1262 lines
118 KiB
XML
1262 lines
118 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.96.51962" ID-GBIF-Dataset="a134fec9-ee97-4d3e-ae3f-eb295c3267db" ID-GBIF-Taxon="164501242" ID-Pensoft-Pub="1860-0743-2-275" ID-Pensoft-UUID="FC312E66E3EA52F4A7706C2C191A8621" ID-ZooBank="4A3A5AE9D26340A08621430C7822CFF3" ModsDocID="1860-0743-2-275" checkinTime="1591887254079" checkinUser="pensoft" docAuthor="Viana, Sarah T. F. L. & Carvalho, Marcelo R. de" docDate="2020" docId="3D2634267D885CBFB191C3749F3B11AB" docLanguage="en" docName="ZoosystEvol 96(2): 275-311" docOrigin="Zoosystematics and Evolution 96 (2)" docSource="http://dx.doi.org/10.3897/zse.96.51962" docTitle="Squalus japonicus Ishikawa 1908" docType="treatment" docVersion="4" id="FC312E66E3EA52F4A7706C2C191A8621" lastPageNumber="275" masterDocId="FC312E66E3EA52F4A7706C2C191A8621" masterDocTitle="Squalus shiraii sp. nov. (Squaliformes, Squalidae), a new species of dogfish shark from Japan with regional nominal species revisited" masterLastPageNumber="311" masterPageNumber="275" pageId="0" pageNumber="275" updateTime="1634711231051" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Squalus shiraii sp. nov. (Squaliformes, Squalidae), a new species of dogfish shark from Japan with regional nominal species revisited</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Viana, Sarah T. F. L.</mods:namePart>
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<mods:affiliation>South African Institute for Aquatic Biodiversity, Private Bag 1015 Grahamstown, 6140, South Africa & Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo. Rua do Matao, Travessa 14, no 101, CEP 05508 - 090, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-6905-9292</mods:nameIdentifier>
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<mods:nameIdentifier type="email">stviana@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Carvalho, Marcelo R. de</mods:namePart>
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<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo. Rua do Matao, Travessa 14, no 101, CEP 05508 - 090, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>96</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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<mods:extent unit="page">
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<mods:start>275</mods:start>
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<mods:end>311</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zse.96.51962</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.96.51962</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-2-275</mods:identifier>
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<mods:identifier type="ZooBank">4A3A5AE9D26340A08621430C7822CFF3</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FC312E66E3EA52F4A7706C2C191A8621</mods:identifier>
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<treatment ID-GBIF-Taxon="164501242" LSID="urn:lsid:plazi:treatment:3D2634267D885CBFB191C3749F3B11AB" httpUri="http://treatment.plazi.org/id/3D2634267D885CBFB191C3749F3B11AB" lastPageNumber="275" pageId="0" pageNumber="275">
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<subSubSection pageId="0" pageNumber="275" type="nomenclature">
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<paragraph pageId="0" pageNumber="275">
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<taxonomicName LSID="3D263426-7D88-5CBF-B191-C3749F3B11AB" authority="Ishikawa, 1908" authorityName="Ishikawa" authorityYear="1908" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus japonicus" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="japonicus">Squalus japonicus Ishikawa, 1908</taxonomicName>
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Upper (A, C, E, G, I) and lower (B, D, F, H, J) teeth of S. shiraii sp. nov. (A, B; HUMZ 80329), S. mitsukurii (C, D; HUMZ 33680), S. japonicus (E, F; HUMZ 189737), S. brevirostris (G, H; HUMZ 189762) and S. suckleyi (I, J; NSMT-P 92640). Scale bars: 1 mm." figureDoi="10.3897/zse.96.51962.figure3" httpUri="https://binary.pensoft.net/fig/420111" pageId="0" pageNumber="275">Figs 3E, F</figureCitation>
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Squalus japonicus: lateral (A-C) and ventral (D, E) views; first (F) and second (G) dorsal fins; pectoral (H) and caudal (I) fins. NSMT-P 44380 (neotype), adult male, 645 mm TL (A, D, F-I); HUMZ 189738, adult male, 535 mm TL (B); HUMZ 189687, juvenile male, 380 mm TL (C, E). Scale bars: 50 mm (A-E); 20 mm (F-I)." figureDoi="10.3897/zse.96.51962.figure4" httpUri="https://binary.pensoft.net/fig/420112" pageId="0" pageNumber="275">, 4</figureCitation>
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scanning electron microscopy of the dermal denticles. Squalus shiraii sp. nov. (A, B): HUMZ 149389 (holotype), adult male, 590 mm TL. S. formosus (C, D): CSIRO H- 6816 - 01 (holotype), adult male, 720 mm TL. S. mitsukurii (E, F): HUMZ 102987, adult male, 970 mm TL. S. japonicus (G, H): NSMT-P 44380 (neotype), adult male, 645 mm TL (G); NSMT-P 91127, juvenile female, 410 mm TL (H). S. brevirostris (I, J): NSMT-P 47378, adult female, 600 mm TL (I); KAUM-I 187, adult female, 578 mm TL (J). S. suckleyi (K, L): NSMT P- 92640, adult female, 970 mm TL. Scale bars: 50 μm (B); 100 μm (D, F, L); 200 μm (A, C, E, G-K)." figureDoi="10.3897/zse.96.51962.figure7" httpUri="https://binary.pensoft.net/fig/420115" pageId="0" pageNumber="275">, 7G, H</figureCitation>
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Map of the North-western Pacific Ocean, showing the geographical distribution of Squalus species. A: S. shiraii sp. nov. (red); B: S. mitsukurii (orange); C: S. japonicus (brown); D: S. brevirostris (yellow); E: S. acutirostris (green); F: S. suckleyi (pink). Star: holotype / neotype; Dotted circle: paratypes; Circle: other material." figureDoi="10.3897/zse.96.51962.figure8" httpUri="https://binary.pensoft.net/fig/420116" pageId="0" pageNumber="275">, 8C</figureCitation>
|
||
;
|
||
<figureCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. External measurements for S. japonicus expressed as percentage of total length (% TL). Morphometric data of S. nasutus is also provided for comparisons. TL is expressed in millimeters. N: number of specimens; x: mean; SD: standard deviation." pageId="0" pageNumber="275">Tables 3</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Table 6" captionStartId="T6" captionText="Table 6. Meristic data for species of Squalus from Japan. H: holotype; P: paratypes; NT: neotype; N: number of specimens; x; mean; m: mode; SD: standard deviation." pageId="0" pageNumber="275">6 Japanese spurdog; Togari-tsunozame (Japanese)</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="275" type="reference_group">
|
||
<paragraph pageId="0" pageNumber="275">
|
||
<taxonomicName authorityName="Ishikawa" authorityYear="1908" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus japonicus" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="japonicus">Squalus japonicus</taxonomicName>
|
||
<bibRefCitation author="Ishikawa, C" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="275" pagination="71 - 73" refId="B41" refString="Ishikawa, C, 1908. Description of a new species of squaloid shark from Japan. Proceedings of the Academy of Natural Sciences of Philadelphia 60: 71 - 73" title="Description of a new species of squaloid shark from Japan." volume="60" year="1908">Ishikawa 1908</bibRefCitation>
|
||
: 71-73 (original description; not illustrated; type locality: Sagami Bay and Kagoshima, Japan; three syntypes cited and deposited in the Imperial Museum of Tokyo are considered lost: ZUMT uncatalogued, juvenile male, unknown TL, adult male, 700 mm TL, female, unknown TL);
|
||
<bibRefCitation author="Chen, CT" journalOrPublisher="Japanese Journal of Ichthyology" pageId="0" pageNumber="275" pagination="26 - 42" refId="B8" refString="Chen, CT, Taniuchi, T, Nose, Y, 1979. Blainville's dogfish, Squalus blainvillei, from Japan, with notes on S. mitsukurii and S. japonicus. Japanese Journal of Ichthyology 26 (1): 26 - 42" title="Blainville's dogfish, Squalus blainvillei, from Japan, with notes on S. mitsukurii and S. japonicus." volume="26" year="1979">Chen et al. 1979</bibRefCitation>
|
||
: 26-42 (re-description; Japan);
|
||
<bibRefCitation author="Parin, N" journalOrPublisher="Journal of Ichthyology" pageId="0" pageNumber="275" pagination="43 - 50" refId="B70" refString="Parin, N, 1987. Species of spiny dogfish of genus Squalus, living on Southeastern Pacific Ocean Seamounts. Journal of Ichthyology 27 (5): 43 - 50" title="Species of spiny dogfish of genus Squalus, living on Southeastern Pacific Ocean Seamounts." volume="27" year="1987">Parin 1987</bibRefCitation>
|
||
: 48-49 (cited; Japan);
|
||
<bibRefCitation author="Compagno, LJV" editor="Carpenter, KE" journalOrPublisher="The living marine resources of the western central pacific. Cephalopods, crustaceans, holothurans and sharks 2. FAO, Senckenbergiana Biologica, Rome" pageId="0" pageNumber="275" pagination="687 - 1396" refId="B16" refString="Compagno, LJV, Niem, VH, 1998. Several family accounts. In: Carpenter, KE, Niem, VH, Eds., FAO species identification guide for fishery purposes. The living marine resources of the western central pacific. Cephalopods, crustaceans, holothurans and sharks 2. FAO, Senckenbergiana Biologica, Rome: 687 - 1396" title="Several family accounts." volumeTitle="FAO species identification guide for fishery purposes." year="1998">Compagno and Niem 1998</bibRefCitation>
|
||
: 1222, 1229 (listed, cited; Western North and Central Pacific Ocean);
|
||
<bibRefCitation author="Zhu, Y" journalOrPublisher="Oceanologia et Limnologia Sinica" pageId="0" pageNumber="275" refId="B103" refString="Zhu, Y, Meng, QW, 2001. Faina Sinica - Cyclostomata, Chondrichthyes. Science Press, Beijing." title="Faina Sinica - Cyclostomata, Chondrichthyes. Science Press, Beijing." year="2001">Zhu and Meng 2001</bibRefCitation>
|
||
: 311, 316-317 (cited, description; Northwest Pacific Ocean);
|
||
<bibRefCitation author="Nakabo, T" journalOrPublisher="Japanese Journal of Ichthyology" pageId="0" pageNumber="275" refId="B61" refString="Nakabo, T, 2002. Fishes of Japan with pictorial keys to the species, English edition. Tokai University Press, Tokyo." title="Fishes of Japan with pictorial keys to the species, English edition. Tokai University Press, Tokyo." year="2002">Nakabo 2002</bibRefCitation>
|
||
: 155 (listed; Ryukyu Islands, Okinawa Trough, Southern Japan);
|
||
<bibRefCitation author="Compagno, LJV" journalOrPublisher="Vol. 1. Introduction, molluscs, crustaceans, hagfishes, sharks, batoid fishes and chimaeras. FAO Species identification guide for fishery purposes and American Society of Ichthyologists Special Publication 5. FAO, Rome" pageId="0" pageNumber="275" refId="B14" refString="Compagno, LJV, Dando, M, Fowler, S, 2005a. Sharks of the World. Princeton University Press, New Jersey." title="Sharks of the World. Princeton University Press, New Jersey." year="2005 a">Compagno et al. 2005a</bibRefCitation>
|
||
: 75-76 (description; Northwest Pacific Ocean);
|
||
<bibRefCitation author="Ward, RD" editor="Last, PR" journalOrPublisher="CSIRO Marine and Atmospheric Research Paper 14, Hobart" pageId="0" pageNumber="275" pagination="117 - 130" refId="B91" refString="Ward, RD, Holmes, BH, Zemlak, TS, Smith, PJ, 2007. DNA barcoding discriminates spurdogs of the genus Squalus. In: Last, PR, White, WT, Pogonoski, JJ, Eds., Descriptions of new dogfishes of the genus Squalus (Squaloidea: Squalidae). CSIRO Marine and Atmospheric Research Paper 14, Hobart: 117 - 130" title="DNA barcoding discriminates spurdogs of the genus Squalus." volumeTitle="Descriptions of new dogfishes of the genus Squalus (Squaloidea: Squalidae)." year="2007">Ward et al. 2007</bibRefCitation>
|
||
: 118, 123, 129, 130 (cited; Taiwan);
|
||
<bibRefCitation DOI="https://doi.org/10.1206/754.1" author="Naylor, GJP" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="275" pagination="1 - 262" refId="B66" refString="Naylor, GJP, Caira, JN, Jensen, K, Rosana, AM, White, WT, Last, PR, 2012. A DNA sequence-based approach to the identification of shark and rays species and its implication of global elasmobranch diversity and parasitology. Bulletin of the American Museum of Natural History 367: 1 - 262, DOI: https://doi.org/10.1206/754.1" title="A DNA sequence-based approach to the identification of shark and rays species and its implication of global elasmobranch diversity and parasitology." url="https://doi.org/10.1206/754.1" volume="367" year="2012">Naylor et al. 2012</bibRefCitation>
|
||
: 58 (cited; Taiwan);
|
||
<bibRefCitation author="Nakabo, T" journalOrPublisher="Japanese Journal of Ichthyology" pageId="0" pageNumber="275" refId="B62" refString="Nakabo, T, 2013. Fishes of Japan with pictorial keys to the species. Tokai University Press, Tokyo." title="Fishes of Japan with pictorial keys to the species. Tokai University Press, Tokyo." year="2013">Nakabo 2013</bibRefCitation>
|
||
: 194 (listed; Ryukyu Islands, Okinawa Trough, Southern Japan);
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3752.1.16" author="Straube, N" journalOrPublisher="Zootaxa" pageId="0" pageNumber="275" pagination="256 - 278" refId="B80" refString="Straube, N, White, WT, HO, H-C, Rochel, E, Corrigan, S, Li, C, Naylor, GJP, 2013. A DNA sequence-based identification checklist for Taiwanese chondrichthyans. Zootaxa 3752: 256 - 278, DOI: https://doi.org/10.11646/zootaxa.3752.1.16" title="A DNA sequence-based identification checklist for Taiwanese chondrichthyans." url="https://doi.org/10.11646/zootaxa.3752.1.16" volume="3752" year="2013">Straube et al. 2013</bibRefCitation>
|
||
: 264-265 (cited; Taiwan);
|
||
<bibRefCitation author="Shinohara, G" editor="Fujita, T" journalOrPublisher="National Museum of Nature and Science Monographs 44, Tokyo" pageId="0" pageNumber="275" pagination="225 - 291" refId="B75" refString="Shinohara, G, Nakae, M, Ueda, Y, Kojima, S, Matsuura, K, 2014. Annotated checklist of deep-sea fishes of the Sea of Japan. In: Fujita, T, Ed., Deep-sea fauna of the Sea of Japan. National Museum of Nature and Science Monographs 44, Tokyo: 225 - 291" title="Annotated checklist of deep-sea fishes of the Sea of Japan." volumeTitle="Deep-sea fauna of the Sea of Japan." year="2014">Shinohara et al. 2014</bibRefCitation>
|
||
: 233 (cited; Japan).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
<taxonomicName baseAuthorityName="Girard" baseAuthorityYear="1854" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus suckleyi" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="suckleyi">Squalus suckleyi</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Fowler, HW" journalOrPublisher="Bulletin United States National Museum" pageId="0" pageNumber="275" pagination="1 - 879" refId="B35" refString="Fowler, HW, 1941. Contributions to the biology of the Philippine archipelago and adjacent regions. The fishes of the groups Elasmocephalii, Holocephali, Isospondyli, and Ostariophysi obtained by the United States Fisheries Steamer "Albatross" in 1907 to 1910, chiefly in the Philippine islands and adjacent seas. Bulletin United States National Museum 100: 1 - 879" title="Contributions to the biology of the Philippine archipelago and adjacent regions. The fishes of the groups Elasmocephalii, Holocephali, Isospondyli, and Ostariophysi obtained by the United States Fisheries Steamer " Albatross " in 1907 to 1910, chiefly in the Philippine islands and adjacent seas." volume="100" year="1941">Fowler 1941</bibRefCitation>
|
||
: 259 (listed; Japan).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="275" type="neotype">
|
||
<paragraph pageId="0" pageNumber="275">Neotype.</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">NSMT-P 44380, adult male, 645 mm TL, Senoumi Bank, Suruga Bay, Honshu, Japan, approximate coordinates 34°43'N and 138.33E, 270-300 m depth. Collected on 13 April 1982, collector Kazunari Yano.</paragraph>
|
||
<caption ID-Table-UUID="90113632AD9FF7A8367D3D5F96752087" httpUri="http://table.plazi.org/id/90113632AD9FF7A8367D3D5F96752087" pageId="0" pageNumber="275" start="Table 3" startId="T3">
|
||
<paragraph pageId="0" pageNumber="275">
|
||
<emphasis bold="true" pageId="0" pageNumber="275">Table 3.</emphasis>
|
||
External measurements for
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
expressed as percentage of total length (% TL). Morphometric data of
|
||
<taxonomicName genus="S." lsidName="S. nasutus" pageId="0" pageNumber="275" rank="species" species="nasutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. nasutus</emphasis>
|
||
</taxonomicName>
|
||
is also provided for comparisons. TL is expressed in millimeters. N: number of specimens;
|
||
<emphasis italics="true" pageId="0" pageNumber="275">x</emphasis>
|
||
: mean; SD: standard deviation.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
<table pageId="0" pageNumber="275">
|
||
<tr pageId="0" pageNumber="275">
|
||
<th colspan="1" isEmpty="true" pageId="0" pageNumber="275" rowspan="2">-</th>
|
||
<th colspan="3" pageId="0" pageNumber="275" rowspan="1">
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="2" pageId="0" pageNumber="275" rowspan="1">
|
||
<taxonomicName genus="S." lsidName="S. nasutus" pageId="0" pageNumber="275" rank="species" species="nasutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. nasutus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="275" rowspan="2">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">x</emphasis>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="275" rowspan="2">SD</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<th colspan="1" pageId="0" pageNumber="275" rowspan="1">Neotype</th>
|
||
<th colspan="1" pageId="0" pageNumber="275" rowspan="1">N</th>
|
||
<th colspan="1" isEmpty="true" pageId="0" pageNumber="275" rowspan="1">-</th>
|
||
<th colspan="1" pageId="0" pageNumber="275" rowspan="1">N</th>
|
||
<th colspan="1" isEmpty="true" pageId="0" pageNumber="275" rowspan="1">-</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Total length (mm)</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">645.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">281.0-560.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">362.0-580.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">444.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">97.2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Precaudal length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">79.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">77.3-80.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">79.6-81.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">79.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pre-second dorsal length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">62.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">59.1-62.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">60.8-64.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">61.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pre-first dorsal length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">31.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">30.0-33.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">31.2-32.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">32.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pre-vent length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">47.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">47.0-50.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">46.1-51.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">48.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Prepelvic length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">43.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">44.8-47.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">44.5-48.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">45.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Prepectoral length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">22.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">22.0-24.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">22.8-25.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">23.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Head length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">23.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">23.0-24.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">23.5-26.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">23.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Prebranchial length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">18.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">19.4-21.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">20.3-22.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">20.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Prespiracular length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.5-15.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.7-15.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">14.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Preorbital length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.3-10.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.2-10.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Prenarial length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.8-7.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.6-7.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.4</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Preoral length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.1-12.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.8-13.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Inner nostril-labial furrow space</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.4-5.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.5-5.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.3</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Mouth width</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.3-7.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.4-7.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Labial furrow length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.9-2.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.9-2.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Internarial space</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.1-5.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.3-5.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.4</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Interorbital space</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.6-9.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.2-9.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Eye length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.8-5.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.2-5.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.4</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Eye height</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.4-2.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.8-2.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Spiracle length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.0-1.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1-1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First gill-slit height</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.3-2.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.3-1.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Fifth gill-slit height</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.5-2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.8-2.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Interdorsal space</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">24.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">18.4-23.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">23.0-25.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">23.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Dorsal-caudal space</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.5-11.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.9-13.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pectoral-pelvic space</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">18.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">18.7-22.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">17.8-23.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">19.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.4</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pelvic-caudal space</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">27.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">23.6-28.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">25.0-28.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">26.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.3</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.0-14.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.8-13.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin anterior margin</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.6-13.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.9-11.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin base length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.2-8.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.8-8.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin height</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.6-10.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.9-7.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin inner margin</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.1-6.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.0-5.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.4</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin posterior margin</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.7-10.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.8-8.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin spine length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">14</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.0-3.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.4-3.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin spine base width</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5-0.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6-0.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.3-16.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.1-13.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin anterior margin</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.5-14.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.1-11.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.3</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin base length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.9-10.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.9-8.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin height</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.4-7.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.8-5.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin inner margin</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.3-5.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.0-4.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.4</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin posterior margin</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.4-6.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.5-5.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin spine length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">14</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.1-5.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.9-4.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin spine base width</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7-1.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7-1.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pectoral-fin anterior margin length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.9-14.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.4-14.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pectoral-fin inner margin length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.8-8.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.2-8.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pectoral-fin base length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.8-4.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.1-5.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.4</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pectoral-fin posterior margin length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.8-10.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.2-8.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pelvic length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.6-12.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.3-12.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pelvic-fin inner margin length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.6-6.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.3-7.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Dorsal caudal margin length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">20.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">19.6-21.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">18.1-19.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">19.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Preventral caudal margin length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.6-11.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.2-12.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.0</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Caudal fork width</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.0-7.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.2-7.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.4</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Head width at nostrils</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.8-8.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.9-8.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Head width at mouth</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.0-11.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.2-11.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Head width</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.0-13.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.2-12.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Trunk width</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.1-10.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.7-12.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Abdomen width</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.9-10.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.3-9.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Head height</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.5-10.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.9-9.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Trunk height</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.3-11.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.9-10.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Abdomen height</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.0</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.7-12.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.3-12.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Clasper outer length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.4-4.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.1-5.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.7</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="275">
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Clasper inner length</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.8-7.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.3-9.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.6</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="275" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="275">Additional material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
CSIRO H 6294-26, juvenile male, 440 mm TL, Tashi fish market, near I-Lan (NE coast), Taiwan; CSIRO H 6294-27, juvenile female, 360 mm TL, locality same as CSIRO H6294-26; CSIRO H 6294-31, adult female, 540 mm TL, locality same as CSIRO H6294-26; HUMZ 40026, juvenile male, 215 mm TL, unknown locality; HUMZ 80223, adult male, unknown TL, near Okinawa, Japan,
|
||
<geoCoordinate degrees="25" direction="north" minutes="33.8" orientation="latitude" precision="92" value="25.563334">25°33.8'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="126" direction="east" minutes="25.2" orientation="longitude" precision="92" value="126.42">126°25.2'E</geoCoordinate>
|
||
, 310 m depth; HUMZ 95213, unknown sex and TL, East China Sea,
|
||
<geoCoordinate degrees="27" direction="north" minutes="46" orientation="latitude" precision="925" value="27.766666">27°46'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="126" direction="east" minutes="15.3" orientation="longitude" precision="92" value="126.255">126°15.3'E</geoCoordinate>
|
||
, 296-465 m depth; HUMZ 189642, adult male, 530 mm TL, East China Sea; HUMZ 189673, juvenile female, 430 mm TL, East China Sea; HUMZ 189675, juvenile female, 277 mm TL, East China Sea; HUMZ 189676, juvenile male, 288 mm TL, East China Sea; HUMZ 189678, adult male, 545 mm TL, East China Sea; HUMZ 189682, juvenile male, 365 mm TL, East China Sea; HUMZ 189685, adult female, 512 mm TL, East China Sea; HUMZ 189687, juvenile male, 380 mm TL, East China Sea; HUMZ 189689, juvenile male, 276 mm TL, East China Sea; HUMZ 189693, juvenile female, 380 mm TL, East China Sea; HUMZ 189695, juvenile female, 290 mm TL, East China Sea; HUMZ 189696, juvenile male, 291 mm TL, East China Sea; HUMZ 189701, juvenile male, 287 mm TL, East China Sea; HUMZ 189705, juvenile male, 282 mm TL, East China Sea; HUMZ 189735, adult male, 518 mm TL, East China Sea; HUMZ 189737, adult male, 560 mm TL, East China Sea; HUMZ 189738, adult male, 535 mm TL, East China Sea; HUMZ 189739, adult male, 530 mm TL, East China Sea; HUMZ 191688, juvenile female, 340 mm TL, Okinawa, Japan; HUMZ 191689, juvenile male, 281 mm TL, Okinawa, Japan; HUMZ 191691, juvenile male, 338 mm TL, Okinawa, Japan; HUMZ 191693, juvenile female, 410 mm TL, Okinawa, Japan; HUMZ 191694, juvenile female, 440 mm TL, Okinawa, Japan; HUMZ 191695, juvenile male, 338 mm TL, Okinawa, Japan; HUMZ 191697, juvenile female, 330 mm TL, Okinawa, Japan; HUMZ 191698, juvenile male, 388 mm TL, Okinawa, Japan; HUMZ 191699, juvenile female, 410 mm TL, Okinawa, Japan; NMMB P 15491, adult male, 593 mm TL, Pingtung county, Taiwan; NMMB P 15691, adult male, 540 mm TL, locality same as NMMB P 15491; NMMB P 15696, adult male, 580 mm TL, locality same as NMMB P 15491; NMMB P 15698, juvenile female, 448 mm TL, locality same as NMMB P 15491; NMMB P 15699, adult male, 513 mm TL, locality same as NMMB P 15491; NMMB P 15700, juvenile female, 443 mm TL, locality same as NMMB P 15491; NSMT-P 47384, juvenile female, 433 mm TL, Central Pacific Ocean; NSMT-P 67530, adult female, 560 mm TL, East China Sea,
|
||
<geoCoordinate degrees="31" direction="north" minutes="21.31" orientation="latitude" precision="9" value="31.355167">31°21.31'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="128" direction="east" minutes="19.12" orientation="longitude" precision="9" value="128.31866">128°19.12'E</geoCoordinate>
|
||
, 254-260 m depth; NSMT-P 91127, juvenile female, 410 mm TL, Mimase Fishing Port, Kochi City, Kochi Prefecture, Shikoku, Japan; UF 44332, adult female, 490 mm TL, Seno-Umi, Honshu Island, Suruga Bay, Japan, 195 m depth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="275" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="275">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
Single value corresponds to neotype and range values include all examined material from which data was taken. Species of
|
||
<taxonomicName class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">Squalus</emphasis>
|
||
</taxonomicName>
|
||
clearly distinct from all its congeners (except with
|
||
<taxonomicName genus="S." lsidName="S. melanurus" pageId="0" pageNumber="275" rank="species" species="melanurus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. melanurus</emphasis>
|
||
</taxonomicName>
|
||
Fourmanoir, 1979 and
|
||
<taxonomicName genus="S." lsidName="S. nasutus" pageId="0" pageNumber="275" rank="species" species="nasutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. nasutus</emphasis>
|
||
</taxonomicName>
|
||
) by a combination of characters: snout obtuse at tip and conspicuously elongate (prenarial length 5.9%-6.8% TL in adults); mouth markedly narrow, its width 1.6, 1.0-1.6 times prenarial length; pectoral fins conspicuously tapered (pectoral-fin posterior margin length 8.6%, 7.8%-9.2% TL in adults); markedly elongate preoral length 10.7%, 10.7%-12.2% TL.
|
||
<taxonomicName authorityName="Ishikawa" authorityYear="1908" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus japonicus" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">Squalus japonicus</emphasis>
|
||
</taxonomicName>
|
||
is differentiated from its regional congeners
|
||
<taxonomicName genus="S." lsidName="S. mitsukurii" pageId="0" pageNumber="275" rank="species" species="mitsukurii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. mitsukurii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName genus="S." lsidName="S. acutirostris" pageId="0" pageNumber="275" rank="species" species="acutirostris">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. acutirostris</emphasis>
|
||
</taxonomicName>
|
||
by head markedly tapered with head width at mouth 9.7%, 9.7%-10.6% TL (vs. 10.9%-12.2% TL for
|
||
<taxonomicName genus="S." lsidName="S. mitsukurii" pageId="0" pageNumber="275" rank="species" species="mitsukurii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. mitsukurii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="275">vs.</emphasis>
|
||
10.7%-12.1% TL for
|
||
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
vs. 11.2%-11.6% TL for
|
||
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. vs. 10.7%-11.6% TL for
|
||
<taxonomicName genus="S." lsidName="S. acutirostris" pageId="0" pageNumber="275" rank="species" species="acutirostris">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. acutirostris</emphasis>
|
||
</taxonomicName>
|
||
) and caudal fin with conspicuous black caudal bar and black upper caudal blotch (vs. black caudal bar and black upper caudal blotch absent). It is separated from
|
||
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by more elongate pre-first dorsal length (31.8%, 31.8%-33.0% TL vs. 29.6%, 28.9%-31.7% TL for
|
||
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
and 30.5%, 29.9%-31.2% TL for
|
||
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.), larger preorbital length (8.7%, 8.7%-9.3% TL vs. 6.7%, 6.5%-7.5% TL for
|
||
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
vs. 7.9%, 7.4%-7.9% TL for
|
||
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.) and from
|
||
<taxonomicName genus="S." lsidName="S. mitsukurii" pageId="0" pageNumber="275" rank="species" species="mitsukurii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. mitsukurii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by smaller first dorsal fin, its base length 6.7%, 6.6%-7.5% TL (vs. 8.2%, 7.6%-9.0% TL for
|
||
<taxonomicName genus="S." lsidName="S. mitsukurii" pageId="0" pageNumber="275" rank="species" species="mitsukurii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. mitsukurii</emphasis>
|
||
</taxonomicName>
|
||
vs. 7.7%, 7.7%-8.7% TL for
|
||
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.). It is also distinct from
|
||
<taxonomicName genus="S." lsidName="S. mitsukurii" pageId="0" pageNumber="275" rank="species" species="mitsukurii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. mitsukurii</emphasis>
|
||
</taxonomicName>
|
||
by larger inner nostril-labial furrow space (4.6%, 4.6%-5.1% TL vs. 4.3%, 3.9%-4.5% TL for
|
||
<taxonomicName genus="S." lsidName="S. mitsukurii" pageId="0" pageNumber="275" rank="species" species="mitsukurii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. mitsukurii</emphasis>
|
||
</taxonomicName>
|
||
) and from
|
||
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
by larger internarial space (4.1%, 4.1%-4.7% TL vs. 3.6%, 3.4%-3.8% TL for
|
||
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
). It differs from
|
||
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by lower first dorsal fin, its height 6.6%, 6.6%-7.5% TL (vs. 8.1%, 7.9%-8.2% TL for
|
||
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.) and lower number of precaudal and total vertebrae (87, 80-88 and 116, 104-116 vs. 93, 91-94 and 122, 120-123 for
|
||
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.).
|
||
<taxonomicName authorityName="Ishikawa" authorityYear="1908" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus japonicus" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">Squalus japonicus</emphasis>
|
||
</taxonomicName>
|
||
also differs from
|
||
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
by dermal denticles tricuspidate, pectoral-fin free rear tips rounded, postventral caudal margins not uniformly white (vs. unicuspid dermal denticles, pectoral-fin free rear tips pointed, postventral caudal margins uniformly white).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="275" type="description">
|
||
<paragraph pageId="0" pageNumber="275">Description</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">. Single values correspond to the neotype and ranges to all other examined material from which data were obtained.</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="275">External morphology.</emphasis>
|
||
Medium sized species (512-645 mm maximum TL in adults), with body fusiform and skinny, somewhat arched from trunk to abdomen, turning slim to caudal fin; head height 0.9 (0.8-1.1) times trunk and abdomen heights (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Squalus japonicus: lateral (A-C) and ventral (D, E) views; first (F) and second (G) dorsal fins; pectoral (H) and caudal (I) fins. NSMT-P 44380 (neotype), adult male, 645 mm TL (A, D, F-I); HUMZ 189738, adult male, 535 mm TL (B); HUMZ 189687, juvenile male, 380 mm TL (C, E). Scale bars: 50 mm (A-E); 20 mm (F-I)." figureDoi="10.3897/zse.96.51962.figure4" httpUri="https://binary.pensoft.net/fig/420112" pageId="0" pageNumber="275">4A-C</figureCitation>
|
||
). Head compressed and very narrow, its width 1.2 (1.1-1.3) times trunk width and 1.3 (1.0-1.7) times broader than abdomen width; head elongate, its length 23.6% (23.0%-24.5% TL). Snout conical and obtuse at the tip, conspicuously elongate (preorbital length 9.4%, 8.3%-10.1% TL); anterior margin of nostrils bi-lobed with first lobe larger than second lobe; nostrils much closer to the mouth than to the tip of the snout, its prenarial length 1.4 (1.2-1.6) times the distance from nostril to upper labial furrow (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Squalus japonicus: lateral (A-C) and ventral (D, E) views; first (F) and second (G) dorsal fins; pectoral (H) and caudal (I) fins. NSMT-P 44380 (neotype), adult male, 645 mm TL (A, D, F-I); HUMZ 189738, adult male, 535 mm TL (B); HUMZ 189687, juvenile male, 380 mm TL (C, E). Scale bars: 50 mm (A-E); 20 mm (F-I)." figureDoi="10.3897/zse.96.51962.figure4" httpUri="https://binary.pensoft.net/fig/420112" pageId="0" pageNumber="275">4D, E</figureCitation>
|
||
). Eyes oval with anterior margin convex and posterior margin notched; eyes elongate, its length 2.5 (2.0-3.5) times greater than its height. Prespiracular length 0.6 (0.6-0.7) times prepectoral length. Spiracles crescent, placed laterally just posterior to the eyes; spiracles small, its length 0.3 (0.2-0.4) times eye length. Prebranchial length 1.4 (1.4-1.5) times larger than prespiracular length. Gill slits convex, located anterior to origin of pectoral fins; gill slits vertical and low with height of fifth gill slit 1.2 (0.9-1.6) times height of first gill slit.
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.96.51962.figure4" httpUri="https://binary.pensoft.net/fig/420112" pageId="0" pageNumber="275" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="275">
|
||
<emphasis bold="true" pageId="0" pageNumber="275">Figure 4.</emphasis>
|
||
<taxonomicName authorityName="Ishikawa" authorityYear="1908" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus japonicus" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">Squalus japonicus</emphasis>
|
||
</taxonomicName>
|
||
: lateral (
|
||
<emphasis bold="true" pageId="0" pageNumber="275">A-C</emphasis>
|
||
) and ventral (
|
||
<emphasis bold="true" pageId="0" pageNumber="275">D, E</emphasis>
|
||
) views; first (
|
||
<emphasis bold="true" pageId="0" pageNumber="275">F</emphasis>
|
||
) and second (
|
||
<emphasis bold="true" pageId="0" pageNumber="275">G</emphasis>
|
||
) dorsal fins; pectoral (
|
||
<emphasis bold="true" pageId="0" pageNumber="275">H</emphasis>
|
||
) and caudal (
|
||
<emphasis bold="true" pageId="0" pageNumber="275">I</emphasis>
|
||
) fins. NSMT-P 44380 (neotype), adult male, 645 mm TL (
|
||
<emphasis bold="true" pageId="0" pageNumber="275">A, D, F-I</emphasis>
|
||
); HUMZ 189738, adult male, 535 mm TL (
|
||
<emphasis bold="true" pageId="0" pageNumber="275">B</emphasis>
|
||
); HUMZ 189687, juvenile male, 380 mm TL (
|
||
<emphasis bold="true" pageId="0" pageNumber="275">C, E</emphasis>
|
||
). Scale bars: 50 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="275">A-E</emphasis>
|
||
); 20 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="275">F-I</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
Preoral length 1.8 (1.4-2.7) times greater than mouth width. Mouth markedly arched and narrow, its width 1.5 (1.0-1.6) times broader than internarial width and 1.0 (0.6-1.3) times prenarial length; upper labial furrow elongate, its length 2.1% (1.9%-2.4% TL), with fold very thin; lower labial furrow also elongate, although without fold. Teeth unicuspid and tiny on both jaws, although upper teeth smaller than lower teeth; cusp short and oblique, thicker on lower teeth than upper teeth; mesial cutting edge convex; mesial heel markedly notched; distal heel rounded; apron thick and short, somewhat conical in lower teeth (Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Upper (A, C, E, G, I) and lower (B, D, F, H, J) teeth of S. shiraii sp. nov. (A, B; HUMZ 80329), S. mitsukurii (C, D; HUMZ 33680), S. japonicus (E, F; HUMZ 189737), S. brevirostris (G, H; HUMZ 189762) and S. suckleyi (I, J; NSMT-P 92640). Scale bars: 1 mm." figureDoi="10.3897/zse.96.51962.figure3" httpUri="https://binary.pensoft.net/fig/420111" pageId="0" pageNumber="275">3E-F</figureCitation>
|
||
). Two series of functional teeth on each jaw; 13-0-13 teeth rows for neotype (13-0-13 for other material) in upper jaw; 10-0-10 (11-0-11) teeth rows in lower jaw.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
Pre-first dorsal length 1.4 (1.3-1.5) times larger than prepectoral length. Origin of first dorsal fin prior to the vertical line traced at pectoral-fin free rear tips. First dorsal fin evidently slender at fin web (broader in young juveniles); first dorsal-fin anterior margin convex; first dorsal-fin posterior margin almost straight, although falcate distally; first dorsal-fin apex markedly rounded; first dorsal-fin free rear tip pointed (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Squalus japonicus: lateral (A-C) and ventral (D, E) views; first (F) and second (G) dorsal fins; pectoral (H) and caudal (I) fins. NSMT-P 44380 (neotype), adult male, 645 mm TL (A, D, F-I); HUMZ 189738, adult male, 535 mm TL (B); HUMZ 189687, juvenile male, 380 mm TL (C, E). Scale bars: 50 mm (A-E); 20 mm (F-I)." figureDoi="10.3897/zse.96.51962.figure4" httpUri="https://binary.pensoft.net/fig/420112" pageId="0" pageNumber="275">4F</figureCitation>
|
||
); first dorsal fin short, its base length 1.0 (0.8-1.1) times height of first dorsal-fin, its length 1.0 (0.8-1.1) times length of second dorsal fin; first dorsal fin low, its height 1.3 (1.1-1.6) times length of first dorsal-fin inner margin; first dorsal-fin inner margin small, its length 5.6% (5.1%-6.6%) TL. First dorsal-fin spine narrow, its base width 0.6% (0.5%-0.8%) TL and very short (its length 2.9%, 2.0%-3.5% TL), comprising 0.4 (0.3-0.5) times first dorsal-fin height. Interdorsal space 1.0 (0.8-1.1) times prepectoral length and 2.2 (1.8-2.4) times larger than dorsal caudal space. Pre-second dorsal length 2.7 (2.6-2.8) times prepectoral length. Second dorsal fin slender at fin web; second dorsal-fin anterior margin concave; second dorsal-fin posterior margin concave and deeply falcate; second dorsal-fin apex rounded and lobe-like; second dorsal-fin free rear tip pointed (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Squalus japonicus: lateral (A-C) and ventral (D, E) views; first (F) and second (G) dorsal fins; pectoral (H) and caudal (I) fins. NSMT-P 44380 (neotype), adult male, 645 mm TL (A, D, F-I); HUMZ 189738, adult male, 535 mm TL (B); HUMZ 189687, juvenile male, 380 mm TL (C, E). Scale bars: 50 mm (A-E); 20 mm (F-I)." figureDoi="10.3897/zse.96.51962.figure4" httpUri="https://binary.pensoft.net/fig/420112" pageId="0" pageNumber="275">4G</figureCitation>
|
||
); second dorsal fin short, its length 2.1 (1.8-2.5) times second dorsal-fin height; second dorsal-fin inner margin small, its length 5.0% (4.3%-5.5%) TL; second dorsal fin low, its height 1.3 (1.2-1.4) times length of second dorsal-fin inner margin. Second dorsal-fin spine slender, its base width 0.8% (0.7%-1.0%) TL; second dorsal-fin spine small, its length 4.9% (4.1%-5.6%) TL, corresponding to 0.8 (0.6-1.0) times height of second dorsal fin, almost reaching its apex; second dorsal-fin spine 1.7 (1.3-2.2) times larger than first dorsal-fin spine.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
Pectoral fins conspicuously narrow (pectoral-fin posterior margin length 7.9%, 6.8%-10.1% TL); pectoral-fin anterior and inner margins convex; pectoral-fin posterior margin concave (deeply concave in juveniles); pectoral-fin anterior margin length 1.6 (1.4-1.7) times length of pectoral-fin posterior margin and 1.5 (1.3-1.7) times length of pectoral-fin inner margin; pectoral-fin apex and free rear tips rounded; pectoral-fin free rear tips lobe-like and reaching the horizontal line traced at pectoral-fin apex (in juveniles, pectoral-fin free rear tips transcend the pectoral-fin apex) (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Squalus japonicus: lateral (A-C) and ventral (D, E) views; first (F) and second (G) dorsal fins; pectoral (H) and caudal (I) fins. NSMT-P 44380 (neotype), adult male, 645 mm TL (A, D, F-I); HUMZ 189738, adult male, 535 mm TL (B); HUMZ 189687, juvenile male, 380 mm TL (C, E). Scale bars: 50 mm (A-E); 20 mm (F-I)." figureDoi="10.3897/zse.96.51962.figure4" httpUri="https://binary.pensoft.net/fig/420112" pageId="0" pageNumber="275">4H</figureCitation>
|
||
). Pectoral-pelvic space 0.8 (0.7-0.8) times pelvic-caudal space. Pelvic-caudal space 1.2 (1.1-1.4) times larger than interdorsal space; pelvic fins in the midline between two dorsal fins, slightly nearer to first dorsal fin. Pelvic fins narrow with margins straight; pelvic-fin free rear tips pointed; pelvic fins short, its length 10.8% (9.6%-12.4%) TL. Clasper thin and small, slightly transcending pelvic-fin free rear tips; clasper inner margin length 0.8 (0.5-1.3) times length of pelvic inner margin; clasper groove dorsal and longitudinal, elongate and open; apopyle rounded, located anteriorly in the clasper groove; hypopyle constricted, located posteriorly in the clasper groove; rhipidion very thin and soft, flap-like, placed medially at distal end of the clasper.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
Caudal keel prominent and lateral since insertion of second dorsal fin to behind origin of caudal fin. Caudal fin small with dorsal caudal margin length 0.9 (0.8-0.9) times head length and 1.9 (1.8-3.1) times length of preventral caudal margin; dorsal and ventral caudal lobes slender (dorsal lobe rectangular in juveniles); dorsal caudal tip pointed; dorsal caudal margin slightly convex; upper postventral caudal margin straight; lower postventral caudal margin convex (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Squalus japonicus: lateral (A-C) and ventral (D, E) views; first (F) and second (G) dorsal fins; pectoral (H) and caudal (I) fins. NSMT-P 44380 (neotype), adult male, 645 mm TL (A, D, F-I); HUMZ 189738, adult male, 535 mm TL (B); HUMZ 189687, juvenile male, 380 mm TL (C, E). Scale bars: 50 mm (A-E); 20 mm (F-I)." figureDoi="10.3897/zse.96.51962.figure4" httpUri="https://binary.pensoft.net/fig/420112" pageId="0" pageNumber="275">4I</figureCitation>
|
||
); preventral caudal margin convex and short, its length 2.0 (1.4-2.4) times greater than length of pelvic-fin inner margin; ventral caudal tip rounded; caudal fork discontinuous, markedly concave and narrow, its width 6.6% (6.0%-7.4%) TL.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
<taxonomicName genus="Squamation" lsidName="Squamation" pageId="0" pageNumber="275" rank="genus">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="275">Squamation</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scanning electron microscopy of the dermal denticles. Squalus shiraii sp. nov. (A, B): HUMZ 149389 (holotype), adult male, 590 mm TL. S. formosus (C, D): CSIRO H- 6816 - 01 (holotype), adult male, 720 mm TL. S. mitsukurii (E, F): HUMZ 102987, adult male, 970 mm TL. S. japonicus (G, H): NSMT-P 44380 (neotype), adult male, 645 mm TL (G); NSMT-P 91127, juvenile female, 410 mm TL (H). S. brevirostris (I, J): NSMT-P 47378, adult female, 600 mm TL (I); KAUM-I 187, adult female, 578 mm TL (J). S. suckleyi (K, L): NSMT P- 92640, adult female, 970 mm TL. Scale bars: 50 μm (B); 100 μm (D, F, L); 200 μm (A, C, E, G-K)." figureDoi="10.3897/zse.96.51962.figure7" httpUri="https://binary.pensoft.net/fig/420115" pageId="0" pageNumber="275">7G, H</figureCitation>
|
||
). Dermal denticles tricuspid and markedly imbricated, its length slightly greater than its width; denticles conspicuously broad at the crown base; cusps posterior, prominent and pointed; lateral cusps shorter than median cusp, forming well-marked concavity with median cusp; median ridge thick and elongate; one to two lateral ridges, very thin, almost equal in length to median ridge; anterior furrow shallow with one small ridge on each side; anterior margin of the crown arrow-shaped. In juveniles, denticles are slender at the crown base, with cusps conspicuously pointed and more elongate and anterior margin of the crown tapered.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="275">Colouration</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Squalus japonicus: lateral (A-C) and ventral (D, E) views; first (F) and second (G) dorsal fins; pectoral (H) and caudal (I) fins. NSMT-P 44380 (neotype), adult male, 645 mm TL (A, D, F-I); HUMZ 189738, adult male, 535 mm TL (B); HUMZ 189687, juvenile male, 380 mm TL (C, E). Scale bars: 50 mm (A-E); 20 mm (F-I)." figureDoi="10.3897/zse.96.51962.figure4" httpUri="https://binary.pensoft.net/fig/420112" pageId="0" pageNumber="275">4</figureCitation>
|
||
). Body grey to light grey dorsally and white laterally from insertion of pectoral fins to origin of caudal fin. Dorsal fins greyish, darker at its apex to the upper half of the posterior margin, whitish at fin base and free rear tips; dorsal-fin spine brown anteriorly, white posteriorly and at the tips. Pectoral fins grey, slightly darker near pectoral-fin anterior margin; pectoral-fin posterior margin strongly white and uniform. Pelvic fins grey dorsally and completely pale ventrally; pelvic-fin posterior margin white. Caudal fin mostly greyish, light grey over vertebral column; dorsal caudal margin whitish proximally; upper postventral caudal margin white, lower postventral caudal margin broadly white; lower caudal lobe mostly white; preventral caudal margin whitish; black caudal stripe conspicuous; black upper caudal blotch conspicuous in the upper caudal lobe, nearest to dorsal caudal tip; black caudal bar oblique, placed in the caudal fork to midline of upper postventral caudal margin; dorsal and ventral caudal tips broadly white. Embryos, neonates and young juveniles exhibit body light grey with caudal bar and upper caudal blotch more conspicuous than in adults.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="275">
|
||
<taxonomicName genus="Vertebral" lsidName="Vertebral" pageId="0" pageNumber="275" rank="genus">Vertebral</taxonomicName>
|
||
counts
|
||
</emphasis>
|
||
(Table
|
||
<tableCitation captionStart="Table 6" captionStartId="T6" captionText="Table 6. Meristic data for species of Squalus from Japan. H: holotype; P: paratypes; NT: neotype; N: number of specimens; x; mean; m: mode; SD: standard deviation." httpUri="http://table.plazi.org/id/2DA11A1AF0C18FEB0512CBB43BD9057C" pageId="0" pageNumber="275" tableUuid="2DA11A1AF0C18FEB0512CBB43BD9057C">6</tableCitation>
|
||
). 87 (neotype) 80-88 (other material) precaudal vertebrae; 73 (65-75) diplospondylous vertebrae; 115 (104-116) total vertebrae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="275" type="distribution">
|
||
<paragraph pageId="0" pageNumber="275">Geographical distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
<taxonomicName authorityName="Ishikawa" authorityYear="1908" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus japonicus" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">Squalus japonicus</emphasis>
|
||
</taxonomicName>
|
||
is a regional endemic species occurring in the North-western Pacific Ocean from Southern Japan, China to Taiwan (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Map of the North-western Pacific Ocean, showing the geographical distribution of Squalus species. A: S. shiraii sp. nov. (red); B: S. mitsukurii (orange); C: S. japonicus (brown); D: S. brevirostris (yellow); E: S. acutirostris (green); F: S. suckleyi (pink). Star: holotype / neotype; Dotted circle: paratypes; Circle: other material." figureDoi="10.3897/zse.96.51962.figure8" httpUri="https://binary.pensoft.net/fig/420116" pageId="0" pageNumber="275">8C</figureCitation>
|
||
). It is found on the upper continental slope of tropical and subtropical areas, ranging between 195-520 m depth, although it also has been reported at 52 m depth only in
|
||
<bibRefCitation DOI="https://doi.org/10.1111/jfb.12874" author="Weigmann, S" journalOrPublisher="Journal of Fish Biology" pageId="0" pageNumber="275" pagination="837 - 1037" refId="B92" refString="Weigmann, S, 2016. Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. Journal of Fish Biology 88 (3): 837 - 1037, DOI: https://doi.org/10.1111/jfb.12874" title="Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity." url="https://doi.org/10.1111/jfb.12874" volume="88" year="2016">Weigmann (2016)</bibRefCitation>
|
||
. This species also occurs in Korean waters (
|
||
<bibRefCitation author="Chen, CT" journalOrPublisher="Japanese Journal of Ichthyology" pageId="0" pageNumber="275" pagination="26 - 42" refId="B8" refString="Chen, CT, Taniuchi, T, Nose, Y, 1979. Blainville's dogfish, Squalus blainvillei, from Japan, with notes on S. mitsukurii and S. japonicus. Japanese Journal of Ichthyology 26 (1): 26 - 42" title="Blainville's dogfish, Squalus blainvillei, from Japan, with notes on S. mitsukurii and S. japonicus." volume="26" year="1979">Chen et al. 1979</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="275" type="remarks">
|
||
<paragraph pageId="0" pageNumber="275">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
<bibRefCitation author="Ishikawa, C" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="275" pagination="71 - 73" refId="B41" refString="Ishikawa, C, 1908. Description of a new species of squaloid shark from Japan. Proceedings of the Academy of Natural Sciences of Philadelphia 60: 71 - 73" title="Description of a new species of squaloid shark from Japan." volume="60" year="1908">Ishikawa (1908)</bibRefCitation>
|
||
mentioned three syntypes of
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
that were deposited at the Zoological University Museum of Tokyo (formerly Imperial Museum of Tokyo). A visit to the fish collection from ZUMT was undertaken in 2013 to verify the existence of the type material. However, it was noticed that there was a single specimen of
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
held in the museum, assuming that syntypes are lost and no longer exist. Dr. M. Nakae (NSMT,
|
||
<emphasis italics="true" pageId="0" pageNumber="275">person. comm</emphasis>
|
||
. 2014) stated that the NSMT fish collection has never been in possession of this material and ratified that the syntypes are lost.
|
||
<taxonomicName authorityName="Ishikawa" authorityYear="1908" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus japonicus" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">Squalus japonicus</emphasis>
|
||
</taxonomicName>
|
||
has been misidentified in waters outside Japan (e.g. Vietnam by
|
||
<bibRefCitation DOI="https://doi.org/10.15625/1859-3097/13/1/3503" author="Quang, VV" journalOrPublisher="Vietnam Journal of Marine Science and Technology" pageId="0" pageNumber="275" pagination="21 - 30" refId="B72" refString="Quang, VV, Tuan, VS, Thao, LTT, Thinh, TC, Uy, Vu NP, Phuong, LM, 2013. Species composition and distribution of sharks in Quy Nhon Bay and neighboring waters. Vietnam Journal of Marine Science and Technology 13 (1): 21 - 30, DOI: https://doi.org/10.15625/1859-3097/13/1/3503" title="Species composition and distribution of sharks in Quy Nhon Bay and neighboring waters." url="https://doi.org/10.15625/1859-3097/13/1/3503" volume="13" year="2013">Quang et al. 2013</bibRefCitation>
|
||
) and many morphological variations were reported earlier in
|
||
<bibRefCitation author="Compagno, LJV" journalOrPublisher="Vol. 1. Introduction, molluscs, crustaceans, hagfishes, sharks, batoid fishes and chimaeras. FAO Species identification guide for fishery purposes and American Society of Ichthyologists Special Publication 5. FAO, Rome" pageId="0" pageNumber="275" refId="B15" refString="Compagno, LJV, Last, PR, Stevens, JD, Alava, MNR, 2005b. Checklist of Philippine Chondrichthyes. CSIRO Marine Laboratories Report 243, Hobart." title="Checklist of Philippine Chondrichthyes. CSIRO Marine Laboratories Report 243, Hobart." year="2005 b">Compagno et al. (2005b)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Last, PR" editor="Last, PR" journalOrPublisher="CSIRO Marine and Atmospheric Research Paper 14, Hobart" pageId="0" pageNumber="275" pagination="83 - 90" refId="B46" refString="Last, PR, Marshall, LJ, White, WT, 2007b. Part 8 - Squalus nasutus sp. nov, a new long-snouted spurdog of the S. japonicus group from the Indian Ocean. In: Last, PR, White, WT, Pogonoski, JJ, Eds., Descriptions of new dogfishes of the genus Squalus (Squaloidea: Squalidae). CSIRO Marine and Atmospheric Research Paper 14, Hobart: 83 - 90" title="Part 8 - Squalus nasutus sp. nov, a new long-snouted spurdog of the S. japonicus group from the Indian Ocean." volumeTitle="Descriptions of new dogfishes of the genus Squalus (Squaloidea: Squalidae)." year="2007 b">Last et al. (2007b)</bibRefCitation>
|
||
for material from Indonesia, Philippines and Western Australia, compromising the correct application of this nominal species. To clarify the taxonomic status of
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
across the Western Pacific Ocean, the designation of a neotype along with a re-description of this species is provided herein. The neotype was collected from Southern Japan (Senoumi Bank, Suruga Bay), which is within the known range distribution of
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
, based on the examined material, the original description of
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
and information in literature. The designation of the neotype was further based on the sex, maturity and total length that were approximate to those provided for a syntype of
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation author="Ishikawa, C" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="275" pagination="71 - 73" refId="B41" refString="Ishikawa, C, 1908. Description of a new species of squaloid shark from Japan. Proceedings of the Academy of Natural Sciences of Philadelphia 60: 71 - 73" title="Description of a new species of squaloid shark from Japan." volume="60" year="1908">Ishikawa (1908)</bibRefCitation>
|
||
. The neotype is securely deposited in the scientific fish collection from the NSMT in Tokyo at the Department of Zoology for preservation and accessibility purposes. This species is easily separated from its congeners through many characteristics of the external morphology, morphometrics and meristic data, including shape of head, snout and pectoral fins, length of snout, width of mouth and preoral length and several other external measurements as shown in the diagnosis above. The current diagnosis and re-description of
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
are congruent with observations of
|
||
<bibRefCitation author="Ishikawa, C" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="275" pagination="71 - 73" refId="B41" refString="Ishikawa, C, 1908. Description of a new species of squaloid shark from Japan. Proceedings of the Academy of Natural Sciences of Philadelphia 60: 71 - 73" title="Description of a new species of squaloid shark from Japan." volume="60" year="1908">Ishikawa (1908)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Chen, CT" journalOrPublisher="Japanese Journal of Ichthyology" pageId="0" pageNumber="275" pagination="26 - 42" refId="B8" refString="Chen, CT, Taniuchi, T, Nose, Y, 1979. Blainville's dogfish, Squalus blainvillei, from Japan, with notes on S. mitsukurii and S. japonicus. Japanese Journal of Ichthyology 26 (1): 26 - 42" title="Blainville's dogfish, Squalus blainvillei, from Japan, with notes on S. mitsukurii and S. japonicus." volume="26" year="1979">Chen et al. (1979)</bibRefCitation>
|
||
with the exception of the presence of two hooks in the clasper. These cartilages of the clasper probably concern the accessory terminal 3 cartilage (spur) and dorsal terminal cartilage (claw) that are commonly noticed in species of
|
||
<taxonomicName class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">Squalus</emphasis>
|
||
</taxonomicName>
|
||
. Differences in morphometrics are observed between the data provided in
|
||
<bibRefCitation author="Ishikawa, C" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="275" pagination="71 - 73" refId="B41" refString="Ishikawa, C, 1908. Description of a new species of squaloid shark from Japan. Proceedings of the Academy of Natural Sciences of Philadelphia 60: 71 - 73" title="Description of a new species of squaloid shark from Japan." volume="60" year="1908">Ishikawa (1908)</bibRefCitation>
|
||
and those from the present analysis, but this is due to divergences of the applied methodology.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
This species together with
|
||
<taxonomicName genus="S." lsidName="S. melanurus" pageId="0" pageNumber="275" rank="species" species="melanurus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. melanurus</emphasis>
|
||
</taxonomicName>
|
||
and the Australian species
|
||
<taxonomicName genus="S." lsidName="S. nasutus" pageId="0" pageNumber="275" rank="species" species="nasutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. nasutus</emphasis>
|
||
</taxonomicName>
|
||
, comprise the
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
subgroup of species whose members are characterised by having a conspicuously elongate and obtuse snout, thin body, mouth very narrow and pectoral fins tapered (
|
||
<bibRefCitation DOI="https://doi.org/10.5252/zoosystema2018v40a9" author="Viana, STFL" journalOrPublisher="Zoosystema" pageId="0" pageNumber="275" pagination="159 - 177" refId="B86" refString="Viana, STFL, Carvalho, MR, 2018a. Squalus rancureli, a new junior synonym of the blacktailed spurdog S. melanurus, and updated diagnosis of S. bucephalus from New Caledonia (Chondrichthyes: Squaliformes: Squalidae). Zoosystema 40 (9): 159 - 177, DOI: https://doi.org/10.5252/zoosystema2018v40a9" title="Squalus rancureli, a new junior synonym of the blacktailed spurdog S. melanurus, and updated diagnosis of S. bucephalus from New Caledonia (Chondrichthyes: Squaliformes: Squalidae)." url="https://doi.org/10.5252/zoosystema2018v40a9" volume="40" year="2018 a">Viana and Carvalho 2018a</bibRefCitation>
|
||
). The Japanese species is more closely related to
|
||
<taxonomicName genus="S." lsidName="S. nasutus" pageId="0" pageNumber="275" rank="species" species="nasutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. nasutus</emphasis>
|
||
</taxonomicName>
|
||
according to morphological and genetic data (
|
||
<bibRefCitation author="Last, PR" editor="Last, PR" journalOrPublisher="CSIRO Marine and Atmospheric Research Paper 14, Hobart" pageId="0" pageNumber="275" pagination="83 - 90" refId="B46" refString="Last, PR, Marshall, LJ, White, WT, 2007b. Part 8 - Squalus nasutus sp. nov, a new long-snouted spurdog of the S. japonicus group from the Indian Ocean. In: Last, PR, White, WT, Pogonoski, JJ, Eds., Descriptions of new dogfishes of the genus Squalus (Squaloidea: Squalidae). CSIRO Marine and Atmospheric Research Paper 14, Hobart: 83 - 90" title="Part 8 - Squalus nasutus sp. nov, a new long-snouted spurdog of the S. japonicus group from the Indian Ocean." volumeTitle="Descriptions of new dogfishes of the genus Squalus (Squaloidea: Squalidae)." year="2007 b">Last et al. 2007b</bibRefCitation>
|
||
). Shape of dermal denticles, dentition and shape of dorsal, pectoral, pelvic and caudal fins are intrinsically similar between these two species. Morphometrics are also mostly congruent, although differences in measurements are apparent when adults are compared: second dorsal fin height 5.4%-6.5% TL for
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
vs. 4.9%-5.3% TL for
|
||
<taxonomicName genus="S." lsidName="S. nasutus" pageId="0" pageNumber="275" rank="species" species="nasutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. nasutus</emphasis>
|
||
</taxonomicName>
|
||
; length of second dorsal-fin anterior margin 10.8%-12.7% TL vs. 8.6%-10.5% TL; length of second dorsal-fin spine 4.7%-5.6% TL vs. 3.9%-4.4% TL; length of dorsal caudal margin 19.6%-20.4% TL vs. 18.1%-19.4% TL. Preoral length, eye length, pectoral-fin inner margin length, trunk height and width overlaps between these two species which contradicts
|
||
<bibRefCitation author="Last, PR" editor="Last, PR" journalOrPublisher="CSIRO Marine and Atmospheric Research Paper 14, Hobart" pageId="0" pageNumber="275" pagination="83 - 90" refId="B46" refString="Last, PR, Marshall, LJ, White, WT, 2007b. Part 8 - Squalus nasutus sp. nov, a new long-snouted spurdog of the S. japonicus group from the Indian Ocean. In: Last, PR, White, WT, Pogonoski, JJ, Eds., Descriptions of new dogfishes of the genus Squalus (Squaloidea: Squalidae). CSIRO Marine and Atmospheric Research Paper 14, Hobart: 83 - 90" title="Part 8 - Squalus nasutus sp. nov, a new long-snouted spurdog of the S. japonicus group from the Indian Ocean." volumeTitle="Descriptions of new dogfishes of the genus Squalus (Squaloidea: Squalidae)." year="2007 b">Last et al. (2007b)</bibRefCitation>
|
||
in supporting these measurements as diagnostic characters for these species.
|
||
<taxonomicName genus="Vertebral" lsidName="Vertebral" pageId="0" pageNumber="275" rank="genus">Vertebral</taxonomicName>
|
||
counts differ between
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName genus="S." lsidName="S. nasutus" pageId="0" pageNumber="275" rank="species" species="nasutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. nasutus</emphasis>
|
||
</taxonomicName>
|
||
as well: monospondylous (41-43), precaudal (84-88) vertebrae (vs. 37-40; 80-83 for those from Australia). These results are congruent with recent molecular analyses of the cytochrome c oxydase 1 (COI) and NADH dehydrogenase 2 (NAD2) mitochondrial genes in
|
||
<bibRefCitation author="Ward, RD" editor="Last, PR" journalOrPublisher="CSIRO Marine and Atmospheric Research Paper 14, Hobart" pageId="0" pageNumber="275" pagination="117 - 130" refId="B91" refString="Ward, RD, Holmes, BH, Zemlak, TS, Smith, PJ, 2007. DNA barcoding discriminates spurdogs of the genus Squalus. In: Last, PR, White, WT, Pogonoski, JJ, Eds., Descriptions of new dogfishes of the genus Squalus (Squaloidea: Squalidae). CSIRO Marine and Atmospheric Research Paper 14, Hobart: 117 - 130" title="DNA barcoding discriminates spurdogs of the genus Squalus." volumeTitle="Descriptions of new dogfishes of the genus Squalus (Squaloidea: Squalidae)." year="2007">Ward et al. (2007)</bibRefCitation>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.1206/754.1" author="Naylor, GJP" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="275" pagination="1 - 262" refId="B66" refString="Naylor, GJP, Caira, JN, Jensen, K, Rosana, AM, White, WT, Last, PR, 2012. A DNA sequence-based approach to the identification of shark and rays species and its implication of global elasmobranch diversity and parasitology. Bulletin of the American Museum of Natural History 367: 1 - 262, DOI: https://doi.org/10.1206/754.1" title="A DNA sequence-based approach to the identification of shark and rays species and its implication of global elasmobranch diversity and parasitology." url="https://doi.org/10.1206/754.1" volume="367" year="2012">Naylor et al. (2012)</bibRefCitation>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3752.1.16" author="Straube, N" journalOrPublisher="Zootaxa" pageId="0" pageNumber="275" pagination="256 - 278" refId="B80" refString="Straube, N, White, WT, HO, H-C, Rochel, E, Corrigan, S, Li, C, Naylor, GJP, 2013. A DNA sequence-based identification checklist for Taiwanese chondrichthyans. Zootaxa 3752: 256 - 278, DOI: https://doi.org/10.11646/zootaxa.3752.1.16" title="A DNA sequence-based identification checklist for Taiwanese chondrichthyans." url="https://doi.org/10.11646/zootaxa.3752.1.16" volume="3752" year="2013">Straube et al. (2013)</bibRefCitation>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.1111/zsc.12224" author="Verissimo, A" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="275" pagination="414 - 428" refId="B84" refString="Verissimo, A, Zaera-Perez, D, Leslie, R, Iglesias, SP, Seret, B, Grigoriou, P, Sterioti, A, Gubili, C, Barria, C, Duffy, C, Hernandez, S, Batjakas, IE, Griffiths, AM, 2017. Molecular diversity and distribution of eastern Atlantic and Mediterranean dogfishes Squalus highlight taxonomic issues in the genus. Zoologica Scripta 46 (4): 414 - 428, DOI: https://doi.org/10.1111/zsc.12224" title="Molecular diversity and distribution of eastern Atlantic and Mediterranean dogfishes Squalus highlight taxonomic issues in the genus." url="https://doi.org/10.1111/zsc.12224" volume="46" year="2017">
|
||
<normalizedToken originalValue="Veríssimo">Verissimo</normalizedToken>
|
||
et al. (2017)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.798.28375" author="Daly-Engel, TS" journalOrPublisher="ZooKeys" pageId="0" pageNumber="275" pagination="135 - 157" refId="B18" refString="Daly-Engel, TS, Koch, A, Anderson, JM, Cotton, CF, Grubbs, RD, 2018. Description of a new deep-water dogfish shark from Hawaii, with comments on the Squalus mitsukurii species complex in the West Pacific. ZooKeys 798: 135 - 157, DOI: https://doi.org/10.3897/zookeys.798.28375" title="Description of a new deep-water dogfish shark from Hawaii, with comments on the Squalus mitsukurii species complex in the West Pacific." url="https://doi.org/10.3897/zookeys.798.28375" volume="798" year="2018">Daly-Engel et al. (2018)</bibRefCitation>
|
||
which support genetic divergence between
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName genus="S." lsidName="S. nasutus" pageId="0" pageNumber="275" rank="species" species="nasutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. nasutus</emphasis>
|
||
</taxonomicName>
|
||
. The latter study also noticed high intraspecific genetic distances in
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
when including data taken from populations from Japan and Taiwan, but variations in the species are not yet supported through morphological evaluation, except for the black caudal marking near the base of the lower lobe in Taiwanese specimens that is absent in other material.
|
||
<bibRefCitation author="Compagno, LJV" journalOrPublisher="Vol. 1. Introduction, molluscs, crustaceans, hagfishes, sharks, batoid fishes and chimaeras. FAO Species identification guide for fishery purposes and American Society of Ichthyologists Special Publication 5. FAO, Rome" pageId="0" pageNumber="275" refId="B15" refString="Compagno, LJV, Last, PR, Stevens, JD, Alava, MNR, 2005b. Checklist of Philippine Chondrichthyes. CSIRO Marine Laboratories Report 243, Hobart." title="Checklist of Philippine Chondrichthyes. CSIRO Marine Laboratories Report 243, Hobart." year="2005 b">Compagno et al. (2005b)</bibRefCitation>
|
||
also noticed variations in this species when examining material from the Philippines and considered it to be a separated species (
|
||
<taxonomicName class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">Squalus</emphasis>
|
||
</taxonomicName>
|
||
sp. 2) due to differences with
|
||
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName genus="S." lsidName="S. nasutus" pageId="0" pageNumber="275" rank="species" species="nasutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. nasutus</emphasis>
|
||
</taxonomicName>
|
||
. In contrast,
|
||
<bibRefCitation author="Last, PR" editor="Last, PR" journalOrPublisher="CSIRO Marine and Atmospheric Research Paper 14, Hobart" pageId="0" pageNumber="275" pagination="83 - 90" refId="B46" refString="Last, PR, Marshall, LJ, White, WT, 2007b. Part 8 - Squalus nasutus sp. nov, a new long-snouted spurdog of the S. japonicus group from the Indian Ocean. In: Last, PR, White, WT, Pogonoski, JJ, Eds., Descriptions of new dogfishes of the genus Squalus (Squaloidea: Squalidae). CSIRO Marine and Atmospheric Research Paper 14, Hobart: 83 - 90" title="Part 8 - Squalus nasutus sp. nov, a new long-snouted spurdog of the S. japonicus group from the Indian Ocean." volumeTitle="Descriptions of new dogfishes of the genus Squalus (Squaloidea: Squalidae)." year="2007 b">Last et al. (2007b)</bibRefCitation>
|
||
considered it to be con-specific with
|
||
<taxonomicName genus="S." lsidName="S. nasutus" pageId="0" pageNumber="275" rank="species" species="nasutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. nasutus</emphasis>
|
||
</taxonomicName>
|
||
as the morphological differences pointed in
|
||
<bibRefCitation author="Compagno, LJV" journalOrPublisher="Vol. 1. Introduction, molluscs, crustaceans, hagfishes, sharks, batoid fishes and chimaeras. FAO Species identification guide for fishery purposes and American Society of Ichthyologists Special Publication 5. FAO, Rome" pageId="0" pageNumber="275" refId="B15" refString="Compagno, LJV, Last, PR, Stevens, JD, Alava, MNR, 2005b. Checklist of Philippine Chondrichthyes. CSIRO Marine Laboratories Report 243, Hobart." title="Checklist of Philippine Chondrichthyes. CSIRO Marine Laboratories Report 243, Hobart." year="2005 b">Compagno et al. (2005b)</bibRefCitation>
|
||
and in their study represent intraspecific variations. Whether the Philippine form represents an undescribed species is still unclear, but we consider it here to be con-specific with
|
||
<taxonomicName genus="S." lsidName="S. nasutus" pageId="0" pageNumber="275" rank="species" species="nasutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. nasutus</emphasis>
|
||
</taxonomicName>
|
||
for nomenclature stability.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="275" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="275">Comparative material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="275">
|
||
<taxonomicName authorityName="Last, Marshall & White" authorityYear="2007" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus nasutus" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="nasutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">Squalus nasutus</emphasis>
|
||
</taxonomicName>
|
||
. AMS I22817-008, juvenile male, 362 mm TL, Northwest Shelf, 240 km north of Port Hedland, Western Australia,
|
||
<geoCoordinate degrees="18" direction="south" minutes="06" orientation="latitude" precision="925" value="-18.1">18°06'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="117" direction="east" minutes="45" orientation="longitude" precision="925" value="117.75">117°45'E</geoCoordinate>
|
||
, 492-520 m depth; CSIRO H 4132-02, adult male, 465 mm TL, Bolinao evening market, Philippines; CSIRO H 4132-03, juvenile female, 475 mm TL, locality same as CSIRO H 4132-02; CSIRO H 4132-04, adult female, 540 mm TL, locality same as CSIRO H 4132-02; CSIRO H 5860-01, adult female, 540 mm TL, Cilacap (South coast, central Java), Indonesia,
|
||
<geoCoordinate degrees="07" direction="south" minutes="40" orientation="latitude" precision="925" value="-7.6666665">07°40'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="109" direction="east" minutes="00" orientation="longitude" precision="925" value="109.0">109°00'E</geoCoordinate>
|
||
; CSIRO H 5860-02, adult female, 570 mm TL, locality same as CSIRO H 5860-01; CSIRO H 5860-03, adult female, 545 mm TL, locality same as CSIRO H 5860-01; CSIRO H 6125-04, adult male, 465 mm TL, Kedonganan, Jimbaran Bay (South coast of Bali), Indonesia,
|
||
<geoCoordinate degrees="08" direction="south" minutes="45" orientation="latitude" precision="925" value="-8.75">08°45'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="115" direction="east" minutes="10" orientation="longitude" precision="925" value="115.166664">115°10'E</geoCoordinate>
|
||
; CSIRO H 6413-01, adult female, 580 mm TL, West of Shark Bay, Western Australia,
|
||
<geoCoordinate degrees="25" direction="south" minutes="03.8" orientation="latitude" precision="92" value="-25.063334">25°03.8'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="112" direction="east" minutes="08.9" orientation="longitude" precision="92" value="112.14833">112°08.9'E</geoCoordinate>
|
||
, 315-340 m depth; CSIRO H 6484-01, adult female, 575 mm TL, locality same as CSIRO H 5860-01. Type material of
|
||
<taxonomicName genus="S." lsidName="S. nasutus" pageId="0" pageNumber="275" rank="species" species="nasutus">
|
||
<emphasis italics="true" pageId="0" pageNumber="275">S. nasutus</emphasis>
|
||
</taxonomicName>
|
||
: CSIRO H2590-12, adult female, 503 mm TL, west of Leander Point, Western Australia,
|
||
<geoCoordinate degrees="29" direction="south" minutes="15" orientation="latitude" precision="925" value="-29.25">29°15'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="113" direction="east" minutes="56" orientation="longitude" precision="925" value="113.933334">113°56'E</geoCoordinate>
|
||
(
|
||
<emphasis italics="true" pageId="0" pageNumber="275">holotype</emphasis>
|
||
);
|
||
<emphasis italics="true" pageId="0" pageNumber="275">Paratypes</emphasis>
|
||
: CSIRO H 2608-15, juvenile female, 411 mm TL, Rottnest Canyon, Western Australia,
|
||
<geoCoordinate degrees="31" direction="south" minutes="57" orientation="latitude" precision="925" value="-31.95">31°57'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="115" direction="east" minutes="08" orientation="longitude" precision="925" value="115.13333">115°08'E</geoCoordinate>
|
||
; CSIRO H 2598-07, juvenile female, 465 mm TL, west of Green Head, Western Australia,
|
||
<geoCoordinate degrees="29" direction="south" minutes="58" orientation="latitude" precision="925" value="-29.966667">29°58'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="114" direction="east" minutes="27" orientation="longitude" precision="925" value="114.45">114°27'E</geoCoordinate>
|
||
; CSIRO H 2567-08, adult male, 470 mm TL, west of Dorre Island, Western Australia,
|
||
<geoCoordinate degrees="25" direction="south" minutes="09" orientation="latitude" precision="925" value="-25.15">25°09'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="112" direction="east" minutes="09" orientation="longitude" precision="925" value="112.15">112°09'E</geoCoordinate>
|
||
; CSIRO H 1652-01, juvenile female, 315 mm TL, northwest of Port Hedland, Western Australia,
|
||
<geoCoordinate degrees="18" direction="south" minutes="25" orientation="latitude" precision="925" value="-18.416666">18°25'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="117" direction="east" minutes="48" orientation="longitude" precision="925" value="117.8">117°48'E</geoCoordinate>
|
||
; CSIRO H 1652-02, juvenile female, 459 mm TL, same locality as CSIRO H 1652-01; CSIRO H 1207-07, adult female, 537 mm TL, northwest of Port Hedland, Western Australia,
|
||
<geoCoordinate degrees="18" direction="south" minutes="20" orientation="latitude" precision="925" value="-18.333334">18°20'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="117" direction="east" minutes="50" orientation="longitude" precision="925" value="117.833336">117°50'E</geoCoordinate>
|
||
; CSIRO H 1207-08, juvenile female, 496 mm TL, same locality as CSIRO H 1207-07; CSIRO CA 3290, adult female, 549 mm TL, southwest of Rowley Shoals, Western Australia,
|
||
<geoCoordinate degrees="18" direction="south" minutes="10" orientation="latitude" precision="925" value="-18.166666">18°10'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="118" direction="east" minutes="20" orientation="longitude" precision="925" value="118.333336">118°20'E</geoCoordinate>
|
||
; CSIRO CA 4055, adult female, 527 mm TL, southwest of Rowley Shoals, Western Australia,
|
||
<geoCoordinate degrees="18" direction="south" minutes="11" orientation="latitude" precision="925" value="-18.183332">18°11'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="118" direction="east" minutes="04" orientation="longitude" precision="925" value="118.066666">118°04'E</geoCoordinate>
|
||
; CSIRO H 2898-07, juvenile male, 452 mm TL, north-northwest of Port Hedland, Western Australia,
|
||
<geoCoordinate degrees="18" direction="south" minutes="07" orientation="latitude" precision="925" value="-18.116667">18°07'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="118" direction="east" minutes="12" orientation="longitude" precision="925" value="118.2">118°12'E</geoCoordinate>
|
||
; CSIRO CA 4110, adult male, 497 mm TL, east of Rowley Shoals, Western Australia,
|
||
<geoCoordinate degrees="17" direction="south" minutes="18" orientation="latitude" precision="925" value="-17.3">17°18'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="120" direction="east" minutes="09" orientation="longitude" precision="925" value="120.15">120°09'E</geoCoordinate>
|
||
; CSIRO H 1693-01, juvenile male, 361 mm TL, Rowley Shoals, Western Australia,
|
||
<geoCoordinate degrees="17" direction="south" minutes="02" orientation="latitude" precision="925" value="-17.033333">17°02'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="120" direction="east" minutes="05" orientation="longitude" precision="925" value="120.083336">120°05'E</geoCoordinate>
|
||
; CSIRO H 1693-02, juvenile female, 306 mm TL, same locality as CSIRO H 1693-01; CSIRO H 1694-01, juvenile female, 425 mm TL, Rowley Shoals, Western Australia,
|
||
<geoCoordinate degrees="16" direction="south" minutes="57" orientation="latitude" precision="925" value="-16.95">16°57'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="120" direction="east" minutes="14" orientation="longitude" precision="925" value="120.23333">120°14'E</geoCoordinate>
|
||
; CSIRO H 2032-01, adult male, 461 mm TL, northeast of Mermaid Reef, Rowley Shoals, Western Australia,
|
||
<geoCoordinate degrees="16" direction="south" minutes="54" orientation="latitude" precision="925" value="-16.9">16°54'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="120" direction="east" minutes="25" orientation="longitude" precision="925" value="120.416664">120°25'E</geoCoordinate>
|
||
; CSIRO H 2032-02, juvenile female, 404 mm TL, same locality as CSIRO H 2032-01; WAM P 28086-006, juvenile male, 380 mm TL; juvenile female, 450 mm TL, Rowley Shoals, Western Australia,
|
||
<geoCoordinate degrees="17" direction="south" minutes="49" orientation="latitude" precision="925" value="-17.816668">17°49'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="118" direction="east" minutes="41" orientation="longitude" precision="925" value="118.683334">118°41'E</geoCoordinate>
|
||
, 308-310 m depth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |