202 lines
23 KiB
XML
202 lines
23 KiB
XML
<document id="44CFBC89BDD1AA145642BF3386124B83" ID-CLB-Dataset="24216" ID-DOI="10.11646/zootaxa.4729.3.1" ID-GBIF-Dataset="f7aee5da-a7ce-4660-b00c-b272b6204a19" ID-ISSN="1175-5326" ID-Zenodo-Dep="3632662" ID-ZooBank="330533E0-25B8-4C66-89F2-B93F3E88DD5A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1580475428538" checkinUser="plazi" docAuthor="Fominykh, Dmitriy D., Zamotajlov, Alexandr S., Khomitskiy, Eugeniy E. & Titarenko, Andrey Yu." docDate="2020" docId="E676063DFFF43861FF70B2A63E74E6AD" docLanguage="en" docName="zootaxa.4729.3.1.pdf" docOrigin="Zootaxa 4729 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Carabus (Lipaster) stjernvalli subsp. bogatshevi Zamotajlov 1989" docType="treatment" docVersion="8" lastPageNumber="309" masterDocId="1A4F7E45FFFC3869FFE7B23C3B0DE566" masterDocTitle="Contribution to the knowledge of Carabus (Lipaster) stjernvalli Mannerheim, 1830 in the Caucasus, with description of two new subspecies (Coleoptera: Carabidae: Carabini)" masterLastPageNumber="328" masterPageNumber="301" pageNumber="309" updateTime="1698779826648" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F9616E2461345782D48E7D35693DA698">Contribution to the knowledge of Carabus (Lipaster) stjernvalli Mannerheim, 1830 in the Caucasus, with description of two new subspecies (Coleoptera: Carabidae: Carabini)</mods:title>
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<mods:namePart id="640637037309196170F0D45BA18429D6">Fominykh, Dmitriy D.</mods:namePart>
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<mods:namePart id="084D14687189B2A9F76D6C5B9A6C64B4">Zamotajlov, Alexandr S.</mods:namePart>
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<mods:namePart id="C4E13883F11FE41C5F23B16FE641F317">Khomitskiy, Eugeniy E.</mods:namePart>
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<mods:namePart id="B66923D24B620908B85C57E2868F2768">Titarenko, Andrey Yu.</mods:namePart>
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<mods:title id="6B22F9AECD3E7B25BFC027C092E38A9F">Zootaxa</mods:title>
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<mods:date id="82E2D814D220380A0720457CA7157238">2020</mods:date>
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<treatment id="E676063DFFF43861FF70B2A63E74E6AD" ID-DOI="http://doi.org/10.5281/zenodo.5658542" ID-GBIF-Taxon="161926261" ID-Zenodo-Dep="5658542" LSID="urn:lsid:plazi:treatment:E676063DFFF43861FF70B2A63E74E6AD" httpUri="http://treatment.plazi.org/id/E676063DFFF43861FF70B2A63E74E6AD" lastPageNumber="309" pageId="8" pageNumber="309">
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<paragraph id="6E60B72BFFF43861FF70B2A63F2DE5D2" blockId="8.[151,1056,153,216]" box="[151,1056,153,180]" pageId="8" pageNumber="309">
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<heading id="35280047FFF43861FF70B2A63F2DE5D2" bold="true" box="[151,1056,153,180]" fontSize="11" level="1" pageId="8" pageNumber="309" reason="1">
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<emphasis id="5CAB6B39FFF43861FF70B2A63F2DE5D2" bold="true" box="[151,1056,153,180]" pageId="8" pageNumber="309">
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5.
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<taxonomicName id="A9DFCCA8FFF43861FF52B2A5388EE5D5" ID-CoL="8M7TX" authority="Zamotajlov, 1989" authorityName="Zamotajlov" authorityYear="1989" box="[181,899,153,180]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="subSpecies" species="stjernvalli" subGenus="Lipaster" subSpecies="bogatshevi">
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<emphasis id="5CAB6B39FFF43861FF52B2A539AFE5D2" bold="true" box="[181,674,153,180]" italics="true" pageId="8" pageNumber="309">Carabus (Lipaster) stjernvalli bogatshevi</emphasis>
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<bibRefCitation id="0A4ECADAFFF43861FD4EB2A5388EE5D5" author="Zamotajlov, A. S." box="[681,899,153,179]" pageId="8" pageNumber="309" pagination="19 - 24" refId="ref10406" refString="Zamotajlov, A. S. (1989) On new carabids of the genus Carabus L. (Coleoptera, Carabidae) from the Caucasus. Entomologica Basiliensia, 13, 19 - 24." type="journal article" year="1989">Zamotajlov, 1989</bibRefCitation>
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</taxonomicName>
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, stat. resurr.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="26C5E4A0FFF43861FF70B2813A2DE5BE" box="[151,288,189,216]" pageId="8" pageNumber="309" type="description">
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<paragraph id="6E60B72BFFF43861FF70B2813A2DE5BE" blockId="8.[151,1056,153,216]" box="[151,288,189,216]" pageId="8" pageNumber="309">
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(
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<figureCitation id="F6E4ABAEFFF43861FF78B2813BE0E5B1" box="[159,237,189,215]" captionStart="FIGURE 1" captionStartId="2.[151,250,1031,1056]" captionTargetBox="[416,1172,414,1002]" captionTargetId="figure@2.[416,1172,414,1002]" captionTargetPageId="2" captionText="FIGURE 1. Collecting sites of Carabus (Lipaster) stjernvalli Mannerheim in the South Caucasus: red square—C. stjernvalli ssp., Gorelovka env.; green square—C. stjernvalli paravanicus subsp. nov.; pink rhombs—C. stjernvalli tabackuriensis Novotný & Vořišek; red rhombus—C. stjernvalli tabackuriensis Novotný & Vořišek, type locality; red triangles—C. stjernvalli kartalinicus subsp. nov.; blue circle—C. stjernvalli bogatshevi Zamotajlov, type locality; violet empty squares—C. stjernvalli stjernvalli Mannerheim; red crosses—C. stjernvalli gvalijai Retezár." figureDoi="http://doi.org/10.5281/zenodo.3632664" httpUri="https://zenodo.org/record/3632664/files/figure.png" pageId="8" pageNumber="309">Figs. 1</figureCitation>
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,
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<figureCitation id="F6E4ABAEFFF43861FF1CB2813A15E5BE" box="[251,280,189,216]" captionStart="FIGURES 7–15" captionStartId="5.[151,264,1784,1809]" captionTargetBox="[160,1430,502,1751]" captionTargetId="figure@5.[151,1436,489,1759]" captionTargetPageId="5" captionText="FIGURES 7–15. Dorsal habitus of Carabus (Lipaster) stjernvalli Mannerheim: 7–10—C. stjernvalli tabackuriensis Novotný & Vořišek, Tsalka distr., 1700 m; 11–14—idem, Tukhmatash Pass env., 2100 m; 15—C. stjernvalli bogatshevi Zamotajlov, holotype, Chareli Mt., 2100 m. Scale bar 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3632670" httpUri="https://zenodo.org/record/3632670/files/figure.png" pageId="8" pageNumber="309">15</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="26C5E4A0FFF43861FF70B3393F9BE425" pageId="8" pageNumber="309" type="materials_examined">
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<paragraph id="6E60B72BFFF43861FF70B3393F9BE425" blockId="8.[151,1437,261,972]" pageId="8" pageNumber="309">
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<emphasis id="5CAB6B39FFF43861FF70B3393A8DE446" bold="true" box="[151,384,261,288]" pageId="8" pageNumber="309">Material examined.</emphasis>
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<materialsCitation id="DEB7BD76FFF43861FE61B3393AFEE425" ID-GBIF-Occurrence="2561484933" collectingDate="1929-07-30" collectionCode="ZISP" collectorName="A. Bogacev" county="Borzhomi" elevation="2100" location="Transcaucasus" municipality="Chareli" pageId="8" pageNumber="309" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="B1640989FFF43861FE61B3393AFCE479" box="[390,497,261,287]" pageId="8" pageNumber="309" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="78D97CA2FFF43861FE1BB33A391EE446" box="[508,531,262,288]" pageId="8" pageNumber="309" type="male">♂</specimenCount>
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, labelled “
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<location id="6B00E1F0FFF43861FD61B3393823E479" LSID="urn:lsid:plazi:treatment:E676063DFFF43861FF70B2A63E74E6AD:6B00E1F0FFF43861FD61B3393823E479" box="[646,814,261,287]" county="Borzhomi" municipality="Chareli" name="Transcaucasus" pageId="8" pageNumber="309">Transcaucasus</location>
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,
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<location id="6B00E1F0FFF43861FCDEB33A38B0E479" LSID="urn:lsid:plazi:treatment:E676063DFFF43861FF70B2A63E74E6AD:6B00E1F0FFF43861FCDEB33A38B0E479" box="[825,957,261,288]" county="Borzhomi" municipality="Chareli" name="mt. Tshareli" pageId="8" pageNumber="309">mt. Tshareli</location>
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(=
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<collectingMunicipality id="8E042D51FFF43861FC3DB3393F3DE446" box="[986,1072,261,288]" pageId="8" pageNumber="309">Chareli</collectingMunicipality>
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) near
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<collectingCounty id="8701CFA7FFF43861FB92B3393FE5E479" box="[1141,1256,261,287]" pageId="8" pageNumber="309">Borzhomi</collectingCounty>
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,
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<quantity id="A9271ACEFFF43861FB15B3393E45E446" box="[1266,1352,261,288]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="309" unit="m" value="2100.0">
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<elevation id="E5F25018FFF43861FB15B3393E45E446" box="[1266,1352,261,288]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="309" unit="m" value="2100.0">2100 m</elevation>
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</quantity>
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,
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<date id="1A6191EBFFF43861FAB3B3393BDFE425" pageId="8" pageNumber="309" value="1929-07-30">
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<collectingDate id="0A256803FFF43861FAB3B3393BDFE425" pageId="8" pageNumber="309" value="1929-07-30">30 VII 1929</collectingDate>
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</date>
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,
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<collectorName id="C32AD2FDFFF43861FF3BB3163A6DE422" box="[220,352,298,324]" pageId="8" pageNumber="309">A. Bogačev</collectorName>
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leg.” (
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<collectionCode id="08CE2FEEFFF43861FE4BB3153AE6E425" box="[428,491,297,323]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="8" pageNumber="309">ZISP</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="DEB7BD76FFF43861FE19B3163F9FE425" ID-GBIF-Occurrence="2561484948" box="[510,1170,297,324]" collectionCode="ZISP, CZAM" pageId="8" pageNumber="309" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
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<typeStatus id="B1640989FFF43861FE19B3163963E422" box="[510,622,298,324]" pageId="8" pageNumber="309" type="paratype">Paratypes</typeStatus>
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,
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<specimenCount id="78D97CA2FFF43861FD9EB31539BEE425" box="[633,691,297,323]" pageId="8" pageNumber="309" type="male">2♂♂</specimenCount>
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, labelled same as
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<typeStatus id="B1640989FFF43861FC9AB31538D2E425" box="[893,991,297,323]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="08CE2FEEFFF43861FC09B3153F24E425" box="[1006,1065,297,323]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="8" pageNumber="309">ZISP</collectionCode>
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,
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<collectionCode id="08CE2FEEFFF43861FBD3B3153F87E425" box="[1076,1162,297,323]" pageId="8" pageNumber="309">CZAM</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="26C5E4A0FFF43861FF20B3723E74E6AD" pageId="8" pageNumber="309" type="discussion">
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<paragraph id="6E60B72BFFF43861FF20B3723F40E49E" blockId="8.[151,1437,261,972]" pageId="8" pageNumber="309">
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<emphasis id="5CAB6B39FFF43861FF20B3723A36E40E" bold="true" box="[199,315,334,360]" pageId="8" pageNumber="309">Remarks.</emphasis>
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Described from
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<specimenCount id="78D97CA2FFF43861FE18B3713956E401" box="[511,603,333,359]" pageId="8" pageNumber="309" type="male">10 male</specimenCount>
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and
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<specimenCount id="78D97CA2FFF43861FD70B3713805E401" box="[663,776,333,359]" pageId="8" pageNumber="309" type="female">17 female</specimenCount>
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specimens, collected by A.V. Bogačev in 1929, and later synonymized with
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<taxonomicName id="A9DFCCA8FFF43861FE97B34E39BDE4ED" authorityName="Novotny & Vorisek, Tsalka" authorityYear="1988" box="[368,688,369,396]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="subSpecies" species="stjernvalli" subSpecies="tabackuriensis">
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<emphasis id="5CAB6B39FFF43861FE97B34E39BDE4ED" box="[368,688,369,396]" italics="true" pageId="8" pageNumber="309">C. stjernvalli tabackuriensis</emphasis>
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</taxonomicName>
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, known from the close locality (
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<figureCitation id="F6E4ABAEFFF43861FBCFB34D3F61E4ED" box="[1064,1132,369,395]" captionStart="FIGURE 1" captionStartId="2.[151,250,1031,1056]" captionTargetBox="[416,1172,414,1002]" captionTargetId="figure@2.[416,1172,414,1002]" captionTargetPageId="2" captionText="FIGURE 1. Collecting sites of Carabus (Lipaster) stjernvalli Mannerheim in the South Caucasus: red square—C. stjernvalli ssp., Gorelovka env.; green square—C. stjernvalli paravanicus subsp. nov.; pink rhombs—C. stjernvalli tabackuriensis Novotný & Vořišek; red rhombus—C. stjernvalli tabackuriensis Novotný & Vořišek, type locality; red triangles—C. stjernvalli kartalinicus subsp. nov.; blue circle—C. stjernvalli bogatshevi Zamotajlov, type locality; violet empty squares—C. stjernvalli stjernvalli Mannerheim; red crosses—C. stjernvalli gvalijai Retezár." figureDoi="http://doi.org/10.5281/zenodo.3632664" httpUri="https://zenodo.org/record/3632664/files/figure.png" pageId="8" pageNumber="309">Fig. 1</figureCitation>
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, red rhombus). Due to the courtesy of Dr. B. Kataev (ZISP), we had the opportunity to reexamine and measure several specimens of the type series, including the
|
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<typeStatus id="B1640989FFF43861FE99B3853AEEE4B5" box="[382,483,441,467]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
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, and found sufficient differences of these individuals from
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<taxonomicName id="A9DFCCA8FFF43861FB92B3863BB8E491" authorityName="Novotny & Vorisek, Tsalka" authorityYear="1988" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="subSpecies" species="stjernvalli" subSpecies="tabackuriensis">
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<emphasis id="5CAB6B39FFF43861FB92B3863BB8E491" italics="true" pageId="8" pageNumber="309">C. stjernvalli tabackuriensis</emphasis>
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</taxonomicName>
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(
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<figureCitation id="F6E4ABAEFFF43861FF23B3E13A29E49E" box="[196,292,477,504]" captionStart="FIGURE 116" captionStartId="20.[151,250,1905,1930]" captionTargetBox="[289,1301,1133,1873]" captionTargetId="figure@20.[279,1308,1127,1880]" captionTargetPageId="20" captionText="FIGURE 116. Distribution of morphometric characters in some Carabus (Lipaster) stjernvalli Mannerheim subspecies constructed using discriminant analysis based on 8 parameters (see text for measurements details): green squares—C. stjernvalli paravanicus subsp. nov.; pink rhombs—C. stjernvalli tabackuriensis Novotný & Vořišek; red triangles—C. stjernvalli kartalinicus subsp. nov.; blue circles—C. stjernvalli bogatshevi Zamotajlov." figureDoi="http://doi.org/10.5281/zenodo.3632700" httpUri="https://zenodo.org/record/3632700/files/figure.png" pageId="8" pageNumber="309">Fig. 116</figureCitation>
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), so we resurrect here the subspecific status of
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<taxonomicName id="A9DFCCA8FFF43861FCD3B3E23F4BE491" box="[820,1094,477,504]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="subSpecies" species="stjernvalli" subSpecies="bogatshevi">
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<emphasis id="5CAB6B39FFF43861FCD3B3E23F4BE491" box="[820,1094,477,504]" italics="true" pageId="8" pageNumber="309">C. stjernvalli bogatshevi</emphasis>
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</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6E60B72BFFF43861FF20B03D38F8E67E" blockId="8.[151,1437,261,972]" pageId="8" pageNumber="309">
|
||
The range of this form is known to occupy a small territory located west-southwards of
|
||
<taxonomicName id="A9DFCCA8FFF43861FB4AB03E3BFDE759" authorityName="Novotny & Vorisek, Tsalka" authorityYear="1988" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="subSpecies" species="stjernvalli" subSpecies="tabackuriensis">
|
||
<emphasis id="5CAB6B39FFF43861FB4AB03E3BFDE759" italics="true" pageId="8" pageNumber="309">C. stjernvalli tabackuriensis</emphasis>
|
||
</taxonomicName>
|
||
’ range, with both subspecies distributed on neighbouring mountains with no visible substantial spatial barriers. This unexpected fact of speciation probably reflects the peripheral position of
|
||
<taxonomicName id="A9DFCCA8FFF43861FB9AB0763E9EE705" box="[1149,1427,585,612]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="subSpecies" species="stjernvalli" subSpecies="bogatshevi">
|
||
<emphasis id="5CAB6B39FFF43861FB9AB0763E9EE705" box="[1149,1427,585,612]" italics="true" pageId="8" pageNumber="309">C. stjernvalli bogatshevi</emphasis>
|
||
</taxonomicName>
|
||
’ geographic range with respect to the large range of
|
||
<taxonomicName id="A9DFCCA8FFF43861FD36B0523F01E7E1" authorityName="Novotny & Vorisek, Tsalka" authorityYear="1988" box="[721,1036,621,648]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="subSpecies" species="stjernvalli" subSpecies="tabackuriensis">
|
||
<emphasis id="5CAB6B39FFF43861FD36B0523F01E7E1" box="[721,1036,621,648]" italics="true" pageId="8" pageNumber="309">C. stjernvalli tabackuriensis</emphasis>
|
||
</taxonomicName>
|
||
’, occupying the vast territory of the Lesser Caucasus, similar in this respect to the small “marginal” ranges observed in both
|
||
<taxonomicName id="A9DFCCA8FFF43861FB9FB0AE3E91E7CD" authorityName="Fominykh & Zamotajlov & Khomitskiy & Titarenko" authorityYear="2020" box="[1144,1436,657,684]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="subSpecies" species="stjernvalli" subSpecies="paravanicus">
|
||
<emphasis id="5CAB6B39FFF43861FB9FB0AE3E91E7CD" box="[1144,1436,657,684]" italics="true" pageId="8" pageNumber="309">C. stjernvalli paravanicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5CAB6B39FFF43861FF70B08A3A11E7B6" bold="true" box="[151,284,694,720]" pageId="8" pageNumber="309">subsp. nov.</emphasis>
|
||
(southernmost extremity of the species’ range in
|
||
<collectingCountry id="16C8F7BBFFF43861FCB6B08938A0E7B6" box="[849,941,693,720]" name="Georgia" pageId="8" pageNumber="309">Georgia</collectingCountry>
|
||
) and
|
||
<taxonomicName id="A9DFCCA8FFF43861FC0AB08A3E02E7A9" authorityName="Fominykh & Zamotajlov & Khomitskiy & Titarenko" authorityYear="2020" box="[1005,1295,693,720]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="subSpecies" species="stjernvalli" subSpecies="kartalinicus">
|
||
<emphasis id="5CAB6B39FFF43861FC0AB08A3E02E7A9" box="[1005,1295,693,720]" italics="true" pageId="8" pageNumber="309">C. stjernvalli kartalinicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5CAB6B39FFF43861FAF0B08A3E91E7B6" bold="true" box="[1303,1436,694,720]" pageId="8" pageNumber="309">subsp. nov.</emphasis>
|
||
(northern-easternmost extremity of the species’ range in
|
||
<collectingCountry id="16C8F7BBFFF43861FCF7B0E53863E792" box="[784,878,729,756]" name="Georgia" pageId="8" pageNumber="309">Georgia</collectingCountry>
|
||
), which also inhabits the Trialeti Mountain Range and its nearest surroundings in the Lesser Caucasus (see Discussion section).
|
||
</paragraph>
|
||
<paragraph id="6E60B72BFFF43861FF20B11E3E74E6AD" blockId="8.[151,1437,261,972]" pageId="8" pageNumber="309">
|
||
<taxonomicName id="A9DFCCA8FFF43861FF20B11E3AD3E65D" box="[199,478,801,828]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="subSpecies" species="stjernvalli" subSpecies="bogatshevi">
|
||
<emphasis id="5CAB6B39FFF43861FF20B11E3AD3E65D" box="[199,478,801,828]" italics="true" pageId="8" pageNumber="309">C. stjernvalli bogatshevi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="A9DFCCA8FFF43861FD9FB11E38B4E65D" authorityName="Novotny & Vorisek, Tsalka" authorityYear="1988" box="[632,953,801,828]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="subSpecies" species="stjernvalli" subSpecies="tabackuriensis">
|
||
<emphasis id="5CAB6B39FFF43861FD9FB11E38B4E65D" box="[632,953,801,828]" italics="true" pageId="8" pageNumber="309">C. stjernvalli tabackuriensis</emphasis>
|
||
</taxonomicName>
|
||
in generally somewhat more robust body, more transverse pronotum with less rounded sides and broader explanate margin distinctly widened basally; frons with less prominent, often somewhat smoothed sculpture; elytra less ovate and more parallel-sided. It is distinguishable from another closely distributed subspecies,
|
||
<taxonomicName id="A9DFCCA8FFF43861FD23B1B238E4E6C1" authorityName="Fominykh & Zamotajlov & Khomitskiy & Titarenko" authorityYear="2020" box="[708,1001,909,936]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="subSpecies" species="stjernvalli" subSpecies="paravanicus">
|
||
<emphasis id="5CAB6B39FFF43861FD23B1B238E4E6C1" box="[708,1001,909,936]" italics="true" pageId="8" pageNumber="309">C. stjernvalli paravanicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5CAB6B39FFF43861FC16B1B23F78E6CE" bold="true" box="[1009,1141,910,936]" pageId="8" pageNumber="309">subsp. nov.</emphasis>
|
||
, in many respects, first of all, in the more transverse pronotum with more emarginate apex and hind angles stronger protruded backwards.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |