163 lines
20 KiB
XML
163 lines
20 KiB
XML
<document id="15FC33EB8F7E1C8AF297E2474E4F5E8D" ID-DOI="10.11646/zootaxa.3669.1.3" ID-GBIF-Dataset="cb79db91-29d7-4a4a-92ac-99f85e03a61e" ID-ISSN="1175-5326" ID-Zenodo-Dep="283874" ID-ZooBank="1BF325AD-E5E9-4C02-AB0F-AF2A8DDE8E57" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460628755008" checkinUser="plazi" docAuthor="Kamiński, Marcin Jan & Iwan, Dariusz" docDate="2013" docId="114D87C2433FFF97FF0BF9CBFB8E5AC4" docLanguage="en" docName="zt03669p036.pdf" docOrigin="Zootaxa 3669 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kochogaster impressicollis Fairmaire 1897" docType="treatment" docVersion="6" lastPageNumber="29" masterDocId="ED74FFBA433EFF95FF9CFF89FFD75E37" masterDocTitle="Taxonomy of the genus Kochogaster Kamiński et Raś, 2011 (Coleoptera: Tenebrionidae: Pedinini), with description of a second known species" masterLastPageNumber="36" masterPageNumber="27" pageNumber="28" updateTime="1698334376538" updateUser="plazi">
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<mods:titleInfo id="2D3564276E45567D49DCCBE2543F52A4">
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<mods:title id="27308BF1A344B5F31FE0CCD08AF33627">Taxonomy of the genus Kochogaster Kamiński et Raś, 2011 (Coleoptera: Tenebrionidae: Pedinini), with description of a second known species</mods:title>
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<mods:namePart id="80852968A1F246B90B3F81DC757BD5C4">Kamiński, Marcin Jan</mods:namePart>
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<mods:roleTerm id="543D66F984786B334FF996FC8F099831">Author</mods:roleTerm>
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<mods:namePart id="50C53FAD131D38816C13C797F3A1DA25">Iwan, Dariusz</mods:namePart>
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<mods:typeOfResource id="69852188901A2A197E5974AFDC4ECEC5">text</mods:typeOfResource>
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<mods:title id="59F67A30FD61B6648ECF25406D8D4CDB">Zootaxa</mods:title>
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<mods:part id="329B21141C6A83ECF6A46AA7B097ABF9">
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<mods:date id="AE76E1F01A09900074FE668FB1EE070E">2013</mods:date>
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<mods:number id="60350766AECB3B78135E44B4E9E01909">3669</mods:number>
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<mods:identifier id="1ABB0133F04F36D4D3D72C1028E3BA2B" type="DOI">10.11646/zootaxa.3669.1.3</mods:identifier>
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<treatment id="114D87C2433FFF97FF0BF9CBFB8E5AC4" ID-DOI="http://doi.org/10.5281/zenodo.6146785" ID-GBIF-Taxon="127664281" ID-Zenodo-Dep="6146785" LSID="urn:lsid:plazi:treatment:114D87C2433FFF97FF0BF9CBFB8E5AC4" httpUri="http://treatment.plazi.org/id/114D87C2433FFF97FF0BF9CBFB8E5AC4" lastPageId="2" lastPageNumber="29" pageId="1" pageNumber="28">
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<subSubSection id="D1FE655F433FFF94FF0BF9CBFBDD58F4" pageId="1" pageNumber="28" type="nomenclature">
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<paragraph id="995B36D4433FFF94FF0BF9CBFD6B586B" blockId="1.[151,700,1602,1661]" box="[151,700,1602,1628]" pageId="1" pageNumber="28">
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<heading id="C21381B8433FFF94FF0BF9CBFD6B586B" bold="true" box="[151,700,1602,1628]" fontSize="11" level="1" pageId="1" pageNumber="28" reason="1">
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<taxonomicName id="5EE44D57433FFF94FF0BF9CBFD6B586B" authority="Fairmaire, 1897" authorityName="Fairmaire" authorityYear="1897" box="[151,700,1602,1628]" class="Insecta" family="Curculionidae" genus="Kochogaster" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="impressicollis">
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<emphasis id="AB90EAC6433FFF94FF0BF9CBFD6B586B" bold="true" box="[151,700,1602,1628]" pageId="1" pageNumber="28">
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<emphasis id="AB90EAC6433FFF94FF0BF9CBFE0E586B" bold="true" box="[151,473,1602,1628]" italics="true" pageId="1" pageNumber="28">Kochogaster impressicollis</emphasis>
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(Fairmaire, 1897)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="995B36D4433FFF94FF0BF9ECFE40584A" blockId="1.[151,700,1602,1661]" box="[151,407,1637,1661]" pageId="1" pageNumber="28">
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(
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<figureCitation id="01DF2A51433FFF94FF03F9ECFEC8584A" box="[159,287,1637,1661]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,269,1966,1990]" captionTargetBox="[277,1337,231,1902]" captionTargetId="figure@3.[212,1378,195,1940]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Primary type labels (1); body structure (2, 3); punctuation of pronotal (4, 5) and elytral disc (6, 7) of Kochogaster gerardi sp. nov. (1, 2, 4, 6) and K. impressicollis (3, 5, 7)." httpUri="https://zenodo.org/record/283875/files/figure.png" pageId="1" pageNumber="28">Figs 3, 5, 7</figureCitation>
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, 8–13, 15)
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</paragraph>
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<paragraph id="995B36D4433FFF94FF0BF924FBDD58F4" blockId="1.[151,1200,1709,1793]" box="[151,1034,1709,1731]" pageId="1" pageNumber="28">
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<taxonomicName id="5EE44D57433FFF94FF0BF924FD7C58F4" authority="Fairmaire, 1897: 121" authorityName="Fairmaire" authorityPageNumber="121" authorityYear="1897" box="[151,683,1709,1731]" class="Insecta" family="Tenebrionidae" genus="Anchophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="impressicollis">
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<emphasis id="AB90EAC6433FFF94FF0BF924FE0558F4" box="[151,466,1709,1731]" italics="true" pageId="1" pageNumber="28">Anchophthalmus impressicollis</emphasis>
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Fairmaire, 1897: 121
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</taxonomicName>
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.—Gebien 1910: 278, 1938b: 299.
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</paragraph>
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</subSubSection>
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<subSubSection id="D1FE655F433FFF94FF0BF945FBD35937" pageId="1" pageNumber="28" type="description">
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<paragraph id="995B36D4433FFF94FF0BF945FBD35937" blockId="1.[151,1200,1709,1793]" pageId="1" pageNumber="28">
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<taxonomicName id="5EE44D57433FFF94FF0BF945FDB258D6" authority="Fairmaire, 1897" authorityName="Fairmaire" authorityYear="1897" box="[151,613,1739,1762]" class="Insecta" family="Curculionidae" genus="Cosmogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="impressicollis">
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<emphasis id="AB90EAC6433FFF94FF0BF945FE7858D5" box="[151,431,1740,1762]" italics="true" pageId="1" pageNumber="28">Cosmogaster impressicollis</emphasis>
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(Fairmaire, 1897)
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</taxonomicName>
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.—Koch 1956: 165 (comb.); Iwan 2002a: 64, 2002b: 261.
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<taxonomicName id="5EE44D57433FFF94FF0BF963FDB75937" authority="Fairmaire, 1897" authorityName="Fairmaire" authorityYear="1897" box="[151,608,1770,1792]" class="Insecta" family="Curculionidae" genus="Kochogaster" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="impressicollis">
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<emphasis id="AB90EAC6433FFF94FF0BF963FE7D5937" box="[151,426,1770,1792]" italics="true" pageId="1" pageNumber="28">Kochogaster impressicollis</emphasis>
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(Fairmaire, 1897)
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</taxonomicName>
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.—Kamiński and Raś 2011: 654 (comb.).
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</paragraph>
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</subSubSection>
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<subSubSection id="D1FE655F433FFF97FF0BF8A4FC395A9C" lastPageId="2" lastPageNumber="29" pageId="1" pageNumber="28" type="materials_examined">
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<paragraph id="995B36D4433FFF94FF0BF8A4FC4559E2" blockId="1.[151,1436,1837,2005]" pageId="1" pageNumber="28">
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<emphasis id="AB90EAC6433FFF94FF0BF8A4FEA65971" bold="true" box="[151,369,1837,1862]" pageId="1" pageNumber="28">Studied material.</emphasis>
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Male (HNHM): “
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<taxonomicName id="5EE44D57433FFF94FDFEF8A4FCC95972" box="[610,798,1837,1861]" class="Insecta" family="Tenebrionidae" genus="Anchophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis id="AB90EAC6433FFF94FDFEF8A4FCC95972" box="[610,798,1837,1861]" italics="true" pageId="1" pageNumber="28">Anchophthalmus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="5EE44D57433FFF94FCD1F8A4FC305972" box="[845,999,1837,1861]" class="Insecta" family="Curculionidae" genus="Kochogaster" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="impressicollis">
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<emphasis id="AB90EAC6433FFF94FCD1F8A4FC305972" box="[845,999,1837,1861]" italics="true" pageId="1" pageNumber="28">impressicollis</emphasis>
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</taxonomicName>
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/ det. dr. Kaszab”, “O. Africa / Luidpoldkette”; male (MNHN): “Afr. or. / Luidpoldkette”; female (FMNH): “CNHM 1955 / Karl Brancsik Colln / ex Eduard Knirsch”, “O. Africa”; female (TMNH): “B. E. Africa /
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<collectingCountry id="E1F37644433FFF94FC1DF8FCFC2259BA" box="[897,1013,1909,1933]" name="Kenya" pageId="1" pageNumber="28">Mombasa</collectingCountry>
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”; female (HNHM): “
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<taxonomicName id="5EE44D57433FFF94FB7CF8FCFA4B59BA" box="[1248,1436,1909,1933]" class="Insecta" family="Tenebrionidae" genus="Anchophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis id="AB90EAC6433FFF94FB7CF8FCFA4B59BA" box="[1248,1436,1909,1933]" italics="true" pageId="1" pageNumber="28">Anchophthalmus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="5EE44D57433FFF94FF3BF810FE965986" box="[167,321,1945,1969]" class="Insecta" family="Curculionidae" genus="Kochogaster" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="impressicollis">
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<emphasis id="AB90EAC6433FFF94FF3BF810FE965986" box="[167,321,1945,1969]" italics="true" pageId="1" pageNumber="28">impressicollis</emphasis>
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</taxonomicName>
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/ Fairmaire”; female (MRAC): “Musée du
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<collectingCountry id="E1F37644433FFF94FCA8F810FCA85986" box="[820,895,1945,1969]" name="Republic of the Congo" pageId="1" pageNumber="28">Congo</collectingCountry>
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/ B.E.A.: Tsavo R. /
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<date id="ED5A1014433FFF94FBEFF810FB1C5986" box="[1139,1227,1945,1969]" pageId="1" pageNumber="28" value="1921-05-04">4/21 V-</date>
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1913 / Dr. Bayer”, “R. Det / 5428”, “
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<taxonomicName id="5EE44D57433FFF94FEFDF834FDCD59E2" box="[353,538,1981,2005]" class="Insecta" family="Tenebrionidae" genus="Anchophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="36" phylum="Arthropoda" rank="genus">
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<emphasis id="AB90EAC6433FFF94FEFDF834FDCD59E2" box="[353,538,1981,2005]" italics="true" pageId="1" pageNumber="28">anchophthalmus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="5EE44D57433FFF94FDACF834FD1C59E2" box="[560,715,1981,2005]" class="Insecta" family="Curculionidae" genus="Kochogaster" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="impressicollis">
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<emphasis id="AB90EAC6433FFF94FDACF834FD1C59E2" box="[560,715,1981,2005]" italics="true" pageId="1" pageNumber="28">impressicollis</emphasis>
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</taxonomicName>
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F. / det. Kueger”.
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</paragraph>
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<paragraph id="995B36D4433CFF97FF5BFF1EFBF45FCE" blockId="2.[151,1437,151,1267]" pageId="2" pageNumber="29">
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<emphasis id="AB90EAC6433CFF97FF5BFF1EFEA35E87" bold="true" box="[199,372,151,176]" pageId="2" pageNumber="29">Redescription.</emphasis>
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Measurements. Body—length (male) = 13.5–14.0 mm; length (female) = 16.0–
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<quantity id="5E1C9B31433CFF97FAB0FF1EFA4F5E98" box="[1324,1432,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="2" pageNumber="29" unit="mm" value="17.5">17.5 mm</quantity>
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; width ratio elytra / pronotum = 1.0–1.1; length ratio elytra / middle of pronotum = 2.3–2.6. Clypeus—clypeal emargination width / depth ratio ca. 9.0. Antennae—length ratio of antennomere 3rd / 2nd ca. 2.0. Maxillary palpratio length of apical segment of maxillary palp/3rd antennal segment = 1.2–1.3 (female), 1.8-1.9 (male). Pronotum—middle of pronotum length / width ratio = 0.6–0.7; length ratio middle of pronotum length/anterior pronotal angles (from tip of anterior pronotal angle to tip of posterior pronotal angle) = 0.8–0.9. Elytra—elytra length / width ratio = 1.3–1.5. Scutellum—width ratio anterior margin of elytra / scutellum at base ca. 3.5. Metaventrite—length ratio cavity of hind coxa/metaventrite between the insertions of mid and hind coxae = 1.9– 2.0. Abdomen—width ratio process of I abdominal ventrite / process of metaventrite ca. 2.0. Both sexes legsprotibiae length / width ratio = 4.6–5.0; profemora length / width ratio = 3.5–3.6.
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</paragraph>
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<paragraph id="995B36D4433CFF97FF5BFD8CFE7E5C2A" blockId="2.[151,1437,151,1267]" box="[199,425,517,541]" pageId="2" pageNumber="29">
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Habitus as in
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<figureCitation id="01DF2A51433CFF97FEFCFD8CFE745C2A" box="[352,419,517,541]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,269,1966,1990]" captionTargetBox="[277,1337,231,1902]" captionTargetId="figure@3.[212,1378,195,1940]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Primary type labels (1); body structure (2, 3); punctuation of pronotal (4, 5) and elytral disc (6, 7) of Kochogaster gerardi sp. nov. (1, 2, 4, 6) and K. impressicollis (3, 5, 7)." httpUri="https://zenodo.org/record/283875/files/figure.png" pageId="2" pageNumber="29">Fig. 3</figureCitation>
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.
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</paragraph>
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<paragraph id="995B36D4433CFF97FF5BFDA0FB615CBE" blockId="2.[151,1437,151,1267]" pageId="2" pageNumber="29">Dorsal side of head shiny, with conspicuous punctures (less than 1 diameter apart). Frontoclypeal suture distinct. Clypeal emargination shallow. Eye slightly constricted in the middle by epistomal canthus (3 facets between gena and temple). Mentum with median part narrow. Submentum with elongate base.</paragraph>
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<paragraph id="995B36D4433CFF97FF5BFD1CFE9E5D0A" blockId="2.[151,1437,151,1267]" pageId="2" pageNumber="29">
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Pronotal disc shiny, with visible punctures (0.5–1 diameters apart). Anterior pronotal angles sharp, strongly protruding towards apex. Anterior border of pronotum interrupted in the middle. Lateral sides marginated; rounded on the whole length. Posterior pronotal angles sharp; protruding towards elytra. Pronotal base with lateral arcuate indentations; marginated laterally (
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<figureCitation id="01DF2A51433CFF97FDABFC88FD455D2E" box="[567,658,769,793]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,269,1916,1940]" captionTargetBox="[231,1354,215,1890]" captionTargetId="figure@5.[225,1362,194,1895]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Structure of the chosen body elements: pronotal (13) and elytral (14) base; upper edge of elytral epipleuron with visible elytral slope (15, 16). Kochogaster impressicollis (13, 15), K. gerardi sp. nov. (14), Anchophthalmops maximus Koch, 1956 (16)." httpUri="https://zenodo.org/record/283876/files/figure.png" pageId="2" pageNumber="29">Fig. 13</figureCitation>
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). Pronotal hypomeron shiny, with conspicuous punctures (0.5–2 diameter apart).
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</paragraph>
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<paragraph id="995B36D4433CFF97FF5BFCC0FB0F5D9B" blockId="2.[151,1437,151,1267]" pageId="2" pageNumber="29">Elytral rows with coarse punctures (1 diameters apart). Intervals shiny, with fine punctures (2–3 diameters apart). Elytral base sinusoidal; intervals from 4th to 6th forms an appendix at elytral base. Elytral humeri rounded, not protruding laterad. Scutellum triangular. Epipleura with visible punctures (2 diameters apart).</paragraph>
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<paragraph id="995B36D4433CFF97FF5BFC31FBE45DE7" blockId="2.[151,1437,151,1267]" box="[199,1075,952,976]" pageId="2" pageNumber="29">
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Intercoxal process of prosternum straight with incomplete bordering (
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<figureCitation id="01DF2A51433CFF97FC4EFC31FBF15DE7" box="[978,1062,952,976]" captionStart="FIGURES 17 – 19" captionStartId="6.[151,269,1914,1938]" captionTargetBox="[177,1409,199,1885]" captionTargetId="figure@6.[170,1417,194,1893]" captionTargetPageId="6" captionText="FIGURES 17 – 19. Structure of the chosen body elements: intercoxal process of prosternum (17), mentum and submentum (18, 19). Kochogaster impressicollis (17, 18), Anchophthalmus muellerae (19)." httpUri="https://zenodo.org/record/283877/files/figure.png" pageId="2" pageNumber="29">Fig. 17</figureCitation>
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).
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</paragraph>
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<paragraph id="995B36D4433CFF97FF5BFC55FF375A2F" blockId="2.[151,1437,151,1267]" pageId="2" pageNumber="29">Female abdominal process without tubercle. Punctuation of to first abdominal ventrites sparse (2 diameters apart).</paragraph>
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<paragraph id="995B36D4433CFF97FF5BFBACFAB45A0B" blockId="2.[151,1437,151,1267]" box="[199,1379,1060,1085]" pageId="2" pageNumber="29">Legs. Pro-, meso- and metatarsi narrow. Legs without denticles. Male mesotibiae flattened dorsoventraly.</paragraph>
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<paragraph id="995B36D4433CFF97FF5BFBC0FAAB5A56" blockId="2.[151,1437,151,1267]" box="[199,1404,1096,1121]" pageId="2" pageNumber="29">Parameres and clavae narrowing evenly towards apex. Parameres equal to the 0.33 rest of aedeagal tegmen.</paragraph>
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<paragraph id="995B36D4433CFF97FF5BFBF9FC395A9C" blockId="2.[151,1437,151,1267]" pageId="2" pageNumber="29">Paraproct slightly longer then coxites. First sclerite wide and short; sclerites 3rd and 4th narrow and long. Bursa copulatrix with ring-shaped sclerite in distal part. Spermatheca as in Fig. 12.</paragraph>
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</subSubSection>
|
||
<subSubSection id="D1FE655F433CFF97FF5BFB3FFB8E5AC4" pageId="2" pageNumber="29" type="distribution">
|
||
<paragraph id="995B36D4433CFF97FF5BFB3FFB8E5AC4" blockId="2.[151,1437,151,1267]" pageId="2" pageNumber="29">
|
||
<emphasis id="AB90EAC6433CFF97FF5BFB3FFEB75AF8" bold="true" box="[199,352,1206,1231]" pageId="2" pageNumber="29">Distribution.</emphasis>
|
||
This species has been collected in the following ecoregions: Northern
|
||
<taxonomicName id="5EE44D57433CFF97FB2DFB3EFB285AF8" box="[1201,1279,1207,1231]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="29" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="AB90EAC6433CFF97FB2DFB3EFB285AF8" box="[1201,1279,1207,1231]" italics="true" pageId="2" pageNumber="29">Acacia</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<taxonomicName id="5EE44D57433CFF97FA94FB3EFA4B5AF8" box="[1288,1436,1207,1231]" class="Magnoliopsida" family="Burseraceae" genus="Commiphora" kingdom="Plantae" order="Sapindales" pageId="2" pageNumber="29" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="AB90EAC6433CFF97FA94FB3EFA4B5AF8" box="[1288,1436,1207,1231]" italics="true" pageId="2" pageNumber="29">Commiphora</emphasis>
|
||
</taxonomicName>
|
||
bushlands and thickets, Northern Zanzibar-Inhambane coastal forest mosaic (
|
||
<figureCitation id="01DF2A51433CFF97FC64FB52FB9B5AC4" box="[1016,1100,1243,1267]" captionStart="FIGURE 20" captionStartId="7.[151,255,1616,1640]" captionTargetBox="[187,1339,249,1615]" captionTargetId="figure@7.[170,1417,181,1615]" captionTargetPageId="7" captionText="FIGURE 20. Distribution of the species of Kochogaster. The division of Afrotropical Realm into ecoregions was adopted after Olson et al. 2001. Different hues of green were used to distinguished the adjacent ecoregions." httpUri="https://zenodo.org/record/283878/files/figure.png" pageId="2" pageNumber="29">Fig. 20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |