242 lines
42 KiB
XML
242 lines
42 KiB
XML
<document id="4478AD0999F477ED121E57372341BDB1" ID-CLB-Dataset="30222" ID-DOI="10.11646/zootaxa.4413.1.9" ID-GBIF-Dataset="038432ac-12c2-4e26-ba94-e5da796df109" ID-ISSN="1175-5326" ID-Zenodo-Dep="1222020" ID-ZooBank="8879B612-4B17-4EF3-8C8B-7AA4A4A35C08" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1524199432412" checkinUser="plazi" docAuthor="Tan, Ming Kai & Wahab, Rodzay Bin Haji Abdul" docDate="2018" docId="A56F7A3CFFF1FFF8FF79351EFC42A701" docLanguage="en" docName="zootaxa.4413.1.9.pdf" docOrigin="Zootaxa 4413 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tapiena paraincisa Tan & Wahab 2018, new species" docType="treatment" docVersion="7" lastPageNumber="196" masterDocId="59560244FFF0FFFBFFEE343CFF97A52D" masterDocTitle="A new species of Tapiena (Orthoptera: Tettigonioidea: Phaneropterinae) from Brunei Darussalam (Borneo, Southeast Asia)" masterLastPageNumber="196" masterPageNumber="193" pageNumber="194" updateTime="1698540648623" updateUser="ExternalLinkService">
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<mods:mods id="6BF51901E1D0278A0E400F2250AE353A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="72DD4CDE8FB6EECFF42AC33802BA79CA">
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<mods:title id="7D519B31EF45F4AD8A425DD746D409E3">A new species of Tapiena (Orthoptera: Tettigonioidea: Phaneropterinae) from Brunei Darussalam (Borneo, Southeast Asia)</mods:title>
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</mods:titleInfo>
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<mods:name id="AD7561508D93CC1EE7643B7B84E6E164" type="personal">
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<mods:roleTerm id="D3B15D65750967F16D11E0362D9A5C77">Author</mods:roleTerm>
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<mods:namePart id="2D03C2DDF624D132E095C4B8727DB11A">Tan, Ming Kai</mods:namePart>
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</mods:name>
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<mods:name id="DD729CF1C2B67D3D0B5F91F14DBDA7CE" type="personal">
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<mods:role id="A19F803D2AC6AF0F00463EB56F4B2039">
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<mods:roleTerm id="AF6E999BA10D06B4C0CD59A48108F479">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2F42919A837389B6722DBB875268F44B">Wahab, Rodzay Bin Haji Abdul</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="138BA6C24214572C66CD6206500DF6F3">text</mods:typeOfResource>
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<mods:relatedItem id="6EA5D1511FEABFC3F0AD8D7EF7E3917C" type="host">
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<mods:titleInfo id="BA28B5466D3B5688389112F4B3A35F1E">
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<mods:title id="FE0D21EE2954B35644F7316624D0F1E8">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="6962E03E6E450C25F3172004769471F0">
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<mods:date id="B9679BB0E6997EA2E38EF907BB03A99E">2018</mods:date>
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<mods:detail id="BF49C381D246DFCD43609B37B2B0BDF5" type="pubDate">
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<mods:number id="6FDC5C66CD4A2B81E9799A4E9EA719D3">2018-04-20</mods:number>
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</mods:detail>
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<mods:detail id="2C08DD289FB63A2B77DE5CD2E624238B" type="volume">
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<mods:number id="5BCD5AC0DC1B370B0F2B14136AA72C67">4413</mods:number>
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</mods:detail>
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<mods:detail id="09037ADFF6976DC7AA19515B5E33A9E2" type="issue">
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<mods:number id="E250DDF6CBF56A316622998D50E930AA">1</mods:number>
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<mods:end id="C4B5328F81C8358E9083D1715A09972C">196</mods:end>
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</mods:extent>
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<mods:classification id="01DAB75F81CDD5168F90154CC2FC6422">journal article</mods:classification>
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<mods:identifier id="0217B5489641D1A96686AF9CF14575C3" type="DOI">10.11646/zootaxa.4413.1.9</mods:identifier>
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<mods:identifier id="E458187B351C5E04C3522AEB7401AA94" type="GBIF-Dataset">038432ac-12c2-4e26-ba94-e5da796df109</mods:identifier>
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<mods:identifier id="AE69858FB8D793D04DDE330EC206F051" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="A56F7A3CFFF1FFF8FF79351EFC42A701" ID-DOI="http://doi.org/10.5281/zenodo.6492223" ID-GBIF-Taxon="143619766" ID-Zenodo-Dep="6492223" LSID="urn:lsid:plazi:treatment:A56F7A3CFFF1FFF8FF79351EFC42A701" httpUri="http://treatment.plazi.org/id/A56F7A3CFFF1FFF8FF79351EFC42A701" lastPageId="3" lastPageNumber="196" pageId="1" pageNumber="194">
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<subSubSection id="65DC98A1FFF1FFFAFF79351EFC64A471" pageId="1" pageNumber="194" type="nomenclature">
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<paragraph id="2D79CB2AFFF1FFFAFF79351EFDEAA411" blockId="1.[151,1011,290,348]" box="[151,637,290,316]" pageId="1" pageNumber="194">
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<heading id="76317C46FFF1FFFAFF79351EFDEAA411" bold="true" box="[151,637,290,316]" fontSize="11" level="1" pageId="1" pageNumber="194" reason="1">
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<emphasis id="1FB21738FFF1FFFAFF79351EFDEAA411" bold="true" box="[151,637,290,316]" pageId="1" pageNumber="194">
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<taxonomicName id="EAC6B0A9FFF1FFFAFF79351EFEE9A411" ID-CoL="54SPQ" authority="Tan & Wahab, 2018" authorityName="Tan & Wahab" authorityYear="2018" box="[151,382,290,316]" class="Insecta" family="Tettigoniidae" genus="Tapiena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="194" phylum="Arthropoda" rank="species" species="paraincisa" status="sp. nov.">
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<emphasis id="1FB21738FFF1FFFAFF79351EFEE9A411" bold="true" box="[151,382,290,316]" italics="true" pageId="1" pageNumber="194">Tapiena paraincisa</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="0481AA43FFF1FFFAFE62351EFD8BA411" box="[396,540,290,316]" pageId="1" pageNumber="194" rank="species">new species</taxonomicNameLabel>
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(
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<figureCitation id="B5FDD7AFFFF1FFFAFDC2351EFDE4A411" box="[556,627,290,316]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="2D79CB2AFFF1FFFAFF793579FC64A471" blockId="1.[151,1011,290,348]" box="[151,1011,325,348]" pageId="1" pageNumber="194">
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http://lsid.speciesfile.org/urn:lsid:
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<taxonomicName id="EAC6B0A9FFF1FFFAFE193579FDE6A471" box="[503,625,325,348]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="194" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
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.speciesfile.org:TaxonName:502556
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</paragraph>
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</subSubSection>
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<subSubSection id="65DC98A1FFF1FFFAFF7935B6FB71A701" pageId="1" pageNumber="194" type="materials_examined">
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<paragraph id="2D79CB2AFFF1FFFAFF7935B6FCFEA4CA" blockId="1.[151,1436,394,2012]" pageId="1" pageNumber="194">
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<materialsCitation id="9DAEC177FFF1FFFAFF7935B6FCFEA4CA" ID-GBIF-Occurrence="1838188858" collectingDate="2016-09-23" collectionCode="IBER" collectorName="M. K. Tan" country="Brunei" latitude="4.54727" location="Kuala Belalong Field Studies Centre" longLatPrecision="1" longitude="115.15804" municipality="Material" pageId="1" pageNumber="194" specimenCount="1" stateProvince="Temburong" typeStatus="holotype">
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<emphasis id="1FB21738FFF1FFFAFF7935B6FEE1A48C" bold="true" box="[151,374,394,417]" pageId="1" pageNumber="194">
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<collectingMunicipality id="CD1D5150FFF1FFFAFF7935B6FF6DA48C" box="[151,250,394,417]" pageId="1" pageNumber="194">Material</collectingMunicipality>
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examined.
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</emphasis>
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<typeStatus id="F27D7588FFF1FFFAFE9335B6FE77A48C" box="[381,480,394,417]" pageId="1" pageNumber="194" type="holotype">Holotype</typeStatus>
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(male, KB.16.23):
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<collectingCountry id="55D18BBAFFF1FFFAFD5D35B6FC07A48C" box="[691,912,394,417]" name="Brunei" pageId="1" pageNumber="194">
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<emphasis id="1FB21738FFF1FFFAFD5D35B6FC07A48C" bold="true" box="[691,912,394,417]" pageId="1" pageNumber="194">Brunei Darussalam</emphasis>
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</collectingCountry>
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, Ulu
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<collectingRegion id="EF0205C8FFF1FFFAFC2135B6FBDEA48C" box="[975,1097,394,417]" country="Brunei" name="Temburong" pageId="1" pageNumber="194">Temburong</collectingRegion>
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National Park,
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<location id="28199DF1FFF1FFFAFB1A35B6FEE1A4E9" LSID="urn:lsid:plazi:treatment:A56F7A3CFFF1FFF8FF79351EFC42A701:28199DF1FFF1FFFAFB1A35B6FEE1A4E9" country="Brunei" latitude="4.54727" longLatPrecision="1" longitude="115.15804" municipality="Material" name="Kuala Belalong Field Studies Centre" pageId="1" pageNumber="194" stateProvince="Temburong">Kuala Belalong Field Studies Centre</location>
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, primary ridge dipterocarp forest,
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<geoCoordinate id="48F2ADEDFFF1FFFAFD043591FCC1A4E9" box="[746,854,429,452]" direction="north" orientation="latitude" pageId="1" pageNumber="194" precision="1" value="4.54727">N4.54727</geoCoordinate>
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,
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<geoCoordinate id="48F2ADEDFFF1FFFAFC8C3591FC71A4E9" box="[866,998,429,452]" direction="east" orientation="longitude" pageId="1" pageNumber="194" precision="1" value="115.15804">E115.15804</geoCoordinate>
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, 115.0±9.0 m.a.s.l., attracted to light,
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<collectingDate id="493C1402FFF1FFFAFA6F3591FED0A4CA" pageId="1" pageNumber="194" value="2016-09-23">
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<date id="5978EDEAFFF1FFFAFA6F3591FED0A4CA" pageId="1" pageNumber="194" value="2016-09-23">23 September 2016</date>
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</collectingDate>
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, 2230 hours, coll.
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<collectorName id="8033AEFCFFF1FFFAFDE235ECFDEFA4CA" box="[524,632,464,487]" pageId="1" pageNumber="194">M. K. Tan</collectorName>
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(dried pinned,
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<collectionCode id="4BD753EFFFF1FFFAFCF235ECFCC9A4CA" box="[796,862,464,487]" pageId="1" pageNumber="194">IBER</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="2D79CB2AFFF1FFFAFF2935CEFB71A701" blockId="1.[151,1436,394,2012]" pageId="1" pageNumber="194">
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<emphasis id="1FB21738FFF1FFFAFF2935CEFEE3A724" bold="true" box="[199,372,498,521]" pageId="1" pageNumber="194">Generic status.</emphasis>
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This species looks like
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<taxonomicName id="EAC6B0A9FFF1FFFAFD6D35CEFD69A724" authorityName="Stål" authorityYear="1873" box="[643,766,498,521]" class="Insecta" family="Tettigoniidae" genus="Holochlora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="194" phylum="Arthropoda" rank="genus">
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<emphasis id="1FB21738FFF1FFFAFD6D35CEFD69A724" box="[643,766,498,521]" italics="true" pageId="1" pageNumber="194">Holochlora</emphasis>
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</taxonomicName>
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in habitus but differs by area around stridulatory file of left tegmen not swollen, tenth abdominal tergite not bilobate, and head and pronotum distinctly punctuated.
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</paragraph>
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</subSubSection>
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<subSubSection id="65DC98A1FFF1FFFAFF293604FF7FA684" pageId="1" pageNumber="194" type="diagnosis">
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<paragraph id="2D79CB2AFFF1FFFAFF293604FEF5A79A" blockId="1.[151,1436,394,2012]" pageId="1" pageNumber="194">
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<emphasis id="1FB21738FFF1FFFAFF293604FEABA762" bold="true" box="[199,316,568,591]" pageId="1" pageNumber="194">Diagnosis.</emphasis>
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This species is typical of the genus but differs from all other congeners by: unicolourous green but slightly red brown on fore femur near knee and fore tibia around tympanal area; tenth abdominal tergite with posterior margin emarginated laterally, swollen in the middle, deeply and narrowly notched; cercus long with two lamellate teeth of subacute apices.
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</paragraph>
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<paragraph id="2D79CB2AFFF1FFFAFF2936FEFF7FA684" blockId="1.[151,1436,394,2012]" pageId="1" pageNumber="194">
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<emphasis id="1FB21738FFF1FFFAFF2936FEFD85A7F4" bold="true" box="[199,530,706,729]" pageId="1" pageNumber="194">Comparison with congeners.</emphasis>
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The male of this new species is most similar to
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<taxonomicName id="EAC6B0A9FFF1FFFAFBDF36FEFAC9A7F4" authority="Karny, 1923" authorityName="Karny" authorityYear="1923" box="[1073,1374,706,729]" class="Insecta" family="Tettigoniidae" genus="Tapiena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="194" phylum="Arthropoda" rank="species" species="incisa">
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<emphasis id="1FB21738FFF1FFFAFBDF36FEFB58A7F4" box="[1073,1231,706,729]" italics="true" pageId="1" pageNumber="194">Tapiena incisa</emphasis>
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<bibRefCitation id="4957B6DBFFF1FFFAFB3636FEFAC9A7F4" box="[1240,1374,706,729]" pageId="1" pageNumber="194" refId="ref2200" refString="Karny, H. H. (1923) On Malaysian Katydids (Gryllacridae and Tettigoniidae), from the Raffles Museum, Singapore. Journal of the Malayan Branch of the Royal Asiatic Society, 1 (1), 116 - 193." type="journal article">Karny, 1923</bibRefCitation>
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</taxonomicName>
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from Gunung Tamabo (= Tama Abu Range), Baram River (
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<collectingRegion id="EF0205C8FFF1FFFAFD3B36D9FCA2A7D1" box="[725,821,741,764]" country="Malaysia" name="Sarawak" pageId="1" pageNumber="194">
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<collectingCountry id="55D18BBAFFF1FFFAFD3B36D9FCA2A7D1" box="[725,821,741,764]" name="Malaysia" pageId="1" pageNumber="194">Sarawak</collectingCountry>
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</collectingRegion>
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, Borneo) by tenth abdominal tergite incised in the middle along the hind margin; but differs by cercus with two lamellate teeth of subacute apices (instead of obtusely flattened with an obtuse tooth at the apex in
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<taxonomicName id="EAC6B0A9FFF1FFFAFDF83716FDE7A66C" authorityName="Karny" authorityYear="1923" box="[534,624,810,833]" class="Insecta" family="Tettigoniidae" genus="Tapiena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="194" phylum="Arthropoda" rank="species" species="incisa">
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<emphasis id="1FB21738FFF1FFFAFDF83716FDE7A66C" box="[534,624,810,833]" italics="true" pageId="1" pageNumber="194">T. incisa</emphasis>
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</taxonomicName>
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), and subgenital plate with shorter styli. The new species is also similar to
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<taxonomicName id="EAC6B0A9FFF1FFFAFF793771FD92A649" authority="Liu & Xia, 1996" authorityName="Liu & Xia" authorityYear="1996" box="[151,517,845,868]" class="Insecta" family="Tettigoniidae" genus="Tapiena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="194" phylum="Arthropoda" rank="species" species="simplicis">
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<emphasis id="1FB21738FFF1FFFAFF793771FEC6A649" box="[151,337,845,868]" italics="true" pageId="1" pageNumber="194">Tapiena simplicis</emphasis>
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Liu & Xia, 1996
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</taxonomicName>
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from
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<collectingRegion id="EF0205C8FFF1FFFAFDAB3771FDEBA649" box="[581,636,845,868]" country="China" name="Xizang" pageId="1" pageNumber="194">Tibet</collectingRegion>
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by tenth abdominal tergite without apical process, cercus with apical hook/ tooth; but differs by cercus not expanded apically, subgenital plate with posterior margin more deeply excised in the middle.
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</paragraph>
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</subSubSection>
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<subSubSection id="65DC98A1FFF1FFFAFF293789FBB1A6C2" pageId="1" pageNumber="194" type="discussion">
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<paragraph id="2D79CB2AFFF1FFFAFF293789FBB1A6C2" blockId="1.[151,1436,394,2012]" pageId="1" pageNumber="194">
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<emphasis id="1FB21738FFF1FFFAFF293789FED1A6E1" bold="true" box="[199,326,949,972]" pageId="1" pageNumber="194">Discussion.</emphasis>
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Although only one specimen was collected, the geographical or morphological discontinuities with similar congeners are distinct, and are unlikely an artefact of intra-specific variations.
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</paragraph>
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</subSubSection>
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<subSubSection id="65DC98A1FFF1FFF8FF2937C6FECEA4CA" lastPageId="3" lastPageNumber="196" pageId="1" pageNumber="194" type="description">
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<paragraph id="2D79CB2AFFF1FFFAFF2937C6FE93A389" blockId="1.[151,1436,394,2012]" pageId="1" pageNumber="194">
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<emphasis id="1FB21738FFF1FFFAFF2937C6FEC5A13C" bold="true" box="[199,338,1018,1041]" pageId="1" pageNumber="194">Description.</emphasis>
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Habitus as shown in
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<figureCitation id="B5FDD7AFFFF1FFFAFDD137C6FD03A13C" box="[575,660,1018,1041]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1A</figureCitation>
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, typical of genus. Head with length circa 0.4 times of pronotum length (
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<figureCitation id="B5FDD7AFFFF1FFFAFF713021FF60A119" box="[159,247,1053,1076]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1B</figureCitation>
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). Fastigium narrowly triangular, short, feebly surpassing anterior apex of eyes in dorsal view; furrowed longitudinally and with a transverse median sulcus in the middle; anterior of sulcus distinctly narrower than antennal scapus and pedicel (uncharacteristic of the genus), posterior of sulcus wider than antennal scapus; with apex roundly truncated (
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<figureCitation id="B5FDD7AFFFF1FFFAFEE630B9FECDA1B1" box="[264,346,1157,1180]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1B</figureCitation>
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). Eye prominent; median ocellus prominent, round and large (
|
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<figureCitation id="B5FDD7AFFFF1FFFAFC0030B9FBD7A1B1" box="[1006,1088,1157,1180]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1C</figureCitation>
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). Frons slightly punctuated (
|
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<figureCitation id="B5FDD7AFFFF1FFFAFA9E30B9FF20A192" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1C</figureCitation>
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); dorsum of head (
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<figureCitation id="B5FDD7AFFFF1FFFAFE603094FE75A192" box="[398,482,1192,1215]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1B</figureCitation>
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) and genae (
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<figureCitation id="B5FDD7AFFFF1FFFAFD9A3094FD5DA192" box="[628,714,1192,1215]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1D</figureCitation>
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) densely punctuated. Pronotum circa 1.3 times longer than wide, densely punctuated (
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<figureCitation id="B5FDD7AFFFF1FFFAFE9C30F6FE6AA1CC" box="[370,509,1226,1249]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Figs. 1B, 1D</figureCitation>
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). Dorsal plate with anterior margin straight; lateral margins straight and nearly parallel (feebly diverging posteriorly); posterior margin broadly rounded, feebly notched in the middle; hind lobe after middle, separated by shallow suture; longitudinal furrow absent (
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<figureCitation id="B5FDD7AFFFF1FFFAFD19312CFCDDA00A" box="[759,842,1296,1319]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1B</figureCitation>
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). Lateral keel present. Lateral lobe circa as tall as long; ventral margin short, and posterior margin substraight, anterior margin slightly sinuous (
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<figureCitation id="B5FDD7AFFFF1FFFAFB84310EFB55A064" box="[1130,1218,1330,1353]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1D</figureCitation>
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). Thoracic auditory spiracle [= thoracic foramen] broadly elongated, covered by pronotal lateral lobe (
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<figureCitation id="B5FDD7AFFFF1FFFAFBD33169FB00A041" box="[1085,1175,1365,1388]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1D</figureCitation>
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). Prosternum unarmed. Mesosternum triangular with subacute apex; metasternum rectangular with anterior angle obtusely rounded, posterior angle 90° and rounded. Tegmen and hind wing fully developed. Tegmen with anterior and posterior margins feebly convex, widens in the middle before tapering to rounded apex; appears punctuated with many small transparent cells. Tegminal venation: Costal vein (Cu) present but not distinct (
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<figureCitation id="B5FDD7AFFFF1FFFAFCC931DCFCEFA0DA" box="[807,888,1504,1527]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1E</figureCitation>
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). Radius sector (Rs) originates before middle, with fork longer than stem (
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<figureCitation id="B5FDD7AFFFF1FFFAFE65323EFE77A334" box="[395,480,1538,1561]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1A</figureCitation>
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). Hind wing with apex distinctively projecting beyond tegmen; exposed apex with dorsal and ventral margins straight and tapering to a subacute apex (
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<figureCitation id="B5FDD7AFFFF1FFFAFCCE3219FCE3A311" box="[800,884,1573,1596]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1A</figureCitation>
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). Legs generally with very short and fine setae (
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<figureCitation id="B5FDD7AFFFF1FFFAFA9E3219FF2EA372" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1D</figureCitation>
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). Fore coxa with distinct, long, slightly curved spine (
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<figureCitation id="B5FDD7AFFFF1FFFAFCCF3274FCEEA372" box="[801,889,1608,1631]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1D</figureCitation>
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). Fore tibia with external tympanum open, oval; internal tympanum slit covered by large conchate swelling (
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<figureCitation id="B5FDD7AFFFF1FFFAFCF33256FCE4A3AC" box="[797,883,1642,1665]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1D</figureCitation>
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). Hind femur with genicular lobe rounded, without any spine.
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</paragraph>
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<paragraph id="2D79CB2AFFF1FFF9FF29328CFDE2A5FC" blockId="1.[151,1436,394,2012]" lastBlockId="2.[151,1435,152,244]" lastPageId="2" lastPageNumber="195" pageId="1" pageNumber="194">
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Male. Stridulatory file of left tegmen crescent-shaped,
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<quantity id="EA3E66CFFFF1FFFAFCE3328CFCC8A3EA" box="[781,863,1712,1735]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="194" unit="mm" value="2.6">2.6 mm</quantity>
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in length (
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<figureCitation id="B5FDD7AFFFF1FFFAFC3F328CFBC1A3EA" box="[977,1110,1712,1735]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Figs. 1E, 1F</figureCitation>
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); area around file not swollen; file with circa 52 per mm teeth in the middle; teeth densely and evenly spaced elongated (more closely packed at the sides than in the middle) (
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<figureCitation id="B5FDD7AFFFF1FFFAFE5E32C9FD96A221" box="[432,513,1781,1804]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Fig. 1F</figureCitation>
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). Tenth abdominal tergite simple, with little or no modification, without tooth or long process apically; with posterior margin emarginated laterally; in the middle swollen, deeply and narrowly notched (
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<figureCitation id="B5FDD7AFFFF1FFFAFA883324FF70A27C" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Figs. 1G, 1H</figureCitation>
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). Epiproct concealed beneath tenth abdominal tergite (
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<figureCitation id="B5FDD7AFFFF1FFFAFCD03306FC1DA27C" box="[830,906,1850,1873]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">
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Fig.
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<date id="5978EDEAFFF1FFFAFC9C3306FC1DA27C" box="[882,906,1850,1873]" pageId="1" pageNumber="194">1I</date>
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</figureCitation>
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). Cercus long, cylindrical and setose (especially towards the apex), modified at the apex, mostly substraight, bent fairly acutely at the apex; apex sclerotized, with two lamellate teeth of subacute apices; larger apical tooth blade-shaped, smaller tooth originating base of apical tooth, triangular (
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<figureCitation id="B5FDD7AFFFF1FFFAFEE0339EFE04A294" box="[270,403,1954,1977]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="1" pageNumber="194">Figs. 1H, 1J</figureCitation>
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). Subgenital plate anteriorly very wide, tapering sharply into a narrow apical half; with anterior margin concave; with posterior margin deeply and roundly excised in the middle, laterally extended into two long and narrow lobes; styli relatively short, longer than half of length apical lobe but shorter than half of length of subgenital plate, tapering into an obtuse apex (
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<figureCitation id="B5FDD7AFFFF2FFF9FDFD3486FDFFA5FC" box="[531,616,186,209]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="2" pageNumber="195">Fig. 1K</figureCitation>
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).
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</paragraph>
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<paragraph id="2D79CB2AFFF2FFF9FF2934E1FE04A5D9" blockId="2.[151,1435,152,244]" box="[199,403,221,244]" pageId="2" pageNumber="195">Female. Unknown.</paragraph>
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<caption id="79B99BA2FFF2FFF9FF7933BFFE9DA2DC" httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="2" pageNumber="195" startId="2.[151,244,1923,1945]" targetBox="[150,1436,298,1900]" targetPageId="2">
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<paragraph id="2D79CB2AFFF2FFF9FF7933BFFE9DA2DC" blockId="2.[151,1436,1923,2033]" pageId="2" pageNumber="195">
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<emphasis id="1FB21738FFF2FFF9FF7933BFFE9BA2B4" bold="true" box="[151,268,1923,1945]" pageId="2" pageNumber="195">FIGURE 1.</emphasis>
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<taxonomicName id="EAC6B0A9FFF2FFF9FEFC33B8FE5DA2B4" authority="Tan & Wahab, 2018" authorityName="Tan & Wahab" authorityYear="2018" box="[274,458,1924,1945]" class="Insecta" family="Tettigoniidae" genus="Tapiena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="195" phylum="Arthropoda" rank="species" species="paraincisa" status="sp. nov.">
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<emphasis id="1FB21738FFF2FFF9FEFC33B8FE5DA2B4" box="[274,458,1924,1945]" italics="true" pageId="2" pageNumber="195">Tapiena paraincisa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="0481AA43FFF2FFF9FE3E33BFFD8BA2B4" box="[464,540,1923,1945]" pageId="2" pageNumber="195" rank="species">
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<emphasis id="1FB21738FFF2FFF9FE3E33BFFD8BA2B4" bold="true" box="[464,540,1923,1945]" pageId="2" pageNumber="195">sp. nov.</emphasis>
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</taxonomicNameLabel>
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male holotype habitus with live colouration (A); head and pronotum in dorsal (B) and profile (D) views; face in anterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars: 1 mm.
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</paragraph>
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</caption>
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<paragraph id="2D79CB2AFFF3FFF8FF2934A4FEC0A48C" blockId="3.[151,1436,152,556]" pageId="3" pageNumber="196">
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Colouration. Generally green when alive (
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<figureCitation id="B5FDD7AFFFF3FFF8FD7534A4FD64A582" box="[667,755,152,175]" captionStart="FIGURE 1" captionStartId="2.[151,244,1923,1945]" captionTargetBox="[150,1436,298,1900]" captionTargetId="figure@2.[150,1436,298,1900]" captionTargetPageId="2" captionText="FIGURE 1. Tapiena paraincisa sp. nov. male holotypehabitus with live colouration(A); head and pronotum indorsal(B)andprofile (D) views; face inanterior view (C); stridulatory area of left tegmen in dorsal view (E); stridulatory file of left tegmen in ventral view (F); abdominal apex in posterior (G, H) and profile (I) views; cercus in profile view (J); subgenital plate in ventral view (K). Scale bars:1 mm." httpUri="https://zenodo.org/record/1222022/files/figure.png" pageId="3" pageNumber="196">Fig. 1A</figureCitation>
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). Head, including frontal rostrum and frons, generally green; dorsum of head with tint of yellow green. Gena pale green; mouth parts, including maxillary palps and mandibles, yellow green. Antennal scapus dark red brown; pedicel red brown. Behind each eye with a yellow stripe that extend to the lateral keel of pronotum. Pronotum green, with yellow stripe along lateral keel. Tegmen green. Hind wings hyalinous, with exposed part green. Fore and middle legs green: fore femur near knee and fore tibia around tympanal area slightly red brown. Hind legs, including tarsi, unicolourous green; knee faintly brown. Thoracic segments green; abdominal segments pale or yellow green; abdominal apex mostly pale green to slightly pale yellow green, subgenital plate and cercus pale green.
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</paragraph>
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<paragraph id="2D79CB2AFFF3FFF8FF293591FECEA4CA" blockId="3.[151,1436,152,556]" pageId="3" pageNumber="196">
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Measurements (
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<typeStatus id="F27D7588FFF3FFF8FE9C3591FE44A4E9" box="[370,467,429,452]" pageId="3" pageNumber="196" type="holotype">holotype</typeStatus>
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, in mm). BL = 22.1; BWL = 37.9; PL = 5.3; PW = 4.1; TL = 30.3; HWT = 3.8; HFL = 14.1; HTL = 14.8.
|
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</paragraph>
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|
</subSubSection>
|
|
<subSubSection id="65DC98A1FFF3FFF8FF2935CEFC42A701" pageId="3" pageNumber="196" type="etymology">
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<paragraph id="2D79CB2AFFF3FFF8FF2935CEFC42A701" blockId="3.[151,1436,152,556]" pageId="3" pageNumber="196">
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|
<emphasis id="1FB21738FFF3FFF8FF2935CEFED0A724" bold="true" box="[199,327,498,521]" pageId="3" pageNumber="196">Etymology.</emphasis>
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|
This species name reflects the similarity with
|
|
<taxonomicName id="EAC6B0A9FFF3FFF8FCD935CEFC44A724" authorityName="Karny" authorityYear="1923" box="[823,979,498,521]" class="Insecta" family="Tettigoniidae" genus="Tapiena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="196" phylum="Arthropoda" rank="species" species="incisa">
|
|
<emphasis id="1FB21738FFF3FFF8FCD935CEFC44A724" box="[823,979,498,521]" italics="true" pageId="3" pageNumber="196">Tapiena incisa</emphasis>
|
|
</taxonomicName>
|
|
in the incised tenth abdominal tergite. The
|
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<typeStatus id="F27D7588FFF3FFF8FF793629FF52A701" box="[151,197,533,556]" pageId="3" pageNumber="196">type</typeStatus>
|
|
localities of both species are also in Borneo in relatively close proximity.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
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</document> |