278 lines
40 KiB
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278 lines
40 KiB
XML
<document id="3D9463B94566E3C5F64F067F91934E4A" ID-CLB-Dataset="3748" ID-DOI="10.5252/geodiversitas2021v43a17" ID-GBIF-Dataset="9f355bf4-4e90-4db3-af0e-c04f11ceb309" ID-ISSN="1638-9395" ID-Zenodo-Dep="5628435" ID-ZooBank="urn:lsid:zoobank.org:pub:E0BAA274-37CB-407B-849A-FD284BBE954B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1635541439614" checkinUser="felipe" docAuthor="Augé, Marc Louis, Dion, Michaël & Phélizon, Alain" docDate="2021" docId="0F23E968FF8BFF93FC4E791CD0A6FEE5" docLanguage="en" docName="Geodiversitas.43.17.645-661.pdf" docOrigin="Geodiversitas 43 (17)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Animalia" docType="treatment" docVersion="9" lastPageNumber="657" masterDocId="F31A9110FF87FF9DFFBA7E5DD435FF94" masterDocTitle="The lizard (Reptilia, Squamata) assemblage from the Paleocene of Montchenot (Paris Basin, MP 6)" masterLastPageNumber="661" masterPageNumber="645" pageNumber="655" updateTime="1699467898079" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:mods id="F4C90133A5FACF4D2546C05F9DAD621C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8B92944E13E7AADDA62253B683893DDD">
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<mods:title id="448857EFA3B3D173E0F86002D7F94EBA">The lizard (Reptilia, Squamata) assemblage from the Paleocene of Montchenot (Paris Basin, MP 6)</mods:title>
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<mods:name id="46A4F0791959882880D8BE93CE48C34D" type="personal">
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<mods:role id="3C5CE9B3E6679DD76F9E9A4728663B4C">
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<mods:roleTerm id="743643686ECECBED906941464E7A052A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F6FCFB559098EC3F59867B810D66AB2B">Augé, Marc Louis</mods:namePart>
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<mods:affiliation id="8BE49474D5A4128B48450D8F8271DF2E">Centre de Recherches sur la Paléobiodiversité et les Paléoenvironnements (MNHN, CNRS, UPMC-Paris 6, Sorbonne Université), Muséum national d’Histoire naturelle, 57 rue Cuvier, case postale 38, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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</mods:name>
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<mods:name id="A91BC514C6BA039753538CF28E84B3C5" type="personal">
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<mods:role id="DD78863FAF7F1707B26F900147B59853">
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<mods:roleTerm id="566AB853C32E4C6B8CB290DC4402BD43">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C34E71F5FA0E3F1A60BA904044845CAA">Dion, Michaël</mods:namePart>
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<mods:affiliation id="FDB7D799300EAFD14466A222BC76E2CD">1 chemin du Cochelet, F- 02220 Mont Saint Martin (France)</mods:affiliation>
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</mods:name>
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<mods:name id="8C6F0B5C05128E3B6BF4134DEBD21965" type="personal">
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<mods:role id="4D2E5DEEBFB6C7CAC2C0C485C96F479B">
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<mods:roleTerm id="4F446AB7FBAFA494C1E3ACE5C4CFB017">Author</mods:roleTerm>
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<mods:namePart id="2FC5328A3D0E26121B76326C7C1CF5A1">Phélizon, Alain</mods:namePart>
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<mods:affiliation id="15EE998099F8AEF343382CFCDEF1B621">Société d’Étude des Sciences naturelles de Reims, 122 bis rue du Barbâtre, F- 51100 Reims (France)</mods:affiliation>
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<mods:nameIdentifier id="C4C6E5D98CA6366B57E1F543DAF91B05" type="email">alain.phelizon@orange.fr</mods:nameIdentifier>
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<mods:typeOfResource id="1546FEF6A7C66CA8725AD6419B813560">text</mods:typeOfResource>
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<mods:titleInfo id="DF81497B11D0B05DB4188AC235AB22FF">
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<mods:title id="05154F9858C054428D95EEE25ACC28DA">Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:part id="C372B1CABA0700DC63469B98B219818E">
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<mods:date id="8A81608D86EA1432EB5D771C4874DD22">2021</mods:date>
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<mods:detail id="C975B50258383ED0383228B11F05633D" type="pubDate">
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<mods:number id="DCA610939221BFD2B50E81F60623823E">2021-09-09</mods:number>
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<mods:detail id="FF02A28003EF7476C5043DA94DDC4156" type="volume">
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<mods:number id="4A9305B970036602057051D74B69452E">43</mods:number>
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<mods:detail id="01D79A91409F343E080518B56F5F5DE5" type="issue">
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<mods:number id="11EA9936EE5295D732900091AC7BC46F">17</mods:number>
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<mods:start id="196541B785660D0D6EBA8DC95AFB3054">645</mods:start>
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<mods:end id="8075763D7717CD9615188F088C6BBFDB">661</mods:end>
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<mods:classification id="9D816F05E35C29F0D9D49FFCE7C3720F">journal article</mods:classification>
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<mods:identifier id="FCEAD2F0D4753E3F2324B3389A795702" type="CLB-Dataset">3748</mods:identifier>
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<mods:identifier id="A052D3F6E912E5B4E423C50D4EB76E73" type="DOI">10.5252/geodiversitas2021v43a17</mods:identifier>
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<mods:identifier id="ABE28F944226DD34BD16B3B43A1AAAE3" type="GBIF-Dataset">9f355bf4-4e90-4db3-af0e-c04f11ceb309</mods:identifier>
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<mods:identifier id="2D745BBD11DC5C4A236C3F72D593B969" type="ISSN">1638-9395</mods:identifier>
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<treatment id="0F23E968FF8BFF93FC4E791CD0A6FEE5" ID-DOI="http://doi.org/10.5281/zenodo.5636447" ID-GBIF-Taxon="189796616" ID-Zenodo-Dep="5636447" LSID="urn:lsid:plazi:treatment:0F23E968FF8BFF93FC4E791CD0A6FEE5" httpUri="http://treatment.plazi.org/id/0F23E968FF8BFF93FC4E791CD0A6FEE5" lastPageId="14" lastPageNumber="657" pageId="12" pageNumber="655">
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<subSubSection id="CF900BF5FF8BFF91FC4E791CD0DDF8CF" box="[1012,1256,1857,1883]" pageId="12" pageNumber="655" type="nomenclature">
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<paragraph id="8735587EFF8BFF91FC4E791CD0DDF8CF" blockId="12.[1012,1256,1857,1883]" box="[1012,1256,1857,1883]" pageId="12" pageNumber="655">
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<taxonomicName id="408A23FDFF8BFF91FC4E791CD0D1F8CF" box="[1012,1252,1857,1883]" kingdom="Animalia" pageId="12" pageNumber="655">Pan-Shinisaurus indet</taxonomicName>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CF900BF5FF8BFF91FC9779D9D7EBF87F" pageId="12" pageNumber="655" type="materials_examined">
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<paragraph id="8735587EFF8BFF91FC9779D9D7EBF87F" blockId="12.[813,1456,1924,2027]" pageId="12" pageNumber="655">
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MATERIAL EXAMINED. —
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<materialsCitation id="37E25223FF8BFF91FBEE79D9D7EBF87F" ID-GBIF-Occurrence="3396082305" pageId="12" pageNumber="655">
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<collectionCode id="E19BC0BBFF8BFF91FBEE79D9D085F808" box="[1108,1200,1924,1948]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="655">MNHN</collectionCode>
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||
.
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<collectionCode id="E19BC0BBFF8BFF91FB0E79D9D0F1F808" box="[1204,1220,1924,1948]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="12" pageNumber="655" type="Herbarium">F</collectionCode>
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.MTC240-MTC242, MTC243, nearly fifty osteoderms, a few complete, most more or less severely damaged by digestive processes or post-burial damages (
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<figureCitation id="1FB144FBFF8BFF91FC29798ED7FBF87F" box="[915,974,2003,2027]" captionStart="FIG" captionStartId="13.[149,160,1769,1786]" captionTargetBox="[261,1327,214,1726]" captionTargetId="figure-154@13.[261,1327,214,1726]" captionTargetPageId="13" captionText="FIG. 9. — Pan-Shinisauridae,osteoderms,MNHN.F.MTC240-MTC242,specimen in:A, dorsal view;B, dorsal view;C, dorsal and ventral views.Scale bar:1 mm." figureDoi="http://doi.org/10.5281/zenodo.5628453" httpUri="https://zenodo.org/record/5628453/files/figure.png" pageId="12" pageNumber="655">Fig. 9</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<caption id="D3F508F6FF8AFF90FF2F78B4D195F96E" ID-DOI="http://doi.org/10.5281/zenodo.5628453" ID-Zenodo-Dep="5628453" box="[149,1440,1769,1786]" httpUri="https://zenodo.org/record/5628453/files/figure.png" pageId="13" pageNumber="656" startId="13.[149,160,1769,1786]" targetBox="[261,1327,214,1726]" targetPageId="13">
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<paragraph id="8735587EFF8AFF90FF2F78B4D195F96E" blockId="13.[149,1440,1769,1786]" box="[149,1440,1769,1786]" pageId="13" pageNumber="656">
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FIG. 9. — Pan-Shinisauridae,osteoderms,MNHN.F.MTC240-MTC242,specimen in:
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<emphasis id="B5FE846CFF8AFF90FCFA78B4D778F96E" bold="true" box="[832,845,1769,1786]" pageId="13" pageNumber="656">A</emphasis>
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, dorsal view;
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<emphasis id="B5FE846CFF8AFF90FC0078B4D7F2F96E" bold="true" box="[954,967,1769,1786]" pageId="13" pageNumber="656">B</emphasis>
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, dorsal view;
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<emphasis id="B5FE846CFF8AFF90FB8F78B4D076F96E" bold="true" box="[1077,1091,1769,1786]" pageId="13" pageNumber="656">C</emphasis>
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, dorsal and ventral views.Scale bar:1 mm.
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</paragraph>
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</caption>
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<subSubSection id="CF900BF5FF8AFF90FF3E796FD02EF87F" pageId="13" pageNumber="656" type="description">
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<paragraph id="8735587EFF8AFF90FF3E796FD520F8DE" blockId="13.[132,777,1842,2028]" box="[132,277,1842,1868]" pageId="13" pageNumber="656">DESCRIPTION</paragraph>
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<paragraph id="8735587EFF8AFF90FF3E790FD02EF87F" blockId="13.[132,777,1842,2028]" lastBlockId="13.[811,1458,1841,2028]" pageId="13" pageNumber="656">
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Measures (mm): maximum width (w) of osteoderms ranges between 1.1 and
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<quantity id="4072F59BFF8AFF90FE85792CD5ACF818" box="[319,409,1905,1932]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="13" pageNumber="656" unit="mm" value="3.9">3.9 mm</quantity>
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. The osteoderms, when complete, are oval, suboval or subrectangular elements with more or less irregular margins. Gliding surfaces are absent. All bear a prominent medial keel that extends the full length of the osteoderm. A slight concavity on the underside of some osteoderms reflects the form of the keel. From the keel a pattern of deep pits or grooves and marked ridges radiates. The dorsal surface of most osteoderms is flat but a few have a rather strongly vaulted shape which certainly reflect different positions on the body.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CF900BF5FF89FF93FF3E7E8AD0A6FEE5" pageId="14" pageNumber="657" type="discussion">
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<paragraph id="8735587EFF89FF93FF3E7E8AD5D6FF64" blockId="14.[131,775,215,2028]" box="[132,483,215,241]" pageId="14" pageNumber="657">COMPARISONS AND DISCUSSION</paragraph>
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<paragraph id="8735587EFF89FF93FF3E7EAAD0A6FEE5" blockId="14.[131,775,215,2028]" lastBlockId="14.[811,1455,215,369]" pageId="14" pageNumber="657">
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||
Some characters are traditionally used to separate anguimorph taxa (and more specifically anguid genera) by their osteoderms (
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<bibRefCitation id="E31B258FFF89FF93FEA87F6AD5ECFEC5" author="HOFFSTETTER R." box="[274,473,311,337]" pageId="14" pageNumber="657" pagination="149 - 157" refId="ref10991" refString="HOFFSTETTER R. 1962 a. - Observations sur les osteodermes et la classification des anguides actuels et fossiles (Reptiles, Sauriens). Bulletin du Museum national d'Histoire naturelle, 2 eme serie, 34: 149 - 157. https: // www. biodiversitylibrary. org / page / 54962884" type="journal article" year="1962">Hoffstetter 1962a</bibRefCitation>
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||
;
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||
<bibRefCitation id="E31B258FFF89FF93FE5D7F6AD6B8FEC5" author="MESZOELY C. A." box="[487,653,311,337]" pageId="14" pageNumber="657" pagination="87 - 149" refId="ref12208" refString="MESZOELY C. A. 1970. - North American Fossil Anguid Lizards. Bulletin of the Museum of Comparative Zoology, Harvard University 139: 87 - 149. https: // www. biodiversitylibrary. org / part / 31376" type="journal article" year="1970">Meszoely 1970</bibRefCitation>
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||
;
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<bibRefCitation id="E31B258FFF89FF93FD217F6AD4CCFEE6" author="BOCHATON C. & DE BUFFRENIL V. & LEMOINE M. & BAILON S. & INEICH I." pageId="14" pageNumber="657" pagination="1333 - 1344" refId="ref8490" refString="BOCHATON C., DE BUFFRENIL V., LEMOINE M., BAILON S. & INEICH I. 2015. - Body location and tail regeneration effects on osteoderms morphology - are they useful tools for systematic, paleontology, and skeletochronology in diploglossine lizards (Squamata, Anguidae)? Journal of Morphology 276 (11): 1333 - 1344. https: // doi. org / 10.1002 / jmor. 20421" type="journal article" year="2015">
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Bochaton
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||
<emphasis id="B5FE846CFF89FF93FF3E7F05D48CFEE5" box="[132,185,343,369]" italics="true" pageId="14" pageNumber="657">et al.</emphasis>
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||
2015
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||
</bibRefCitation>
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||
,
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||
<bibRefCitation id="E31B258FFF89FF93FEBC7F0AD575FEE5" author="BOCHATON C. & BOISTEL R. & CASAGRANDE F. & GROUARD S. & BAILON S." box="[262,320,343,369]" pageId="14" pageNumber="657" pagination="1 - 12" refId="ref8567" refString="BOCHATON C., BOISTEL R., CASAGRANDE F., GROUARD S. & BAILON S. 2016. - A fossil Diploglossus (Squamata, Anguidae) lizard from Basse-Terre and Grande-Terre Islands (Guadeloupe, French West Indies). Scientific Reports 6 (28475): 1 - 12. https: // doi. org / 10.1038 / srep 28475" type="journal article" year="2016">2016</bibRefCitation>
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), including the presence of a gliding surface and of a keel. For example,
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<bibRefCitation id="E31B258FFF89FF93FE627F2AD6B9FE05" author="GAUTHIER J." box="[472,652,375,401]" pageId="14" pageNumber="657" pagination="7 - 54" refId="ref10409" refString="GAUTHIER J. 1982. - Fossil xenosaurid and anguid lizards from the early Eocene formation Wasatch, southeast Wyoming, and a revision of the Anguioidea. Contribution to Geology of the University of Wyoming 21 (1): 7 - 54." type="journal article" year="1982">Gauthier (1982)</bibRefCitation>
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considered keeled body osteoderms to be the plesiomorphic state for
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<taxonomicName id="408A23FDFF89FF93FF3E7FEBD510FE44" authorityName="Furbringer" authorityYear="1900" box="[132,293,438,464]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="653" phylum="Chordata" rank="subOrder" subOrder="Anguimorpha">Anguimorpha</taxonomicName>
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. Referral of the osteoderms from Montchenot to Pan
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<emphasis id="B5FE846CFF89FF93FEB17F8BD527FE64" box="[267,274,470,496]" italics="true" pageId="14" pageNumber="657">-</emphasis>
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||
<taxonomicName id="408A23FDFF89FF93FEAB7F8BD732FE65" authority="(sensu Smith & Gauthier 2013)" baseAuthorityName="sensu Smith & Gauthier" baseAuthorityYear="2013" box="[273,775,470,497]" class="Reptilia" family="Shinisauridae" genus="Shinisaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="genus">
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<emphasis id="B5FE846CFF89FF93FEAB7F8BD5A7FE64" box="[273,402,470,496]" italics="true" pageId="14" pageNumber="657">Shinisaurus</emphasis>
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||
(
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<emphasis id="B5FE846CFF89FF93FE1C7F8AD5E8FE64" box="[422,477,471,496]" italics="true" pageId="14" pageNumber="657">sensu</emphasis>
|
||
<bibRefCitation id="E31B258FFF89FF93FE5C7F8BD6C8FE64" author="SMITH K. T. & GAUTHIER J. A." box="[486,765,470,496]" pageId="14" pageNumber="657" pagination="135 - 230" refId="ref13306" refString="SMITH K. T. & GAUTHIER J. A. 2013. - Early Eocene Lizards of the Wasatch Formation near Bitter Creek, Wyoming: Diversity and Paleoenvironment during an Interval of Global Warming. Bulletin of the Peabody Museum of Natural History 54 (2): 135 - 230. https: // doi. org / 10.3374 / 014.054.0205" type="journal article" year="2013">Smith & Gauthier 2013</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
follows from the combination of the features described above. These osteoderms are similar in shape to those of other fossil pan-shinisaurs, particularly
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<taxonomicName id="408A23FDFF89FF93FDE87C6AD5D4FDE4" authority="Smith & Gauthier, 2013" authorityName="Smith & Gauthier" authorityYear="2013" class="Reptilia" family="Necrosauridae" genus="Provaranosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="species" species="fatuus">
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||
<emphasis id="B5FE846CFF89FF93FDE87C6AD4F3FDE4" italics="true" pageId="14" pageNumber="657">Provaranosaurus fatuus</emphasis>
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||
<bibRefCitation id="E31B258FFF89FF93FF777C0BD5D4FDE4" author="SMITH K. T. & GAUTHIER J. A." box="[205,481,598,624]" pageId="14" pageNumber="657" pagination="135 - 230" refId="ref13306" refString="SMITH K. T. & GAUTHIER J. A. 2013. - Early Eocene Lizards of the Wasatch Formation near Bitter Creek, Wyoming: Diversity and Paleoenvironment during an Interval of Global Warming. Bulletin of the Peabody Museum of Natural History 54 (2): 135 - 230. https: // doi. org / 10.3374 / 014.054.0205" type="journal article" year="2013">Smith & Gauthier, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="E31B258FFF89FF93FE4B7C0BD6CAFDE4" author="SMITH K. T. & GAUTHIER J. A." box="[497,767,598,624]" pageId="14" pageNumber="657" pagination="135 - 230" refId="ref13306" refString="SMITH K. T. & GAUTHIER J. A. 2013. - Early Eocene Lizards of the Wasatch Formation near Bitter Creek, Wyoming: Diversity and Paleoenvironment during an Interval of Global Warming. Bulletin of the Peabody Museum of Natural History 54 (2): 135 - 230. https: // doi. org / 10.3374 / 014.054.0205" type="journal article" year="2013">Smith & Gauthier 2013</bibRefCitation>
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||
, early Eocene of the Wasatch Formation,
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||
<collectingRegion id="454E969CFF89FF93FD847C2BD69AFD04" box="[574,687,630,656]" country="United States of America" name="Wyoming" pageId="14" pageNumber="657">Wyoming</collectingRegion>
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||
,
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<collectingCountry id="FF9D18EEFF89FF93FD037C2BD4FDFD24" name="United States of America" pageId="14" pageNumber="657">United States</collectingCountry>
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||
),
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<taxonomicName id="408A23FDFF89FF93FF617CCBD6BCFD24" authority="Klembara, 2008" authorityName="Klembara" authorityYear="2008" box="[219,649,662,688]" class="Reptilia" genus="Merkurosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="species" species="ornatus">
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<emphasis id="B5FE846CFF89FF93FF617CCBD5E7FD3B" box="[219,466,662,688]" italics="true" pageId="14" pageNumber="657">Merkurosaurus ornatus</emphasis>
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<bibRefCitation id="E31B258FFF89FF93FE6D7CCBD6BCFD24" author="KLEMBARA J." box="[471,649,662,688]" pageId="14" pageNumber="657" pagination="81 - 94" refId="ref11527" refString="KLEMBARA J. 2008. - A new Anguimorph lizard from the Lower Miocene of North-West Bohemia, Czech Republic. Palaeontology 51 (1): 81 - 94. https: // doi. org / 10.1111 / j. 1475 - 4983.2007.00732. x" type="journal article" year="2008">Klembara, 2008</bibRefCitation>
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||
</taxonomicName>
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||
(
|
||
<bibRefCitation id="E31B258FFF89FF93FD217CCBD48BFD44" author="KLEMBARA J." pageId="14" pageNumber="657" pagination="81 - 94" refId="ref11527" refString="KLEMBARA J. 2008. - A new Anguimorph lizard from the Lower Miocene of North-West Bohemia, Czech Republic. Palaeontology 51 (1): 81 - 94. https: // doi. org / 10.1111 / j. 1475 - 4983.2007.00732. x" type="journal article" year="2008">Klembara 2008</bibRefCitation>
|
||
, early Miocene, Orleanian, MN3,
|
||
<collectingCountry id="FF9D18EEFF89FF93FDFD7CE8D698FD44" box="[583,685,693,720]" name="Czech Republic" pageId="14" pageNumber="657">Bohemia</collectingCountry>
|
||
) and an indeterminate pan-shinisaur from Messel (
|
||
<bibRefCitation id="E31B258FFF89FF93FDCC7C88D6CAFD64" author="SMITH K. T." box="[630,767,725,752]" pageId="14" pageNumber="657" pagination="1313743" refId="ref13262" refString="SMITH K. T. 2017. - First Crocodile-Tailed Lizard (Squamata: Pan-Shinisaurus) from the Paleogene of Europe. Journal of Vertebrate Paleontology 37 (3): e 1313743. https: // doi. org / 10.1080 / 02724634.2017.1313743" type="journal article" year="2017">Smith 2017</bibRefCitation>
|
||
, middle Eocene,
|
||
<collectingCountry id="FF9D18EEFF89FF93FE957CA8D5ACFC9B" box="[303,409,757,783]" name="Germany" pageId="14" pageNumber="657">Germany</collectingCountry>
|
||
). Crocodile-tailed lizards (Chinese crocodile lizard) or shinisaurs are represented by a single living species,
|
||
<taxonomicName id="408A23FDFF89FF93FEA67D68D6BBFCDB" authority="Ahl, 1930" authorityName="Ahl" authorityYear="1930" box="[284,654,821,847]" class="Reptilia" family="Shinisauridae" genus="Shinisaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="species" species="crocodilurus">
|
||
<emphasis id="B5FE846CFF89FF93FEA67D68D62DFCDB" box="[284,536,821,847]" italics="true" pageId="14" pageNumber="657">Shinisaurus crocodilurus</emphasis>
|
||
Ahl, 1930
|
||
</taxonomicName>
|
||
. It is worth noting that similar osteoderms have already been reported in the European
|
||
<taxonomicName id="408A23FDFF89FF93FEFE7D28D587FC1B" authority="Montchenot" authorityName="Montchenot" box="[324,434,885,911]" class="Reptilia" family="Shinisauridae" genus="Paleocene" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="645" phylum="Chordata" rank="genus">Paleocene</taxonomicName>
|
||
and early Eocene, in particular in the localities of Cernay (MP6,
|
||
<bibRefCitation id="E31B258FFF89FF93FD8F7DC8D6C3FC3B" author="HOFFSTETTER R." box="[565,758,917,943]" pageId="14" pageNumber="657" pagination="134 - 141" refId="ref10919" refString="HOFFSTETTER R. 1943. - Varanidae et Necrosauridae fossiles. Bulletin du Museum national d'Histoire naturelle, 2 eme serie, 15: 134 - 141. https: // www. biodiversitylibrary. org / page / 52906476" type="journal article" year="1943">Hoffstetter 1943</bibRefCitation>
|
||
), Dormaal and Le Quesnoy (early Eocene, MP7,
|
||
<bibRefCitation id="E31B258FFF89FF93FD197DE8D576FC7A" author="HECHT M. K. & HOFFSTETTER R." pageId="14" pageNumber="657" pagination="1 - 30" refId="ref10875" refString="HECHT M. K. & HOFFSTETTER R. 1962. - Note preliminaire sur les Amphibiens et les Squamates du Landenien superieur et du Tongrien de Belgique. Bulletin de l'Institut royal des Sciences naturelles de Belgique 39: 1 - 30." type="journal article" year="1962">Hecht & Hoffstetter 1962</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E31B258FFF89FF93FEE97D88D5FBFC7B" author="AUGE M." box="[339,462,981,1007]" pageId="14" pageNumber="657" pagination="161 - 173" refId="ref7979" refString="AUGE M. 1990. - La faune de lezards et d'amphisbenes (Reptilia, Squamata) dugisement de Dormaal (Belgique, Eocene inferieur). Bulletin de l'Institut royal des Sciences naturelles de Belgique, Sciences de la Terre 60: 161 - 173." type="journal article" year="1990">Augé 1990</bibRefCitation>
|
||
) and perhaps at Rivecourt- Petit Pâtis (
|
||
<bibRefCitation id="E31B258FFF89FF93FEBB7DA9D589FB9B" author="SMITH T. & QUESNEL F. & DE PLOEG G. & DE FRANCESCHI D. & METAIS G. & DE BAST E. & SOLE F. & FOLIE A. & BOURA A. & CLAUDE C. & DUPUIS C. & GAGNAISON C. & IAKOVLEVA A. & MARTIN J. & MAUBERT F. & PRIEUR J. & ROCHE E. & STORME J. - Y. & THOMAS R. & TONG H. & YANS J. & BUFFETAUT E." box="[257,444,1012,1039]" pageId="14" pageNumber="657" pagination="93249" refId="ref13433" refString="SMITH T., QUESNEL F., DE PLOEG G., DE FRANCESCHI D., METAIS G., DE BAST E., SOLE F., FOLIE A., BOURA A., CLAUDE C., DUPUIS C., GAGNAISON C., IAKOVLEVA A., MARTIN J., MAUBERT F., PRIEUR J., ROCHE E., STORME J. - Y., THOMAS R., TONG H., YANS J. & BUFFETAUT E. 2014. - First Clarkforkian Equivalent Land Mammal Age in the Latest Paleocene Basal Sparnacian Facies of Europe: Fauna, Flora, Paleoenvironment and (Bio) stratigraphy. Plos One 9 (3): e 93249. https: // doi. org / 10.1371 / journal. pone. 0086229" type="journal article" year="2014">
|
||
Smith
|
||
<emphasis id="B5FE846CFF89FF93FEF07DA8D548FB9A" box="[330,381,1012,1038]" italics="true" pageId="14" pageNumber="657">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="E31B258FFF89FF93FE697DA9D732FB9B" author="HECHT M. K. & HOFFSTETTER R." box="[467,775,1012,1039]" pageId="14" pageNumber="657" pagination="1 - 30" refId="ref10875" refString="HECHT M. K. & HOFFSTETTER R. 1962. - Note preliminaire sur les Amphibiens et les Squamates du Landenien superieur et du Tongrien de Belgique. Bulletin de l'Institut royal des Sciences naturelles de Belgique 39: 1 - 30." type="journal article" year="1962">Hecht & Hoffstetter (1962)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="E31B258FFF89FF93FF027A49D572FBBB" author="AUGE M." box="[184,327,1044,1071]" pageId="14" pageNumber="657" refId="ref8109" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Museum national d'Histoire naturelle, Paris, 369 p. (Memoires du Museum national d'Histoire naturelle; 192)." type="book" year="2005">Augé (2005)</bibRefCitation>
|
||
suggested that these osteoderms could be attributed to the genus
|
||
<taxonomicName id="408A23FDFF89FF93FE797A68D67CFBDA" authorityName="Filhol" authorityYear="1876" box="[451,585,1077,1102]" class="Reptilia" family="Necrosauridae" genus="Necrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="genus">
|
||
<emphasis id="B5FE846CFF89FF93FE797A68D67CFBDA" box="[451,585,1077,1102]" italics="true" pageId="14" pageNumber="657">Necrosaurus</emphasis>
|
||
</taxonomicName>
|
||
as they are also similar to those of
|
||
<taxonomicName id="408A23FDFF89FF93FEE87A09D60AFBFA" authorityName="Zittel" authorityYear="1887" box="[338,575,1108,1134]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="species" species="cayluxi">
|
||
<emphasis id="B5FE846CFF89FF93FEE87A09D60AFBFA" box="[338,575,1108,1134]" italics="true" pageId="14" pageNumber="657">Palaeovaranus cayluxi</emphasis>
|
||
</taxonomicName>
|
||
(Ex
|
||
<taxonomicName id="408A23FDFF89FF93FDC87A08D6C1FBFA" authorityName="Filhol" authorityYear="1876" box="[626,756,1109,1134]" class="Reptilia" family="Necrosauridae" genus="Necrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="genus">
|
||
<emphasis id="B5FE846CFF89FF93FDC87A08D6C1FBFA" box="[626,756,1109,1134]" italics="true" pageId="14" pageNumber="657">Necrosaurus</emphasis>
|
||
</taxonomicName>
|
||
), see figs. in
|
||
<bibRefCitation id="E31B258FFF89FF93FEBF7A28D548FB1A" author="RAGE J. - C." box="[261,381,1140,1167]" pageId="14" pageNumber="657" pagination="167 - 326" refId="ref12641" refString="RAGE J. - C. 1978. - Squamates, in GEZE B., RAGE J. - C., VERGNAUD- GRAZZINI C., DE BROIN F., BUFFETAUT E., MOURER- CHAUVIRE C., CROCHET J. Y., SIGE B., SUDRE J., REMY J. A., LANGE- BADRE B., BONIS L. DE, HARTENBERGER J. L. & VIANEY- LIAUD M. (eds), La Poche a Phosphate de Ste-Neboule (Lot) et sa faune de vertebres du Ludien superieur. Palaeovertebrata 8: 167 - 326." type="journal article" year="1978">Rage 1978</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E31B258FFF89FF93FE377A28D63DFB1A" author="ESTES R." box="[397,520,1140,1167]" pageId="14" pageNumber="657" refId="ref9665" refString="ESTES R. 1983. - Sauria terrestria, Amphisbaenia, in KUHN O. & WELLNHOFER P. (eds), Handbuch der Palaoherpetologie, Teil 10 A, G. Fischer Verlag, 249 p." type="book" year="1983">Estes 1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E31B258FFF89FF93FDA27A29D6CAFB1A" author="AUGE M. & SMITH R." box="[536,767,1140,1166]" pageId="14" pageNumber="657" pagination="148 - 170" refId="ref8207" refString="AUGE M. & SMITH R. 2009. - An assemblage of early oligocene lizards (Squamata) from the locality of Boutersem (Belgium), with comments on the Eocene-Oligocene transition. Zoological Journal Linnean Society 155: 148 - 170. https: // doi. org / 10.1111 / j. 1096 - 3642.2008.00435. x" type="journal article" year="2009">Augé & Smith 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E31B258FFF89FF93FF3E7AC9D5A8FB3A" author="KLEMBARA J. & GREEN B." box="[132,413,1172,1198]" pageId="14" pageNumber="657" pagination="97 - 129" refId="ref11625" refString="KLEMBARA J. & GREEN B. 2010. - Anguimorph lizards (Squamata, Anguimorpha) from the Middle and Late Eocene of the Hampshire Basin of southern England. Journal of Systematic Palaeontology 8 (1): 97 - 129. https: // doi. org / 10.1080 / 14772011003603531" type="journal article" year="2010">Klembara & Green 2010</bibRefCitation>
|
||
. However, the taxonomic status and phylogenetic affinities of these lizards are a complex matter. Georgalis (2017) pointed out that the name
|
||
<taxonomicName id="408A23FDFF89FF93FD057A89D4FDFA99" authorityName="Filhol" authorityYear="1876" class="Reptilia" family="Necrosauridae" genus="Necrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="genus">
|
||
<emphasis id="B5FE846CFF89FF93FD057A89D4FDFA99" italics="true" pageId="14" pageNumber="657">Necrosaurus</emphasis>
|
||
</taxonomicName>
|
||
, as established by
|
||
<bibRefCitation id="E31B258FFF89FF93FE1A7AAED602FA9A" author="FILHOL H." box="[416,567,1267,1294]" pageId="14" pageNumber="657" pagination="27 - 28" refId="ref10069" refString="FILHOL H. 1876. - Sur les reptiles des Phosphorites du Quercy. Bulletin de la Societe philomatique de Paris, serie 6, 13: 27 - 28. https: // www. biodiversitylibrary. org / page / 31643639" type="journal article" year="1876">Filhol (1876)</bibRefCitation>
|
||
is a nomina nuda and that
|
||
<bibRefCitation id="E31B258FFF89FF93FF517B4ED540FABA" author="ZITTEL K. A." box="[235,373,1299,1326]" pageId="14" pageNumber="657" refId="ref13891" refString="ZITTEL K. A. 1887 - 1890. - Handbuch der Palaontologie. Palaeozoologie. III. Pisces, Amphibia, Reptilia, Aves. Druck und Verlag von R. Oldenbourg, Munchen / Leipzig. https: // doi. org / 10.5962 / bhl. title. 34265" type="book" year="1887">Zittel (1887</bibRefCitation>
|
||
-1890) was the first author to make the name
|
||
<taxonomicName id="408A23FDFF89FF93FF407B6ED5DBFAD9" authorityName="Zittel" authorityYear="1887" box="[250,494,1331,1357]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="species" species="cayluxi">
|
||
<emphasis id="B5FE846CFF89FF93FF407B6ED5DBFAD9" box="[250,494,1331,1357]" italics="true" pageId="14" pageNumber="657">Palaeovaranus cayluxi</emphasis>
|
||
</taxonomicName>
|
||
available. The phylogenetic affinities of
|
||
<taxonomicName id="408A23FDFF89FF93FEF67B0ED5D3FAF9" box="[332,486,1363,1389]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="genus">
|
||
<emphasis id="B5FE846CFF89FF93FEF67B0ED5D3FAF9" box="[332,486,1363,1389]" italics="true" pageId="14" pageNumber="657">Palaeovaranus</emphasis>
|
||
</taxonomicName>
|
||
are a moot point: briefly,
|
||
<bibRefCitation id="E31B258FFF89FF93FF3F7B2ED582FA1A" author="MCDOWELL S. B. & BOGERT C. M." box="[133,439,1395,1422]" pageId="14" pageNumber="657" pagination="1 - 142" refId="ref12152" refString="MCDOWELL S. B. & BOGERT C. M. 1954. - The systematic position of Lanthanotus and the affinities of the Anguinomorphan lizards. Bulletin of the American Museum of Natural History 105 (1): 1 - 142. http: // hdl. handle. net / 2246 / 1146" type="journal article" year="1954">McDowell & Bogert (1954)</bibRefCitation>
|
||
noted significant morphological differences between
|
||
<taxonomicName id="408A23FDFF89FF93FE307BCED611FA39" box="[394,548,1427,1453]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="genus">
|
||
<emphasis id="B5FE846CFF89FF93FE307BCED611FA39" box="[394,548,1427,1453]" italics="true" pageId="14" pageNumber="657">Palaeovaranus</emphasis>
|
||
</taxonomicName>
|
||
and members of the Platynota (
|
||
<emphasis id="B5FE846CFF89FF93FEBA7BE9D502FA59" box="[256,311,1460,1485]" italics="true" pageId="14" pageNumber="657">sensu</emphasis>
|
||
<bibRefCitation id="E31B258FFF89FF93FE847BEED637FA59" author="PREGILL G. K. & GAUTHIER J. A. & GREENE H. W." box="[318,514,1459,1485]" pageId="14" pageNumber="657" pagination="167 - 202" refId="ref12573" refString="PREGILL G. K., GAUTHIER J. A. & GREENE H. W. 1986. - The evolution of helodermatid squamates, with description of a new taxon and an overview of Varanoidea. Transactions of the San Diego Society of Natural History 21 (11): 167 - 202. https: // www. biodiversitylibrary. org / part / 24582" type="journal article" year="1986">
|
||
Pregill
|
||
<emphasis id="B5FE846CFF89FF93FE377BE9D5F7FA59" box="[397,450,1459,1485]" italics="true" pageId="14" pageNumber="657">et al.</emphasis>
|
||
1986
|
||
</bibRefCitation>
|
||
) and they referred it to xenosaurid lizards, an option first adopted by
|
||
<bibRefCitation id="E31B258FFF89FF93FD317B8ED4E0F999" author="HOFFSTETTER R." pageId="14" pageNumber="657" pagination="422 - 424" refId="ref10959" refString="HOFFSTETTER R. 1954. - Sur la position systematique de Necrosaurus: Saurien de l'Eocene europeen. Comptes Rendus de la Societe geologique de France 16: 422 - 424." type="journal article" year="1954">Hoffstetter (1954)</bibRefCitation>
|
||
. Later this author returned
|
||
<taxonomicName id="408A23FDFF89FF93FDA77BAED68CF999" box="[541,697,1523,1549]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="genus">
|
||
<emphasis id="B5FE846CFF89FF93FDA77BAED68CF999" box="[541,697,1523,1549]" italics="true" pageId="14" pageNumber="657">Palaeovaranus</emphasis>
|
||
</taxonomicName>
|
||
to the Platynota (
|
||
<bibRefCitation id="E31B258FFF89FF93FEBF784FD5E3F9B8" author="HOFFSTETTER R." box="[261,470,1554,1580]" pageId="14" pageNumber="657" pagination="243 - 279" refId="ref11045" refString="HOFFSTETTER R. 1962 b. - Revue des recentes acquisitions concernant l'histoire et la systematique des squamates. Problemes actuels de Paleontologie (evolution des vertebres). Colloques internationaux du CNRS 104: 243 - 279." type="journal article" year="1962">Hoffstetter 1962b</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="E31B258FFF89FF93FE4B784ED645F9B9" author="LEE M. S. Y." box="[497,624,1555,1581]" pageId="14" pageNumber="657" pagination="53 - 91" refId="ref11957" refString="LEE M. S. Y. 1997. - The phylogeny of varanoid lizards and the affinities of snakes. Philosophical Transactions of the Royal Society B: Biological Sciences 352: 53 - 91. https: // doi. org / 10.1098 / rstb. 1997.0005" type="journal article" year="1997">Lee (1997)</bibRefCitation>
|
||
rejected this taxon as paraphyletic. The phylogenetic position of
|
||
<taxonomicName id="408A23FDFF89FF93FD67786FD4CEF9F8" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="genus">
|
||
<emphasis id="B5FE846CFF89FF93FD67786FD4CEF9F8" italics="true" pageId="14" pageNumber="657">Palaeovaranus</emphasis>
|
||
</taxonomicName>
|
||
is still a matter of discussion, although several derived characters suggest Platynotan relationships (see discussion in
|
||
<bibRefCitation id="E31B258FFF89FF93FEA678CFD594F938" author="SMITH K. T." box="[284,417,1682,1708]" pageId="14" pageNumber="657" pagination="1313743" refId="ref13262" refString="SMITH K. T. 2017. - First Crocodile-Tailed Lizard (Squamata: Pan-Shinisaurus) from the Paleogene of Europe. Journal of Vertebrate Paleontology 37 (3): e 1313743. https: // doi. org / 10.1080 / 02724634.2017.1313743" type="journal article" year="2017">Smith 2017</bibRefCitation>
|
||
). In contrast, the attribution of
|
||
<taxonomicName id="408A23FDFF89FF93FF3E78EED54AF958" authorityName="Smith & Gauthier" authorityYear="2013" box="[132,383,1714,1740]" class="Reptilia" family="Necrosauridae" genus="Provaranosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="species" species="fatuus">
|
||
<emphasis id="B5FE846CFF89FF93FF3E78EED54AF958" box="[132,383,1714,1740]" italics="true" pageId="14" pageNumber="657">Provaranosaurus fatuus</emphasis>
|
||
</taxonomicName>
|
||
and specimen SMF ME 11403 (an autotomized tail) from Messel to pan-shinisaur is a settled matter as they show no Platynotan derived characters (
|
||
<bibRefCitation id="E31B258FFF89FF93FF34794FD594F8B8" author="SMITH K. T. & GAUTHIER J. A." box="[142,417,1810,1836]" pageId="14" pageNumber="657" pagination="135 - 230" refId="ref13306" refString="SMITH K. T. & GAUTHIER J. A. 2013. - Early Eocene Lizards of the Wasatch Formation near Bitter Creek, Wyoming: Diversity and Paleoenvironment during an Interval of Global Warming. Bulletin of the Peabody Museum of Natural History 54 (2): 135 - 230. https: // doi. org / 10.3374 / 014.054.0205" type="journal article" year="2013">Smith & Gauthier 2013</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E31B258FFF89FF93FE0B794FD602F8B8" author="SMITH K. T." box="[433,567,1810,1836]" pageId="14" pageNumber="657" pagination="1313743" refId="ref13262" refString="SMITH K. T. 2017. - First Crocodile-Tailed Lizard (Squamata: Pan-Shinisaurus) from the Paleogene of Europe. Journal of Vertebrate Paleontology 37 (3): e 1313743. https: // doi. org / 10.1080 / 02724634.2017.1313743" type="journal article" year="2017">Smith 2017</bibRefCitation>
|
||
). The fossils from Dormaal (osteoderms, vertebrae and an undescribed dentary) previously attributed to
|
||
<taxonomicName id="408A23FDFF89FF93FE6A790FD732F8F8" authority="" box="[464,775,1873,1900]" class="Reptilia" family="Necrosauridae" genus="Necrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="subGenus" subGenus="Palaeovaranus">
|
||
<emphasis id="B5FE846CFF89FF93FE6A790FD667F8FF" box="[464,594,1874,1899]" italics="true" pageId="14" pageNumber="657">Necrosaurus</emphasis>
|
||
(
|
||
<emphasis id="B5FE846CFF89FF93FDD9790CD6C8F8FF" box="[611,765,1873,1899]" italics="true" pageId="14" pageNumber="657">Palaeovaranus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
show no Platynotan features and are nearly identical to the material of
|
||
<taxonomicName id="408A23FDFF89FF93FEB379CFD633F83F" authorityName="Smith & Gauthier" authorityYear="2013" box="[265,518,1937,1963]" class="Reptilia" family="Necrosauridae" genus="Provaranosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="species" species="fatuus">
|
||
<emphasis id="B5FE846CFF89FF93FEB379CFD633F83F" box="[265,518,1937,1963]" italics="true" pageId="14" pageNumber="657">Provaranosaurus fatuus</emphasis>
|
||
</taxonomicName>
|
||
described by
|
||
<bibRefCitation id="E31B258FFF89FF93FD1F79CCD51AF85F" author="SMITH K. T. & GAUTHIER J. A." pageId="14" pageNumber="657" pagination="135 - 230" refId="ref13306" refString="SMITH K. T. & GAUTHIER J. A. 2013. - Early Eocene Lizards of the Wasatch Formation near Bitter Creek, Wyoming: Diversity and Paleoenvironment during an Interval of Global Warming. Bulletin of the Peabody Museum of Natural History 54 (2): 135 - 230. https: // doi. org / 10.3374 / 014.054.0205" type="journal article" year="2013">Smith & Gauthier 2013</bibRefCitation>
|
||
. In particular,
|
||
<taxonomicName id="408A23FDFF89FF93FE5979EFD6ACF85F" authorityName="Gilmore" authorityYear="1942" box="[483,665,1970,1995]" class="Reptilia" family="Necrosauridae" genus="Provaranosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="genus">
|
||
<emphasis id="B5FE846CFF89FF93FE5979EFD6ACF85F" box="[483,665,1970,1995]" italics="true" pageId="14" pageNumber="657">Provaranosaurus</emphasis>
|
||
</taxonomicName>
|
||
has both rectangular and oval osteoderms, as in the material from Monchenot, while
|
||
<taxonomicName id="408A23FDFF89FF93FBBD7E8AD094FF65" box="[1031,1185,215,241]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="genus">
|
||
<emphasis id="B5FE846CFF89FF93FBBD7E8AD094FF65" box="[1031,1185,215,241]" italics="true" pageId="14" pageNumber="657">Palaeovaranus</emphasis>
|
||
</taxonomicName>
|
||
bears only ovoid osteoderms. On the basis of these resemblances, the osteoderms from Monchenot may be referred to pan-
|
||
<taxonomicName id="408A23FDFF89FF93FABA7F4AD14BFEA5" baseAuthorityName="sensu Smith & Gauthier" baseAuthorityYear="2013" box="[1280,1406,279,305]" class="Reptilia" family="Shinisauridae" genus="Shinisaurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="genus">
|
||
<emphasis id="B5FE846CFF89FF93FABA7F4AD14BFEA5" box="[1280,1406,279,305]" italics="true" pageId="14" pageNumber="657">Shinisaurus</emphasis>
|
||
</taxonomicName>
|
||
and the presence of rectangular osteoderms seems to exclude an attribution to
|
||
<taxonomicName id="408A23FDFF89FF93FC487F0AD0B9FEE5" box="[1010,1164,343,369]" class="Reptilia" family="Palaeovaranidae" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="657" phylum="Chordata" rank="genus">
|
||
<emphasis id="B5FE846CFF89FF93FC487F0AD0B9FEE5" box="[1010,1164,343,369]" italics="true" pageId="14" pageNumber="657">Palaeovaranus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |