treatments-xml/data/AE/04/87/AE0487C8F538FFD30986A77E657AF8D9.xml
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<document ID-DOI="10.11646/zootaxa.5256.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7745281" ID-ZooBank="CE0AC7CD-E55D-484C-9346-86F16C985DBE" approvalRequired="29" approvalRequired_for_taxonomicNames="9" approvalRequired_for_textStreams="19" approvalRequired_for_treatments="1" checkinTime="1679054310250" checkinUser="plazi" docAuthor="Serrano, Artur R. M." docDate="2023" docId="AE0487C8F538FFD30986A77E657AF8D9" docLanguage="en" docName="zootaxa.5256.1.1.pdf" docOrigin="Zootaxa 5256 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrocheilus basilewskyi Serrano 2000" docType="treatment" docVersion="2" lastPageNumber="12" masterDocId="523DFFB0F533FFD80911A31C612BFFA8" masterDocTitle="Afrotropical ground beetles of Macrocheilus Hope, 1838 (Coleoptera, Carabidae) description of four new species and faunistic notes" masterLastPageNumber="35" masterPageNumber="1" pageNumber="12" updateTime="1679055118150" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Afrotropical ground beetles of Macrocheilus Hope, 1838 (Coleoptera, Carabidae) description of four new species and faunistic notes</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Serrano, Artur R. M.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-4282-6073</mods:nameIdentifier>
<mods:affiliation>cE 3 c-Center for Ecology, Evolution and Environmental Changes &amp; CHANGE - Global Change and Sustainability Institute, Departamento de Biologia Animal, Faculdade de Ciências da Universidade de Lisboa, R. Ernesto de Vasconcelos, Ed. C 2, Campo Grande, 1749 - 016 Lisboa, PORTUGAL. aserrano @ fc. ul. pt; http: // orcid. org / 0000 - 0003 - 4282 - 6073</mods:affiliation>
<mods:nameIdentifier type="email">aserrano@fc.ul.pt</mods:nameIdentifier>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-16</mods:number>
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<mods:detail type="volume">
<mods:number>5256</mods:number>
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<mods:identifier type="DOI">10.11646/zootaxa.5256.1.1</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7745281</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:AE0487C8F538FFD30986A77E657AF8D9" httpUri="http://treatment.plazi.org/id/AE0487C8F538FFD30986A77E657AF8D9" lastPageNumber="12" pageId="11" pageNumber="12">
<subSubSection box="[151,644,1122,1149]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph blockId="11.[151,644,1122,1185]" box="[151,644,1122,1149]" pageId="11" pageNumber="12">
<heading bold="true" box="[151,644,1122,1149]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName ID-CoL="72SB5" authority="Serrano, 2000" authorityName="Serrano" authorityYear="2000" box="[151,644,1122,1149]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis bold="true" box="[151,644,1122,1149]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[151,462,1122,1149]" italics="true" pageId="11" pageNumber="12">Macrocheilus basilewskyi</emphasis>
<bibRefCitation author="Serrano, A. R. M." box="[469,644,1122,1148]" pageId="11" pageNumber="12" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano, 2000</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,321,1159,1185]" pageId="11" pageNumber="12" type="description">
<paragraph blockId="11.[151,644,1122,1185]" box="[151,321,1159,1185]" pageId="11" pageNumber="12">
<figureCitation box="[151,243,1159,1185]" captionStart="FIGURE 1" captionStartId="7.[152,255,1916,1940]" captionTargetBox="[165,1321,160,1875]" captionTargetId="figure-80@7.[151,1340,151,1890]" captionTargetPageId="7" captionText="FIGURE 1. Habitus of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (holotype); d, M. schuelei n. sp. (holotype); e, M. allardi (holotype, plus labels); f, M. alluaudi (holotype, plus labels); g, M. alluaudi, CON (PSC); h, M. basilewskyi (holotype, plus labels); i, M. biguttatus (MRAC, plus labels)." figureDoi="http://doi.org/10.5281/zenodo.7745283" httpUri="https://zenodo.org/record/7745283/files/figure.png" pageId="11" pageNumber="12">Figs. 1h</figureCitation>
,
<figureCitation box="[255,282,1159,1185]" captionStart="FIGURE 2" captionStartId="8.[152,255,1463,1487]" captionTargetBox="[162,1421,162,1440]" captionTargetId="figure-282@8.[151,1436,151,1453]" captionTargetPageId="8" captionText="FIGURE 2. Labrum of Macrocheilus (dorsal view). a, M. assmanni n. sp. (holotype); b, M. kudrnai n. sp. (holotype); c, M. philippeorum n. sp. (paratype); d, M. schuelei n. sp. (holotype); e, M. alluaudi, CON (PSC); f, M. basilewskyi, TAN (TAC); g, M. biguttatus, SEN (ACC); h, M. bimaculatus, RCA (TAC); i, M. clasispilus, RCI (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745285" httpUri="https://zenodo.org/record/7745285/files/figure.png" pageId="11" pageNumber="12">2f</figureCitation>
,
<figureCitation box="[293,321,1159,1185]" captionStart="FIGURE 4" captionStartId="13.[152,255,1917,1941]" captionTargetBox="[203,1362,163,1877]" captionTargetId="figure-76@13.[197,1378,151,1890]" captionTargetPageId="13" captionText="FIGURE 4. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. assmanni (holotype); b, M. philippeorum (paratype); c, M. schuelei (paratype); d, M. basilewskyi, ZIM (TAC); e, M. biguttatus, ETI (TAC); f, M. biguttatus, SEN (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745289" httpUri="https://zenodo.org/record/7745289/files/figure.png" pageId="11" pageNumber="12">4d</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="materials_examined">
<paragraph blockId="11.[151,1437,1230,1905]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[151,361,1230,1256]" pageId="11" pageNumber="12">Studied material.</emphasis>
<specimenCount box="[371,454,1231,1257]" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
: “
<collectingCountry box="[477,580,1231,1257]" name="Tanzania" pageId="11" pageNumber="12">Tanzania</collectingCountry>
3/
<date box="[612,734,1231,1257]" pageId="11" pageNumber="12" value="1997-12-06">6.XII.1997</date>
\ near Babati\ leg. Werner &amp; Lizler” \\ “Collection\ Th. Assmann\ Bleckede, CAB” (yellow label) \\ “
<taxonomicName authorityName="Serrano" authorityYear="2023" box="[635,788,1267,1293]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis box="[635,788,1267,1293]" italics="true" pageId="11" pageNumber="12">Macrocheilus</emphasis>
</taxonomicName>
\
<taxonomicName authorityName="Serrano" authorityYear="2000" box="[805,934,1267,1293]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis box="[805,934,1267,1293]" italics="true" pageId="11" pageNumber="12">basilewskyi</emphasis>
</taxonomicName>
\
<bibRefCitation author="Serrano, A. R. M." box="[951,1112,1267,1293]" pageId="11" pageNumber="12" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano, 2000</bibRefCitation>
” [h]\ “A.Serrano det. 2014” [t] [h];
<specimenCount box="[235,339,1303,1329]" pageId="11" pageNumber="12" type="female">1 female</specimenCount>
: “
<collectingCountry box="[361,464,1303,1329]" name="Tanzania" pageId="11" pageNumber="12">Tanzania</collectingCountry>
10/
<date box="[509,643,1303,1329]" pageId="11" pageNumber="12" value="1999-12-16">16.XII.1999</date>
\ Near Mitundo,
<collectingRegion box="[837,1000,1303,1329]" country="Tanzania" name="Dodoma" pageId="11" pageNumber="12">Dodoma Prov.</collectingRegion>
\ leg. Werner &amp; Lizler” \\ “Collection\ Th. Assmann\ Bleckede, CAB” (yellow label)\\ “
<taxonomicName authorityName="Serrano" authorityYear="2023" box="[709,862,1339,1365]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis box="[709,862,1339,1365]" italics="true" pageId="11" pageNumber="12">Macrocheilus</emphasis>
</taxonomicName>
cf.\
<taxonomicName authority="Serrano" authorityName="Serrano" authorityYear="2000" box="[915,1143,1339,1365]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis box="[915,1044,1339,1365]" italics="true" pageId="11" pageNumber="12">basilewskyi</emphasis>
Serrano
</taxonomicName>
” [h] \ “det. Th. Assmann, 2006” [t]\\ “
<taxonomicName authorityName="Serrano" authorityYear="2023" box="[293,446,1375,1401]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis box="[293,446,1375,1401]" italics="true" pageId="11" pageNumber="12">Macrocheilus</emphasis>
</taxonomicName>
\
<taxonomicName authorityName="Serrano" authorityYear="2000" box="[473,602,1375,1401]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis box="[473,602,1375,1401]" italics="true" pageId="11" pageNumber="12">basilewskyi</emphasis>
</taxonomicName>
\
<bibRefCitation author="Serrano, A. R. M." box="[619,781,1375,1401]" pageId="11" pageNumber="12" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano, 2000</bibRefCitation>
” [h]\ A.Serrano det. 2014” [t] [h];
<specimenCount box="[1187,1269,1375,1401]" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
: “
<collectingCountry box="[1292,1351,1375,1401]" name="South Africa" pageId="11" pageNumber="12">RSA</collectingCountry>
, 25/
<date pageId="11" pageNumber="12" value="2001-11-27">27. XI.2001</date>
\ Thabazimbi, N. Pr.\ leg. Werner &amp; Lizler” \\ “Collection\ Th. Assmann\ Bleckede, CAB” (yellow label) \\ “
<taxonomicName authorityName="Serrano" authorityYear="2023" box="[187,340,1447,1473]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis box="[187,340,1447,1473]" italics="true" pageId="11" pageNumber="12">Macrocheilus</emphasis>
</taxonomicName>
\
<taxonomicName authorityName="Klug" authorityYear="1834" box="[356,446,1447,1473]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis box="[356,446,1447,1473]" italics="true" pageId="11" pageNumber="12">dorsalis</emphasis>
</taxonomicName>
\or\
<taxonomicName authority="Serrano" authorityName="Serrano" authorityYear="2000" box="[485,712,1447,1473]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis box="[485,614,1447,1473]" italics="true" pageId="11" pageNumber="12">basilewskyi</emphasis>
Serrano
</taxonomicName>
” [h] \ det. Th. Assmann, 2006” [t]\\“
<taxonomicName authorityName="Serrano" authorityYear="2023" box="[1131,1284,1447,1473]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis box="[1131,1284,1447,1473]" italics="true" pageId="11" pageNumber="12">Macrocheilus</emphasis>
</taxonomicName>
\
<taxonomicName authorityName="Serrano" authorityYear="2000" box="[1299,1428,1447,1473]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis box="[1299,1428,1447,1473]" italics="true" pageId="11" pageNumber="12">basilewskyi</emphasis>
</taxonomicName>
\
<bibRefCitation author="Serrano, A. R. M." box="[151,310,1483,1509]" pageId="11" pageNumber="12" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano, 2000</bibRefCitation>
” [h]\ “A.Serrano det. 2014” [t] [h];
<specimenCount box="[714,792,1483,1509]" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
,
<specimenCount box="[804,906,1483,1509]" pageId="11" pageNumber="12" type="female">1 female</specimenCount>
: “
<collectingCountry box="[926,1048,1483,1509]" name="Zimbabwe" pageId="11" pageNumber="12">Zimbabwe</collectingCountry>
W\
<collectingRegion box="[1096,1212,1483,1509]" country="Zimbabwe" name="Bulawayo" pageId="11" pageNumber="12">Bulawayo</collectingRegion>
, Shangani\ Naletale ruins env\
<date box="[268,396,1519,1545]" pageId="11" pageNumber="12" value="2011-12-20">20.12.2011</date>
, Snižek” \\ “Coll Wrase\ Berlin” (green labels) \\ “
<taxonomicName authorityName="Serrano" authorityYear="2023" box="[957,1110,1519,1545]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis box="[957,1110,1519,1545]" italics="true" pageId="11" pageNumber="12">Macrocheilus</emphasis>
</taxonomicName>
\
<taxonomicName authorityName="Serrano" authorityYear="2000" box="[1124,1253,1519,1545]" class="Insecta" family="Carabidae" genus="Macrocheilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis box="[1124,1253,1519,1545]" italics="true" pageId="11" pageNumber="12">basilewskyi</emphasis>
</taxonomicName>
\
<bibRefCitation author="Serrano, A. R. M." box="[1267,1426,1519,1545]" pageId="11" pageNumber="12" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano, 2000</bibRefCitation>
” [h]\ “A.Serrano det. 2014” [t] [h].
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="12" type="discussion">
<paragraph blockId="11.[151,1437,1230,1905]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[199,315,1591,1617]" pageId="11" pageNumber="12">Remarks.</emphasis>
The species was described by
<bibRefCitation author="Serrano, A. R. M." box="[669,840,1591,1617]" pageId="11" pageNumber="12" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano (2000)</bibRefCitation>
based on
<specimenCount box="[957,1125,1591,1617]" pageId="11" pageNumber="12" type="generic">two specimens</specimenCount>
from
<collectingCountry box="[1195,1281,1591,1617]" name="Angola" pageId="11" pageNumber="12">Angola</collectingCountry>
. The material now examined allows extending its distribution to central and eastern Africa, being first reported for the
<collectingCountry name="South Africa" pageId="11" pageNumber="12">Republic of South Africa</collectingCountry>
,
<collectingCountry box="[337,438,1663,1689]" name="Tanzania" pageId="11" pageNumber="12">Tanzania</collectingCountry>
and
<collectingCountry box="[492,615,1663,1689]" name="Zimbabwe" pageId="11" pageNumber="12">Zimbabwe</collectingCountry>
.
</paragraph>
<paragraph blockId="11.[151,1437,1230,1905]" pageId="11" pageNumber="12">
<emphasis box="[199,365,1699,1725]" italics="true" pageId="11" pageNumber="12">Male genitalia</emphasis>
. Description and drawings of median lobe in lateral and dorsal views and left paramere shape were provided by
<bibRefCitation author="Serrano, A. R. M." box="[354,516,1735,1761]" pageId="11" pageNumber="12" pagination="137 - 145" refId="ref25943" refString="Serrano, A. R. M. (2000) Beetle species of Macrocheilus Hope, 1838 from Africa: Faunistic and description of new species (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie, Nouvelle Serie, 17 (2), 137 - 145." type="journal article" year="2000">Serrano (2000</bibRefCitation>
,
<figureCitation box="[528,620,1735,1761]" captionStart="FIGURE 13" captionStartId="27.[152,255,872,896]" captionTargetBox="[370,1262,159,835]" captionTargetId="figure-117@27.[365,1272,151,843]" captionTargetPageId="27" captionText="FIGURE 13. Aedeagus of Macrocheilus: median lobe and right paramere (left, lateral view) and median lobe (right, dorsal view). a, M. persimilis, ZAM (TAC); b, M. scapularis (MRAC); c, M. spectandus, ZAM (TAC); d, M. viduatus katanganus, ZAM (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745311" httpUri="https://zenodo.org/record/7745311/files/figure.png" pageId="11" pageNumber="12">Figs. 13</figureCitation>
,
<figureCitation box="[632,663,1735,1761]" captionStart="FIGURE 14" captionStartId="27.[152,255,1916,1940]" captionTargetBox="[421,1245,991,1878]" captionTargetId="figure-120@27.[413,1253,983,1890]" captionTargetPageId="27" captionText="FIGURE 14. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. proximus, ETI (TAC);; b, M. quadrinotatus, ANG (ASC); c, M. taedatus, TAN (TAC); d, M. viduatus katanganus, RSA (PSC)." figureDoi="http://doi.org/10.5281/zenodo.7745313" httpUri="https://zenodo.org/record/7745313/files/figure.png" pageId="11" pageNumber="12">14</figureCitation>
).
</paragraph>
<paragraph blockId="11.[151,1437,1230,1905]" pageId="11" pageNumber="12">
<emphasis box="[199,390,1771,1797]" italics="true" pageId="11" pageNumber="12">Female genitalia</emphasis>
(
<figureCitation box="[405,489,1771,1797]" captionStart="FIGURE 4" captionStartId="13.[152,255,1917,1941]" captionTargetBox="[203,1362,163,1877]" captionTargetId="figure-76@13.[197,1378,151,1890]" captionTargetPageId="13" captionText="FIGURE 4. Right gonopods (gonocoxite 1 and 2) of female genitalia of Macrocheilus: gonocoxite 2 (left, dorsal view) and gonocoxites 1 and 2 (right, ventral view). a, M. assmanni (holotype); b, M. philippeorum (paratype); c, M. schuelei (paratype); d, M. basilewskyi, ZIM (TAC); e, M. biguttatus, ETI (TAC); f, M. biguttatus, SEN (ACC)." figureDoi="http://doi.org/10.5281/zenodo.7745289" httpUri="https://zenodo.org/record/7745289/files/figure.png" pageId="11" pageNumber="12">Fig. 4d</figureCitation>
). Gonocoxite 1 subtriangular; gonocoxite 2 largely sickle shaped, apex hardly point- ed; dorsal surface concave, proximal half covered with numerous setae, subapical inner margin with four setae; ventral surface slightly convex, some long and minor setae near outer margin in apical half.
</paragraph>
</subSubSection>
<subSubSection box="[199,1105,1878,1905]" pageId="11" pageNumber="12" type="distribution">
<paragraph blockId="11.[151,1437,1230,1905]" box="[199,1105,1878,1905]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[199,355,1878,1904]" pageId="11" pageNumber="12">Distribution:</emphasis>
<collectingCountry box="[362,447,1879,1905]" name="Angola" pageId="11" pageNumber="12">Angola</collectingCountry>
,
<collectingCountry box="[458,557,1879,1905]" name="Namibia" pageId="11" pageNumber="12">Namibia</collectingCountry>
,
<collectingCountry box="[569,852,1879,1905]" name="South Africa" pageId="11" pageNumber="12">Republic of South Africa</collectingCountry>
,
<collectingCountry box="[863,967,1879,1905]" name="Tanzania" pageId="11" pageNumber="12">Tanzania</collectingCountry>
,
<collectingCountry box="[977,1100,1879,1905]" name="Zimbabwe" pageId="11" pageNumber="12">Zimbabwe</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>