]" captionTargetBox="[192,1397,1222,1886]" captionTargetId="figure@11.[192,1400,1222,1886]" captionTargetPageId="11" captionText="Fig. 8. Endophallus of Intybia spp., A. Intybia histrio (Kiesenwetter, 1874). B. I. pelegrini pelegrini (Pic, 1910). C. I. takaraensis (Nakane, 1955). D. I. kishiii (Nakane, 1955). Abbreviations: gp = gonoporal piece; l = ligula; sgp = semi-gonoporal piece; sp = spinous plate. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829794" httpUri="https://zenodo.org/record/3829794/files/figure.png" pageId="3" pageNumber="4">Figs 8C</figureCitation>
,
<figureCitation id="5ACFDDC64500FFE1FB9BBD15FB06FE34" box="[1149,1162,368,394]" captionStart="Fig" captionStartId="12.[190,233,1893,1919]" captionTargetBox="[191,1394,543,1850]" captionTargetId="figure@12.[189,1398,538,1851]" captionTargetPageId="12" captionText="Fig. 9. Spinous area and spinous plate of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica(Lewis,1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955).Abbreviations: sa = spinous area; sp = spinous plate. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829796" httpUri="https://zenodo.org/record/3829796/files/figure.png" pageId="3" pageNumber="4">9</figureCitation>
E–F)…………… 5
</keyLead>
</paragraph>
<paragraph id="C24BC1434500FFE1FF27BDF1FAFAFE6E" blockId="3.[189,1399,368,465]" pageId="3" pageNumber="4">
<keyLead id="790562764500FFE1FF27BDF1FAFAFE6E" pageId="3" pageNumber="4">
– Body elongate; femur yellow; endophallus with two spinous plates (
<figureCitation id="5ACFDDC64500FFE1FBE8BDF6FBE3FE13" box="[1038,1135,403,429]" captionStart="Fig" captionStartId="11.[190,233,1928,1954]" captionTargetBox="[192,1397,1222,1886]" captionTargetId="figure@11.[192,1400,1222,1886]" captionTargetPageId="11" captionText="Fig. 8. Endophallus of Intybia spp., A. Intybia histrio (Kiesenwetter, 1874). B. I. pelegrini pelegrini (Pic, 1910). C. I. takaraensis (Nakane, 1955). D. I. kishiii (Nakane, 1955). Abbreviations: gp = gonoporal piece; l = ligula; sgp = semi-gonoporal piece; sp = spinous plate. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829794" httpUri="https://zenodo.org/record/3829794/files/figure.png" pageId="3" pageNumber="4">Figs 8B</figureCitation>
,
<figureCitation id="5ACFDDC64500FFE1FB9DBDF6FB12FE13" box="[1147,1182,403,429]" captionStart="Fig" captionStartId="12.[190,233,1893,1919]" captionTargetBox="[191,1394,543,1850]" captionTargetId="figure@12.[189,1398,538,1851]" captionTargetPageId="12" captionText="Fig. 9. Spinous area and spinous plate of Intybia spp. A. Intybia histrio (Kiesenwetter, 1874). B. I. tsushimensis (Satô &amp; Ohbayashi, 1968). C. I. donan sp. nov. D. I. pelegrini pelegrini (Pic, 1910). E. I. niponica(Lewis,1895). F. I. takaraensis (Nakane, 1955). G. I. kishiii (Nakane, 1955).Abbreviations: sa = spinous area; sp = spinous plate. Scales = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3829796" httpUri="https://zenodo.org/record/3829796/files/figure.png" pageId="3" pageNumber="4">9D</figureCitation>
)………………… ……………………………………………………………………
<taxonomicName id="05F4BAC04500FFE1FC08BDD2FAFAFE6E" authority="pelegrini (Pic, 1910)" authorityName="pelegrini (Pic" authorityYear="1910" baseAuthorityName="Pic" baseAuthorityYear="1910" box="[1006,1398,438,465]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="pelegrini" subSpecies="pelegrini">
<emphasis id="F0801D514500FFE1FC08BDD2FB66FE6F" bold="true" box="[1006,1258,438,465]" italics="true" pageId="3" pageNumber="4">I. pelegrini pelegrini</emphasis>
(
<bibRefCitation id="A665BCB24500FFE1FB1DBDD3FAE2FE6E" author="Pic M." box="[1275,1390,438,464]" pageId="3" pageNumber="4" pagination="60 - 63" refId="ref17992" refString="Pic M. 1910. Coleopteres exotiques nouveaux ou peu connus. L' Echange, Revue Linneenne 26: 60 - 63." type="journal article" year="1910">Pic, 1910</bibRefCitation>
)
</taxonomicName>
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</paragraph>
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<keyStep id="7900D9E64500FFE1FF5BBD9DFAFAFDC2" pageId="3" pageNumber="4">
<paragraph id="C24BC1434500FFE1FF5BBD9DFAFAFD88" blockId="3.[189,1398,504,636]" pageId="3" pageNumber="4">
<keyLead id="790562764500FFE1FF5BBD9DFAFAFD88" pageId="3" pageNumber="4">
5. Elytral fascia clearly widened at suture; body black; male antennomere III broad bean-like shaped; female apterous……………………………………………………………
<taxonomicName id="05F4BAC04500FFE1FBAABE7EFAFAFD88" authority="(Lewis, 1895)" baseAuthorityName="Lewis" baseAuthorityYear="1895" box="[1100,1398,539,566]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="niponica">
<emphasis id="F0801D514500FFE1FBAABE7EFB42FD8B" bold="true" box="[1100,1230,539,565]" italics="true" pageId="3" pageNumber="4">I. niponica</emphasis>
(
<bibRefCitation id="A665BCB24500FFE1FB3ABE7EFAE2FD88" author="Lewis G." box="[1244,1390,539,566]" pageId="3" pageNumber="4" pagination="98 - 122" refId="ref17589" refString="Lewis G. 1895. On the Daschillidae and malacoderm Coleoptera of Japan. The Annals and Magazine of Natural History (6) 16: 98 - 122, pl. 6. https: // doi. org / 10.1080 / 00222939508680233" type="journal article" year="1895">Lewis, 1895</bibRefCitation>
)
</taxonomicName>
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</paragraph>
<paragraph id="C24BC1434500FFE1FF27BE5AFAFAFDC2" blockId="3.[189,1398,504,636]" pageId="3" pageNumber="4">
<keyLead id="790562764500FFE1FF27BE5AFAFAFDC2" pageId="3" pageNumber="4">
– Elytral fascia slightly widened at suture; body black with bluish luster; male antennomere III heartlike shaped; female hind wing present ………………………………
<taxonomicName id="05F4BAC04500FFE1FBF3BE07FAFAFDC2" authority="(Nakane, 1955)" baseAuthorityName="Nakane" baseAuthorityYear="1955" box="[1045,1398,609,636]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="takaraensis">
<emphasis id="F0801D514500FFE1FBF3BE07FB36FDC2" bold="true" box="[1045,1210,609,636]" italics="true" pageId="3" pageNumber="4">I. takaraensis</emphasis>
(Nakane, 1955)
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</paragraph>
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<paragraph id="C24BC1434500FFE1FF5BBEC6FAFAFD03" blockId="3.[189,1398,675,737]" box="[189,1398,675,701]" pageId="3" pageNumber="4">
<keyLead id="790562764500FFE1FF5BBEC6FAFAFD03" box="[189,1398,675,701]" pageId="3" pageNumber="4">6. Fore legs fully yellow; elytra with a yellow or divided into two spots by suture ………………………7</keyLead>
</paragraph>
<paragraph id="C24BC1434500FFE1FF27BEA2FAFAFD5F" blockId="3.[189,1398,675,737]" box="[193,1398,710,737]" pageId="3" pageNumber="4">
<keyLead id="790562764500FFE1FF27BEA2FAFAFD5F" box="[193,1398,710,737]" pageId="3" pageNumber="4">
– Fore legs fully black; elytra with a fascia ………
<taxonomicName id="05F4BAC04500FFE1FC8EBEA3FAFAFD5F" authority="(Sato &amp; Ohbayashi, 1968)" baseAuthorityName="Sato &amp; Ohbayashi" baseAuthorityYear="1968" box="[872,1398,710,737]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="tsushimensis">
<emphasis id="F0801D514500FFE1FC8EBEA3FBAFFD5F" bold="true" box="[872,1059,710,737]" italics="true" pageId="3" pageNumber="4">I. tsushimensis</emphasis>
(Satô &amp; Ohbayashi, 1968)
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="7900D9E64500FFE1FF5BBF6DFAFAFC11" pageId="3" pageNumber="4">
<paragraph id="C24BC1434500FFE1FF5BBF6DFAFAFCD6" blockId="3.[189,1398,775,943]" pageId="3" pageNumber="4">
<keyLead id="790562764500FFE1FF5BBF6DFAFAFCD6" pageId="3" pageNumber="4">
7. Elytral fascia varies in development (
<figureCitation id="5ACFDDC64500FFE1FD71BF62FD50FC9C" box="[663,732,775,802]" captionStart="Fig" captionStartId="6.[190,233,2000,2026]" captionTargetBox="[203,1388,1517,1947]" captionTargetId="figure@6.[189,1398,1502,1959]" captionTargetPageId="6" captionText="Fig. 3. Elytral pattern variation of Intybia histrio (Kiesenwetter, 1874)." figureDoi="http://doi.org/10.5281/zenodo.3829784" httpUri="https://zenodo.org/record/3829784/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
), sometimes interrupted by suture; antennae yellow; male antennomere III fusiform, with an oval funnel-shaped on the outer edge, with a twisted projection near base; male frons yellow in apical half ………………………
<taxonomicName id="05F4BAC04500FFE1FBEFBF2BFAFAFCD6" authority="(Kiesenwetter, 1874)" baseAuthorityName="Kiesenwetter" baseAuthorityYear="1874" box="[1033,1398,845,872]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="histrio">
<emphasis id="F0801D514500FFE1FBEFBF2BFBFFFCD6" bold="true" box="[1033,1139,845,872]" italics="true" pageId="3" pageNumber="4">I. histrio</emphasis>
(
<bibRefCitation id="A665BCB24500FFE1FB63BF2BFAE2FCD6" author="Kiesenwetter E. A. H. von" box="[1157,1390,846,872]" pageId="3" pageNumber="4" pagination="241 - 288" refId="ref17457" refString="Kiesenwetter E. A. H. von. 1874. Die Malacodermen Japans nach dem Ergebnisse der Sammulungen des Herrn G. Lewis wahread der Jahre 1869 - 1871. Berliner Entomologische Zeitschrift 18: 241 - 288. https: // doi. org / 10.1002 / mmnd. 18740180302" type="journal article" year="1874">Kiesenwetter, 1874</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C24BC1434500FFE1FF27BF17FAFAFC11" blockId="3.[189,1398,775,943]" pageId="3" pageNumber="4">
<keyLead id="790562764500FFE1FF27BF17FAFAFC11" pageId="3" pageNumber="4">
– Elytral fascia interrupted by suture; antennomeres V–XI infuscate; male antennomere III ear-like shaped; male frons fully black ………………………………………
<taxonomicName id="05F4BAC04500FFE1FBC6BFF0FAFAFC11" authority="(Nakane, 1956)" baseAuthorityName="Nakane" baseAuthorityYear="1956" box="[1056,1398,916,943]" class="Insecta" family="Melyridae" genus="Intybia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="kawasakii">
<emphasis id="F0801D514500FFE1FBC6BFF0FB39FC11" bold="true" box="[1056,1205,916,943]" italics="true" pageId="3" pageNumber="4">I. kawasakii</emphasis>
(
<bibRefCitation id="A665BCB24500FFE1FB20BFF1FAE2FC11" author="Nakane T." box="[1222,1390,916,943]" pageId="3" pageNumber="4" pagination="159 - 174" refId="ref17827" refString="Nakane T. 1956. New or little-known Coleoptera from Japan and its abject regions, XIII. The scientific Reports of the Saikyo University (Natural Science and Living Science) (A) 2: 29 - 44, 159 - 174." type="journal article" year="1956">Nakane, 1956</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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