378 lines
44 KiB
XML
378 lines
44 KiB
XML
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<mods:title id="D476E921C56BC09E64EFF03CC5BE0A99">Two new species of Laonice (Norgensia) (Spionidae, Polychaeta) from subtropical Atlantic and subequatorial Pacific coasts of North America</mods:title>
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<mods:namePart id="5C27AD49739AFD3AF5B9F7183B5663F1">Sikorski, Andrey</mods:namePart>
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<mods:namePart id="12358D256E042867757AFAD2843673CE">Pavlova, Lyudmila</mods:namePart>
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<mods:title id="2823A3BB5A7B880465EC80E13C05DCC6">Zootaxa</mods:title>
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<mods:date id="03E57038A4CA75708F47B3DE0C886C30">2018</mods:date>
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<taxonomicName id="F7EC5FAE24229F1AF4FD301C0B5A950D" ID-CoL="3S888" authority="Sikorski & Pavlova, 2018" authorityName="Sikorski & Pavlova" authorityYear="2018" box="[151,562,1055,1082]" class="Polychaeta" family="Spionidae" genus="Laonice" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spionida" pageId="2" pageNumber="173" phylum="Annelida" rank="species" species="costaricensis" status="sp. nov." subGenus="Norgensia">
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<emphasis id="0298F83F24229F1AF4FD301C0991950D" bold="true" box="[151,249,1055,1081]" italics="true" pageId="2" pageNumber="173">Laonice</emphasis>
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(
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<emphasis id="0298F83F24229F1AF560301C08EF950D" bold="true" box="[266,391,1055,1081]" italics="true" pageId="2" pageNumber="173">Norgensia</emphasis>
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)
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<emphasis id="0298F83F24229F1AF5F2301C0B5A950D" bold="true" box="[408,562,1055,1081]" italics="true" pageId="2" pageNumber="173">costaricensis</emphasis>
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<taxonomicNameLabel id="19AB454424229F1AF65330230BFF950E" box="[569,663,1056,1082]" pageId="2" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="3053242D24229F1AF4FD3041086F956F" blockId="2.[151,663,1055,1115]" box="[151,263,1090,1115]" pageId="2" pageNumber="173">
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(
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<figureCitation id="A8D738A824229F1AF4F430410997956F" box="[158,255,1090,1115]" captionStart="FIGURE 2" captionStartId="3.[151,244,1934,1956]" captionTargetBox="[166,1426,455,1887]" captionTargetId="figure@3.[151,1435,445,1914]" captionTargetPageId="3" captionText="FIGURE 2. Laonice (Norgensia) costaricensis sp. nov. A. Anterior end, dorsal view; B. Posterior end with pygidium, ventral view; C–F. Parapodia of chaetigers 2, 7, 24 and 42; G–H. Hooded hook from chaetiger 42, side and frontal view. A–B—Holotype (MCZ 101143); C–G—Paratype (MZUCR 348-05).Scale:A–B, D–F—0.5 mm; C—0.3 mm; G—0.05 mm." httpUri="https://zenodo.org/record/1245062/files/figure.png" pageId="2" pageNumber="173">Figure 2</figureCitation>
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)
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</paragraph>
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<subSubSection id="78F677A624229F1AF4FD30880B549594" box="[151,572,1162,1184]" pageId="2" pageNumber="173" type="reference_group">
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<paragraph id="3053242D24229F1AF4FD30880B549594" blockId="2.[151,572,1162,1184]" box="[151,572,1162,1184]" pageId="2" pageNumber="173">
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<taxonomicName id="F7EC5FAE24229F1AF4FD308808859594" ID-CoL="3S88N" authority="Dean et al., 2012" authorityName="Dean et al." authorityYear="2012" box="[151,493,1162,1184]" class="Polychaeta" family="Spionidae" genus="Laonice" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spionida" pageId="2" pageNumber="173" phylum="Annelida" rank="species" species="cirrata">
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<emphasis id="0298F83F24229F1AF4FD3088085D9594" box="[151,309,1163,1184]" italics="true" pageId="2" pageNumber="173">Laonice cirrata</emphasis>
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<bibRefCitation id="547D59DC24229F1AF551308808859594" box="[315,493,1162,1184]" pageId="2" pageNumber="173" refId="ref3728" refString="Dean, H. K., Sibaja-Cordero, J. A. & Cortes, J. (2012) Polychaetes (Annelida: Polychaeta) of Cocos Island National Park, Pacific Costa Rica. Pacific Science, 66 (3), 347 - 386. https: // doi. org / 10.2984 / 66.3.8" type="journal article">
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Dean
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<emphasis id="0298F83F24229F1AF51D308808C29594" box="[375,426,1163,1184]" italics="true" pageId="2" pageNumber="173">et al.</emphasis>
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, 2012
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</bibRefCitation>
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</taxonomicName>
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: p. 366.
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</paragraph>
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<materialsCitation id="80842E7024229F1AF4FD30CE0DE895D2" ID-GBIF-Occurrence="1844422637" box="[151,1152,1229,1254]" collectingDate="2007-10-07" collectionCode="MCZ" elevation="28" latitude="5.55" location="Bahia Chatham" longLatPrecision="783" longitude="-87.05" pageId="2" pageNumber="173" specimenCode="MCZ 101143" specimenCount="1" typeStatus="holotype">
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<typeStatus id="EF579A8F24229F1AF4FD30CE086095D2" box="[151,264,1229,1254]" pageId="2" pageNumber="173" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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(
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), algae,
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,
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<date id="445202ED24229F1AF72E30CD0AA095D2" box="[836,968,1230,1254]" pageId="2" pageNumber="173" value="2007-10-07">7 Oct. 2007</date>
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</collectingDate>
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(
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MCZ
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).
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</paragraph>
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<paragraph id="3053242D24229F1AF4AD30F20A179442" blockId="2.[151,1437,1229,2010]" pageId="2" pageNumber="173">
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<typeStatus id="EF579A8F24229F1AF4AD30F2082B943E" box="[199,323,1265,1290]" pageId="2" pageNumber="173" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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(
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), algae,
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,
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<date id="445202ED24229F1AF7E430F10D7D943E" box="[910,1045,1266,1290]" pageId="2" pageNumber="173" value="2007-10-07">7 Oct. 2007</date>
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</collectingDate>
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(
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<specimenCode id="604A8C5624229F1AF04F30F10DAC943E" box="[1061,1220,1265,1290]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="2" pageNumber="173">MCZ 141960</specimenCode>
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,
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<specimenCode id="604A8C5624229F1AF0BA30F10DB5943E" box="[1232,1245,1266,1290]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="2" pageNumber="173">1</specimenCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="80842E7024229F1AF09D30F20DE5941A" ID-GBIF-Occurrence="1844422638" collectingDate="2007-10-07" collectionCode="MZUCR" collectorName="J. A. Sibaja" country="Iceland" elevation="28" location="Coco" municipality="Bahia Chatham" pageId="2" pageNumber="173" specimenCode="MZUCR 246-02, 1" specimenCount="1" typeStatus="paratype">
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<emphasis id="0298F83F24229F1AF09D30F20831941A" bold="true" pageId="2" pageNumber="173">
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<collectingCountry id="48FB64BD24229F1AF09D30F20C4B943E" box="[1271,1315,1265,1290]" name="Iceland" pageId="2" pageNumber="173">Isla</collectingCountry>
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del
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<location id="353372F624229F1AF13230F20CFE943E" LSID="urn:lsid:plazi:treatment:B845953B24229F1CF4FD301C0BF593C4:353372F624229F1AF13230F20CFE943E" box="[1368,1430,1265,1290]" country="Iceland" municipality="Bahia Chatham" name="Coco" pageId="2" pageNumber="173">Coco</location>
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,
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<collectingMunicipality id="D037BE5724229F1AF4FD31160831941A" box="[151,345,1301,1326]" pageId="2" pageNumber="173">Bahia Chatham</collectingMunicipality>
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</emphasis>
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, grab,
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<quantity id="F71489C824229F1AF5DA311508989419" box="[432,496,1302,1326]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="173" unit="m" value="28.0">28 m</quantity>
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</elevation>
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,
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<collectingDate id="5416FB0524229F1AF66B31150BF9941A" box="[513,657,1302,1326]" pageId="2" pageNumber="173" value="2007-10-07">
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<date id="445202ED24229F1AF66B31150BF9941A" box="[513,657,1302,1326]" pageId="2" pageNumber="173" value="2007-10-07">7 Oct. 2007</date>
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</collectingDate>
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, collector
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<collectorName id="9D1941FB24229F1AF76531150AE6941A" box="[783,910,1301,1326]" pageId="2" pageNumber="173">J.A. Sibaja</collectorName>
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(
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<specimenCode id="604A8C5624229F1AF7CB31150D0D941A" box="[929,1125,1301,1326]" collectionCode="MZUCR" pageId="2" pageNumber="173">MZUCR 246-02</specimenCode>
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,
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<specimenCode id="604A8C5624229F1AF01E31150DE9941A" box="[1140,1153,1302,1326]" collectionCode="MZUCR" pageId="2" pageNumber="173">1</specimenCode>
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)
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</materialsCitation>
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;
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<materialsCitation id="80842E7024229F1AF0F531160D709466" ID-GBIF-Occurrence="1844422545" collectingDate="2008-04" collectionCode="MZUCR" collectorName="J. A. Sibaja" country="Iceland" elevation="24" location="Coco" municipality="Bahia Chatham" pageId="2" pageNumber="173" specimenCode="MZUCR 223-01, 1" specimenCount="1" typeStatus="paratype">
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<emphasis id="0298F83F24229F1AF0F53116086F9466" bold="true" pageId="2" pageNumber="173">
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<collectingCountry id="48FB64BD24229F1AF0F531160DA3941A" box="[1183,1227,1301,1326]" name="Iceland" pageId="2" pageNumber="173">Isla</collectingCountry>
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del
|
||
<location id="353372F624229F1AF16C31160C2C941A" LSID="urn:lsid:plazi:treatment:B845953B24229F1CF4FD301C0BF593C4:353372F624229F1AF16C31160C2C941A" box="[1286,1348,1301,1326]" country="Iceland" municipality="Bahia Chatham" name="Coco" pageId="2" pageNumber="173">Coco</location>
|
||
,
|
||
<collectingMunicipality id="D037BE5724229F1AF13C3116086F9466" pageId="2" pageNumber="173">Bahia Chatham</collectingMunicipality>
|
||
</emphasis>
|
||
, trawl,
|
||
<elevation id="BBC1C31E24229F1AF50B313908F69465" box="[353,414,1338,1362]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="173" unit="m" value="24.0">
|
||
<quantity id="F71489C824229F1AF50B313908F69465" box="[353,414,1338,1362]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="173" unit="m" value="24.0">24 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="5416FB0524229F1AF5C6313A0B439466" box="[428,555,1337,1362]" pageId="2" pageNumber="173" value="2008-04">
|
||
<date id="445202ED24229F1AF5C6313A0B439466" box="[428,555,1337,1362]" pageId="2" pageNumber="173" value="2008-04">April 2008</date>
|
||
</collectingDate>
|
||
, collector
|
||
<collectorName id="9D1941FB24229F1AF6CE31390A489466" box="[676,800,1337,1362]" pageId="2" pageNumber="173">J.A. Sibaja</collectorName>
|
||
(
|
||
<specimenCode id="604A8C5624229F1AF75B31390A999466" box="[817,1009,1338,1362]" collectionCode="MZUCR" pageId="2" pageNumber="173">MZUCR 223-01</specimenCode>
|
||
,
|
||
<specimenCode id="604A8C5624229F1AF79531390D649466" box="[1023,1036,1338,1362]" collectionCode="MZUCR" pageId="2" pageNumber="173">1</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="80842E7024229F1AF04D313A0A129442" ID-GBIF-Occurrence="1844422721" collectingDate="2008-04" collectionCode="MZUCR" collectorName="J. A. Sibaja" country="Iceland" elevation="65" location="Coco" municipality="Bahia Chatham" pageId="2" pageNumber="173" specimenCode="MZUCR 348-05, 1" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="0298F83F24229F1AF04D313A0CFD9466" bold="true" box="[1063,1429,1337,1362]" pageId="2" pageNumber="173">
|
||
<collectingCountry id="48FB64BD24229F1AF04D313A0D3B9466" box="[1063,1107,1337,1362]" name="Iceland" pageId="2" pageNumber="173">Isla</collectingCountry>
|
||
del
|
||
<location id="353372F624229F1AF0E3313A0DAF9466" LSID="urn:lsid:plazi:treatment:B845953B24229F1CF4FD301C0BF593C4:353372F624229F1AF0E3313A0DAF9466" box="[1161,1223,1337,1362]" country="Iceland" municipality="Bahia Chatham" name="Coco" pageId="2" pageNumber="173">Coco</location>
|
||
,
|
||
<collectingMunicipality id="D037BE5724229F1AF0BC313A0CFD9466" box="[1238,1429,1337,1362]" pageId="2" pageNumber="173">Bahia Chatham</collectingMunicipality>
|
||
</emphasis>
|
||
, grab,
|
||
<elevation id="BBC1C31E24229F1AF4BC315E087A9441" box="[214,274,1373,1398]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="173" unit="m" value="65.0">
|
||
<quantity id="F71489C824229F1AF4BC315E087A9441" box="[214,274,1373,1398]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="173" unit="m" value="65.0">65 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="5416FB0524229F1AF577315E08F29442" box="[285,410,1373,1398]" pageId="2" pageNumber="173" value="2008-04">
|
||
<date id="445202ED24229F1AF577315E08F29442" box="[285,410,1373,1398]" pageId="2" pageNumber="173" value="2008-04">April 2008</date>
|
||
</collectingDate>
|
||
, collector
|
||
<collectorName id="9D1941FB24229F1AF665315D0BE19442" box="[527,649,1373,1398]" pageId="2" pageNumber="173">J.A. Sibaja</collectorName>
|
||
(
|
||
<specimenCode id="604A8C5624229F1AF6FD315D0A3D9442" box="[663,853,1373,1398]" collectionCode="MZUCR" pageId="2" pageNumber="173">MZUCR 348-05</specimenCode>
|
||
,
|
||
<specimenCode id="604A8C5624229F1AF70B315D0A069442" box="[865,878,1374,1398]" collectionCode="MZUCR" pageId="2" pageNumber="173">1</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78F677A624229F1AF4AD31820A459732" pageId="2" pageNumber="173" type="description">
|
||
<paragraph id="3053242D24229F1AF4AD31820A459732" blockId="2.[151,1437,1229,2010]" pageId="2" pageNumber="173">
|
||
<emphasis id="0298F83F24229F1AF4AD318208A494AE" bold="true" box="[199,460,1409,1434]" pageId="2" pageNumber="173">
|
||
Description.
|
||
<typeStatus id="EF579A8F24229F1AF535318208A494AE" box="[351,460,1409,1434]" pageId="2" pageNumber="173" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
: Complete specimen
|
||
<quantity id="F71489C824229F1AF6D531810A7D94AD" box="[703,789,1410,1434]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="173" unit="mm" value="0.3">0.3 mm</quantity>
|
||
wide,
|
||
<quantity id="F71489C824229F1AF70A31810AC694AD" box="[864,942,1410,1434]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="173" unit="mm" value="10.0">10 mm</quantity>
|
||
long for 66 chaetigers. Neuropodial hooded hooks appear from chaetiger 17, up to 5 per neuropodium. Nuchal organ extending to chaetiger 11. Chaetiger 45 is the last one carrying branchiae. Sabre chaetae present from chaetiger 11, one per fascicle. Genital pouches appear on chaetiger 12 (genital pouches absent after chaetiger 21).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78F677A624229F1CF4AD32120A639184" lastPageId="4" lastPageNumber="175" pageId="2" pageNumber="173" type="materials_examined">
|
||
<paragraph id="3053242D24229F1AF4AD32120AC0977A" blockId="2.[151,1437,1229,2010]" pageId="2" pageNumber="173">
|
||
<emphasis id="0298F83F24229F1AF4AD321208B7971E" bold="true" box="[199,479,1553,1578]" pageId="2" pageNumber="173">All material examined:</emphasis>
|
||
One
|
||
<typeStatus id="EF579A8F24229F1AF64832110BEA971E" box="[546,642,1554,1578]" pageId="2" pageNumber="173" type="paratype">paratype</typeStatus>
|
||
is a complete specimen
|
||
<quantity id="F71489C824229F1AF7C832110A93971D" box="[930,1019,1554,1578]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="173" unit="mm" value="0.3">0.3 mm</quantity>
|
||
wide,
|
||
<quantity id="F71489C824229F1AF02132110DE7971D" box="[1099,1167,1554,1578]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="173" unit="mm" value="9.0">9 mm</quantity>
|
||
long for 63 chaetigers. Three
|
||
<typeStatus id="EF579A8F24229F1AF4B532350822977A" box="[223,330,1590,1614]" pageId="2" pageNumber="173" type="paratype">paratypes</typeStatus>
|
||
are anterior fragments between 0.5 and
|
||
<quantity id="F71489C824229F1AF76732350A0B9779" box="[781,867,1590,1614]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="173" unit="mm" value="0.8">0.8 mm</quantity>
|
||
wide.
|
||
</paragraph>
|
||
<paragraph id="3053242D24229F1BF4AD325A0C679074" blockId="2.[151,1437,1229,2010]" lastBlockId="3.[151,1437,151,392]" lastPageId="3" lastPageNumber="174" pageId="2" pageNumber="173">
|
||
Prostomium blunt, anteriorly rounded, entire or very slightly indented, without frontal or lateral horns. Caruncle following nuchal organ extending to chaetiger 11–20. One pair of large eyes. Long, finger-like occipital antenna present on posterior part of prostomium, behind eyes. Peristomium not fused to prostomium (
|
||
<figureCitation id="A8D738A824229F1AF16632A20C0E978E" box="[1292,1382,1697,1722]" captionStart="FIGURE 2" captionStartId="3.[151,244,1934,1956]" captionTargetBox="[166,1426,455,1887]" captionTargetId="figure@3.[151,1435,445,1914]" captionTargetPageId="3" captionText="FIGURE 2. Laonice (Norgensia) costaricensis sp. nov. A. Anterior end, dorsal view; B. Posterior end with pygidium, ventral view; C–F. Parapodia of chaetigers 2, 7, 24 and 42; G–H. Hooded hook from chaetiger 42, side and frontal view. A–B—Holotype (MCZ 101143); C–G—Paratype (MZUCR 348-05).Scale:A–B, D–F—0.5 mm; C—0.3 mm; G—0.05 mm." httpUri="https://zenodo.org/record/1245062/files/figure.png" pageId="2" pageNumber="173">Fig. 2A</figureCitation>
|
||
). No palps remain. Branchiae present from chaetiger 2 on about ¾ of the following chaetigers. Branchiae free, separate from notopodial lamellae throughout; longest on anterior third of specimen, first branchiae approximately the same length as notopodial lobe (
|
||
<figureCitation id="A8D738A824229F1AF5AC330E0B489612" box="[454,544,1805,1830]" captionStart="FIGURE 2" captionStartId="3.[151,244,1934,1956]" captionTargetBox="[166,1426,455,1887]" captionTargetId="figure@3.[151,1435,445,1914]" captionTargetPageId="3" captionText="FIGURE 2. Laonice (Norgensia) costaricensis sp. nov. A. Anterior end, dorsal view; B. Posterior end with pygidium, ventral view; C–F. Parapodia of chaetigers 2, 7, 24 and 42; G–H. Hooded hook from chaetiger 42, side and frontal view. A–B—Holotype (MCZ 101143); C–G—Paratype (MZUCR 348-05).Scale:A–B, D–F—0.5 mm; C—0.3 mm; G—0.05 mm." httpUri="https://zenodo.org/record/1245062/files/figure.png" pageId="2" pageNumber="173">Fig. 2C</figureCitation>
|
||
), by about chaetiger 7 approximately twice the length of notopodial lobe (
|
||
<figureCitation id="A8D738A824229F1AF105330E09D4967E" captionStart="FIGURE 2" captionStartId="3.[151,244,1934,1956]" captionTargetBox="[166,1426,455,1887]" captionTargetId="figure@3.[151,1435,445,1914]" captionTargetPageId="3" captionText="FIGURE 2. Laonice (Norgensia) costaricensis sp. nov. A. Anterior end, dorsal view; B. Posterior end with pygidium, ventral view; C–F. Parapodia of chaetigers 2, 7, 24 and 42; G–H. Hooded hook from chaetiger 42, side and frontal view. A–B—Holotype (MCZ 101143); C–G—Paratype (MZUCR 348-05).Scale:A–B, D–F—0.5 mm; C—0.3 mm; G—0.05 mm." httpUri="https://zenodo.org/record/1245062/files/figure.png" pageId="2" pageNumber="173">Fig. 2D</figureCitation>
|
||
), and in posterior half up to four times the length of the lobes (
|
||
<figureCitation id="A8D738A824229F1AF71833320A8F967E" box="[882,999,1841,1866]" captionStart="FIGURE 2" captionStartId="3.[151,244,1934,1956]" captionTargetBox="[166,1426,455,1887]" captionTargetId="figure@3.[151,1435,445,1914]" captionTargetPageId="3" captionText="FIGURE 2. Laonice (Norgensia) costaricensis sp. nov. A. Anterior end, dorsal view; B. Posterior end with pygidium, ventral view; C–F. Parapodia of chaetigers 2, 7, 24 and 42; G–H. Hooded hook from chaetiger 42, side and frontal view. A–B—Holotype (MCZ 101143); C–G—Paratype (MZUCR 348-05).Scale:A–B, D–F—0.5 mm; C—0.3 mm; G—0.05 mm." httpUri="https://zenodo.org/record/1245062/files/figure.png" pageId="2" pageNumber="173">Fig. 2E–F</figureCitation>
|
||
). Notopodial postchaetal lobes reduced to small subtriangular projection on chaetiger 1; then leaf-like and slightly pointed until about chaetiger 7 (
|
||
<figureCitation id="A8D738A824229F1AF105335609B596A6" captionStart="FIGURE 2" captionStartId="3.[151,244,1934,1956]" captionTargetBox="[166,1426,455,1887]" captionTargetId="figure@3.[151,1435,445,1914]" captionTargetPageId="3" captionText="FIGURE 2. Laonice (Norgensia) costaricensis sp. nov. A. Anterior end, dorsal view; B. Posterior end with pygidium, ventral view; C–F. Parapodia of chaetigers 2, 7, 24 and 42; G–H. Hooded hook from chaetiger 42, side and frontal view. A–B—Holotype (MCZ 101143); C–G—Paratype (MZUCR 348-05).Scale:A–B, D–F—0.5 mm; C—0.3 mm; G—0.05 mm." httpUri="https://zenodo.org/record/1245062/files/figure.png" pageId="2" pageNumber="173">Fig. 2C–D</figureCitation>
|
||
), thereafter becoming more rounded and gradually reduced to small round lobes (
|
||
<figureCitation id="A8D738A824229F1AF0EE337A0D9496A6" box="[1156,1276,1913,1938]" captionStart="FIGURE 2" captionStartId="3.[151,244,1934,1956]" captionTargetBox="[166,1426,455,1887]" captionTargetId="figure@3.[151,1435,445,1914]" captionTargetPageId="3" captionText="FIGURE 2. Laonice (Norgensia) costaricensis sp. nov. A. Anterior end, dorsal view; B. Posterior end with pygidium, ventral view; C–F. Parapodia of chaetigers 2, 7, 24 and 42; G–H. Hooded hook from chaetiger 42, side and frontal view. A–B—Holotype (MCZ 101143); C–G—Paratype (MZUCR 348-05).Scale:A–B, D–F—0.5 mm; C—0.3 mm; G—0.05 mm." httpUri="https://zenodo.org/record/1245062/files/figure.png" pageId="2" pageNumber="173">Fig. 2E–F</figureCitation>
|
||
). Neuropodial postchaetal lobes leaf-like from chaetiger 1; ventrally rounded and dorsally pointed until about chaetiger 6 (
|
||
<figureCitation id="A8D738A824229F1AF105339E09D396EE" captionStart="FIGURE 2" captionStartId="3.[151,244,1934,1956]" captionTargetBox="[166,1426,455,1887]" captionTargetId="figure@3.[151,1435,445,1914]" captionTargetPageId="3" captionText="FIGURE 2. Laonice (Norgensia) costaricensis sp. nov. A. Anterior end, dorsal view; B. Posterior end with pygidium, ventral view; C–F. Parapodia of chaetigers 2, 7, 24 and 42; G–H. Hooded hook from chaetiger 42, side and frontal view. A–B—Holotype (MCZ 101143); C–G—Paratype (MZUCR 348-05).Scale:A–B, D–F—0.5 mm; C—0.3 mm; G—0.05 mm." httpUri="https://zenodo.org/record/1245062/files/figure.png" pageId="2" pageNumber="173">Fig. 2C</figureCitation>
|
||
), becoming gradually rounded and reduced in size after about chaetiger 15 (
|
||
<figureCitation id="A8D738A824229F1AF05133C20DDD96EE" box="[1083,1205,1985,2010]" captionStart="FIGURE 2" captionStartId="3.[151,244,1934,1956]" captionTargetBox="[166,1426,455,1887]" captionTargetId="figure@3.[151,1435,445,1914]" captionTargetPageId="3" captionText="FIGURE 2. Laonice (Norgensia) costaricensis sp. nov. A. Anterior end, dorsal view; B. Posterior end with pygidium, ventral view; C–F. Parapodia of chaetigers 2, 7, 24 and 42; G–H. Hooded hook from chaetiger 42, side and frontal view. A–B—Holotype (MCZ 101143); C–G—Paratype (MZUCR 348-05).Scale:A–B, D–F—0.5 mm; C—0.3 mm; G—0.05 mm." httpUri="https://zenodo.org/record/1245062/files/figure.png" pageId="2" pageNumber="173">Fig. 2E–F</figureCitation>
|
||
). Transversal dorsal membranes not pronounced—only on the largest specimen low membranes between branchiae present. Genital pouches from chaetiger 10–17. Anterior capillaries in two rows only on one
|
||
<typeStatus id="EF579A8F24239F1BF79034BE0D3291E1" box="[1018,1114,189,213]" pageId="3" pageNumber="174" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="56FDBCE824239F1BF00034BF0DA791E0" box="[1130,1231,188,212]" pageId="3" pageNumber="174">MZUCR</collectionCode>
|
||
246-02), all other specimens have capillaries arranged in three rows from chaetiger 5 to 10–11. Neuropodial hooded hooks appear from chaetiger 15–20, 5–7 per fascicle, bi- or tridentate (with two paired apical teeth above the main fang arranged side by side) (
|
||
<figureCitation id="A8D738A824239F1BF55E352408DA9074" box="[308,434,295,320]" captionStart="FIGURE 2" captionStartId="3.[151,244,1934,1956]" captionTargetBox="[166,1426,455,1887]" captionTargetId="figure@3.[151,1435,445,1914]" captionTargetPageId="3" captionText="FIGURE 2. Laonice (Norgensia) costaricensis sp. nov. A. Anterior end, dorsal view; B. Posterior end with pygidium, ventral view; C–F. Parapodia of chaetigers 2, 7, 24 and 42; G–H. Hooded hook from chaetiger 42, side and frontal view. A–B—Holotype (MCZ 101143); C–G—Paratype (MZUCR 348-05).Scale:A–B, D–F—0.5 mm; C—0.3 mm; G—0.05 mm." httpUri="https://zenodo.org/record/1245062/files/figure.png" pageId="3" pageNumber="174">Fig. 2G–H</figureCitation>
|
||
). Neuropodial sabre chaetae first present on chaetiger 9–12, 1–2 per fascicle.
|
||
</paragraph>
|
||
<paragraph id="3053242D24239F1BF4AD354F0B5090BC" blockId="3.[151,1437,151,392]" pageId="3" pageNumber="174">
|
||
The only pygidium remaining in the material belongs to the
|
||
<typeStatus id="EF579A8F24239F1BF7EF354F0A839051" box="[901,1003,332,357]" pageId="3" pageNumber="174" type="holotype">holotype</typeStatus>
|
||
; it has two large ventral lobes but no dorsal cirri exist or remain (
|
||
<figureCitation id="A8D738A824239F1BF5A5356C0B4390BC" box="[463,555,367,392]" captionStart="FIGURE 2" captionStartId="3.[151,244,1934,1956]" captionTargetBox="[166,1426,455,1887]" captionTargetId="figure@3.[151,1435,445,1914]" captionTargetPageId="3" captionText="FIGURE 2. Laonice (Norgensia) costaricensis sp. nov. A. Anterior end, dorsal view; B. Posterior end with pygidium, ventral view; C–F. Parapodia of chaetigers 2, 7, 24 and 42; G–H. Hooded hook from chaetiger 42, side and frontal view. A–B—Holotype (MCZ 101143); C–G—Paratype (MZUCR 348-05).Scale:A–B, D–F—0.5 mm; C—0.3 mm; G—0.05 mm." httpUri="https://zenodo.org/record/1245062/files/figure.png" pageId="3" pageNumber="174">Figs 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="649374A524239F1BF4FD338D0D0A96EB" httpUri="https://zenodo.org/record/1245062/files/figure.png" pageId="3" pageNumber="174" startId="3.[151,244,1934,1956]" targetBox="[166,1426,455,1887]" targetPageId="3">
|
||
<paragraph id="3053242D24239F1BF4FD338D0D0A96EB" blockId="3.[151,1436,1934,2015]" pageId="3" pageNumber="174">
|
||
<emphasis id="0298F83F24239F1BF4FD338D0BD09690" bold="true" box="[151,696,1934,1956]" pageId="3" pageNumber="174">
|
||
FIGURE 2.
|
||
<taxonomicName id="F7EC5FAE24239F1BF579338D0B0D9690" authority="Sikorski & Pavlova, 2018" authorityName="Sikorski & Pavlova" authorityYear="2018" box="[275,613,1934,1956]" class="Polychaeta" family="Spionidae" genus="Laonice" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spionida" pageId="3" pageNumber="174" phylum="Annelida" rank="species" species="costaricensis" status="sp. nov." subGenus="Norgensia">
|
||
<emphasis id="0298F83F24239F1BF579338D0B0D9690" bold="true" box="[275,613,1934,1956]" italics="true" pageId="3" pageNumber="174">Laonice (Norgensia) costaricensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19AB454424239F1BF601338D0BD09690" box="[619,696,1934,1956]" pageId="3" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Anterior end, dorsal view; B. Posterior end with pygidium, ventral view; C–F. Parapodia of chaetigers 2, 7, 24 and 42; G–H. Hooded hook from chaetiger 42, side and frontal view. A–B—Holotype (MCZ 101143); C–G—Paratype (MZUCR 348-05). Scale: A–B, D–F—0.5 mm; C—0.3 mm; G—0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3053242D24249F1CF4AD34940A639184" blockId="4.[151,1436,151,752]" box="[199,779,151,176]" pageId="4" pageNumber="175">
|
||
<emphasis id="0298F83F24249F1CF4AD349408059184" bold="true" box="[199,365,151,176]" pageId="4" pageNumber="175">Methyl green.</emphasis>
|
||
No special staining pattern detected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78F677A624249F1CF4AD34BF0A4791E0" box="[199,815,188,213]" pageId="4" pageNumber="175" type="distribution">
|
||
<paragraph id="3053242D24249F1CF4AD34BF0A4791E0" blockId="4.[151,1436,151,752]" box="[199,815,188,213]" pageId="4" pageNumber="175">
|
||
<emphasis id="0298F83F24249F1CF4AD34BF083191E1" bold="true" box="[199,345,188,213]" pageId="4" pageNumber="175">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="48FB64BD24249F1CF50D34BF08E791E1" box="[359,399,188,213]" name="Iceland" pageId="4" pageNumber="175">Isla</collectingCountry>
|
||
del Coco, Bahia Chatham,
|
||
<quantity id="F71489C824249F1CF6AE34BF0A4291E0" box="[708,810,188,213]" metricMagnitude="1" metricUnit="m" metricValue="4.45" metricValueMax="6.5" metricValueMin="2.4" pageId="4" pageNumber="175" unit="m" value="44.5" valueMax="65.0" valueMin="24.0">
|
||
24–
|
||
<specimenCount id="26EAEFA424249F1CF68534BF0A4291E0" box="[751,810,188,213]" pageId="4" pageNumber="175" type="generic">65 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78F677A624249F1CF4AD34DC0AA291CC" box="[199,970,223,248]" pageId="4" pageNumber="175" type="etymology">
|
||
<paragraph id="3053242D24249F1CF4AD34DC0AA291CC" blockId="4.[151,1436,151,752]" box="[199,970,223,248]" pageId="4" pageNumber="175">
|
||
<emphasis id="0298F83F24249F1CF4AD34DC082691CC" bold="true" box="[199,334,223,248]" pageId="4" pageNumber="175">Etymology.</emphasis>
|
||
Name is given after the name of the country
|
||
<collectingCountry id="48FB64BD24249F1CF72234E30AAE91CC" box="[840,966,223,248]" name="Costa Rica" pageId="4" pageNumber="175">Costa Rica</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78F677A624249F1CF4AD35070BF593C4" pageId="4" pageNumber="175" type="discussion">
|
||
<paragraph id="3053242D24249F1CF4AD350708E29051" blockId="4.[151,1436,151,752]" pageId="4" pageNumber="175">
|
||
<emphasis id="0298F83F24249F1CF4AD350708539029" bold="true" box="[199,315,260,285]" pageId="4" pageNumber="175">Remarks.</emphasis>
|
||
This species is morphologically close to
|
||
<emphasis id="0298F83F24249F1CF77535060D0F9028" box="[799,1127,260,284]" italics="true" pageId="4" pageNumber="175">
|
||
<taxonomicName id="F7EC5FAE24249F1CF77535060A9A9028" box="[799,1010,260,284]" class="Polychaeta" family="Spionidae" genus="Laonice" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spionida" pageId="4" pageNumber="175" phylum="Annelida" rank="subSpecies" species="lemniscata" subSpecies="give">L. lemniscata give</taxonomicName>
|
||
authority,
|
||
</emphasis>
|
||
described from Southwest
|
||
<collectingCountry id="48FB64BD24249F1CF4FD352408699074" box="[151,257,295,320]" name="Australia" pageId="4" pageNumber="175">Australia</collectingCountry>
|
||
. The latter species was described on a basis of
|
||
<specimenCount id="26EAEFA424249F1CF77C35240ACB9074" box="[790,931,295,320]" pageId="4" pageNumber="175" type="generic">5 specimens</specimenCount>
|
||
, of a similar size to the
|
||
<typeStatus id="EF579A8F24249F1CF0D8352B0D8A9074" box="[1202,1250,296,320]" pageId="4" pageNumber="175">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="F7EC5FAE24249F1CF1EF352A08419050" authority="Sikorski & Pavlova, 2018" authorityName="Sikorski & Pavlova" authorityYear="2018" class="Polychaeta" family="Spionidae" genus="Laonice" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spionida" pageId="4" pageNumber="175" phylum="Annelida" rank="species" species="costaricensis" status="sp. nov.">
|
||
<emphasis id="0298F83F24249F1CF1EF352A08419050" italics="true" pageId="4" pageNumber="175">L. costaricensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19AB454424249F1CF55A354F08E29051" box="[304,394,332,357]" pageId="4" pageNumber="175" rank="species">
|
||
<emphasis id="0298F83F24249F1CF55A354F08E29051" bold="true" box="[304,394,332,357]" pageId="4" pageNumber="175">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="3053242D24249F1CF4AD35730BF593C4" blockId="4.[151,1436,151,752]" pageId="4" pageNumber="175">
|
||
However, according to the original description
|
||
<taxonomicName id="F7EC5FAE24249F1CF68135720AEE90BC" box="[747,902,368,392]" class="Polychaeta" family="Spionidae" genus="Laonice" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spionida" pageId="4" pageNumber="175" phylum="Annelida" rank="species" species="lemniscata">
|
||
<emphasis id="0298F83F24249F1CF68135720AEE90BC" box="[747,902,368,392]" italics="true" pageId="4" pageNumber="175">L. lemniscata</emphasis>
|
||
</taxonomicName>
|
||
does not have anterior capillaries arranged in more than two vertical rows. Nevertheless, communication with Dr Robin Wilson (Senior curator of the marine invertebrate collection in Museum
|
||
<collectingRegion id="F228EACF24249F1CF65E35B40BF990E4" box="[564,657,439,464]" country="Australia" name="Victoria" pageId="4" pageNumber="175">Victoria</collectingRegion>
|
||
, Melbourne,
|
||
<collectingCountry id="48FB64BD24249F1CF74535B40AF090E4" box="[815,920,439,464]" name="Australia" pageId="4" pageNumber="175">Australia</collectingCountry>
|
||
) allowed us to partially clarify the original diagnosis of
|
||
<taxonomicName id="F7EC5FAE24249F1CF54E35DE08D290C0" box="[292,442,476,500]" class="Polychaeta" family="Spionidae" genus="Laonice" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spionida" pageId="4" pageNumber="175" phylum="Annelida" rank="species" species="lemniscata">
|
||
<emphasis id="0298F83F24249F1CF54E35DE08D290C0" box="[292,442,476,500]" italics="true" pageId="4" pageNumber="175">L. lemniscata</emphasis>
|
||
</taxonomicName>
|
||
. Wilson found that in the largest specimen (
|
||
<typeStatus id="EF579A8F24249F1CF7C935DF0D6190C1" box="[931,1033,476,501]" pageId="4" pageNumber="175" type="holotype">holotype</typeStatus>
|
||
) of
|
||
<taxonomicName id="F7EC5FAE24249F1CF05E35DE0DA290C0" box="[1076,1226,476,500]" class="Polychaeta" family="Spionidae" genus="Laonice" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spionida" pageId="4" pageNumber="175" phylum="Annelida" rank="species" species="lemniscata">
|
||
<emphasis id="0298F83F24249F1CF05E35DE0DA290C0" box="[1076,1226,476,500]" italics="true" pageId="4" pageNumber="175">L. lemniscata</emphasis>
|
||
</taxonomicName>
|
||
the notochaetae of chaetigers 3–8 and the neurochaetae on chaetigers 6–8 were arranged in three rows. He allowed the possibility that this is an example of size variability; however almost all smaller specimens of
|
||
<taxonomicName id="F7EC5FAE24249F1CF07436260DB89308" authority="Sikorski & Pavlova, 2018" authorityName="Sikorski & Pavlova" authorityYear="2018" box="[1054,1232,549,572]" class="Polychaeta" family="Spionidae" genus="Laonice" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spionida" pageId="4" pageNumber="175" phylum="Annelida" rank="species" species="costaricensis" status="sp. nov.">
|
||
<emphasis id="0298F83F24249F1CF07436260DB89308" box="[1054,1232,549,572]" italics="true" pageId="4" pageNumber="175">L. costaricensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19AB454424249F1CF0B236270C5C9309" box="[1240,1332,548,573]" pageId="4" pageNumber="175" rank="species">
|
||
<emphasis id="0298F83F24249F1CF0B236270C5C9309" bold="true" box="[1240,1332,548,573]" pageId="4" pageNumber="175">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
have the anterior capillaries arranged in 3 rows. Moreover, the presence of transdorsal crests is clearly mentioned in the original description of
|
||
<taxonomicName id="F7EC5FAE24249F1CF5C7366E0B2393B0" box="[429,587,620,644]" class="Polychaeta" family="Spionidae" genus="Laonice" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spionida" pageId="4" pageNumber="175" phylum="Annelida" rank="species" species="lemniscata">
|
||
<emphasis id="0298F83F24249F1CF5C7366E0B2393B0" box="[429,587,620,644]" italics="true" pageId="4" pageNumber="175">L. lemniscata</emphasis>
|
||
</taxonomicName>
|
||
, whereas specimens of
|
||
<taxonomicName id="F7EC5FAE24249F1CF71E366E0D4493B0" authority="Sikorski & Pavlova, 2018" authorityName="Sikorski & Pavlova" authorityYear="2018" box="[884,1068,621,644]" class="Polychaeta" family="Spionidae" genus="Laonice" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spionida" pageId="4" pageNumber="175" phylum="Annelida" rank="species" species="costaricensis" status="sp. nov.">
|
||
<emphasis id="0298F83F24249F1CF71E366E0D4493B0" box="[884,1068,621,644]" italics="true" pageId="4" pageNumber="175">L. costaricensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19AB454424249F1CF050366F0DF393B1" box="[1082,1179,620,645]" pageId="4" pageNumber="175" rank="species">
|
||
<emphasis id="0298F83F24249F1CF050366F0DF393B1" bold="true" box="[1082,1179,620,645]" pageId="4" pageNumber="175">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
have no pronounced transdorsal crests. The absence of well-pronounced transdorsal crests is the main distinguishing feature from other species in this subgenus. Paired apical teeth above the main fang in hooded hooks of
|
||
<taxonomicName id="F7EC5FAE24249F1CF00736B60C7793F8" authorityName="Sikorski & Pavlova" authorityYear="2018" box="[1133,1311,693,716]" class="Polychaeta" family="Spionidae" genus="Laonice" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spionida" pageId="4" pageNumber="175" phylum="Annelida" rank="species" species="costaricensis">
|
||
<emphasis id="0298F83F24249F1CF00736B60C7793F8" box="[1133,1311,693,716]" italics="true" pageId="4" pageNumber="175">L. costaricensis</emphasis>
|
||
</taxonomicName>
|
||
is also the character distinguishing it from
|
||
<emphasis id="0298F83F24249F1CF59436DA0BF593C4" box="[510,669,728,752]" italics="true" pageId="4" pageNumber="175">
|
||
<taxonomicName id="F7EC5FAE24249F1CF59436DA0BF193C4" box="[510,665,728,752]" class="Polychaeta" family="Spionidae" genus="Laonice" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spionida" pageId="4" pageNumber="175" phylum="Annelida" rank="species" species="lemniscata">L. lemniscata</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |