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<document id="A4A651561CD6DC0257C38604BD498BEB" ID-DOI="10.1649/0010-065X-75.4.723" ID-ISSN="1938-4394" ID-Zenodo-Dep="13252037" ID-ZooBank="urn:lsid:zoobank.org:pub:9766A693-00DD-41CF-A7D3-5715CF0C03FC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723001874209" checkinUser="felipe" docAuthor="Barney, Robert J." docDate="2021" docId="C436412BFFE5FF8742CFFA80FE3AFE0E" docLanguage="en" docName="TheColeopBull.75.4.723-748.pdf" docOrigin="The Coleopterists Bulletin 75 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-75.4.723" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Pachybrachis ephedra Barney 2021, new species" docType="treatment" docUuid="09221B9C-7B99-46B2-B6C6-983A0451B38B" docUuidSource="ZooBank" docVersion="2" lastPageNumber="739" masterDocId="380F3953FFEAFF964272FFF5FF9EFF9D" masterDocTitle="Definition and Revision of the cylindricus Species-Group of North American Pachybrachis Chevrolat (Coleoptera: Chrysomelidae: Cryptocephalinae), Including Descriptions of Six New Species" masterLastPageNumber="748" masterPageNumber="723" pageNumber="737" updateTime="1723004090703" updateUser="ExternalLinkService">
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<mods:title id="166C8B58F00CE2862CAD8934A1B0D8A4">Definition and Revision of the cylindricus Species-Group of North American Pachybrachis Chevrolat (Coleoptera: Chrysomelidae: Cryptocephalinae), Including Descriptions of Six New Species</mods:title>
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<mods:namePart id="39DBF43C1A483331ACF483D1B63FDE69">Barney, Robert J.</mods:namePart>
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<mods:title id="3D668078C97288A475A2C1E3640C1A4D">The Coleopterists Bulletin</mods:title>
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<mods:date id="B3B100AF9E85FCC87090FFDE77A97EB0">2021</mods:date>
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<mods:number id="381401E59CDD839C812CC91E81B36C68">2021-12-31</mods:number>
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<mods:number id="43E5E707A1C81A182C482B70306FD094">75</mods:number>
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<mods:classification id="12C74A5C071947B73F076694C3B8CD86">journal article</mods:classification>
<mods:identifier id="6B5D28527E5B6AAF2C4EBCB277DD183C" type="DOI">10.1649/0010-065X-75.4.723</mods:identifier>
<mods:identifier id="C9312DD51C59D33253B787DE8F429EBF" type="ISSN">1938-4394</mods:identifier>
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</mods:mods>
<treatment id="C436412BFFE5FF8742CFFA80FE3AFE0E" LSID="urn:lsid:zoobank.org:act:09221B9C-7B99-46B2-B6C6-983A0451B38B" httpUri="http://treatment.plazi.org/id/C436412BFFE5FF8742CFFA80FE3AFE0E" lastPageId="17" lastPageNumber="739" pageId="15" pageNumber="737">
<subSubSection id="0485A3B6FFE5FF9942CFFA80FE7CFA47" pageId="15" pageNumber="737" type="nomenclature">
<paragraph id="4C20F03DFFE5FF9942CFFA80FE7CFA47" blockId="15.[189,643,1397,1499]" pageId="15" pageNumber="737">
<emphasis id="7EEB2C2FFFE5FF9942CFFA80FD1DFA16" bold="true" box="[189,643,1397,1419]" pageId="15" pageNumber="737">
8.
<taxonomicName id="8B9F8BBEFFE5FF9942A7FA80FD9DFA17" authority="Barney" authorityName="Barney" authorityYear="2021" box="[213,515,1397,1418]" class="Insecta" family="Chrysomelidae" genus="Pachybrachis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="723" phylum="Arthropoda" rank="species" species="ephedra" status="sp. nov.">
<emphasis id="7EEB2C2FFFE5FF9942A7FA80FE2CFA17" bold="true" box="[213,434,1397,1418]" italics="true" pageId="15" pageNumber="737">PACHYBRACHIS epHeDRA</emphasis>
Barney
</taxonomicName>
,
<taxonomicNameLabel id="65D89154FFE5FF99407FFA80FD1DFA16" box="[525,643,1397,1419]" pageId="15" pageNumber="737" rank="species">new species</taxonomicNameLabel>
</emphasis>
zoobank.org/
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urn:lsid:zoobank.org:act:
<uuid id="3839CAE8FFE5FF9942CFFA5FFD1CFA5D" box="[189,642,1450,1472]" pageId="15" pageNumber="737">09221B9C-7B99-46B2-B6C6-983A0451B38B</uuid>
</uri>
(
<figureCitation id="D4A4ECB8FFE5FF994316FA30FE36FA47" box="[356,424,1477,1499]" captionStart="Fig" captionStartId="16.[176,207,1594,1613]" captionTargetBox="[160,1165,202,1565]" captionTargetId="figure-6@16.[154,1168,193,1567]" captionTargetPageId="16" captionText="Fig. 8. Pachybrachis ephedra, new species, Washington Co., Utah, holotype.A) Dorsal habitus, B) Lateral habitus, C) Face, D) Pygidium, E) Median lobe of aedeagus, lateral view, F) Median lobe of aedeagus, en-face view, G) Median lobe of aedeagus, sub view." figureDoi="http://doi.org/10.5281/zenodo.13252053" httpUri="https://zenodo.org/record/13252053/files/figure.png" pageId="15" pageNumber="737">Figs. 8</figureCitation>
,
<figureCitation id="D4A4ECB8FFE5FF9943C1FA30FE45FA47" box="[435,475,1477,1498]" captionStart="Fig" captionStartId="25.[197,228,1297,1316]" captionTargetBox="[171,1195,193,1270]" captionTargetId="figure-207@25.[171,1195,193,1270]" captionTargetPageId="25" captionText="Fig. 14. Distribution of Pachybrachis species, based only upon specimens examined in this study. A) P. hybridus (black), P. turgidicollis (red), P. texasensis, new species (green), P. chaoticus (blue), P. pinguescens (orange), P. crassus (purple), B) P. saguaro, new species (black), P. peninsularis (purple), P. cylindricus (red), P. huachuca, new species (orange), P. hurricane, new species (gray), P. ephedra, new species (blue), and P. ashfork, new species (green)." figureDoi="http://doi.org/10.5281/zenodo.13252067" httpUri="https://zenodo.org/record/13252067/files/figure.png" pageId="15" pageNumber="737">14B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0485A3B6FFE5FF9942B7FA07FBFAFEBA" pageId="15" pageNumber="737" type="materials_examined">
<paragraph id="4C20F03DFFE5FF9942B7FA07FDF2F95E" blockId="15.[173,660,1522,1731]" pageId="15" pageNumber="737">
<materialsCitation id="FCF7FA60FFE5FF9942B7FA07FDF2F95E" collectingDate="2015-05-17" collectionCode="BYUC" collectorName="S. M. Clark &amp; E. G. Riley" latitude="37.1442" location="Male" longLatPrecision="6" longitude="-113.254" pageId="15" pageNumber="737" specimenCount="2" stateProvince="Utah" typeStatus="holotype">
<emphasis id="7EEB2C2FFFE5FF9942B7FA07FEB6F99A" bold="true" box="[197,296,1522,1543]" pageId="15" pageNumber="737">
<typeStatus id="93244E9FFFE5FF9942B7FA07FEBAF99A" box="[197,292,1522,1543]" pageId="15" pageNumber="737" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="4940A6E6FFE5FF994347FA07FEF8F995" LSID="urn:lsid:plazi:treatment:C436412BFFE5FF8742CFFA80FE3AFE0E:4940A6E6FFE5FF994347FA07FEF8F995" box="[309,358,1522,1544]" latitude="37.1442" longLatPrecision="6" longitude="-113.254" name="Male" pageId="15" pageNumber="737" stateProvince="Utah">Male</location>
labeled “
<collectingRegion id="8E5B3EDFFFE5FF9943BEFA07FD8DF99A" box="[460,531,1522,1543]" country="United States of America" name="Utah" pageId="15" pageNumber="737">UTAH</collectingRegion>
:
<collectingRegion id="8E5B3EDFFFE5FF994053FA07FD0DF995" box="[545,659,1522,1544]" country="United States of America" name="Washington" pageId="15" pageNumber="737">Washington</collectingRegion>
Co./
<locationDeviation id="5D92E91FFFE5FF9942A7F9F8FE2CF9BE" box="[213,434,1549,1571]" location="Hurricane" pageId="15" pageNumber="737">
<quantity id="8B675DD8FFE5FF9942A7F9F8FE80F9BE" box="[213,286,1549,1571]" metricMagnitude="3" metricUnit="m" metricValue="4.7" pageId="15" pageNumber="737" unit="km" value="4.7">4.7 km</quantity>
SE Hurricane
</locationDeviation>
/
<geoCoordinate id="29AB96FAFFE5FF9943C8F9F8FDBFF9BE" box="[442,545,1549,1571]" degrees="37.1442" direction="north" orientation="latitude" pageId="15" pageNumber="737" precision="5" value="37.1442">37.1442°N</geoCoordinate>
,
<geoCoordinate id="29AB96FAFFE5FF994042F9F8FF58F9A0" degrees="113.2540" direction="west" orientation="longitude" pageId="15" pageNumber="737" precision="5" value="-113.254">113.2540° W</geoCoordinate>
/
<date id="3821D6FDFFE5FF9942BCF9D2FEB2F9A0" box="[206,300,1575,1597]" pageId="15" pageNumber="737" value="2015-05-17">
<collectingDate id="28652F15FFE5FF9942BCF9D2FEB2F9A0" box="[206,300,1575,1597]" pageId="15" pageNumber="737" value="2015-05-17">17.v.2015</collectingDate>
</date>
,
<collectorName id="E16A95EBFFE5FF994344F9DDFE37F9A3" box="[310,425,1576,1598]" pageId="15" pageNumber="737">S. M. Clark</collectorName>
&amp;
<collectorName id="E16A95EBFFE5FF9943B9F9DDFDA7F9A3" box="[459,569,1576,1598]" pageId="15" pageNumber="737">E. G. Riley</collectorName>
[printed, white paper]//
<typeStatus id="93244E9FFFE5FF994341F9B7FE2DF9CA" box="[307,435,1602,1623]" pageId="15" pageNumber="737" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="8B9F8BBEFFE5FF9943CBF9B7FDA3F9CA" box="[441,573,1602,1623]" class="Insecta" family="Chrysomelidae" genus="Pachybrachis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="737" phylum="Arthropoda" rank="genus">
<emphasis id="7EEB2C2FFFE5FF9943CBF9B7FDA3F9CA" box="[441,573,1602,1623]" italics="true" pageId="15" pageNumber="737">Pachybrachis</emphasis>
</taxonomicName>
/
<taxonomicName id="8B9F8BBEFFE5FF994036F9B7FF6AF9EF" authority="Barney" authorityName="Barney" authorityYear="2021" class="Insecta" family="Chrysomelidae" genus="Pachybrachis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="737" phylum="Arthropoda" rank="species" species="ephedra">
<emphasis id="7EEB2C2FFFE5FF994036F9B7FD0DF9CA" box="[580,659,1602,1623]" italics="true" pageId="15" pageNumber="737">ephedra</emphasis>
Barney
</taxonomicName>
/R. J.
<bibRefCitation id="280E8DCCFFE5FF994346F9A8FE2AF9EF" author="Barney, R. J." box="[308,436,1629,1650]" pageId="15" pageNumber="737" pagination="485 - 496" refId="ref15382" refString="Barney, R. J. 2021. Redescription of five species of Pachybrachis Chevrolat (Coleoptera: Chrysomelidae: Cryptocephalinae) of the western USA, including new synonymies, records of greatly expanded ranges into the eastern USA, and new plant associations. The Coleopterists Bulletin 75 (2): 485 - 496." type="journal article" year="2021">Barney 2021</bibRefCitation>
[printed, red paper]”. Deposited in
<collectionCode id="2A8E68F8FFE5FF994355F98DFEF4F910" box="[295,362,1656,1677]" pageId="15" pageNumber="737">BYUC</collectionCode>
. The specimen is point-mounted with the genitalia affixed to a second point. It is in excellent condition with all appendages intact.
</materialsCitation>
</paragraph>
<paragraph id="4C20F03DFFE5FF9940A9FF37FBFAFEBA" blockId="15.[707,1193,194,1731]" pageId="15" pageNumber="737">
<emphasis id="7EEB2C2FFFE5FF9940A9FF37FCD9FF4A" bold="true" box="[731,839,194,215]" pageId="15" pageNumber="737">
<typeStatus id="93244E9FFFE5FF9940A9FF37FCDDFF4A" box="[731,835,194,215]" pageId="15" pageNumber="737" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="5A993BB4FFE5FF99413FFF37FC71FF45" box="[845,1007,194,216]" count="3" pageId="15" pageNumber="737" type="generic">Three specimens</specimenCount>
(
<specimenCount id="5A993BB4FFE5FF99418EFF37FBB2FF4A" box="[1020,1068,194,215]" count="2" pageId="15" pageNumber="737" type="male">2♂♂</specimenCount>
<specimenCount id="5A993BB4FFE5FF994640FF37FBD0FF4A" box="[1074,1102,194,215]" count="1" pageId="15" pageNumber="737" type="female">1♀</specimenCount>
), including two (
<specimenCount id="5A993BB4FFE5FF99416FFF28FCA4FF6F" box="[797,826,221,242]" count="1" pageId="15" pageNumber="737" type="male">1♂</specimenCount>
<specimenCount id="5A993BB4FFE5FF994130FF28FCC0FF6F" box="[834,862,221,242]" count="1" pageId="15" pageNumber="737" type="female">1♀</specimenCount>
) from the
<typeStatus id="93244E9FFFE5FF9941B8FF28FB80FF6E" box="[970,1054,221,243]" pageId="15" pageNumber="737" type="holotype">holotype</typeStatus>
series, are labeled “
<typeStatus id="93244E9FFFE5FF994171FF02FCE5FE91" box="[771,891,247,268]" pageId="15" pageNumber="737" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="8B9F8BBEFFE5FF9941F0FF02FB9AFE91" box="[898,1028,247,268]" class="Insecta" family="Chrysomelidae" genus="Pachybrachis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="737" phylum="Arthropoda" rank="genus">
<emphasis id="7EEB2C2FFFE5FF9941F0FF02FB9AFE91" box="[898,1028,247,268]" italics="true" pageId="15" pageNumber="737">Pachybrachis</emphasis>
</taxonomicName>
/
<taxonomicName id="8B9F8BBEFFE5FF994679FF02FB3FFE91" authority="Barney" authorityName="Barney" authorityYear="2021" box="[1035,1185,247,268]" class="Insecta" family="Chrysomelidae" genus="Pachybrachis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="737" phylum="Arthropoda" rank="species" species="ephedra">
<emphasis id="7EEB2C2FFFE5FF994679FF02FBC7FE91" box="[1035,1113,247,268]" italics="true" pageId="15" pageNumber="737">ephedra</emphasis>
Barney
</taxonomicName>
/ R. J.
<bibRefCitation id="280E8DCCFFE5FF994086FEE6FCF1FEBA" author="Barney, R. J." box="[756,879,274,296]" pageId="15" pageNumber="737" pagination="485 - 496" refId="ref15382" refString="Barney, R. J. 2021. Redescription of five species of Pachybrachis Chevrolat (Coleoptera: Chrysomelidae: Cryptocephalinae) of the western USA, including new synonymies, records of greatly expanded ranges into the eastern USA, and new plant associations. The Coleopterists Bulletin 75 (2): 485 - 496." type="journal article" year="2021">Barney 2021</bibRefCitation>
[printed, yellow paper]”.
</paragraph>
</subSubSection>
<subSubSection id="0485A3B6FFE5FF9940A9FEDAFCC4FB48" pageId="15" pageNumber="737" type="description">
<paragraph id="4C20F03DFFE5FF9940A9FEDAFCC4FB48" blockId="15.[707,1193,194,1731]" pageId="15" pageNumber="737">
<emphasis id="7EEB2C2FFFE5FF9940A9FEDAFC06FED8" bold="true" box="[731,920,303,325]" pageId="15" pageNumber="737">Description. Male.</emphasis>
Large, whitish yellow with black maculae and black pronotal M (
<figureCitation id="D4A4ECB8FFE5FF99462BFEBFFB04FEFD" box="[1113,1178,330,352]" captionStart="Fig" captionStartId="16.[176,207,1594,1613]" captionTargetBox="[160,1165,202,1565]" captionTargetId="figure-6@16.[154,1168,193,1567]" captionTargetPageId="16" captionText="Fig. 8. Pachybrachis ephedra, new species, Washington Co., Utah, holotype.A) Dorsal habitus, B) Lateral habitus, C) Face, D) Pygidium, E) Median lobe of aedeagus, lateral view, F) Median lobe of aedeagus, en-face view, G) Median lobe of aedeagus, sub view." figureDoi="http://doi.org/10.5281/zenodo.13252053" httpUri="https://zenodo.org/record/13252053/files/figure.png" pageId="15" pageNumber="737">Fig. 8</figureCitation>
); L =
<quantity id="8B675DD8FFE5FF994086FE90FC19FEE6" box="[756,903,357,379]" metricMagnitude="-3" metricUnit="m" metricValue="3.855" metricValueMax="3.94" metricValueMin="3.77" pageId="15" pageNumber="737" unit="mm" value="3.855" valueMax="3.94" valueMin="3.77">3.77–3.94 mm</quantity>
(mean =
<quantity id="8B675DD8FFE5FF994183FE90FBD1FEE6" box="[1009,1103,357,379]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" pageId="15" pageNumber="737" unit="mm" value="3.85">3.85 mm</quantity>
,
<emphasis id="7EEB2C2FFFE5FF99462CFE93FBF4FEE6" box="[1118,1130,358,379]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 3); W =
<quantity id="8B675DD8FFE5FF994089FE75FC12FE0B" box="[763,908,384,406]" metricMagnitude="-3" metricUnit="m" metricValue="2.08" metricValueMax="2.17" metricValueMin="1.99" pageId="15" pageNumber="737" unit="mm" value="2.08" valueMax="2.17" valueMin="1.99">1.99–2.17 mm</quantity>
(mean =
<quantity id="8B675DD8FFE5FF994187FE75FBCFFE0B" box="[1013,1105,384,406]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="737" unit="mm" value="2.1">2.10 mm</quantity>
,
<emphasis id="7EEB2C2FFFE5FF99462DFE74FBF5FE0B" box="[1119,1131,385,406]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 3); L/W = 1.78–1.91 (mean = 1.84,
<emphasis id="7EEB2C2FFFE5FF994188FE69FB98FE2C" box="[1018,1030,412,433]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 3).
<emphasis id="7EEB2C2FFFE5FF99464CFE6EFBE0FE2D" bold="true" box="[1086,1150,411,432]" pageId="15" pageNumber="737">Head:</emphasis>
Not wider than thoracic apex, HW =
<quantity id="8B675DD8FFE5FF994665FE43FB39FE56" box="[1047,1191,438,459]" metricMagnitude="-3" metricUnit="m" metricValue="1.3399999999999999" metricValueMax="1.38" metricValueMin="1.3" pageId="15" pageNumber="737" unit="mm" value="1.3399999999999999" valueMax="1.38" valueMin="1.3">1.30–1.38 mm</quantity>
(mean =
<quantity id="8B675DD8FFE5FF99416EFE24FCE8FE7B" box="[796,886,465,487]" metricMagnitude="-3" metricUnit="m" metricValue="1.36" pageId="15" pageNumber="737" unit="mm" value="1.36">1.36 mm</quantity>
,
<emphasis id="7EEB2C2FFFE5FF9941F3FE24FC13FE7B" box="[897,909,465,486]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 3); eyes moderately separated, without ocular lines, IOD =
<quantity id="8B675DD8FFE5FF99466BFE1EFB39FD9C" box="[1049,1191,491,513]" metricMagnitude="-4" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="3.5999999999999996" metricValueMin="3.1" pageId="15" pageNumber="737" unit="mm" value="0.33499999999999996" valueMax="0.36" valueMin="0.31">0.31–0.36 mm</quantity>
(mean =
<quantity id="8B675DD8FFE5FF99416FFDF3FCE9FD81" box="[797,887,518,540]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="15" pageNumber="737" unit="mm" value="0.33">0.33 mm</quantity>
,
<emphasis id="7EEB2C2FFFE5FF9941F1FDF2FC11FD81" box="[899,911,519,540]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 3); IOD/HW = 0.22–0.26 (mean = 0.24,
<emphasis id="7EEB2C2FFFE5FF99413DFDD7FCC5FDAA" box="[847,859,546,567]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 3); face yellow with large black macula descending from the vertex and between the upper lobes of eyes, punctation dense in dark areas, antennae progressing from yellow to brown, not reaching elytral declivity.
<emphasis id="7EEB2C2FFFE5FF994198FD79FBC2FD3C" bold="true" box="[1002,1116,652,673]" pageId="15" pageNumber="737">Pronotum:</emphasis>
Yellow with large M-shaped black macula, densely punctate, less so at margins and median line, strongly narrowed apically; PL =
<quantity id="8B675DD8FFE5FF9941CFFD28FBD3FD6F" box="[957,1101,733,754]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.37" metricValueMin="1.18" pageId="15" pageNumber="737" unit="mm" value="1.275" valueMax="1.37" valueMin="1.18">1.18–1.37 mm</quantity>
(mean =
<quantity id="8B675DD8FFE5FF9940B1FD0DFC80FC90" box="[707,798,760,781]" metricMagnitude="-3" metricUnit="m" metricValue="1.34" pageId="15" pageNumber="737" unit="mm" value="1.34">1.34 mm</quantity>
,
<emphasis id="7EEB2C2FFFE5FF994159FD0DFCA9FC90" box="[811,823,760,781]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 3); PW =
<quantity id="8B675DD8FFE5FF9941CEFD0DFBD3FC90" box="[956,1101,760,782]" metricMagnitude="-3" metricUnit="m" metricValue="1.865" metricValueMax="1.96" metricValueMin="1.77" pageId="15" pageNumber="737" unit="mm" value="1.865" valueMax="1.96" valueMin="1.77">1.77–1.96 mm</quantity>
(mean =
<quantity id="8B675DD8FFE5FF9940B1FCE6FC87FCB5" box="[707,793,787,808]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="15" pageNumber="737" unit="mm" value="1.9">1.90 mm</quantity>
,
<emphasis id="7EEB2C2FFFE5FF994153FCE6FCB3FCB5" box="[801,813,787,808]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 3); PL/PW = 0.67–0.72 (mean = 0.70,
<emphasis id="7EEB2C2FFFE5FF9940B1FCDBFD51FCDE" box="[707,719,814,835]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 3).
<emphasis id="7EEB2C2FFFE5FF994167FCD8FCC1FCDF" bold="true" box="[789,863,813,834]" pageId="15" pageNumber="737">Elytra:</emphasis>
Whitish yellow with prominent black maculae on each elytron prior to declivity, thus appearing as a trans-elytral stripe, densely punctate in baso-sutural area, striae irregular.
<emphasis id="7EEB2C2FFFE5FF9940B1FC6DFCB1FC33" bold="true" box="[707,815,920,942]" pageId="15" pageNumber="737">Pygidium:</emphasis>
Black with 2 oval conjoined yellow spots.
<emphasis id="7EEB2C2FFFE5FF994171FC46FCCEFC55" bold="true" box="[771,848,947,968]" pageId="15" pageNumber="737">Venter:</emphasis>
Black with yellow side and apical margins.
<emphasis id="7EEB2C2FFFE5FF994150FC3BFCC5FC7E" bold="true" box="[802,859,974,995]" pageId="15" pageNumber="737">Legs:</emphasis>
Yellow with brown/black spots, front claws of male greatly enlarged.
<emphasis id="7EEB2C2FFFE5FF99464DFC1CFB39FC62" bold="true" box="[1087,1191,1001,1023]" pageId="15" pageNumber="737">Genitalia:</emphasis>
Median lobe in en-face view shows a large, oval OS, with a wide projecting terminus, bounded on each side with wisps at the ALM. The lateral view reveals a large, darkened, sharp-edged keel, projecting from near the base to the T. A sub view of the median lobe shows that the keel ridge is not grooved or split.
<specimenCount id="5A993BB4FFE5FF994114FB50FC45FB26" box="[870,987,1189,1211]" count="3" pageId="15" pageNumber="737" type="male">Three males</specimenCount>
from two
<collectingCountry id="3488B0ADFFE5FF99464CFB50FBF3FB27" box="[1086,1133,1189,1210]" name="United States of America" pageId="15" pageNumber="737">USA</collectingCountry>
states were dissected.
</paragraph>
</subSubSection>
<subSubSection id="0485A3B6FFE5FF9940A9FB28FBB0FA90" pageId="15" pageNumber="737" type="etymology">
<paragraph id="4C20F03DFFE5FF9940A9FB28FBB0FA90" blockId="15.[707,1193,194,1731]" pageId="15" pageNumber="737">
<emphasis id="7EEB2C2FFFE5FF9940A9FB28FCD0FB6F" bold="true" box="[731,846,1245,1266]" pageId="15" pageNumber="737">Etymology.</emphasis>
This species is named for the presumed host plant,
<emphasis id="7EEB2C2FFFE5FF994103FB0DFBB6FA93" box="[881,1064,1272,1294]" italics="true" pageId="15" pageNumber="737">Ephedra torreyana</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="0485A3B6FFE5FF9940A9FAE0FB04F9CF" pageId="15" pageNumber="737" type="discussion">
<paragraph id="4C20F03DFFE5FF9940A9FAE0FB04F9CF" blockId="15.[707,1193,194,1731]" pageId="15" pageNumber="737">
<emphasis id="7EEB2C2FFFE5FF9940A9FAE0FCA0FAB7" bold="true" box="[731,830,1301,1322]" pageId="15" pageNumber="737">Remarks.</emphasis>
The
<taxonomicName id="8B9F8BBEFFE5FF994100FAE3FC44FAB7" authorityName="Barney" authorityYear="2021" box="[882,986,1301,1323]" class="Insecta" family="Chrysomelidae" genus="Pachybrachis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="737" phylum="Arthropoda" rank="species" species="ephedra">
<emphasis id="7EEB2C2FFFE5FF994100FAE3FC44FAB7" box="[882,986,1301,1323]" italics="true" pageId="15" pageNumber="737">P. ephedra</emphasis>
</taxonomicName>
specimens were part of a series of four (
<specimenCount id="5A993BB4FFE5FF994107FAC5FC38FAD8" box="[885,934,1328,1349]" count="3" pageId="15" pageNumber="737" type="male">3♂♂</specimenCount>
<specimenCount id="5A993BB4FFE5FF9941D9FAC5FC59FAD8" box="[939,967,1328,1349]" count="1" pageId="15" pageNumber="737" type="female">1♀</specimenCount>
) beetles collected from
<emphasis id="7EEB2C2FFFE5FF9940B1FAB9FCA4FAFC" box="[707,826,1356,1377]" italics="true" pageId="15" pageNumber="737">E. torreyana</emphasis>
in
<collectingRegion id="8E5B3EDFFFE5FF994124FABEFC16FAFC" box="[854,904,1355,1377]" country="United States of America" name="Utah" pageId="15" pageNumber="737">Utah</collectingRegion>
. I initially assumed these were all the same species, but dissection of the males revealed two distinct median lobes. The median lobe of
<taxonomicName id="8B9F8BBEFFE5FF9940AEFA69FCDFFA2D" authorityName="Barney" authorityYear="2021" box="[732,833,1435,1457]" class="Insecta" family="Chrysomelidae" genus="Pachybrachis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="737" phylum="Arthropoda" rank="species" species="ephedra">
<emphasis id="7EEB2C2FFFE5FF9940AEFA69FCDFFA2D" box="[732,833,1435,1457]" italics="true" pageId="15" pageNumber="737">P. ephedra</emphasis>
</taxonomicName>
has a distinct sharp-edged keel, while the median lobe of
<taxonomicName id="8B9F8BBEFFE5FF9941E3FA42FB93FA56" authorityName="Barney" authorityYear="2021" box="[913,1037,1462,1484]" class="Insecta" family="Chrysomelidae" genus="Pachybrachis" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="737" phylum="Arthropoda" rank="species" species="hurricane">
<emphasis id="7EEB2C2FFFE5FF9941E3FA42FB93FA56" box="[913,1037,1462,1484]" italics="true" pageId="15" pageNumber="737">P. hurricane</emphasis>
</taxonomicName>
is smooth and rounded. The singleton female cannot be assigned to either species at this time, but measurements are here provided: L =
<quantity id="8B675DD8FFE5FF994109F9F2FC4EF981" box="[891,976,1543,1564]" metricMagnitude="-3" metricUnit="m" metricValue="4.12" pageId="15" pageNumber="737" unit="mm" value="4.12">4.12 mm</quantity>
(
<emphasis id="7EEB2C2FFFE5FF9941ACF9F2FC74F981" box="[990,1002,1543,1564]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 1); W =
<quantity id="8B675DD8FFE5FF994621F9F2FB36F981" box="[1107,1192,1543,1564]" metricMagnitude="-3" metricUnit="m" metricValue="2.22" pageId="15" pageNumber="737" unit="mm" value="2.22">2.22 mm</quantity>
(
<emphasis id="7EEB2C2FFFE5FF9940B8F9D7FD48F9AA" box="[714,726,1570,1591]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 1); L/W = 1.85 (
<emphasis id="7EEB2C2FFFE5FF9941F5F9D7FC0DF9AA" box="[903,915,1570,1591]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 1); HW =
<quantity id="8B675DD8FFE5FF994678F9D4FBC3F9AA" box="[1034,1117,1569,1591]" metricMagnitude="-3" metricUnit="m" metricValue="1.51" pageId="15" pageNumber="737" unit="mm" value="1.51">1.51 mm</quantity>
(
<emphasis id="7EEB2C2FFFE5FF994618F9D7FBE8F9AA" box="[1130,1142,1570,1591]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 1); IOD =
<quantity id="8B675DD8FFE5FF994175F9C9FCC2F9CF" box="[775,860,1596,1618]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="15" pageNumber="737" unit="mm" value="0.54">0.54 mm</quantity>
(
<emphasis id="7EEB2C2FFFE5FF994118F9C8FCE8F9CF" box="[874,886,1597,1618]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 1); IOD/HW = 0.36 (
<emphasis id="7EEB2C2FFFE5FF994629F9C8FBF9F9CF" box="[1115,1127,1597,1618]" italics="true" pageId="15" pageNumber="737">n</emphasis>
= 1).
</paragraph>
</subSubSection>
<subSubSection id="0485A3B6FFE5FF8740A9F9AFFE3AFE0E" lastPageId="17" lastPageNumber="739" pageId="15" pageNumber="737" type="distribution">
<paragraph id="4C20F03DFFE5FF9940A9F9AFFCE1F917" blockId="15.[707,1193,194,1731]" pageId="15" pageNumber="737">
<emphasis id="7EEB2C2FFFE5FF9940A9F9AFFCC9F9ED" bold="true" box="[731,855,1626,1648]" pageId="15" pageNumber="737">Distribution.</emphasis>
This species was found in
<collectingRegion id="8E5B3EDFFFE5FF99463EF9AFFB37F9ED" box="[1100,1193,1626,1648]" country="United States of America" name="California" pageId="15" pageNumber="737">California</collectingRegion>
and
<collectingRegion id="8E5B3EDFFFE5FF99409BF980FC88F916" box="[745,790,1653,1675]" country="United States of America" name="Utah" pageId="15" pageNumber="737">Utah</collectingRegion>
(
<figureCitation id="D4A4ECB8FFE5FF994153F980FCEAF917" box="[801,884,1653,1675]" captionStart="Fig" captionStartId="25.[197,228,1297,1316]" captionTargetBox="[171,1195,193,1270]" captionTargetId="figure-207@25.[171,1195,193,1270]" captionTargetPageId="25" captionText="Fig. 14. Distribution of Pachybrachis species, based only upon specimens examined in this study. A) P. hybridus (black), P. turgidicollis (red), P. texasensis, new species (green), P. chaoticus (blue), P. pinguescens (orange), P. crassus (purple), B) P. saguaro, new species (black), P. peninsularis (purple), P. cylindricus (red), P. huachuca, new species (orange), P. hurricane, new species (gray), P. ephedra, new species (blue), and P. ashfork, new species (green)." figureDoi="http://doi.org/10.5281/zenodo.13252067" httpUri="https://zenodo.org/record/13252067/files/figure.png" pageId="15" pageNumber="737">Fig. 14B</figureCitation>
).
</paragraph>
<paragraph id="4C20F03DFFE5FF9940A9F967FC50F95E" blockId="15.[707,1193,194,1731]" pageId="15" pageNumber="737">
<emphasis id="7EEB2C2FFFE5FF9940A9F967FC1BF93A" bold="true" box="[731,901,1682,1704]" pageId="15" pageNumber="737">Biological Notes.</emphasis>
This species was collected on
<emphasis id="7EEB2C2FFFE5FF9940B1F958FCE7F95E" box="[707,889,1709,1731]" italics="true" pageId="15" pageNumber="737">Ephedra torreyana</emphasis>
in
<collectingRegion id="8E5B3EDFFFE5FF9941EAF958FC54F95E" box="[920,970,1709,1731]" country="United States of America" name="Utah" pageId="15" pageNumber="737">Utah</collectingRegion>
.
</paragraph>
<caption id="18E0A0B5FFFAFF8642C2F9CFFE1AF9E0" ID-DOI="http://doi.org/10.5281/zenodo.13252053" ID-Zenodo-Dep="13252053" httpUri="https://zenodo.org/record/13252053/files/figure.png" pageId="16" pageNumber="738" startId="16.[176,207,1594,1613]" targetBox="[160,1165,202,1565]" targetPageId="16" targetType="figure">
<paragraph id="4C20F03DFFFAFF8642C2F9CFFE1AF9E0" blockId="16.[152,1172,1594,1661]" pageId="16" pageNumber="738">
<emphasis id="7EEB2C2FFFFAFF8642C2F9CFFF79F9D0" bold="true" box="[176,231,1594,1613]" pageId="16" pageNumber="738">Fig. 8.</emphasis>
<taxonomicName id="8B9F8BBEFFFAFF86428FF9CFFE23F9D0" authorityName="Barney" authorityYear="2021" box="[253,445,1594,1613]" class="Insecta" family="Chrysomelidae" genus="Pachybrachis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="738" phylum="Arthropoda" rank="species" species="ephedra" status="sp. nov.">
<emphasis id="7EEB2C2FFFFAFF86428FF9CFFE23F9D0" box="[253,445,1594,1613]" italics="true" pageId="16" pageNumber="738">Pachybrachis ephedra</emphasis>
</taxonomicName>
,
<emphasis id="7EEB2C2FFFFAFF8643B5F9CFFDAEF9D0" bold="true" box="[455,560,1594,1613]" pageId="16" pageNumber="738">
<taxonomicNameLabel id="65D89154FFFAFF8643B5F9CFFDAEF9D0" box="[455,560,1594,1613]" pageId="16" pageNumber="738" rank="species">new species</taxonomicNameLabel>
</emphasis>
, Washington Co., Utah, holotype.A) Dorsal habitus, B) Lateral habitus, C) Face, D) Pygidium, E) Median lobe of aedeagus, lateral view, F) Median lobe of aedeagus, en-face view, G) Median lobe of aedeagus, sub view.
</paragraph>
</caption>
<paragraph id="4C20F03DFFFBFF8742B7FF34FE3AFE0E" blockId="17.[173,660,193,403]" pageId="17" pageNumber="739">
<emphasis id="7EEB2C2FFFFBFF8742B7FF34FE3BFF4A" bold="true" box="[197,421,193,215]" pageId="17" pageNumber="739">Specimens Examined.</emphasis>
<materialsCitation id="FCF7FA60FFFBFF8743DEFF37FE3AFE0E" collectingDate="1941-05-28" collectingDateMax="2015-05-17" collectingDateMin="1941-05-28" collectionCode="OSUC, BYUC, RJBC" collectorName="D. J. and J. N. Knull &amp; S. M. Clark &amp; E. G. Riley &amp; Ephedra &amp; R. J. Barney" country="United States of America" county="San Bernardino Co." latitude="37.1442" location="Morongo Valley" longLatPrecision="6" longitude="-113.254" pageId="17" pageNumber="739" specimenCount="5" specimenCount-male="3" stateProvince="California" typeStatus="holotype">
<typeStatus id="93244E9FFFFBFF8743DEFF37FD9BFF45" box="[428,517,194,216]" pageId="17" pageNumber="739" type="holotype">Holotype</typeStatus>
+
<specimenCount id="5A993BB4FFFBFF87406DFF37FD0EFF4A" box="[543,656,194,215]" count="2" pageId="17" pageNumber="739" type="generic" typeStatus="paratype">2 paratypes</specimenCount>
.
<emphasis id="7EEB2C2FFFFBFF8742DFFF29FE02FF6F" bold="true" box="[173,412,220,242]" pageId="17" pageNumber="739">
<collectingCountry id="3488B0ADFFFBFF8742DFFF29FF7FFF6F" box="[173,225,220,242]" name="United States of America" pageId="17" pageNumber="739">USA</collectingCountry>
:
<collectingRegion id="8E5B3EDFFFFBFF874288FF29FE06FF6F" box="[250,408,220,242]" country="United States of America" name="California" pageId="17" pageNumber="739">CALIFORNIA</collectingRegion>
:
</emphasis>
<collectingCounty id="A54188B1FFFBFF8743C3FF28FD11FF6F" box="[433,655,220,242]" pageId="17" pageNumber="739">San Bernardino Co.</collectingCounty>
,
<location id="4940A6E6FFFBFF8742DFFF0DFED2FE90" LSID="urn:lsid:plazi:treatment:C436412BFFE5FF8742CFFA80FE3AFE0E:4940A6E6FFFBFF8742DFFF0DFED2FE90" box="[173,332,247,269]" country="United States of America" county="San Bernardino Co." latitude="37.1442" longLatPrecision="6" longitude="-113.254" name="Morongo Valley" pageId="17" pageNumber="739" stateProvince="California">Morongo Valley</location>
,
<date id="3821D6FDFFFBFF87432AFF0DFE26FE90" box="[344,440,248,269]" pageId="17" pageNumber="739" value="1941-05-28">
<collectingDate id="28652F15FFFBFF87432AFF0DFE26FE90" box="[344,440,248,269]" pageId="17" pageNumber="739" value="1941-05-28">28.v.1941</collectingDate>
</date>
,
<collectorName id="E16A95EBFFFBFF8743B6FF0DFD0DFE90" box="[452,659,247,269]" pageId="17" pageNumber="739">D. J. and J. N. Knull</collectorName>
[
<specimenCount id="5A993BB4FFFBFF8742C7FEE7FF4BFEBA" box="[181,213,274,295]" count="1" pageId="17" pageNumber="739" type="male">1♂</specimenCount>
,
<collectionCode id="2A8E68F8FFFBFF874292FEE7FEBBFEBA" box="[224,293,274,295]" country="USA" name="Oregon State University" pageId="17" pageNumber="739">OSUC</collectionCode>
].
<emphasis id="7EEB2C2FFFFBFF874346FEE7FE1EFEBA" bold="true" box="[308,384,274,295]" pageId="17" pageNumber="739">
<collectingRegion id="8E5B3EDFFFFBFF874346FEE7FEE5FEBA" box="[308,379,274,295]" country="United States of America" name="Utah" pageId="17" pageNumber="739">UTAH</collectingRegion>
:
</emphasis>
<collectingRegion id="8E5B3EDFFFFBFF8743F4FEE7FE66FEB5" box="[390,504,274,296]" country="United States of America" name="Washington" pageId="17" pageNumber="739">Washington</collectingRegion>
Co.,
<locationDeviation id="5D92E91FFFFBFF87405FFEE7FE91FEDE" location="Hurricane" pageId="17" pageNumber="739">
<quantity id="8B675DD8FFFBFF87405FFEE7FDEFFEB5" box="[557,625,274,296]" metricMagnitude="3" metricUnit="m" metricValue="4.7" pageId="17" pageNumber="739" unit="km" value="4.7">4.7 km</quantity>
SE Hurricane
</locationDeviation>
,
<geoCoordinate id="29AB96FAFFFBFF87436EFED8FE1BFEDF" box="[284,389,301,322]" degrees="37.1442" direction="north" orientation="latitude" pageId="17" pageNumber="739" precision="5" value="37.1442">37.1442°N</geoCoordinate>
<geoCoordinate id="29AB96FAFFFBFF8743FFFED8FD98FEDE" box="[397,518,301,323]" degrees="113.2540" direction="west" orientation="longitude" pageId="17" pageNumber="739" precision="5" value="-113.254">113.2540°W</geoCoordinate>
,
<date id="3821D6FDFFFBFF874061FED8FDEDFEDE" box="[531,627,301,323]" pageId="17" pageNumber="739" value="2015-05-17">
<collectingDate id="28652F15FFFBFF874061FED8FDEDFEDE" box="[531,627,301,323]" pageId="17" pageNumber="739" value="2015-05-17">17.v.2015</collectingDate>
</date>
,
<collectorName id="E16A95EBFFFBFF87400DFED8FE92FEC3" pageId="17" pageNumber="739">S. M. Clark</collectorName>
and
<collectorName id="E16A95EBFFFBFF874338FEBDFE58FEC3" box="[330,454,328,350]" pageId="17" pageNumber="739">E. G. Riley</collectorName>
[
<specimenCount id="5A993BB4FFFBFF8743A9FEBDFE65FEC0" box="[475,507,328,349]" count="1" pageId="17" pageNumber="739" type="male">1♂</specimenCount>
,
<typeStatus id="93244E9FFFFBFF87407CFEBDFD11FEC0" box="[526,655,328,349]" pageId="17" pageNumber="739" type="holotype">HOLOTYPE</typeStatus>
,
<collectionCode id="2A8E68F8FFFBFF8742DFFE96FF6BFEE5" box="[173,245,355,376]" pageId="17" pageNumber="739">BYUC</collectionCode>
]; same data, except ex.
<emphasis id="7EEB2C2FFFFBFF8743A4FE96FD13FEE4" box="[470,653,355,377]" italics="true" pageId="17" pageNumber="739">
<collectorName id="E16A95EBFFFBFF8743A4FE96FDB7FEE5" box="[470,553,355,376]" pageId="17" pageNumber="739">Ephedra</collectorName>
torreyana
</emphasis>
,
<collectorName id="E16A95EBFFFBFF8742DFFE8BFEBAFE0E" box="[173,292,382,403]" pageId="17" pageNumber="739">R. J. Barney</collectorName>
[
<specimenCount id="5A993BB4FFFBFF874343FE8BFECFFE0E" box="[305,337,382,403]" count="1" pageId="17" pageNumber="739" type="male">1♂</specimenCount>
,
<collectionCode id="2A8E68F8FFFBFF87432EFE8BFE05FE0E" box="[348,411,382,403]" pageId="17" pageNumber="739">RJBC</collectionCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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