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<document id="5AAC4779B949CD7D7B3DE870CC9F6BBC" ID-CLB-Dataset="274469" ID-DOI="10.11646/zootaxa.5357.4.5" ID-GBIF-Dataset="b71b135c-e7a8-4f91-add9-d9132736e878" ID-ISSN="1175-5326" ID-Zenodo-Dep="10064893" ID-ZooBank="122F5432-1468-4BE6-9566-73083903A050" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1697788260247" checkinUser="plazi" docAuthor="Jałoszyński, Paweł, Maruyama, Munetoshi &amp; Klimaszewski, Jan" docDate="2023" docId="336587DDB93B0030F9E7F89F51D70513" docLanguage="en" docName="zootaxa.5357.4.5.pdf" docOrigin="Zootaxa 5357 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5357.4.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wowini Jaloszynski, Maruyama &amp; Klimaszewski 2023, trib. n." docType="treatment" docVersion="5" lastPageNumber="575" masterDocId="CF5CFFA5B93A0032F970FF91503E064E" masterDocTitle="Wow assingi gen. and spec. n., with description of a new tribe of Aleocharinae (Coleoptera, Staphylinidae)" masterLastPageNumber="586" masterPageNumber="573" pageNumber="574" updateTime="1698876130098" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A2A579C5F89E2F2D8FD0CD3B17B46BEE">Wow assingi gen. and spec. n., with description of a new tribe of Aleocharinae (Coleoptera, Staphylinidae)</mods:title>
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<mods:namePart id="EE46F43EF31A2D9A31BB3FB5FF4F99E8">Jałoszyński, Paweł</mods:namePart>
<mods:affiliation id="783CA7E47FCFBD7522761932B647E97E">Museum of Natural History, University of Wrocław, Sienkiewicza 21, 50 - 335 Wrocław, Poland</mods:affiliation>
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<mods:namePart id="1B31F4FD89341BA38A9DDEAB068AD7C2">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="E2305B41129B29DACE6D2972C22C6494">The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581 Japan</mods:affiliation>
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<mods:namePart id="902862B4CE99A423B61AB00C18A64DCF">Klimaszewski, Jan</mods:namePart>
<mods:affiliation id="966119103BF0F3059402D27E4C865CE0">3407 Sparrowhawk Ave., Victoria, BC, Canada V 9 C 0 L 9 (emeritus scientist: Natural Resources Canada, Pacific Forestry Centre, 506 West Burnside Road, Victoria, BC, Canada V 8 Z)</mods:affiliation>
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<mods:title id="2633142FE401975076D998B973519874">Zootaxa</mods:title>
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<mods:date id="F8DD43508D895D09DD1D7FA55B806306">2023</mods:date>
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<mods:number id="826861B1F2016FCB4526A95A4FCB6462">2023-10-20</mods:number>
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<treatment id="336587DDB93B0030F9E7F89F51D70513" ID-GBIF-Taxon="214288137" LSID="urn:lsid:plazi:treatment:336587DDB93B0030F9E7F89F51D70513" httpUri="http://treatment.plazi.org/id/336587DDB93B0030F9E7F89F51D70513" lastPageId="2" lastPageNumber="575" pageId="1" pageNumber="574">
<subSubSection id="F3D66540B93B0033F9E7F89F53920167" box="[151,940,1806,1833]" pageId="1" pageNumber="574" type="nomenclature">
<paragraph id="BB7336CBB93B0033F9E7F89F53920167" blockId="1.[151,1111,1806,1869]" box="[151,940,1806,1833]" pageId="1" pageNumber="574">
<heading id="E03B81A7B93B0033F9E7F89F53920167" bold="true" box="[151,940,1806,1833]" fontSize="11" level="1" pageId="1" pageNumber="574" reason="1">
<emphasis id="89B8EAD9B93B0033F9E7F89F53920167" bold="true" box="[151,940,1806,1833]" pageId="1" pageNumber="574">
Tribe
<taxonomicName id="7CCC4D48B93B0033F992F89F53760166" authority="Jaloszynski, Maruyama &amp; Klimaszewski" authorityName="Jaloszynski, Maruyama &amp; Klimaszewski" authorityYear="2023" box="[226,840,1806,1833]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="574" phylum="Arthropoda" rank="tribe" status="trib. n." tribe="Wowini">Wowini Jałoszyński, Maruyama &amp; Klimaszewski</taxonomicName>
,
<taxonomicNameLabel id="928B57A2B93B0033FA26F89F53920167" box="[854,940,1806,1833]" pageId="1" pageNumber="574" rank="tribe">trib. n.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="F3D66540B93B0033F9E7F8A254690103" box="[151,1111,1843,1869]" pageId="1" pageNumber="574" type="materials_examined">
<paragraph id="BB7336CBB93B0033F9E7F8A254690103" blockId="1.[151,1111,1806,1869]" box="[151,1111,1843,1869]" pageId="1" pageNumber="574">
<typeStatus id="64778869B93B0033F9E7F8A250F10103" box="[151,207,1843,1869]" pageId="1" pageNumber="574">Type</typeStatus>
genus:
<taxonomicName id="7CCC4D48B93B0033F856F8A2530E0103" authority="Jaloszynski, Maruyama &amp; Klimaszewski" authorityName="Jaloszynski, Maruyama &amp; Klimaszewski" authorityYear="2023" box="[294,816,1843,1869]" class="Insecta" family="Staphylinidae" genus="Wow" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="574" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis id="89B8EAD9B93B0033F856F8A251650102" box="[294,347,1843,1868]" italics="true" pageId="1" pageNumber="574">Wow</emphasis>
Jałoszyński, Maruyama &amp; Klimaszewski
</taxonomicName>
,
<taxonomicNameLabel id="928B57A2B93B0033FA4BF8A253B90103" box="[827,903,1843,1869]" pageId="1" pageNumber="574" rank="genus">gen. n.</taxonomicNameLabel>
(here designated).
</paragraph>
</subSubSection>
<subSubSection id="F3D66540B93B0030F9E7F8EB5485072B" lastPageId="2" lastPageNumber="575" pageId="1" pageNumber="574" type="diagnosis">
<paragraph id="BB7336CBB93B0030F9E7F8EB5485072B" blockId="1.[151,1436,1914,2013]" lastBlockId="2.[151,1437,151,861]" lastPageId="2" lastPageNumber="575" pageId="1" pageNumber="574">
<emphasis id="89B8EAD9B93B0033F9E7F8EB512C01DA" bold="true" box="[151,274,1914,1940]" pageId="1" pageNumber="574">Diagnosis.</emphasis>
The following unique combination of character states defines this tribe: maxillary palpus with pseudosegment (
<figureCitation id="23F72A4EB93B0033F825F80E518001F7" box="[341,446,1951,1977]" captionStart="FIGURES 6–11" captionStartId="5.[151,264,1709,1733]" captionTargetBox="[151,1435,525,1685]" captionTargetId="figure-175@5.[151,1435,525,1685]" captionTargetPageId="5" captionText="FIGURES 6–11. Wow assingi sp. n., holotype male. Mouthparts in dorsal (6) and ventral (9) views; right maxilla in ventral view (7, 8); labium in ventral view (10, 11). Abbreviations: aps, apical ‘pseudosegment’; atp, anterior tentorial pit; bst, basistipes; cd, cardo; cl, clypeus; cmb, clypeolabral connecting membrane; gal, galea; hr, hypostomal ridge; hs, hypostomal suture; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; mdb, mandible; mn, mentum; msc, median sclerotization of prelabium; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pat, preapical tooth; ppg, palpiger; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10064901" httpUri="https://zenodo.org/record/10064901/files/figure.png" pageId="1" pageNumber="574">Figs 7–8</figureCitation>
); labial palpus trimerous with palpomere 3 large, apically dilated (
<figureCitation id="23F72A4EB93B0033FDB1F80E557B01F7" box="[1217,1349,1951,1977]" captionStart="FIGURES 6–11" captionStartId="5.[151,264,1709,1733]" captionTargetBox="[151,1435,525,1685]" captionTargetId="figure-175@5.[151,1435,525,1685]" captionTargetPageId="5" captionText="FIGURES 6–11. Wow assingi sp. n., holotype male. Mouthparts in dorsal (6) and ventral (9) views; right maxilla in ventral view (7, 8); labium in ventral view (10, 11). Abbreviations: aps, apical ‘pseudosegment’; atp, anterior tentorial pit; bst, basistipes; cd, cardo; cl, clypeus; cmb, clypeolabral connecting membrane; gal, galea; hr, hypostomal ridge; hs, hypostomal suture; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; mdb, mandible; mn, mentum; msc, median sclerotization of prelabium; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pat, preapical tooth; ppg, palpiger; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10064901" httpUri="https://zenodo.org/record/10064901/files/figure.png" pageId="1" pageNumber="574">Figs 10–11</figureCitation>
); ligula undivided, broadly subconical with broadly rounded apex, strongly sclerotized (
<figureCitation id="23F72A4EB93B0033FD69F85254A40193" box="[1049,1178,1987,2013]" captionStart="FIGURES 6–11" captionStartId="5.[151,264,1709,1733]" captionTargetBox="[151,1435,525,1685]" captionTargetId="figure-175@5.[151,1435,525,1685]" captionTargetPageId="5" captionText="FIGURES 6–11. Wow assingi sp. n., holotype male. Mouthparts in dorsal (6) and ventral (9) views; right maxilla in ventral view (7, 8); labium in ventral view (10, 11). Abbreviations: aps, apical ‘pseudosegment’; atp, anterior tentorial pit; bst, basistipes; cd, cardo; cl, clypeus; cmb, clypeolabral connecting membrane; gal, galea; hr, hypostomal ridge; hs, hypostomal suture; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; mdb, mandible; mn, mentum; msc, median sclerotization of prelabium; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pat, preapical tooth; ppg, palpiger; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10064901" httpUri="https://zenodo.org/record/10064901/files/figure.png" pageId="1" pageNumber="574">Figs 10–11</figureCitation>
); lateral lobes of labial apodeme (component “r” of premental sclerite in
<bibRefCitation id="DF5D4B3AB9380030FBACFF0653F606FF" author="Weide, D. &amp; Thayer, M. K. &amp; Betz, O." box="[732,968,151,177]" pageId="2" pageNumber="575" pagination="84 - 110" refId="ref9036" refString="Weide, D., Thayer, M. K. &amp; Betz, O. (2014) Comparative morphology of the tentorium and hypopharyngeal-premental sclerites in sporophagous and non-sporophagous adult Aleocharinae (Coleoptera: Staphylinidae). Acta Zoologica, 95, 84 - 110. https: // doi. org / 10.1111 / azo. 12011" type="journal article" year="2014">
Weide
<emphasis id="89B8EAD9B9380030FA5DFF06535706FF" box="[813,873,151,177]" italics="true" pageId="2" pageNumber="575">et al.</emphasis>
(2014))
</bibRefCitation>
short (
<figureCitation id="23F72A4EB9380030FD6DFF06549F06FF" box="[1053,1185,151,177]" captionStart="FIGURES 6–11" captionStartId="5.[151,264,1709,1733]" captionTargetBox="[151,1435,525,1685]" captionTargetId="figure-175@5.[151,1435,525,1685]" captionTargetPageId="5" captionText="FIGURES 6–11. Wow assingi sp. n., holotype male. Mouthparts in dorsal (6) and ventral (9) views; right maxilla in ventral view (7, 8); labium in ventral view (10, 11). Abbreviations: aps, apical ‘pseudosegment’; atp, anterior tentorial pit; bst, basistipes; cd, cardo; cl, clypeus; cmb, clypeolabral connecting membrane; gal, galea; hr, hypostomal ridge; hs, hypostomal suture; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; mdb, mandible; mn, mentum; msc, median sclerotization of prelabium; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pat, preapical tooth; ppg, palpiger; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10064901" httpUri="https://zenodo.org/record/10064901/files/figure.png" pageId="2" pageNumber="575">Figs 10–11</figureCitation>
); prelabium with two pairs of medial setae inserted on inversely drop-shaped median premental sclerotization (
<figureCitation id="23F72A4EB9380030FDD2FF2A551A069B" box="[1186,1316,187,213]" captionStart="FIGURES 6–11" captionStartId="5.[151,264,1709,1733]" captionTargetBox="[151,1435,525,1685]" captionTargetId="figure-175@5.[151,1435,525,1685]" captionTargetPageId="5" captionText="FIGURES 6–11. Wow assingi sp. n., holotype male. Mouthparts in dorsal (6) and ventral (9) views; right maxilla in ventral view (7, 8); labium in ventral view (10, 11). Abbreviations: aps, apical ‘pseudosegment’; atp, anterior tentorial pit; bst, basistipes; cd, cardo; cl, clypeus; cmb, clypeolabral connecting membrane; gal, galea; hr, hypostomal ridge; hs, hypostomal suture; lac, lacinia; lbr, labrum; lig, ligula; llh, lateral lobe of hypopharynx; lp1‒3, labial palpomere 1‒3; mdb, mandible; mn, mentum; msc, median sclerotization of prelabium; mst, mediostipes; mxp1‒4, maxillary palpomere 1‒4; pat, preapical tooth; ppg, palpiger; pst, prostheca; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10064901" httpUri="https://zenodo.org/record/10064901/files/figure.png" pageId="2" pageNumber="575">Figs 10–11</figureCitation>
) (possible autapomorphy); prothoracic hypomeron fully exposed in posterior half in lateral view; mesocoxal rests not separated, with mesoventral and anterior metaventral intermesocoxal processes both short, subtriangular and pointed at apices (
<figureCitation id="23F72A4EB9380030F9EFFEB650C8070F" box="[159,246,295,321]" captionStart="FIGURES 12–18" captionStartId="6.[151,264,1659,1683]" captionTargetBox="[151,1435,489,1636]" captionTargetId="figure-103@6.[151,1435,489,1636]" captionTargetPageId="6" captionText="FIGURES 12–18. Wow assingi sp. n., holotype male. Elytra in dorsal view (12); pterothorax in ventral view (13); protarsus in ventral view (14); metatarsus in lateral view (15); exposed portion of abdomen in dorsal view (16); abdominal apex in dorsal (17) and ventral (18) views. Abbreviations: aest3, metanepisternum; msvp, mesoventral process; scs, scutellar shield; sIII‒VIII, sternite III‒VIII; tII‒X, tergite 2‒10; v2, mesoventrite; v3, metaventrite; 1‒4, tarsomere 1‒4. Arrowhead indicates macroseta." figureDoi="http://doi.org/10.5281/zenodo.10064903" httpUri="https://zenodo.org/record/10064903/files/figure.png" pageId="2" pageNumber="575">Fig. 13</figureCitation>
); tarsal formula 4-4-4 (
<figureCitation id="23F72A4EB9380030FB7CFEB652AD070F" box="[524,659,294,321]" captionStart="FIGURES 12–18" captionStartId="6.[151,264,1659,1683]" captionTargetBox="[151,1435,489,1636]" captionTargetId="figure-103@6.[151,1435,489,1636]" captionTargetPageId="6" captionText="FIGURES 12–18. Wow assingi sp. n., holotype male. Elytra in dorsal view (12); pterothorax in ventral view (13); protarsus in ventral view (14); metatarsus in lateral view (15); exposed portion of abdomen in dorsal view (16); abdominal apex in dorsal (17) and ventral (18) views. Abbreviations: aest3, metanepisternum; msvp, mesoventral process; scs, scutellar shield; sIII‒VIII, sternite III‒VIII; tII‒X, tergite 2‒10; v2, mesoventrite; v3, metaventrite; 1‒4, tarsomere 1‒4. Arrowhead indicates macroseta." figureDoi="http://doi.org/10.5281/zenodo.10064903" httpUri="https://zenodo.org/record/10064903/files/figure.png" pageId="2" pageNumber="575">Figs 14–15</figureCitation>
); median lobe of aedeagus (
<figureCitation id="23F72A4EB9380030FA91FEB6545B070F" box="[993,1125,295,321]" captionStart="FIGURES 19–26" captionStartId="8.[151,264,1620,1644]" captionTargetBox="[164,1424,186,1592]" captionTargetId="figure-20@8.[158,1430,180,1598]" captionTargetPageId="8" captionText="FIGURES 19–26. Wow assingi sp. n., holotype male. Aedeagus in parameral (19, 21) and lateral (20, 22) views; paramere in lateral view (23); tergite VIII (24); sternite VIII 25); tergites IX and X in dorsal view (26). Abbreviations: apl, apical lobe; con, condylite; hz, hinge zone; par, paramerite; tIX‒X, tergite IX‒X; vel, velum; vst, ventral strut." figureDoi="http://doi.org/10.5281/zenodo.10064905" httpUri="https://zenodo.org/record/10064905/files/figure.png" pageId="2" pageNumber="575">Figs 19–20</figureCitation>
) lacking ‘athetine bridge’; paramere with velar sac completely connected (fused) with apical lobe of paramerite (
<figureCitation id="23F72A4EB9380030FD2AFEDA5491072B" box="[1114,1199,331,357]" captionStart="FIGURES 19–26" captionStartId="8.[151,264,1620,1644]" captionTargetBox="[164,1424,186,1592]" captionTargetId="figure-20@8.[158,1430,180,1598]" captionTargetPageId="8" captionText="FIGURES 19–26. Wow assingi sp. n., holotype male. Aedeagus in parameral (19, 21) and lateral (20, 22) views; paramere in lateral view (23); tergite VIII (24); sternite VIII 25); tergites IX and X in dorsal view (26). Abbreviations: apl, apical lobe; con, condylite; hz, hinge zone; par, paramerite; tIX‒X, tergite IX‒X; vel, velum; vst, ventral strut." figureDoi="http://doi.org/10.5281/zenodo.10064905" httpUri="https://zenodo.org/record/10064905/files/figure.png" pageId="2" pageNumber="575">Fig. 23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F3D66540B9380030F9B7FEFF541004E7" pageId="2" pageNumber="575" type="description">
<paragraph id="BB7336CBB9380030F9B7FEFF541004E7" blockId="2.[151,1437,151,861]" pageId="2" pageNumber="575">
<emphasis id="89B8EAD9B9380030F9B7FEFF516707C6" bold="true" box="[199,345,366,392]" pageId="2" pageNumber="575">Description.</emphasis>
Body shape generalized (not limuloid or myrmecoid).Head with postoccipital collar (= ‘neck’) short and narrower than 1/3 of its width, postoccipital ‘suture’ and subocular ridge (= occipital ‘suture’) not discernible in ventral view. Maxillary palpus tetramerous and generalized, except for developed apical pseudosegment. Prementum with median elongate sclerotization bearing two pairs of medial setae. Labial palpus trimerous, with palpomere 3 dilated distally; ligula undivided, subconical with broadly rounded apex, strongly sclerotized. Prothoracic hypomera in posterior half fully exposed in lateral view. Mesocoxal rests adjacent at middle. Mesoventral and anterior metaventral intermesocoxal processes both short subtriangular, apically pointed. Exposed portion of mesoscutellar shield broadly subtriangular and setose. Abdominal paratergites bipartite. Tarsal formula 4-4-4. Median lobe of aedeagus lacking ‘athetine bridge’. Velar sac fused with apical lobe of paramerite.
</paragraph>
</subSubSection>
<subSubSection id="F3D66540B9380030F9B7FD2251D70513" pageId="2" pageNumber="575" type="discussion">
<paragraph id="BB7336CBB9380030F9B7FD2251D70513" blockId="2.[151,1437,151,861]" pageId="2" pageNumber="575">
<emphasis id="89B8EAD9B9380030F9B7FD2251050483" bold="true" box="[199,315,691,717]" pageId="2" pageNumber="575">Remarks.</emphasis>
The bizarre structure of the head and antennae in
<taxonomicName id="7CCC4D48B9380030FAFDFD2254250482" authority="Jałoszyński &amp; Maruyama &amp; Klimaszewski, 2023" authorityName="Jałoszyński &amp; Maruyama &amp; Klimaszewski" authorityYear="2023" box="[909,1051,691,716]" class="Insecta" family="Staphylinidae" genus="Wow" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="species" species="assingi" status="sp. n.">
<emphasis id="89B8EAD9B9380030FAFDFD2254250482" box="[909,1051,691,716]" italics="true" pageId="2" pageNumber="575">Wow assingi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="928B57A2B9380030FD56FD22545B0483" box="[1062,1125,691,717]" pageId="2" pageNumber="575" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="23F72A4EB9380030FD07FD2254E00483" box="[1143,1246,691,717]" captionStart-0="FIGURES 1–3" captionStart-1="FIGURES 4–5" captionStartId-0="3.[151,264,1279,1303]" captionStartId-1="4.[151,264,1612,1636]" captionTargetBox-0="[161,1435,273,1256]" captionTargetBox-1="[151,1435,489,1589]" captionTargetId-0="figure-49@3.[151,1435,273,1256]" captionTargetId-1="figure-164@4.[151,1435,489,1589]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1–3. Wow assingi sp. n., holotype male. Dorsal habitus, (1); head, pronotum and anterior region of elytra in dorsolateral view (2); head in anterodorsal view (3)." captionText-1="FIGURES 4–5. Wow assingi sp. n., holotype male. Head and prothorax in dorsal (4) and ventral (5) views.Abbreviations: acxc, anterior coxal carina; atp, anterior tentorial pit; bst, basisternal region of prosternum; cl clypeus; cmb, clypeolabral connecting membrane; cs, cervical sclerite; fr, frons; gal, galea; gen, gena; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hs, hypostomal suture; lbr, labrum; mdb, mandible; mn, mentum; mxp1‒4, maxillary palpomere 1‒4; nss, notosternal suture; pcxc, posterior coxal carina; prl, prelabium; pst, prostheca; ptp, posterior tentorial pit; sat, supraantennal tubercle; sc, scape; smn, submentum; trt, trochantin; vt, vertex. Arrowheads indicate macrosetae." figureDoi-0="http://doi.org/10.5281/zenodo.10064897" figureDoi-1="http://doi.org/10.5281/zenodo.10064899" httpUri-0="https://zenodo.org/record/10064897/files/figure.png" httpUri-1="https://zenodo.org/record/10064899/files/figure.png" pageId="2" pageNumber="575">Figs 2–4</figureCitation>
) is not included in the tribal diagnosis and description, as it can be an autapomorphy of this peculiar genus. A similar cephalic transformation is known in males and females of a single genus of
<taxonomicName id="7CCC4D48B9380030FA0FFD6A5446055B" authority="Fleming" authorityName="Fleming" box="[895,1144,763,789]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="tribe" tribe="Lomechusini">Lomechusini Fleming</taxonomicName>
, a tribe with all remaining (over 220) genera having unmodified heads. This interesting parallelism and extraordinary head structure are commented in the Discussion.
</paragraph>
</subSubSection>
</treatment>
</document>
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